557 lines
62 KiB
XML
557 lines
62 KiB
XML
<document id="D3D36E3E044185E3556A91B539D07F2C" ID="10.11646/zootaxa.4248.1.1SLASH10685" ID-CLB-Dataset="33324" ID-DOI="10.5281/zenodo.439444" ID-GBIF-Dataset="cf486ccc-d298-4179-8ccf-e29a9c894ab4" ID-ISSN="1175-5326" ID-Zenodo-Dep="439444" ID-ZooBank="71F0AC03-C1FB-40AB-8532-99A638FC91E9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1490945156927" checkinUser="plazi" docAuthor="Génier, François & Moretto, Philippe" docDate="2017" docId="3D5987B70745FFAFFF17A15602D4FF3D" docLanguage="en" docName="zootaxa.4248.1.1SLASH10685.pdf" docOrigin="Zootaxa 4248 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Digitonthophagus dilatatus Genier, new species" docType="treatment" docUuid="28F9FCFD-2012-4B76-B546-B444F54327D9" docUuidSource="ZooBank" docVersion="15" lastPageNumber="61" masterDocId="C160FFCF077FFF93FF80A249006FFFCA" masterDocTitle="Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="110" masterPageNumber="1" pageNumber="59" updateTime="1698438591535" updateUser="ExternalLinkService">
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<mods:title id="C48B6FA790DB1927E364B924625E226D">Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="C7B816D9B8E71C90823F4421D297FDB1">Génier, François</mods:namePart>
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<mods:namePart id="BDADF31FB4918DBA2035C27DEEB7C71C">Moretto, Philippe</mods:namePart>
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<mods:typeOfResource id="7CE242BB7BAB5892BB449C1ADF13D06B">text</mods:typeOfResource>
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<mods:title id="09D9760873C2E90FF91D14F232EA0F56">Zootaxa</mods:title>
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<mods:part id="D6012808761FFCC97373D7F662D5DC59">
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<mods:date id="22B050142089F85794DF27718E077B92">2017</mods:date>
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<mods:detail id="D89E82B875E84CDB9F7E781BC3F0874D" type="volume">
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<mods:number id="09DB5813060D79429C72516F4C98E2B4">4248</mods:number>
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<mods:number id="453A6B46B35270217AF29CAB708FEB77">1</mods:number>
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<mods:identifier id="39988157C94316465BEC655071600FE4" type="DOI">10.5281/zenodo.439444</mods:identifier>
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<treatment id="3D5987B70745FFAFFF17A15602D4FF3D" ID-DOI="http://doi.org/10.5281/zenodo.5490090" ID-GBIF-Taxon="128090247" ID-Zenodo-Dep="5490090" LSID="urn:lsid:zoobank.org:act:28F9FCFD-2012-4B76-B546-B444F54327D9" httpUri="http://treatment.plazi.org/id/3D5987B70745FFAFFF17A15602D4FF3D" lastPageId="60" lastPageNumber="61" pageId="58" pageNumber="59">
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<subSubSection id="FDEA652A0745FFA9FF17A1560347FCB5" pageId="58" pageNumber="59" type="nomenclature">
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<paragraph id="B54F36A10745FFA9FF17A15602B5FCF0" blockId="58.[151,1167,799,895]" box="[151,730,799,826]" pageId="58" pageNumber="59">
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<heading id="EE0781CD0745FFA9FF17A15602B5FCF0" bold="true" box="[151,730,799,826]" fontSize="11" level="1" pageId="58" pageNumber="59" reason="1">
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<emphasis id="8784EAB30745FFA9FF17A15602B5FCF0" bold="true" box="[151,730,799,826]" pageId="58" pageNumber="59">
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<taxonomicName id="72F04D220745FFA9FF17A1560253FCF0" ID-CoL="363WJ" authority="Genier" authorityName="Genier" box="[151,572,799,826]" class="Insecta" family="Scarabaeidae" genus="Digitonthophagus" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="dilatatus" status="sp. nov.">
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<emphasis id="8784EAB30745FFA9FF17A156018FFCF3" bold="true" box="[151,480,799,825]" italics="true" pageId="58" pageNumber="59">Digitonthophagus dilatatus</emphasis>
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Génier
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</taxonomicName>
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,
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<taxonomicNameLabel id="9CB757C80745FFA9FDCAA16902B5FCF0" box="[586,730,800,826]" pageId="58" pageNumber="59" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="B54F36A10745FFA9FF17A10B0347FCB5" blockId="58.[151,1167,799,895]" pageId="58" pageNumber="59">
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http://zoobank.org/urn:lsid:zoobank.org:act:28F9FCFD-2012-4B76-B546-B444
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<accessionNumber id="AAA3AB420745FFA9FB95A10B0401FC91" box="[1045,1134,834,859]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/F54327" pageId="58" pageNumber="59" type="EnaNcbi">F54327</accessionNumber>
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D9 (
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<figureCitation id="2DCB2A240745FFA9FF1FA12F0093FCB5" box="[159,252,870,895]" captionStart="FIGURES 15 – 18" captionStartId="89.[151,264,1942,1964]" captionTargetBox="[218,1372,259,1904]" captionTargetId="figure@89.[206,1381,193,1921]" captionTargetPageId="89" captionText="FIGURES 15 – 18. Male dorsal habitus. 15. D. petilus, holotype; 16. D. viridicollis, holotype; 17. D. aksumensis, holotype; 18. D. dilatatus, holotype." httpUri="https://zenodo.org/record/439453/files/figure.png" pageId="58" pageNumber="59">Figs. 18</figureCitation>
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,
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<figureCitation id="2DCB2A240745FFA9FE88A12F013EFCB5" box="[264,337,870,895]" captionStart="FIGURES 39 – 46. 39, 41, 43, 45" captionStartId="93.[151,264,1959,1981]" captionTargetBox="[200,1387,193,1938]" captionTargetId="figure@93.[200,1387,193,1938]" captionTargetPageId="93" captionText="FIGURES 39 – 46. 39, 41, 43, 45. Male head, dorsal view; 40, 42, 44, 46. Male head, frontal view. 39 – 40. Digitonthophagus petilus, holotype; 41 – 42. D. viridicollis, holotype; 43 – 44. D. aksumensis, holotype; 45 – 46. D. dilatatus, holotype." httpUri="https://zenodo.org/record/439457/files/figure.png" pageId="58" pageNumber="59">45–46</figureCitation>
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,
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<figureCitation id="2DCB2A240745FFA9FEDDA12F0114FCB5" box="[349,379,870,895]" captionStart="FIGURES 63 – 70" captionStartId="96.[151,264,1926,1948]" captionTargetBox="[211,1375,193,1905]" captionTargetId="figure@96.[211,1375,193,1905]" captionTargetPageId="96" captionText="FIGURES 63 – 70. Male pronotum, dorsal view. 63. D. petilus, holotype; 64. Digitonthophagus viridicollis, holotype; 65. D. aksumensis, holotype; 66. D. dilatatus, holotype; 67. D. fimator, holotype; 68. D. lusinganus, lectotype; 69. D. ulcerosus, holotype 70. D. namaquensis, holotype." httpUri="https://zenodo.org/record/439460/files/figure.png" pageId="58" pageNumber="59">66</figureCitation>
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,
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<figureCitation id="2DCB2A240745FFA9FE06A12E01CBFCB5" box="[390,420,871,895]" captionStart="FIGURES 79 – 86" captionStartId="98.[151,264,1926,1948]" captionTargetBox="[211,1375,195,1905]" captionTargetId="figure@98.[211,1375,193,1905]" captionTargetPageId="98" captionText="FIGURES 79 – 86. Female head, dorsal view. 79. Digitonthophagus petilus, allotype; 80. D. viridicollis, allotype; 81. D. aksumensis, allotype; 82. D. dilatatus, allotype; 83. D. fimator, allotype; 84. D. lusinganus, paralectotype; 85. D. ulcerosus, allotype 86. D. namaquensis, allotype." httpUri="https://zenodo.org/record/439462/files/figure.png" pageId="58" pageNumber="59">82</figureCitation>
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, 109–110, 130, 168–170; Map 13)
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</paragraph>
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</subSubSection>
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<subSubSection id="FDEA652A0745FFA9FF17A1E704C0FC0C" box="[151,1199,942,967]" pageId="58" pageNumber="59" type="materials_examined">
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<paragraph id="B54F36A10745FFA9FF17A1E704C0FC0C" blockId="58.[151,1437,942,2011]" box="[151,1199,942,967]" pageId="58" pageNumber="59">
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<materialsCitation id="05983CFC0745FFA9FF17A1E704C0FC0C" ID-GBIF-Occurrence="1455384600" box="[151,1199,942,967]" collectorName="Foret de Sorobouli" country="Burkina Faso" elevation="270" latitude="11.795556" location="Sanguie" longLatPrecision="21" longitude="-2.8902776" pageId="58" pageNumber="59" specimenCount="1" typeStatus="holotype">
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<emphasis id="8784EAB30745FFA9FF17A1E70142FC0D" bold="true" box="[151,301,942,967]" pageId="58" pageNumber="59">
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<typeStatus id="6A4B88030745FFA9FF17A1E700BDFC0D" box="[151,210,942,967]" pageId="58" pageNumber="59">Type</typeStatus>
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locality
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</emphasis>
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.
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<collectorName id="180553770745FFA9FEBCA1E60261FC0D" box="[316,526,942,967]" pageId="58" pageNumber="59">Forêt de Sorobouli</collectorName>
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(
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<geoCoordinate id="D0C450660745FFA9FD9DA1E602C9FC0D" box="[541,678,943,967]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="15" value="11.795556">11°47’44”N</geoCoordinate>
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<geoCoordinate id="D0C450660745FFA9FD2DA1E7032FFC0D" box="[685,832,942,967]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="15" value="-2.8902776">02°53’25”W</geoCoordinate>
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),
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<quantity id="72089B440745FFA9FCD4A1E603F2FC0C" box="[852,925,943,967]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="58" pageNumber="59" unit="m" value="270.0">
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<elevation id="3EDDD1920745FFA9FCD4A1E603F2FC0C" box="[852,925,943,967]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="58" pageNumber="59" unit="m" value="270.0">270 m</elevation>
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</quantity>
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,
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<location id="B02F607A0745FFA9FC29A1E70469FC0D" LSID="urn:lsid:plazi:treatment:3D5987B70745FFAFFF17A15602D4FF3D:B02F607A0745FFA9FC29A1E70469FC0D" box="[937,1030,942,967]" country="Burkina Faso" latitude="11.795556" longLatPrecision="21" longitude="-2.8902776" name="Sanguie" pageId="58" pageNumber="59">Sanguié</location>
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,
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||
<collectingCountry id="CDE776310745FFA9FB91A1E704C5FC0C" box="[1041,1194,942,967]" name="Burkina Faso" pageId="58" pageNumber="59">Burkina Faso</collectingCountry>
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.
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||
</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="FDEA652A0745FFA9FF47A19B04CBFBF9" pageId="58" pageNumber="59" type="diagnosis">
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||
<paragraph id="B54F36A10745FFA9FF47A19B04CBFBF9" blockId="58.[151,1437,942,2011]" pageId="58" pageNumber="59">
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||
<emphasis id="8784EAB30745FFA9FF47A19B0155FC21" bold="true" box="[199,314,978,1003]" pageId="58" pageNumber="59">Diagnosis</emphasis>
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. Male cephalic horns granulate internally at apex (
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<figureCitation id="2DCB2A240745FFA9FC12A19B0385FC21" box="[914,1002,978,1003]" captionStart="FIGURES 39 – 46. 39, 41, 43, 45" captionStartId="93.[151,264,1959,1981]" captionTargetBox="[200,1387,193,1938]" captionTargetId="figure@93.[200,1387,193,1938]" captionTargetPageId="93" captionText="FIGURES 39 – 46. 39, 41, 43, 45. Male head, dorsal view; 40, 42, 44, 46. Male head, frontal view. 39 – 40. Digitonthophagus petilus, holotype; 41 – 42. D. viridicollis, holotype; 43 – 44. D. aksumensis, holotype; 45 – 46. D. dilatatus, holotype." httpUri="https://zenodo.org/record/439457/files/figure.png" pageId="58" pageNumber="59">Fig. 46</figureCitation>
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); males with pronotal anterior angles surface concave, produced anteriorly (
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<figureCitation id="2DCB2A240745FFA9FDD0A1BF02C8FBC5" box="[592,679,1014,1039]" captionStart="FIGURES 39 – 46. 39, 41, 43, 45" captionStartId="93.[151,264,1959,1981]" captionTargetBox="[200,1387,193,1938]" captionTargetId="figure@93.[200,1387,193,1938]" captionTargetPageId="93" captionText="FIGURES 39 – 46. 39, 41, 43, 45. Male head, dorsal view; 40, 42, 44, 46. Male head, frontal view. 39 – 40. Digitonthophagus petilus, holotype; 41 – 42. D. viridicollis, holotype; 43 – 44. D. aksumensis, holotype; 45 – 46. D. dilatatus, holotype." httpUri="https://zenodo.org/record/439457/files/figure.png" pageId="58" pageNumber="59">Fig. 45</figureCitation>
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); FLP sclerite ventral portion with a single carina (
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<figureCitation id="2DCB2A240745FFA9FB7CA1BF050EFBC5" box="[1276,1377,1014,1039]" captionStart="FIGURES 159 – 170" captionStartId="106.[151,264,1926,1948]" captionTargetBox="[232,1322,208,1882]" captionTargetId="figure@106.[211,1375,193,1905]" captionTargetPageId="106" captionText="FIGURES 159 – 170. Internal sac sclerites (FLP, A + SA). 159, 162, 165, 168. Dorsal view; 160, 163, 166, 169. Ventral view. 161, 164, 167, 170. Right lateral view. 159 – 161. Digitonthophagus petilus, holotype; 162 – 164. D. viridicollis, holotype; 165 – 167. D. aksumensis, holotype; 168 – 170. D. dilatatus, holotype." httpUri="https://zenodo.org/record/439466/files/figure.png" pageId="58" pageNumber="59">Fig. 170</figureCitation>
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); SA sclerite apex interrupted much before apex of apical portion of FLP sclerite (
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<figureCitation id="2DCB2A240745FFA9FC72A65304F8FBF9" box="[1010,1175,1050,1075]" captionStart="FIGURES 159 – 170" captionStartId="106.[151,264,1926,1948]" captionTargetBox="[232,1322,208,1882]" captionTargetId="figure@106.[211,1375,193,1905]" captionTargetPageId="106" captionText="FIGURES 159 – 170. Internal sac sclerites (FLP, A + SA). 159, 162, 165, 168. Dorsal view; 160, 163, 166, 169. Ventral view. 161, 164, 167, 170. Right lateral view. 159 – 161. Digitonthophagus petilus, holotype; 162 – 164. D. viridicollis, holotype; 165 – 167. D. aksumensis, holotype; 168 – 170. D. dilatatus, holotype." httpUri="https://zenodo.org/record/439466/files/figure.png" pageId="58" pageNumber="59">Figs. 168–169</figureCitation>
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).
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="FDEA652A0745FFA8FF47A67702BDFB87" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59" type="description">
|
||
<paragraph id="B54F36A10745FFA9FF47A677035AF859" blockId="58.[151,1437,942,2011]" pageId="58" pageNumber="59">
|
||
<emphasis id="8784EAB30745FFA9FF47A677013EFB9D" bold="true" box="[199,337,1086,1111]" pageId="58" pageNumber="59">Description</emphasis>
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.
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<typeStatus id="6A4B88030745FFA9FEE3A67701A4FB9D" box="[355,459,1086,1111]" pageId="58" pageNumber="59" type="holotype">Holotype</typeStatus>
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<specimenCount id="A3F6FD280745FFA9FE55A6760185FB9C" box="[469,490,1087,1110]" pageId="58" pageNumber="59" type="male">♂</specimenCount>
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(
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<figureCitation id="2DCB2A240745FFA9FE7DA677023CFB9D" box="[509,595,1086,1111]" captionStart="FIGURES 15 – 18" captionStartId="89.[151,264,1942,1964]" captionTargetBox="[218,1372,259,1904]" captionTargetId="figure@89.[206,1381,193,1921]" captionTargetPageId="89" captionText="FIGURES 15 – 18. Male dorsal habitus. 15. D. petilus, holotype; 16. D. viridicollis, holotype; 17. D. aksumensis, holotype; 18. D. dilatatus, holotype." httpUri="https://zenodo.org/record/439453/files/figure.png" pageId="58" pageNumber="59">Fig. 18</figureCitation>
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||
).
|
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<emphasis id="8784EAB30745FFA9FDEBA6770375FB9D" bold="true" box="[619,794,1086,1111]" pageId="58" pageNumber="59">Measurements</emphasis>
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. Length 12.0 mm, width 7.0 mm.
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<emphasis id="8784EAB30745FFA9FB31A677049EFB9D" bold="true" box="[1201,1265,1086,1111]" pageId="58" pageNumber="59">Head</emphasis>
|
||
(
|
||
<figureCitation id="2DCB2A240745FFA9FA83A67705FFFB9D" box="[1283,1424,1086,1111]" captionStart="FIGURES 39 – 46. 39, 41, 43, 45" captionStartId="93.[151,264,1959,1981]" captionTargetBox="[200,1387,193,1938]" captionTargetId="figure@93.[200,1387,193,1938]" captionTargetPageId="93" captionText="FIGURES 39 – 46. 39, 41, 43, 45. Male head, dorsal view; 40, 42, 44, 46. Male head, frontal view. 39 – 40. Digitonthophagus petilus, holotype; 41 – 42. D. viridicollis, holotype; 43 – 44. D. aksumensis, holotype; 45 – 46. D. dilatatus, holotype." httpUri="https://zenodo.org/record/439457/files/figure.png" pageId="58" pageNumber="59">Figs. 45–46</figureCitation>
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). Anterior clypeal edge straight on median seventh and appearing semicircular in dorsal view; clypeofrontal carina broadly arcuate and interrupted at gena; vertex lacking median tubercle, surface with punctures fine to small, separated by one to four diameters. Horns rather long, distinctly arcuate and divergent in frontal view, parallel sided on basal four-fifths and abruptly tapering externally and narrower on apical sixth; posterointernal edge unmodified basally; apicointernal surface with few, coarse granules; genal edge slightly upturned and simply arcuate on anterior third, in line with clypeal edge.
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||
<emphasis id="8784EAB30745FFA9FD4FA75F0328FAE5" bold="true" box="[719,839,1302,1327]" pageId="58" pageNumber="59">Pronotum</emphasis>
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||
(
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||
<figureCitation id="2DCB2A240745FFA9FCD7A75F03C4FAE5" box="[855,939,1302,1327]" captionStart="FIGURES 63 – 70" captionStartId="96.[151,264,1926,1948]" captionTargetBox="[211,1375,193,1905]" captionTargetId="figure@96.[211,1375,193,1905]" captionTargetPageId="96" captionText="FIGURES 63 – 70. Male pronotum, dorsal view. 63. D. petilus, holotype; 64. Digitonthophagus viridicollis, holotype; 65. D. aksumensis, holotype; 66. D. dilatatus, holotype; 67. D. fimator, holotype; 68. D. lusinganus, lectotype; 69. D. ulcerosus, holotype 70. D. namaquensis, holotype." httpUri="https://zenodo.org/record/439460/files/figure.png" pageId="58" pageNumber="59">Fig. 66</figureCitation>
|
||
). Surface with granulate punctures restricted to anterior half medially, with distinct umbilicate punctures on posterior half of disc; punctures distinct posterolaterally, with distinct minute punctures throughout. Anteromedian tubercle atrophied, forming an obtuse angle in lateral view; median longitudinal sulcus well-defined but narrow; surface behind the eyes slightly concave, surface of anterior angles concave with anterior angles produced forwardly; anterior half of lateral edge sinuous in dorsal and lateral view; posterior angles unmodified and simply arcuate in dorsal view. Anterior hypomeral ridge straight on most of its length and abruptly arcuate on anterior third, anterior hypomeral depression surface darker in color.
|
||
<emphasis id="8784EAB30745FFA9FF60A45B0142F9E1" bold="true" box="[224,301,1554,1579]" pageId="58" pageNumber="59">Elytra</emphasis>
|
||
(
|
||
<figureCitation id="2DCB2A240745FFA9FEBEA45B01FBF9E1" box="[318,404,1554,1579]" captionStart="FIGURES 15 – 18" captionStartId="89.[151,264,1942,1964]" captionTargetBox="[218,1372,259,1904]" captionTargetId="figure@89.[206,1381,193,1921]" captionTargetPageId="89" captionText="FIGURES 15 – 18. Male dorsal habitus. 15. D. petilus, holotype; 16. D. viridicollis, holotype; 17. D. aksumensis, holotype; 18. D. dilatatus, holotype." httpUri="https://zenodo.org/record/439453/files/figure.png" pageId="58" pageNumber="59">Fig. 18</figureCitation>
|
||
). Intervals 2 and 4 lacking fine granules from base to apex.
|
||
<emphasis id="8784EAB30745FFA9FBD6A45B04E1F9E1" bold="true" box="[1110,1166,1554,1579]" pageId="58" pageNumber="59">Legs</emphasis>
|
||
. Protibial apicointernal tooth enlarged, with dorsal ridge extending to apex.
|
||
<emphasis id="8784EAB30745FFA9FD5FA47F033DF985" bold="true" box="[735,850,1590,1615]" pageId="58" pageNumber="59">Aedeagus</emphasis>
|
||
(
|
||
<figureCitation id="2DCB2A240745FFA9FCE2A47F03ABF985" box="[866,964,1590,1615]" captionStart="FIGURES 119 – 134" captionStartId="103.[151,264,1876,1898]" captionTargetBox="[225,1345,208,1820]" captionTargetId="figure@103.[220,1366,193,1878]" captionTargetPageId="103" captionText="FIGURES 119 – 134. Parameres, dorsal view. 119. D. bonasus, alloreferent (Tamil Nadu State, India); 120. Digitonthophagus uks, holotype; 121. D. sahelicus, holotype; 122. D. catta, alloreferent (West Bengal, India); 123. D. eucatta, holotype; 124. D. falciger, holotype; 125. D. biflagellatus, holotype; 126. D. gazella, non type (Antisiranana Province, Madagascar); 127. D. petillus, holotype; 128. D. viridicollis, holotype; 129. D. aksumensis, holotype; 130. D. dilatatus, holotype; 131. D. fimator, holotype; 132. D. lusinganus, lectotype; 133. D. ulcerosus, holotype 134. D. namaquensis, holotype." httpUri="https://zenodo.org/record/439463/files/figure.png" pageId="58" pageNumber="59">Fig. 130</figureCitation>
|
||
). Parameres with dorsal and ventral edges slightly diverging toward apex in lateral view.
|
||
<emphasis id="8784EAB30745FFA9FD24A41303F5F9B9" bold="true" box="[676,922,1626,1651]" pageId="58" pageNumber="59">Internal sac sclerites</emphasis>
|
||
(
|
||
<figureCitation id="2DCB2A240745FFA9FC2AA413043EF9B9" box="[938,1105,1626,1651]" captionStart="FIGURES 159 – 170" captionStartId="106.[151,264,1926,1948]" captionTargetBox="[232,1322,208,1882]" captionTargetId="figure@106.[211,1375,193,1905]" captionTargetPageId="106" captionText="FIGURES 159 – 170. Internal sac sclerites (FLP, A + SA). 159, 162, 165, 168. Dorsal view; 160, 163, 166, 169. Ventral view. 161, 164, 167, 170. Right lateral view. 159 – 161. Digitonthophagus petilus, holotype; 162 – 164. D. viridicollis, holotype; 165 – 167. D. aksumensis, holotype; 168 – 170. D. dilatatus, holotype." httpUri="https://zenodo.org/record/439466/files/figure.png" pageId="58" pageNumber="59">Figs. 168–170</figureCitation>
|
||
). Axial sclerite short, angular and extending into a point apically. Subaxial sclerite, large basally and abruptly reduced, extending straight and approximately in line with apex of right lateral fold of frontolateral peripheral sclerite, with fine and scattered villi. Frontolateral peripheral sclerite basoventral apophysis well developed; a single rather large medioventral carina present; right lateral fold produced into a rather small everted and open apically conical process with irregular apical edge located behind oblique right lateral edge of apical lobe; left lateral lobe membranous, slightly developed; subapicodorsal lobe membranous, not reaching anterior edge, apex set on left side in dorsal view; apical lobe truncate apically and directed obliquely on left side, apical villi regular in shape; subapicoventral lobe small, acute apically, short not reaching apical edge of apical lobe.
|
||
</paragraph>
|
||
<paragraph id="B54F36A10745FFA8FF47A5D70487FED7" blockId="58.[151,1437,942,2011]" lastBlockId="59.[151,1437,151,429]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
|
||
<emphasis id="8784EAB30745FFA9FF47A5D70150F87D" bold="true" box="[199,319,1950,1975]" pageId="58" pageNumber="59">Variation.</emphasis>
|
||
Measurements (
|
||
<specimenCount id="A3F6FD280745FFA9FE77A5D60227F87C" box="[503,584,1951,1975]" pageId="58" pageNumber="59" type="male">42 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="A3F6FD280745FFA9FDD4A5D602CEF87D" box="[596,673,1951,1975]" pageId="58" pageNumber="59" type="female">83 ♀♀</specimenCount>
|
||
). Length: male 8.5–12.0 mm (10.5 ±
|
||
<quantity id="72089B440745FFA9FBC7A5D604CEF87D" box="[1095,1185,1951,1975]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="58" pageNumber="59" unit="mm" value="0.9">0.9 mm</quantity>
|
||
), female 7.5–12.0 mm (10.1 ±
|
||
<quantity id="72089B440745FFA9FF74A58A013DF811" box="[244,338,1987,2011]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="58" pageNumber="59" unit="mm" value="0.8">0.8 mm</quantity>
|
||
). Female
|
||
<typeStatus id="6A4B88030745FFA9FE44A58B024DF811" box="[452,546,1986,2011]" pageId="58" pageNumber="59" type="allotype">allotype</typeStatus>
|
||
. Cephalic outline in dorsal view as in
|
||
<figureCitation id="2DCB2A240745FFA9FC65A58B0455F811" box="[997,1082,1986,2011]" captionStart="FIGURES 79 – 86" captionStartId="98.[151,264,1926,1948]" captionTargetBox="[211,1375,195,1905]" captionTargetId="figure@98.[211,1375,193,1905]" captionTargetPageId="98" captionText="FIGURES 79 – 86. Female head, dorsal view. 79. Digitonthophagus petilus, allotype; 80. D. viridicollis, allotype; 81. D. aksumensis, allotype; 82. D. dilatatus, allotype; 83. D. fimator, allotype; 84. D. lusinganus, paralectotype; 85. D. ulcerosus, allotype 86. D. namaquensis, allotype." httpUri="https://zenodo.org/record/439462/files/figure.png" pageId="58" pageNumber="59">Fig. 82</figureCitation>
|
||
; vertex with a broadly arcuate transverse carina, dorsal edge broadly arcuate in frontal view, lateral portion gradually sloping down posteriorly; anterior pronotal tubercles well developed, external lateral edges slightly widening toward apex in dorsal view, anterolateral surface simply convex, anterosuperior edge slightly arcuate in dorsal view (Fig. 109), lateral portion of anterosuperior edge slightly upturned (Fig. 110). Protibia short, with external teeth more robust.
|
||
</paragraph>
|
||
<paragraph id="B54F36A10744FFA8FF47A36E02F2FE67" blockId="59.[151,1437,151,429]" pageId="59" pageNumber="60">
|
||
<materialsCitation id="05983CFC0744FFA8FF47A36E02F2FE67" ID-GBIF-Occurrence="1455384581" collectingDate="2005" collectingDateMax="2014" collectingDateMin="2005" collectionCode="CMNC" collectorName="Foret de Sorobouli & F. Genier" country="Burkina Faso" elevation="270" location="Foret de Sorobouli" pageId="59" pageNumber="60" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
|
||
<emphasis id="8784EAB30744FFA8FF47A36E01C2FE8A" bold="true" box="[199,429,295,320]" pageId="59" pageNumber="60">
|
||
Primary
|
||
<typeStatus id="6A4B88030744FFA8FEB8A36E0103FE8A" box="[312,364,295,320]" pageId="59" pageNumber="60">type</typeStatus>
|
||
data
|
||
</emphasis>
|
||
.
|
||
<typeStatus id="6A4B88030744FFA8FE40A36E0247FE8A" box="[448,552,295,320]" pageId="59" pageNumber="60" type="holotype">Holotype</typeStatus>
|
||
<specimenCount id="A3F6FD280744FFA8FDB4A3610226FEF5" box="[564,585,296,319]" pageId="59" pageNumber="60" type="male">♂</specimenCount>
|
||
(
|
||
<collectionCode id="D3E1AE640744FFA8FDDDA36102D9FE8A" box="[605,694,296,320]" pageId="59" pageNumber="60">CMNC</collectionCode>
|
||
): [
|
||
<collectingCountry id="CDE776310744FFA8FD55A36103C1FE8A" box="[725,942,296,320]" name="Burkina Faso" pageId="59" pageNumber="60">BURKINA FASO</collectingCountry>
|
||
: SANGUIÉ /
|
||
<location id="B02F607A0744FFA8FBD9A3610557FE8A" LSID="urn:lsid:plazi:treatment:3D5987B70745FFAFFF17A15602D4FF3D:B02F607A0744FFA8FBD9A3610557FE8A" box="[1113,1336,295,320]" country="Burkina Faso" name="Foret de Sorobouli" pageId="59" pageNumber="60">
|
||
<collectorName id="180553770744FFA8FBD9A3610557FE8A" box="[1113,1336,295,320]" pageId="59" pageNumber="60">Forêt de Sorobouli</collectorName>
|
||
</location>
|
||
,
|
||
<quantity id="72089B440744FFA8FAC8A36105E7FE8A" box="[1352,1416,296,320]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="59" pageNumber="60" unit="m" value="270.0">
|
||
<elevation id="3EDDD1920744FFA8FAC8A36105E7FE8A" box="[1352,1416,296,320]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="59" pageNumber="60" unit="m" value="270.0">270m</elevation>
|
||
</quantity>
|
||
/ 11̊47’44”N 0 0 2̊53’25”O /
|
||
<date id="C14E10610744FFA8FE52A3050208FEAF" box="[466,615,332,357]" pageId="59" pageNumber="60" value="2005-08-13">
|
||
<collectingDate id="D10AE9890744FFA8FE52A3050208FEAF" box="[466,615,332,357]" pageId="59" pageNumber="60" value="2005-08-13">13.VIII.2005</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="180553770744FFA8FDFAA3040287FEAF" box="[634,744,332,357]" pageId="59" pageNumber="60">F. Génier</collectorName>
|
||
/ zone soudanienne sud / piège excr. humain,
|
||
<collectingDate id="D10AE9890744FFA8FAAEA30505FEFEAF" box="[1326,1425,332,357]" pageId="59" pageNumber="60" value="2005" valueMax="2014">
|
||
<date id="C14E10610744FFA8FAAEA30505FEFEAF" box="[1326,1425,332,357]" pageId="59" pageNumber="60" value="2005" valueMax="2014">2005-14</date>
|
||
</collectingDate>
|
||
]; [COLÉOPTÈRES / DU /
|
||
<collectingCountry id="CDE776310744FFA8FE44A33902FCFE42" box="[452,659,368,392]" name="Burkina Faso" pageId="59" pageNumber="60">BURKINA FASO</collectingCountry>
|
||
/ BF015361] barcode label; [
|
||
<typeStatus id="6A4B88030744FFA8FC6CA33904ECFE42" box="[1004,1155,368,392]" pageId="59" pageNumber="60" type="holotype">HOLOTYPE</typeStatus>
|
||
<specimenCount id="A3F6FD280744FFA8FB0EA33904CCFE4D" box="[1166,1187,368,391]" pageId="59" pageNumber="60" type="male">♂</specimenCount>
|
||
/
|
||
<taxonomicName id="72F04D220744FFA8FB3FA33905E6FE42" box="[1215,1417,368,392]" class="Insecta" family="Scarabaeidae" genus="Digitonthophagus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="8784EAB30744FFA8FB3FA33905E6FE42" box="[1215,1417,368,392]" italics="true" pageId="59" pageNumber="60">Digitonthophagus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<emphasis id="8784EAB30744FFA8FF17A3DD015AFE66" box="[151,309,404,428]" italics="true" pageId="59" pageNumber="60">
|
||
<taxonomicName id="72F04D220744FFA8FF17A3DD0096FE66" box="[151,249,404,428]" class="Insecta" family="Scarabaeidae" genus="Digitonthophagus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="dilatatus" status="sp. nov.">dilatatus</taxonomicName>
|
||
<taxonomicNameLabel id="9CB757C80744FFA8FE80A3DC015AFE66" box="[256,309,405,428]" pageId="59" pageNumber="60" rank="species">n.sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
/ des.
|
||
<collectorName id="180553770744FFA8FEFDA3DC0188FE67" box="[381,487,404,429]" pageId="59" pageNumber="60">F. Génier</collectorName>
|
||
, 2016] red card.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="B54F36A10744FFA8FF17A67F02BDFB87" blockId="59.[151,722,1078,1101]" box="[151,722,1078,1101]" pageId="59" pageNumber="60">
|
||
<emphasis id="8784EAB30744FFA8FF17A67F0094FB87" bold="true" box="[151,251,1078,1101]" pageId="59" pageNumber="60">MAP 13.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="72F04D220744FFA8FE26A67F02A4FB87" box="[422,715,1078,1101]" class="Insecta" family="Scarabaeidae" genus="Digitonthophagus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="dilatatus">
|
||
<emphasis id="8784EAB30744FFA8FE26A67F02A4FB87" box="[422,715,1078,1101]" italics="true" pageId="59" pageNumber="60">Digitonthophagus dilatatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FDEA652A0744FFA8FF47A63304B6F919" pageId="59" pageNumber="60" type="materials_examined">
|
||
<paragraph id="B54F36A10744FFA8FF47A63304B6F919" blockId="59.[151,1437,1146,1963]" pageId="59" pageNumber="60">
|
||
<materialsCitation id="05983CFC0744FFA8FF47A633043DFB59" ID-GBIF-Occurrence="1455384586" box="[199,1106,1146,1171]" collectorName="Map" country="Burkina Faso" location="Material" pageId="59" pageNumber="60" specimenCount="150" specimenCount-female="93" specimenCount-male="57">
|
||
<emphasis id="8784EAB30744FFA8FF47A63301C5FB59" bold="true" box="[199,426,1146,1171]" pageId="59" pageNumber="60">
|
||
<location id="B02F607A0744FFA8FF47A6330140FB59" LSID="urn:lsid:plazi:treatment:3D5987B70745FFAFFF17A15602D4FF3D:B02F607A0744FFA8FF47A6330140FB59" box="[199,303,1146,1171]" country="Burkina Faso" name="Material" pageId="59" pageNumber="60">Material</location>
|
||
examined
|
||
</emphasis>
|
||
(
|
||
<specimenCount id="A3F6FD280744FFA8FE39A6330265FB58" box="[441,522,1146,1171]" pageId="59" pageNumber="60" type="male">57 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="A3F6FD280744FFA8FD97A632020BFB59" box="[535,612,1147,1171]" pageId="59" pageNumber="60" type="female">93 ♀♀</specimenCount>
|
||
), distribution (
|
||
<collectorName id="180553770744FFA8FC8DA632032DFB59" box="[781,834,1147,1171]" pageId="59" pageNumber="60">Map</collectorName>
|
||
13):
|
||
<emphasis id="8784EAB30744FFA8FCFEA633043DFB59" bold="true" box="[894,1106,1146,1171]" pageId="59" pageNumber="60">
|
||
<collectingCountry id="CDE776310744FFA8FCFEA633043DFB59" box="[894,1106,1146,1171]" name="Burkina Faso" pageId="59" pageNumber="60">BURKINA FASO</collectingCountry>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
:
|
||
<materialsCitation id="05983CFC0744FFA8FBE2A632022CFB11" ID-GBIF-Occurrence="1455384577" collectingDate="2005-08-10" collectingDateMax="2005-08-15" collectingDateMin="2005-08-10" collectionCode="FGIC, PMOC" collectorName="Foret de Sorobouli & Foret Classee de Baporo" country="Burkina Faso" elevation="270" latitude="11.795556" location="Foret Classee de Baporo" longLatPrecision="21" longitude="-2.8902776" municipality="Foret de Sorobouli" pageId="59" pageNumber="60" specimenCount="216" specimenCount-female="78" specimenCount-male="30" stateProvince="CENTRE-OUEST">
|
||
<collectingRegion id="7734F8430744FFA8FBE2A632055CFB59" box="[1122,1331,1147,1171]" country="Burkina Faso" name="Centre-Ouest" pageId="59" pageNumber="60">CENTRE-OUEST</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="552BACDB0744FFA8FABEA632016AFB7D" pageId="59" pageNumber="60">
|
||
<collectorName id="180553770744FFA8FABEA632016AFB7D" pageId="59" pageNumber="60">Forêt de Sorobouli</collectorName>
|
||
</collectingMunicipality>
|
||
[=
|
||
<location id="B02F607A0744FFA8FEA3A6D6022DFB7D" LSID="urn:lsid:plazi:treatment:3D5987B70745FFAFFF17A15602D4FF3D:B02F607A0744FFA8FEA3A6D6022DFB7D" box="[291,578,1182,1207]" country="Burkina Faso" latitude="11.795556" longLatPrecision="21" longitude="-2.8902776" municipality="Foret de Sorobouli" name="Foret Classee de Baporo" pageId="59" pageNumber="60" stateProvince="CENTRE-OUEST">
|
||
<collectorName id="180553770744FFA8FEA3A6D6022DFB7D" box="[291,578,1182,1207]" pageId="59" pageNumber="60">Forêt Classée de Baporo</collectorName>
|
||
</location>
|
||
],
|
||
<quantity id="72089B440744FFA8FDD4A6D602F3FB7D" box="[596,668,1183,1207]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="59" pageNumber="60" unit="m" value="270.0">
|
||
<elevation id="3EDDD1920744FFA8FDD4A6D602F3FB7D" box="[596,668,1183,1207]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="59" pageNumber="60" unit="m" value="270.0">270 m</elevation>
|
||
</quantity>
|
||
(
|
||
<geoCoordinate id="D0C450660744FFA8FD2CA6D6035EFB7D" box="[684,817,1183,1207]" direction="north" orientation="latitude" pageId="59" pageNumber="60" precision="15" value="11.795556">11°47'44''N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D0C450660744FFA8FCBEA6D703D2FB7D" box="[830,957,1182,1207]" direction="west" orientation="longitude" pageId="59" pageNumber="60" precision="15" value="-2.8902776">2°53'25''W</geoCoordinate>
|
||
),
|
||
<date id="C14E10610744FFA8FC51A6D704EAFB7D" box="[977,1157,1182,1207]" pageId="59" pageNumber="60" value="2005-08-10" valueMax="2005-08-15" valueMin="2005-08-10">
|
||
<collectingDate id="D10AE9890744FFA8FC51A6D704EAFB7D" box="[977,1157,1182,1207]" pageId="59" pageNumber="60" value="2005-08-10" valueMax="2005-08-15" valueMin="2005-08-10">10–15.viii.2005</collectingDate>
|
||
</date>
|
||
, P. Moretto—
|
||
<specimenCount id="A3F6FD280744FFA8FAA5A6D6051DFB7D" box="[1317,1394,1183,1207]" pageId="59" pageNumber="60" type="female">78 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="A3F6FD280744FFA8FA00A6D600AEFB10" pageId="59" pageNumber="60" type="male">30 ♂♂</specimenCount>
|
||
(
|
||
<specimenCount id="A3F6FD280744FFA8FF54A68A0117FB11" box="[212,376,1219,1243]" pageId="59" pageNumber="60" type="generic" typeStatus="paratypes">108 paratypes</specimenCount>
|
||
) (
|
||
<collectionCode id="D3E1AE640744FFA8FE12A68A01B9FB11" box="[402,470,1219,1243]" country="0" httpUri="http://grbio.org/cool/y3g8-69h7" name="Francois Genier" pageId="59" pageNumber="60">FGIC</collectionCode>
|
||
,
|
||
<collectionCode id="D3E1AE640744FFA8FE66A68A0254FB11" box="[486,571,1219,1243]" pageId="59" pageNumber="60">PMOC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
; BALÉ, Boromo,
|
||
<quantity id="72089B440744FFA8FC9BA68B0309FB10" box="[795,870,1218,1243]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="59" pageNumber="60" unit="m" value="250.0">250 m</quantity>
|
||
(11°45'6''N, 2°51'58''W),
|
||
<date id="C14E10610744FFA8FB17A68B054EFB11" box="[1175,1313,1218,1243]" pageId="59" pageNumber="60" value="2005-08-10">10.viii.2005</date>
|
||
, F. Génier (
|
||
<date id="C14E10610744FFA8FF1FA6AF016FFB35" box="[159,256,1254,1279]" pageId="59" pageNumber="60" value="2005" valueMax="2010">2005-10</date>
|
||
)—1 ♀, 1 ♂ (paratypes) (FGIC); KOMPIENGA, Pama,
|
||
<quantity id="72089B440744FFA8FC15A6AE038FFB35" box="[917,992,1255,1279]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="59" pageNumber="60" unit="m" value="230.0">230 m</quantity>
|
||
(11°17'N, 0°42'59''E),
|
||
<date id="C14E10610744FFA8FB6EA6AF0516FB35" box="[1262,1401,1254,1279]" pageId="59" pageNumber="60" value="2005-08-24">24.viii.2005</date>
|
||
, F. Génier (
|
||
<date id="C14E10610744FFA8FF73A743013DFAE9" box="[243,338,1290,1315]" pageId="59" pageNumber="60" value="2005" valueMax="2044">2005-44</date>
|
||
)—3 ♂♂ (3 paratypes) (FGIC); SANGUIÉ, Forêt de Sorobouli,
|
||
<quantity id="72089B440744FFA8FBA8A742041FFAE8" box="[1064,1136,1291,1315]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="59" pageNumber="60" unit="m" value="270.0">270 m</quantity>
|
||
(11°47'44''N, 2°53'25''W),
|
||
<date id="C14E10610744FFA8FF17A767014FFA8D" box="[151,288,1326,1351]" pageId="59" pageNumber="60" value="2005-08-13">13.viii.2005</date>
|
||
, F. Génier (
|
||
<date id="C14E10610744FFA8FE41A767024DFA8D" box="[449,546,1326,1351]" pageId="59" pageNumber="60" value="2005" valueMax="2014">2005-14</date>
|
||
)—1 ♀, 1 ♂ (holotype, allotype) (CMNC); Forêt de Sorobouli,
|
||
<quantity id="72089B440744FFA8FACBA76605F4FA8D" box="[1355,1435,1327,1351]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="59" pageNumber="60" unit="m" value="270.0">270 m</quantity>
|
||
(11°47'44''N, 2°53'25''W),
|
||
<date id="C14E10610744FFA8FE44A71B0222FAA1" box="[452,589,1362,1387]" pageId="59" pageNumber="60" value="2005-08-13">13.viii.2005</date>
|
||
, F. Génier (
|
||
<date id="C14E10610744FFA8FD53A71B035FFAA1" box="[723,816,1362,1387]" pageId="59" pageNumber="60" value="2005" valueMax="2015">2005-15</date>
|
||
)—2 ♀♀, 1 ♂ (paratypes) (FGIC); Forêt de Sorobouli,
|
||
<quantity id="72089B440744FFA8FF17A73E00B1FA45" box="[151,222,1399,1423]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="59" pageNumber="60" unit="m" value="270.0">270 m</quantity>
|
||
(11°47'44''N, 2°53'25''W),
|
||
<date id="C14E10610744FFA8FD91A73F02FCFA45" box="[529,659,1398,1423]" pageId="59" pageNumber="60" value="2006-07-28">28.vii.2006</date>
|
||
, F. & S. Génier (
|
||
<date id="C14E10610744FFA8FCD2A73F03DCFA45" box="[850,947,1398,1423]" pageId="59" pageNumber="60" value="2006" valueMax="2083">2006-83</date>
|
||
)—1 ♂ (paratype) (FGIC).
|
||
<materialsCitation id="05983CFC0744FFA8FB5FA73E0464FA79" ID-GBIF-Occurrence="1455384585" collectingDate="1978-07-04" collectionCode="EXTREME" collectorName="Maroua" country="Cameroon" latitude="10.583333" location="NORTH" longLatPrecision="1297" longitude="14.316667" pageId="59" pageNumber="60" specimenCount="1" stateProvince="NORTH">
|
||
;
|
||
<emphasis id="8784EAB30744FFA8FB6BA73F05FAFA45" bold="true" box="[1259,1429,1398,1423]" pageId="59" pageNumber="60">
|
||
<collectingCountry id="CDE776310744FFA8FB6BA73F05FAFA45" box="[1259,1429,1398,1423]" name="Cameroon" pageId="59" pageNumber="60">CAMEROON</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<collectionCode id="D3E1AE640744FFA8FF17A7D20174FA78" box="[151,283,1435,1458]" pageId="59" pageNumber="60">EXTREME</collectionCode>
|
||
<collectingRegion id="7734F8430744FFA8FEA8A7D201E5FA79" box="[296,394,1435,1459]" country="Cameroon" name="North" pageId="59" pageNumber="60">NORTH</collectingRegion>
|
||
,
|
||
<collectorName id="180553770744FFA8FE1BA7D2019DFA78" box="[411,498,1435,1458]" pageId="59" pageNumber="60">Maroua</collectorName>
|
||
(
|
||
<geoCoordinate id="D0C450660744FFA8FD87A7D30209FA79" box="[519,614,1434,1459]" direction="north" orientation="latitude" pageId="59" pageNumber="60" precision="925" value="10.583333">10°35'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D0C450660744FFA8FDF9A7D202BBFA79" box="[633,724,1435,1459]" direction="east" orientation="longitude" pageId="59" pageNumber="60" precision="925" value="14.316667">14°19'E</geoCoordinate>
|
||
),
|
||
<date id="C14E10610744FFA8FD6FA7D3030CFA79" box="[751,867,1434,1459]" pageId="59" pageNumber="60" value="1978-07-04">
|
||
<collectingDate id="D10AE9890744FFA8FD6FA7D3030CFA79" box="[751,867,1434,1459]" pageId="59" pageNumber="60" value="1978-07-04">4.vii.1978</collectingDate>
|
||
</date>
|
||
, [anonymous]
|
||
</materialsCitation>
|
||
—1 ♂ (paratype) (MNHN);
|
||
<materialsCitation id="05983CFC0744FFA8FADFA7D204CEFA1D" ID-GBIF-Occurrence="1455384592" collectingDate="2006-08-18" collectionCode="FGIC, PMOC" country="Cameroon" latitude="11.4" location="Waza" longLatPrecision="1295" longitude="14.566667" pageId="59" pageNumber="60" specimenCount="4" specimenCount-male="2" stateProvince="NORTH">
|
||
<location id="B02F607A0744FFA8FADFA7D205F3FA78" LSID="urn:lsid:plazi:treatment:3D5987B70745FFAFFF17A15602D4FF3D:B02F607A0744FFA8FADFA7D205F3FA78" box="[1375,1436,1435,1458]" country="Cameroon" latitude="11.4" longLatPrecision="1295" longitude="14.566667" name="Waza" pageId="59" pageNumber="60" stateProvince="NORTH">Waza</location>
|
||
(
|
||
<geoCoordinate id="D0C450660744FFA8FF1FA7F60092FA1D" box="[159,253,1471,1495]" direction="north" orientation="latitude" pageId="59" pageNumber="60" precision="925" value="11.4">11°24'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D0C450660744FFA8FE8FA7F60103FA1D" box="[271,364,1471,1495]" direction="east" orientation="longitude" pageId="59" pageNumber="60" precision="925" value="14.566667">14°34'E</geoCoordinate>
|
||
),
|
||
<date id="C14E10610744FFA8FE05A7F7027FFA1D" box="[389,528,1470,1495]" pageId="59" pageNumber="60" value="2006-08-18">
|
||
<collectingDate id="D10AE9890744FFA8FE05A7F7027FFA1D" box="[389,528,1470,1495]" pageId="59" pageNumber="60" value="2006-08-18">18.viii.2006</collectingDate>
|
||
</date>
|
||
, C. Vanderbergh—
|
||
<specimenCount id="A3F6FD280744FFA8FD73A7F70359FA1D" box="[755,822,1470,1495]" pageId="59" pageNumber="60" type="male">2 ♂♂</specimenCount>
|
||
(
|
||
<specimenCount id="A3F6FD280744FFA8FCCBA7F603BCFA1D" box="[843,979,1471,1495]" pageId="59" pageNumber="60" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
|
||
) (
|
||
<collectionCode id="D3E1AE640744FFA8FC70A7F6045BFA1D" box="[1008,1076,1471,1495]" country="0" httpUri="http://grbio.org/cool/y3g8-69h7" name="Francois Genier" pageId="59" pageNumber="60">FGIC</collectionCode>
|
||
,
|
||
<collectionCode id="D3E1AE640744FFA8FBC5A7F604F6FA1D" box="[1093,1177,1471,1495]" pageId="59" pageNumber="60">PMOC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="05983CFC0744FFA8FB31A7890541FA31" ID-GBIF-Occurrence="1455384576" collectingDate="2006-08-18" collectionCode="FGIC, PMOC" country="Cameroon" latitude="11.25" location="Waza National Park" longLatPrecision="1295" longitude="14.65" pageId="59" pageNumber="60" specimenCount="20" specimenCount-female="5" specimenCount-male="5" stateProvince="NORTH">
|
||
<location id="B02F607A0744FFA8FB31A78905F3FA1D" LSID="urn:lsid:plazi:treatment:3D5987B70745FFAFFF17A15602D4FF3D:B02F607A0744FFA8FB31A78905F3FA1D" box="[1201,1436,1470,1495]" country="Cameroon" latitude="11.25" longLatPrecision="1295" longitude="14.65" name="Waza National Park" pageId="59" pageNumber="60" stateProvince="NORTH">Waza National Park</location>
|
||
(
|
||
<geoCoordinate id="D0C450660744FFA8FF20A7AB0092FA31" box="[160,253,1506,1531]" direction="north" orientation="latitude" pageId="59" pageNumber="60" precision="925" value="11.25">11°15'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D0C450660744FFA8FE93A7AA0100FA31" box="[275,367,1507,1531]" direction="east" orientation="longitude" pageId="59" pageNumber="60" precision="925" value="14.65">14°39'E</geoCoordinate>
|
||
),
|
||
<date id="C14E10610744FFA8FE0CA7AB0278FA31" box="[396,535,1506,1531]" pageId="59" pageNumber="60" value="2006-08-18">
|
||
<collectingDate id="D10AE9890744FFA8FE0CA7AB0278FA31" box="[396,535,1506,1531]" pageId="59" pageNumber="60" value="2006-08-18">18.viii.2006</collectingDate>
|
||
</date>
|
||
, C. Vanderbergh—
|
||
<specimenCount id="A3F6FD280744FFA8FC82A7AB0326FA30" box="[770,841,1506,1531]" pageId="59" pageNumber="60" type="female">5 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="A3F6FD280744FFA8FCDEA7AB03C8FA30" box="[862,935,1506,1531]" pageId="59" pageNumber="60" type="male">5 ♂♂</specimenCount>
|
||
(
|
||
<specimenCount id="A3F6FD280744FFA8FC3FA7AA0436FA31" box="[959,1113,1507,1531]" pageId="59" pageNumber="60" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
|
||
) (
|
||
<collectionCode id="D3E1AE640744FFA8FBF9A7AA04D2FA31" box="[1145,1213,1507,1531]" country="0" httpUri="http://grbio.org/cool/y3g8-69h7" name="Francois Genier" pageId="59" pageNumber="60">FGIC</collectionCode>
|
||
,
|
||
<collectionCode id="D3E1AE640744FFA8FB51A7AA0549FA31" box="[1233,1318,1507,1531]" pageId="59" pageNumber="60">PMOC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="05983CFC0744FFA8FAC2A7AB04B0F9D5" ID-GBIF-Occurrence="1455384583" collectingDate="1968-06" collectionCode="MNHN" country="Chad" latitude="12.116667" location="Farcha" longLatPrecision="1293" longitude="14.983334" pageId="59" pageNumber="60" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||
<emphasis id="8784EAB30744FFA8FAC2A7AB05FAFA31" bold="true" box="[1346,1429,1506,1531]" pageId="59" pageNumber="60">
|
||
<collectingCountry id="CDE776310744FFA8FAC2A7AB05FAFA31" box="[1346,1429,1506,1531]" name="Chad" pageId="59" pageNumber="60">CHAD</collectingCountry>
|
||
</emphasis>
|
||
: N’DJAMENA,
|
||
<location id="B02F607A0744FFA8FEC8A44F01FBF9D5" LSID="urn:lsid:plazi:treatment:3D5987B70745FFAFFF17A15602D4FF3D:B02F607A0744FFA8FEC8A44F01FBF9D5" box="[328,404,1542,1567]" country="Chad" latitude="12.116667" longLatPrecision="1293" longitude="14.983334" name="Farcha" pageId="59" pageNumber="60">Farcha</location>
|
||
(
|
||
<geoCoordinate id="D0C450660744FFA8FE24A44E0199F9D5" box="[420,502,1543,1567]" direction="north" orientation="latitude" pageId="59" pageNumber="60" precision="925" value="12.116667">12°7'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D0C450660744FFA8FD81A44F0231F9D5" box="[513,606,1542,1567]" direction="east" orientation="longitude" pageId="59" pageNumber="60" precision="925" value="14.983334">14°59'E</geoCoordinate>
|
||
),
|
||
<date id="C14E10610744FFA8FDF2A44F02A6F9D5" box="[626,713,1542,1567]" pageId="59" pageNumber="60" value="1968-06">
|
||
<collectingDate id="D10AE9890744FFA8FDF2A44F02A6F9D5" box="[626,713,1542,1567]" pageId="59" pageNumber="60" value="1968-06">vi.1968</collectingDate>
|
||
</date>
|
||
, [anonymous]—
|
||
<specimenCount id="A3F6FD280744FFA8FC0AA44E03C0F9D5" box="[906,943,1543,1567]" pageId="59" pageNumber="60" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount id="A3F6FD280744FFA8FC3DA44E0388F9D5" box="[957,999,1543,1567]" pageId="59" pageNumber="60" type="male">1 ♂</specimenCount>
|
||
(
|
||
<typeStatus id="6A4B88030744FFA8FC76A44E040AF9D5" box="[1014,1125,1543,1567]" pageId="59" pageNumber="60" type="paratype">paratypes</typeStatus>
|
||
) (
|
||
<collectionCode id="D3E1AE640744FFA8FBFBA44E04B8F9D5" box="[1147,1239,1543,1567]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="59" pageNumber="60">MNHN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="05983CFC0744FFA8FB6AA44F039EF989" ID-GBIF-Occurrence="1455384584" collectingDate="1967-06-28" collectionCode="MNHN" country="Chad" latitude="12.116667" location="Farcha" longLatPrecision="1293" longitude="14.983334" pageId="59" pageNumber="60" specimenCount="6" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
|
||
<location id="B02F607A0744FFA8FB6AA44F0559F9D5" LSID="urn:lsid:plazi:treatment:3D5987B70745FFAFFF17A15602D4FF3D:B02F607A0744FFA8FB6AA44F0559F9D5" box="[1258,1334,1542,1567]" country="Chad" latitude="12.116667" longLatPrecision="1293" longitude="14.983334" name="Farcha" pageId="59" pageNumber="60">Farcha</location>
|
||
(
|
||
<geoCoordinate id="D0C450660744FFA8FAC6A44E05F8F9D5" box="[1350,1431,1543,1567]" direction="north" orientation="latitude" pageId="59" pageNumber="60" precision="925" value="12.116667">12°7'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D0C450660744FFA8FF17A463009CF989" box="[151,243,1578,1603]" direction="east" orientation="longitude" pageId="59" pageNumber="60" precision="925" value="14.983334">14°59'E</geoCoordinate>
|
||
),
|
||
<date id="C14E10610744FFA8FE8DA46301E7F989" box="[269,392,1578,1603]" pageId="59" pageNumber="60" value="1967-06-28">
|
||
<collectingDate id="D10AE9890744FFA8FE8DA46301E7F989" box="[269,392,1578,1603]" pageId="59" pageNumber="60" value="1967-06-28">28.vi.1967</collectingDate>
|
||
</date>
|
||
, [anonymous]—
|
||
<specimenCount id="A3F6FD280744FFA8FDCEA4620216F988" box="[590,633,1579,1603]" pageId="59" pageNumber="60" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount id="A3F6FD280744FFA8FD0DA46202BDF988" box="[653,722,1579,1603]" pageId="59" pageNumber="60" type="male">2 ♂♂</specimenCount>
|
||
(
|
||
<specimenCount id="A3F6FD280744FFA8FD68A462031FF989" box="[744,880,1579,1603]" pageId="59" pageNumber="60" type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
|
||
) (
|
||
<collectionCode id="D3E1AE640744FFA8FC0CA4620387F989" box="[908,1000,1579,1603]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="59" pageNumber="60">MNHN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="05983CFC0744FFA8FB81A462039FF9AD" ID-GBIF-Occurrence="1455384607" collectingDate="1958-07-17" collectionCode="FETC" country="Ivory Coast" latitude="12.116667" location="N'Djamena" longLatPrecision="1293" longitude="15.05" municipality="Fort Lamy" pageId="59" pageNumber="60" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<collectingMunicipality id="552BACDB0744FFA8FB81A4620411F989" box="[1025,1150,1579,1603]" pageId="59" pageNumber="60">Fort Lamy</collectingMunicipality>
|
||
[=
|
||
<location id="B02F607A0744FFA8FB21A4630545F989" LSID="urn:lsid:plazi:treatment:3D5987B70745FFAFFF17A15602D4FF3D:B02F607A0744FFA8FB21A4630545F989" box="[1185,1322,1578,1603]" country="Ivory Coast" latitude="12.116667" longLatPrecision="1293" longitude="15.05" municipality="Fort Lamy" name="N'Djamena" pageId="59" pageNumber="60">N’Djamena</location>
|
||
] (
|
||
<geoCoordinate id="D0C450660744FFA8FAC6A46205F8F989" box="[1350,1431,1579,1603]" direction="north" orientation="latitude" pageId="59" pageNumber="60" precision="925" value="12.116667">12°7'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D0C450660744FFA8FF17A407008BF9AD" box="[151,228,1614,1639]" direction="east" orientation="longitude" pageId="59" pageNumber="60" precision="925" value="15.05">15°3'E</geoCoordinate>
|
||
),
|
||
<date id="C14E10610744FFA8FF7AA4070114F9AD" box="[250,379,1614,1639]" pageId="59" pageNumber="60" value="1958-07-17">
|
||
<collectingDate id="D10AE9890744FFA8FF7AA4070114F9AD" box="[250,379,1614,1639]" pageId="59" pageNumber="60" value="1958-07-17">17.vii.1958</collectingDate>
|
||
</date>
|
||
, J. Mateu—
|
||
<specimenCount id="A3F6FD280744FFA8FD8AA4060241F9AD" box="[522,558,1615,1639]" pageId="59" pageNumber="60" type="male">1 ♂</specimenCount>
|
||
(
|
||
<typeStatus id="6A4B88030744FFA8FDBDA40602CEF9AD" box="[573,673,1615,1639]" pageId="59" pageNumber="60" type="paratype">paratype</typeStatus>
|
||
) (
|
||
<collectionCode id="D3E1AE640744FFA8FD37A406036DF9AD" box="[695,770,1615,1639]" pageId="59" pageNumber="60">FETC</collectionCode>
|
||
);
|
||
<emphasis id="8784EAB30744FFA8FC95A407039FF9AD" bold="true" box="[789,1008,1614,1639]" pageId="59" pageNumber="60">
|
||
<collectingCountry id="CDE776310744FFA8FC95A407039FF9AD" box="[789,1008,1614,1639]" name="Ivory Coast" pageId="59" pageNumber="60">CÔTE D’IVOIRE</collectingCountry>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
:
|
||
<materialsCitation id="05983CFC0744FFA8FC7FA406054EF941" ID-GBIF-Occurrence="1455384589" collectingDate="2013-07-13" collectingDateMax="2013-07-15" collectingDateMin="2013-07-13" collectionCode="FGIC, PMOC" country="Ivory Coast" elevation="395" latitude="9.569555" location="Boundiali" longLatPrecision="1" longitude="-6.682361" pageId="59" pageNumber="60" specimenCount="14" specimenCount-female="2" specimenCount-male="5" stateProvince="SAVANES" typeStatus="paratype">
|
||
<collectingRegion id="7734F8430744FFA8FC7FA4060413F9AD" box="[1023,1148,1615,1639]" country="Ivory Coast" name="Savanes" pageId="59" pageNumber="60">SAVANES</collectingRegion>
|
||
, M’banto,
|
||
<location id="B02F607A0744FFA8FB75A4070509F9AD" LSID="urn:lsid:plazi:treatment:3D5987B70745FFAFFF17A15602D4FF3D:B02F607A0744FFA8FB75A4070509F9AD" box="[1269,1382,1614,1639]" country="Ivory Coast" latitude="9.569555" longLatPrecision="1" longitude="-6.682361" name="Boundiali" pageId="59" pageNumber="60" stateProvince="SAVANES">Boundiali</location>
|
||
,
|
||
<quantity id="72089B440744FFA8FAF2A40700C2F940" metricMagnitude="2" metricUnit="m" metricValue="3.95" pageId="59" pageNumber="60" unit="m" value="395.0">
|
||
<elevation id="3EDDD1920744FFA8FAF2A40700C2F940" metricMagnitude="2" metricUnit="m" metricValue="3.95" pageId="59" pageNumber="60" unit="m" value="395.0">395 m</elevation>
|
||
</quantity>
|
||
(
|
||
<geoCoordinate id="D0C450660744FFA8FF3CA43A0126F941" box="[188,329,1651,1675]" direction="north" orientation="latitude" pageId="59" pageNumber="60" precision="1" value="9.569555">9°34'10.4''N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D0C450660744FFA8FED4A43B0188F941" box="[340,487,1650,1675]" direction="west" orientation="longitude" pageId="59" pageNumber="60" precision="1" value="-6.682361">6°40'56.5''W</geoCoordinate>
|
||
),
|
||
<date id="C14E10610744FFA8FE7BA43B02C8F941" box="[507,679,1650,1675]" pageId="59" pageNumber="60" value="2013-07-13" valueMax="2013-07-15" valueMin="2013-07-13">
|
||
<collectingDate id="D10AE9890744FFA8FE7BA43B02C8F941" box="[507,679,1650,1675]" pageId="59" pageNumber="60" value="2013-07-13" valueMax="2013-07-15" valueMin="2013-07-13">13–15.vii.2013</collectingDate>
|
||
</date>
|
||
, P. Moretto—
|
||
<specimenCount id="A3F6FD280744FFA8FCC4A43A03EDF940" box="[836,898,1651,1675]" pageId="59" pageNumber="60" type="female">2 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="A3F6FD280744FFA8FC0FA43B03A1F940" box="[911,974,1650,1675]" pageId="59" pageNumber="60" type="male">5 ♂♂</specimenCount>
|
||
(
|
||
<specimenCount id="A3F6FD280744FFA8FC5DA43A040FF941" box="[989,1120,1651,1675]" pageId="59" pageNumber="60" type="generic" typeStatus="paratypes">7 paratypes</specimenCount>
|
||
) (
|
||
<collectionCode id="D3E1AE640744FFA8FBF6A43A04D5F941" box="[1142,1210,1651,1675]" country="0" httpUri="http://grbio.org/cool/y3g8-69h7" name="Francois Genier" pageId="59" pageNumber="60">FGIC</collectionCode>
|
||
,
|
||
<collectionCode id="D3E1AE640744FFA8FB45A43A0576F941" box="[1221,1305,1651,1675]" pageId="59" pageNumber="60">PMOC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="05983CFC0744FFA8FAACA43B05F7F965" ID-GBIF-Occurrence="1455384570" collectingDate="2002-11-20" collectionCode="OMOC" country="Ghana" latitude="10.783334" location="Bolgatanga" longLatPrecision="1297" longitude="-0.083333336" pageId="59" pageNumber="60" specimenCount="8" specimenCount-female="2" specimenCount-male="2" stateProvince="UPPER EAST" typeStatus="paratype">
|
||
<emphasis id="8784EAB30744FFA8FAACA43B05FBF941" bold="true" box="[1324,1428,1650,1675]" pageId="59" pageNumber="60">
|
||
<collectingCountry id="CDE776310744FFA8FAACA43B05FBF941" box="[1324,1428,1650,1675]" name="Ghana" pageId="59" pageNumber="60">GHANA</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<collectingRegion id="7734F8430744FFA8FF17A4D1012AF965" box="[151,325,1687,1711]" country="Ghana" name="Upper East" pageId="59" pageNumber="60">
|
||
UPPER
|
||
<collectingRegion id="7734F8430744FFA8FF7EA4DE012AF965" box="[254,325,1687,1711]" country="Cameroon" name="East" pageId="59" pageNumber="60">EAST</collectingRegion>
|
||
</collectingRegion>
|
||
,
|
||
<location id="B02F607A0744FFA8FED9A4DF01B6F965" LSID="urn:lsid:plazi:treatment:3D5987B70745FFAFFF17A15602D4FF3D:B02F607A0744FFA8FED9A4DF01B6F965" box="[345,473,1686,1711]" country="Ghana" latitude="10.783334" longLatPrecision="1297" longitude="-0.083333336" name="Bolgatanga" pageId="59" pageNumber="60" stateProvince="UPPER EAST">Bolgatanga</location>
|
||
(
|
||
<geoCoordinate id="D0C450660744FFA8FE72A4DE023EF965" box="[498,593,1687,1711]" direction="north" orientation="latitude" pageId="59" pageNumber="60" precision="925" value="10.783334">10°47'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D0C450660744FFA8FDE6A4DF02DFF965" box="[614,688,1686,1711]" direction="west" orientation="longitude" pageId="59" pageNumber="60" precision="925" value="-0.083333336">0°5'W</geoCoordinate>
|
||
),
|
||
<date id="C14E10610744FFA8FD4DA4DF0327F965" box="[717,840,1686,1711]" pageId="59" pageNumber="60" value="2002-11-20">
|
||
<collectingDate id="D10AE9890744FFA8FD4DA4DF0327F965" box="[717,840,1686,1711]" pageId="59" pageNumber="60" value="2002-11-20">20.xi.2002</collectingDate>
|
||
</date>
|
||
, Bowku—
|
||
<specimenCount id="A3F6FD280744FFA8FC4AA4DF047DF965" box="[970,1042,1686,1711]" pageId="59" pageNumber="60" type="female">2 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="A3F6FD280744FFA8FBA9A4DE041EF965" box="[1065,1137,1687,1711]" pageId="59" pageNumber="60" type="male">2 ♂♂</specimenCount>
|
||
(
|
||
<specimenCount id="A3F6FD280744FFA8FB09A4DE0579F965" box="[1161,1302,1687,1711]" pageId="59" pageNumber="60" type="generic" typeStatus="paratypes">4 paratypes</specimenCount>
|
||
) (
|
||
<collectionCode id="D3E1AE640744FFA8FAB6A4DE05FFF965" box="[1334,1424,1687,1711]" pageId="59" pageNumber="60">OMOC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="05983CFC0744FFA8FF17A4F304BAF919" ID-GBIF-Occurrence="1455384591" box="[151,1237,1722,1747]" collectingDate="1962-05-04" collectionCode="CNC" country="Nigeria" latitude="8.8566675" location="Bussa" longLatPrecision="21" longitude="5.2191668" pageId="59" pageNumber="60" specimenCount="1" specimenCount-male="1" stateProvince="KWARA" typeStatus="paratype">
|
||
<emphasis id="8784EAB30744FFA8FF17A4F3017DF919" bold="true" box="[151,274,1722,1747]" pageId="59" pageNumber="60">
|
||
<collectingCountry id="CDE776310744FFA8FF17A4F3017DF919" box="[151,274,1722,1747]" name="Nigeria" pageId="59" pageNumber="60">NIGERIA</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<collectingRegion id="7734F8430744FFA8FEA0A4F201E5F919" box="[288,394,1723,1747]" country="Nigeria" name="Kwara" pageId="59" pageNumber="60">KWARA</collectingRegion>
|
||
,
|
||
<location id="B02F607A0744FFA8FE15A4F201B7F918" LSID="urn:lsid:plazi:treatment:3D5987B70745FFAFFF17A15602D4FF3D:B02F607A0744FFA8FE15A4F201B7F918" box="[405,472,1723,1746]" country="Nigeria" latitude="8.8566675" longLatPrecision="21" longitude="5.2191668" name="Bussa" pageId="59" pageNumber="60" stateProvince="KWARA">Bussa</location>
|
||
(
|
||
<geoCoordinate id="D0C450660744FFA8FE68A4F30230F919" box="[488,607,1722,1747]" direction="north" orientation="latitude" pageId="59" pageNumber="60" precision="15" value="8.8566675">8°51'24''N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D0C450660744FFA8FDEBA4F302BFF919" box="[619,720,1722,1747]" direction="east" orientation="longitude" pageId="59" pageNumber="60" precision="15" value="5.2191668">5°13'9''E</geoCoordinate>
|
||
),
|
||
<date id="C14E10610744FFA8FD65A4F30327F919" box="[741,840,1722,1747]" pageId="59" pageNumber="60" value="1962-05-04">
|
||
<collectingDate id="D10AE9890744FFA8FD65A4F30327F919" box="[741,840,1722,1747]" pageId="59" pageNumber="60" value="1962-05-04">4.v.1962</collectingDate>
|
||
</date>
|
||
, D.C. Eidt—
|
||
<specimenCount id="A3F6FD280744FFA8FC60A4F3046BF918" box="[992,1028,1722,1747]" pageId="59" pageNumber="60" type="male">1 ♂</specimenCount>
|
||
(
|
||
<typeStatus id="6A4B88030744FFA8FB93A4F20418F919" box="[1043,1143,1723,1747]" pageId="59" pageNumber="60" type="paratype">paratype</typeStatus>
|
||
) (
|
||
<collectionCode id="D3E1AE640744FFA8FB0DA4F204A2F919" box="[1165,1229,1723,1747]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="59" pageNumber="60">CNC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FDEA652A0744FFA8FF47A49701DDF861" pageId="59" pageNumber="60" type="etymology">
|
||
<paragraph id="B54F36A10744FFA8FF47A49703BDF8D1" blockId="59.[151,1437,1146,1963]" pageId="59" pageNumber="60">
|
||
<emphasis id="8784EAB30744FFA8FF47A4970129F93D" bold="true" box="[199,326,1758,1783]" pageId="59" pageNumber="60">Etymology</emphasis>
|
||
.
|
||
<taxonomicName id="72F04D220744FFA8FED4A49601D2F93D" box="[340,445,1759,1783]" class="Insecta" family="Scarabaeidae" genus="Digitonthophagus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="107" phylum="Arthropoda" rank="species" species="dilatatus">
|
||
<emphasis id="8784EAB30744FFA8FED4A49601D2F93D" box="[340,445,1759,1783]" italics="true" pageId="59" pageNumber="60">Dilatatus</emphasis>
|
||
</taxonomicName>
|
||
is a Latin adjective pertaining to the development of the anterior pronotal angles of welldeveloped males of this species. This development is unique in the genus.
|
||
</paragraph>
|
||
<paragraph id="B54F36A10744FFA8FF47A56F01DDF861" blockId="59.[151,1437,1146,1963]" pageId="59" pageNumber="60">
|
||
<emphasis id="8784EAB30744FFA8FF47A56F01EFF8F5" bold="true" box="[199,384,1830,1855]" pageId="59" pageNumber="60">Natural history</emphasis>
|
||
. The known distribution pattern of this species suggests that it is endemic to the Sudanese savanna. Interestingly, all specimens with data were collected at light or in pitfall traps baited with human dung, no specimens were collected in ruminant dung. Additional observations are required to confirm that it is a nonruminant dung specialist.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FDEA652A0743FFAFFF47A2DE02D4FF3D" pageId="60" pageNumber="61" type="discussion">
|
||
<paragraph id="B54F36A10743FFAFFF47A2DE02D4FF3D" blockId="60.[151,1436,151,248]" pageId="60" pageNumber="61">
|
||
<emphasis id="8784EAB30743FFAFFF47A2DE015BFF7A" bold="true" box="[199,308,151,176]" pageId="60" pageNumber="61">Remarks</emphasis>
|
||
. Externally, this species can be confused with
|
||
<taxonomicName id="72F04D220743FFAFFCBEA2D003DDFF7A" box="[830,946,152,176]" class="Insecta" family="Scarabaeidae" genus="Digitonthophagus" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="fimator">
|
||
<emphasis id="8784EAB30743FFAFFCBEA2D003DDFF7A" box="[830,946,152,176]" italics="true" pageId="60" pageNumber="61">D. fimator</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingRegion id="7734F8430743FFAFFC56A2D10462FF65" box="[982,1037,152,175]" country="Cameroon" name="West" pageId="60" pageNumber="61">West</collectingRegion>
|
||
Africa. However, in addition to the characters given, the parameres are smaller and approximately parallel sided in lateral view as opposed to being wider and divergent in lateral view in
|
||
<taxonomicName id="72F04D220743FFAFFDC1A2A802DBFF32" box="[577,692,224,248]" class="Insecta" family="Scarabaeidae" genus="Digitonthophagus" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="fimator">
|
||
<emphasis id="8784EAB30743FFAFFDC1A2A802DBFF32" box="[577,692,224,248]" italics="true" pageId="60" pageNumber="61">D. fimator</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |