treatments-xml/data/25/2C/87/252C87918B362C05FF20F8C5BFCB3D4E.xml

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<mods:title id="F9C1A01FD9C81625AC0829C1C4F32EF8">Descriptions of two new shell-dwelling species of Metriaclima (Cichlidae) from Lake Malaŵi, Africa</mods:title>
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<mods:namePart id="FCE03DF1C052489D147066ED7922DFA2">Miller, Torin A.</mods:namePart>
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<mods:affiliation id="A97143E7C449F8095E1A7C100551E6ED">Cichlid Press, PO Box 13608, El Paso, TX 79913.</mods:affiliation>
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<mods:affiliation id="AC0BC52946D6C1E20489CDDFF1AAB2F8">Penn State University, 432 Forest Resources Building, University Park, Pennsylvania 16802. &amp; Honorary Research Associate, South African Institute for Aquatic Biodiversity, Makhanda, RSA.</mods:affiliation>
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<heading id="F67281EB8B362C09FF20F8C5B96D3848" bold="true" box="[151,584,1869,1896]" fontSize="11" level="1" pageId="6" pageNumber="425" reason="1">
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<taxonomicName id="6A854D048B362C09FF20F8C5BA8F3847" authority="Miller &amp; Konings &amp; Stauffer, 2021" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[151,426,1869,1896]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="gallireyae" status="sp. nov.">
<emphasis id="9FF1EA958B362C09FF20F8C5BA8F3847" bold="true" box="[151,426,1869,1896]" italics="true" pageId="6" pageNumber="425">Metriaclima gallireyae</emphasis>
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, 
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<paragraph id="AD3A36878B362C09FF20F8F9BA8138A3" blockId="6.[151,584,1869,1932]" box="[151,420,1905,1932]" pageId="6" pageNumber="425">
<figureCitation id="35BE2A028B362C09FF20F8F9BA2738A3" box="[151,258,1905,1932]" captionStart="FIGURE 5" captionStartId="7.[151,250,1469,1493]" captionTargetBox="[163,1374,777,1446]" captionTargetId="figure-272@7.[151,1436,770,1446]" captionTargetPageId="7" captionText="FIGURE 5. Metriaclima gallireyae. Holotype, PSU 13370, adult male, 54.4 mm SL, Gallireya Reef, Lake Malaŵi." figureDoi="http://doi.org/10.5281/zenodo.5572228" httpUri="https://zenodo.org/record/5572228/files/figure.png" pageId="6" pageNumber="425">Figures 5</figureCitation>
, 
<figureCitation id="35BE2A028B362C09FEB9F8F9BA3B38A4" box="[270,286,1905,1931]" captionStart="FIGURE 6" captionStartId="8.[151,250,1439,1463]" captionTargetBox="[196,1392,489,1415]" captionTargetId="figure-166@8.[196,1392,489,1415]" captionTargetPageId="8" captionText="FIGURE 6. CT scan of the cranium of Metriaclima gallireyae. Holotype, PSU 13370, adult male, 54.4 mm SL, Gallireya Reef, Lake Malaŵi; angle of ethmovomerine block is 49.6° with parasphenoid." figureDoi="http://doi.org/10.5281/zenodo.5572230" httpUri="https://zenodo.org/record/5572230/files/figure.png" pageId="6" pageNumber="425">6</figureCitation>
, 
<figureCitation id="35BE2A028B362C09FE9DF8FABA1F38A3" box="[298,314,1906,1932]" captionStart="FIGURE 7" captionStartId="10.[151,250,1571,1595]" captionTargetBox="[176,1412,633,1548]" captionTargetId="figure-225@10.[176,1412,633,1548]" captionTargetPageId="10" captionText="FIGURE 7. A male Metriaclima gallireyae at Gallireya Reef (10.500153 S, 34.233362 E), Lake Malaŵi." figureDoi="http://doi.org/10.5281/zenodo.5572232" httpUri="https://zenodo.org/record/5572232/files/figure.png" pageId="6" pageNumber="425">7</figureCitation>
; 
<tableCitation id="E007033C8B362C09FEF1F8F9BA8138A3" box="[326,420,1905,1932]" captionStart="TABLE 2" captionStartId="9.[151,239,151,175]" captionTargetBox="[166,1350,228,2023]" captionTargetPageId="9" captionText="TABLE 2. Morphological and meristic data for Metriaclima gallireyae from Gallireya Reef, Lake Malaŵi, PSU 13370, 13371, n=20. Ranges include holotype." pageId="6" pageNumber="425">Tables 2</tableCitation>
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<paragraph id="AD3A36878B362C09FF20F83EB9BA38E1" blockId="6.[151,678,1974,2029]" box="[151,671,1974,1998]" pageId="6" pageNumber="425">
<taxonomicName id="6A854D048B362C09FF20F83EBA8E38E1" authorityName="Burgess" authorityYear="1976" box="[151,427,1974,1998]" class="Actinopterygii" family="Cichlidae" genus="Pseudotropheus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="lanisticola">
<emphasis id="9FF1EA958B362C09FF20F83EBA8E38E1" box="[151,427,1974,1998]" italics="true" pageId="6" pageNumber="425">Pseudotropheus lanisticola</emphasis>
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(in part), 
<bibRefCitation id="C9144B768B362C09FDA7F83EB9BA38E1" author="Konings, A. F." box="[528,671,1974,1998]" pageId="6" pageNumber="425" refId="ref6746" refString="Konings, A. F. (1989) Malawi cichlids in their natural habitat. Verduijn Cichlids &amp; Lake Fish Movies, Zevenhuizen, 304 pp." type="book" year="1989">Konings 1989</bibRefCitation>
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<taxonomicName id="6A854D048B362C09FF20F85DBA9738C2" baseAuthorityName="Boulenger" baseAuthorityYear="1899" box="[151,434,2005,2029]" class="Actinopterygii" family="Cichlidae" genus="Pseudotropheus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="livingstonii">
<emphasis id="9FF1EA958B362C09FF20F85DBA9738C2" box="[151,434,2005,2029]" italics="true" pageId="6" pageNumber="425">Pseudotropheus livingstonii</emphasis>
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(in part), 
<bibRefCitation id="C9144B768B362C09FDA0F85DB98338C3" author="Konings, A. F." box="[535,678,2004,2029]" pageId="6" pageNumber="425" refId="ref6774" refString="Konings, A. F. (1995) Malaw ˆ i cichlids in their natural habitat. 2 nd Edition. Cichlid Press, El Paso, Texas, 303 pp." type="book" year="1995">Konings 1995</bibRefCitation>
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<taxonomicName id="6A854D048B372C08FF20FF1FBAA83F80" baseAuthorityName="Boulenger" baseAuthorityYear="1899" box="[151,397,151,175]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="livingstonii">
<emphasis id="9FF1EA958B372C08FF20FF1FBAA83F80" box="[151,397,151,175]" italics="true" pageId="7" pageNumber="426">Metriaclima livingstonii</emphasis>
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(in part), 
<bibRefCitation id="C9144B768B372C08FE45FF1FB9A43F80" author="Konings, A." box="[498,641,151,175]" pageId="7" pageNumber="426" refId="ref6807" refString="Konings, A. (2001) Malawi cichlids in their natural habitat. 3 rd Edition. Cichlid Press, El Paso, Texas, 352 pp." type="book" year="2001">Konings 2001</bibRefCitation>
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<taxonomicName id="6A854D048B372C08FF20FF3DBA1D3FE2" box="[151,312,181,205]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="9FF1EA958B372C08FF20FF3DBA313FE2" box="[151,276,181,205]" italics="true" pageId="7" pageNumber="426">Metriaclima</emphasis>
sp.
</taxonomicName>
‘lanisticola north’ 
<bibRefCitation id="C9144B768B372C08FE40FF3DB9A33FE2" author="Konings, A." box="[503,646,181,205]" pageId="7" pageNumber="426" refId="ref6836" refString="Konings, A. (2007) Malawi cichlids in their natural habitat. 4 th Edition. Cichlid Press, El Paso, Texas, 424 pp." type="book" year="2007">Konings 2007</bibRefCitation>
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<typeStatus id="723E88258B372C08FF20FF74BA233E39" box="[151,262,252,278]" pageId="7" pageNumber="426" type="holotype">Holotype</typeStatus>
.
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, 
<specimenCount id="BB83FD0E8B372C08FE2CFF73B9343E3A" box="[411,529,251,277]" pageId="7" pageNumber="426" type="adult">adult male</specimenCount>
, 
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SL, (
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<location id="A85A605C8B372C08FC45FF73BFBD3E3A" LSID="urn:lsid:plazi:treatment:252C87918B362C05FF20F8C5BFCB3D4E:A85A605C8B372C08FC45FF73BFBD3E3A" box="[1010,1176,251,277]" country="Malawi" county="Africa" latitude="-10.500153" longLatPrecision="1" longitude="34.23336" municipality="Lake" name="Gallireya Reef" pageId="7" pageNumber="426">Gallireya Reef</location>
, 
<location id="A85A605C8B372C08FB16FF73BE133E3A" LSID="urn:lsid:plazi:treatment:252C87918B362C05FF20F8C5BFCB3D4E:A85A605C8B372C08FB16FF73BE133E3A" box="[1185,1334,251,277]" country="Malawi" county="Africa" latitude="-10.500153" longLatPrecision="1" longitude="34.23336" municipality="Lake" name="Lake Malawi" pageId="7" pageNumber="426">Lake Malaŵi</location>
, 
<collectingCountry id="D59276178B372C08FAF7FF73BEBD3E3A" box="[1344,1432,251,277]" name="Malawi" pageId="7" pageNumber="426">Malaŵi</collectingCountry>
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<collectingDate id="C97FE9AF8B372C08FF5AFE97BA5B3E15" box="[237,382,287,314]" pageId="7" pageNumber="426" value="2008-01-15">15 Jan. 2008</collectingDate>
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, 
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&amp; 
<collectorName id="007053518B372C08FDFFFEA8B9D93E15" box="[584,764,287,314]" pageId="7" pageNumber="426">J. R. Stauffer Jr.</collectorName>
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<paragraph id="AD3A36878B372C08FF70FECCB8F03E72" blockId="7.[151,1437,251,746]" box="[199,981,323,350]" pageId="7" pageNumber="426">
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<emphasis id="9FF1EA958B372C08FF70FECCBA633E71" bold="true" box="[199,326,324,350]" pageId="7" pageNumber="426">
<typeStatus id="723E88258B372C08FF70FECCBA643E71" box="[199,321,324,350]" pageId="7" pageNumber="426" type="paratype">Paratypes</typeStatus>
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<specimenCode id="FD239EFC8B372C08FEFAFECCBAF53E71" box="[333,464,324,350]" pageId="7" pageNumber="426">PSU 13371</specimenCode>
, 20, (
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SL), data as for holotype
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.
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<subSubSection id="E59F650C8B372C08FF70FEEFBAE83DC6" pageId="7" pageNumber="426" type="diagnosis">
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<emphasis id="9FF1EA958B372C08FF70FEEFBA673EAE" bold="true" box="[199,322,359,385]" pageId="7" pageNumber="426">Diagnosis.</emphasis>
The moderately sloped vomer (49.6º in 
<typeStatus id="723E88258B372C08FCA5FEEFB8533EAD" box="[786,886,359,386]" pageId="7" pageNumber="426" type="holotype">holotype</typeStatus>
) with a swollen rostral tip (
<figureCitation id="35BE2A028B372C08FB01FEEFBFDE3EAD" box="[1206,1275,359,386]" captionStart="FIGURE 6" captionStartId="8.[151,250,1439,1463]" captionTargetBox="[196,1392,489,1415]" captionTargetId="figure-166@8.[196,1392,489,1415]" captionTargetPageId="8" captionText="FIGURE 6. CT scan of the cranium of Metriaclima gallireyae. Holotype, PSU 13370, adult male, 54.4 mm SL, Gallireya Reef, Lake Malaŵi; angle of ethmovomerine block is 49.6° with parasphenoid." figureDoi="http://doi.org/10.5281/zenodo.5572230" httpUri="https://zenodo.org/record/5572230/files/figure.png" pageId="7" pageNumber="426">Fig. 6</figureCitation>
) and bicuspid teeth in the anterior portion of the outer row of both upper and lower jaws place this species in 
<taxonomicName id="6A854D048B372C08FB6FFE03BE473E8A" authorityName="Stauffer, Bowers, Kellogg &amp; McKaye" authorityYear="1997" box="[1240,1378,395,421]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="426" phylum="Chordata" rank="genus">
<emphasis id="9FF1EA958B372C08FB6FFE03BE473E8A" box="[1240,1378,395,421]" italics="true" pageId="7" pageNumber="426">Metriaclima</emphasis>
</taxonomicName>
. The presence of a black submarginal band in the dorsal and anal fin, and 4–5 bars below the dorsal distinguishes 
<taxonomicName id="6A854D048B372C08FAC9FE38BA2D3EC2" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B372C08FAC9FE38BA2D3EC2" italics="true" pageId="7" pageNumber="426">M. gallireyae</emphasis>
</taxonomicName>
from most other species in 
<taxonomicName id="6A854D048B372C08FDFDFE5BB9F13EC2" authorityName="Stauffer, Bowers, Kellogg &amp; McKaye" authorityYear="1997" box="[586,724,467,493]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="426" phylum="Chordata" rank="genus">
<emphasis id="9FF1EA958B372C08FDFDFE5BB9F13EC2" box="[586,724,467,493]" italics="true" pageId="7" pageNumber="426">Metriaclima</emphasis>
</taxonomicName>
, except for 
<emphasis id="9FF1EA958B372C08FCE8FE5CB8F83EC2" box="[863,989,468,493]" italics="true" pageId="7" pageNumber="426">
<taxonomicName id="6A854D048B372C08FCE8FE5CB8FF3EC2" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[863,986,468,493]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="ngarae">M. ngarae</taxonomicName>
.
</emphasis>
It differs from 
<taxonomicName id="6A854D048B372C08FB27FE5CBE223EC2" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[1168,1287,468,493]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B372C08FB27FE5CBE223EC2" box="[1168,1287,468,493]" italics="true" pageId="7" pageNumber="426">M. ngarae</emphasis>
</taxonomicName>
by a smaller interorbital width (30.7–37.4% vs. 23.6–29.7% HL) and by a smaller ratio of the snout length in the distance between snout tip and pelvic fin origin (20.6–27.8% vs. 28.7–32.8% SNP2). On average the ratio of premaxillary pedicel length in snout length is greater in 
<taxonomicName id="6A854D048B372C08FDCDFDC8B8343D76" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[634,785,575,601]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B372C08FDCDFDC8B8343D76" box="[634,785,575,601]" italics="true" pageId="7" pageNumber="426">M. gallireyae</emphasis>
</taxonomicName>
at 44.6% vs. 35.1 %SNL in 
<taxonomicName id="6A854D048B372C08FBE2FDC8BFEF3D76" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[1109,1226,576,601]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B372C08FBE2FDC8BFEF3D76" box="[1109,1226,576,601]" italics="true" pageId="7" pageNumber="426">M. ngarae</emphasis>
</taxonomicName>
(ranges 33.1–62.7 vs. 20.3–45.2 %SNL). Mature males of 
<taxonomicName id="6A854D048B372C08FDD2FDECB9D83D52" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[613,765,611,637]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B372C08FDD2FDECB9D83D52" box="[613,765,611,637]" italics="true" pageId="7" pageNumber="426">M. gallireyae</emphasis>
</taxonomicName>
have a dark brown to blue coloration on flank (
<figureCitation id="35BE2A028B372C08FA9EFDEBBE4A3D51" box="[1321,1391,611,638]" captionStart="FIGURE 7" captionStartId="10.[151,250,1571,1595]" captionTargetBox="[176,1412,633,1548]" captionTargetId="figure-225@10.[176,1412,633,1548]" captionTargetPageId="10" captionText="FIGURE 7. A male Metriaclima gallireyae at Gallireya Reef (10.500153 S, 34.233362 E), Lake Malaŵi." figureDoi="http://doi.org/10.5281/zenodo.5572232" httpUri="https://zenodo.org/record/5572232/files/figure.png" pageId="7" pageNumber="426">Fig. 7</figureCitation>
) on which no bar pattern is visible. Male 
<taxonomicName id="6A854D048B372C08FD87FD00B9863D8E" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[560,675,648,673]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B372C08FD87FD00B9863D8E" box="[560,675,648,673]" italics="true" pageId="7" pageNumber="426">M. ngarae</emphasis>
</taxonomicName>
resemble females and display 4–5 dark bars on a light-colored flank. Female 
<taxonomicName id="6A854D048B372C08FF47FD24BAA13DEA" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[240,388,683,709]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B372C08FF47FD24BAA13DEA" box="[240,388,683,709]" italics="true" pageId="7" pageNumber="426">M. gallireyae</emphasis>
</taxonomicName>
have a similar color pattern to those of 
<taxonomicName id="6A854D048B372C08FC8FFD24B88E3DEA" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[824,939,684,709]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B372C08FC8FFD24B88E3DEA" box="[824,939,684,709]" italics="true" pageId="7" pageNumber="426">M. ngarae</emphasis>
</taxonomicName>
but the bars on the flank are more prominent and extend to the abdomen.
</paragraph>
</subSubSection>
<caption id="F9FA660F8B372C08FF20FA35BE0E3AFA" ID-DOI="http://doi.org/10.5281/zenodo.5572228" ID-Zenodo-Dep="5572228" box="[151,1323,1469,1494]" httpUri="https://zenodo.org/record/5572228/files/figure.png" pageId="7" pageNumber="426" startId="7.[151,250,1469,1493]" targetBox="[163,1374,777,1446]" targetPageId="7">
<paragraph id="AD3A36878B372C08FF20FA35BE0E3AFA" blockId="7.[151,1323,1469,1494]" box="[151,1323,1469,1494]" pageId="7" pageNumber="426">
<emphasis id="9FF1EA958B372C08FF20FA35BA363AF9" bold="true" box="[151,275,1469,1494]" pageId="7" pageNumber="426">FIGURE 5.</emphasis>
<taxonomicName id="6A854D048B372C08FEAEFA35B9263AFA" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[281,515,1469,1493]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B372C08FEAEFA35B9263AFA" box="[281,515,1469,1493]" italics="true" pageId="7" pageNumber="426">Metriaclima gallireyae</emphasis>
</taxonomicName>
. Holotype, PSU 13370, adult male, 54.4 mm SL, Gallireya Reef, Lake Malaŵi.
</paragraph>
</caption>
<subSubSection id="E59F650C8B372C07FF70F98BBA9B3EFE" lastPageId="8" lastPageNumber="427" pageId="7" pageNumber="426" type="description">
<paragraph id="AD3A36878B372C08FF70F98BB8D839D9" blockId="7.[151,1437,1539,2034]" pageId="7" pageNumber="426">
<emphasis id="9FF1EA958B372C08FF70F98BBA7C3932" bold="true" box="[199,345,1539,1565]" pageId="7" pageNumber="426">Description.</emphasis>
Morphometric and meristic data in 
<tableCitation id="E007033C8B372C08FD5BF98CB8673931" box="[748,834,1540,1566]" captionStart="TABLE 2" captionStartId="9.[151,239,151,175]" captionTargetBox="[166,1350,228,2023]" captionTargetPageId="9" captionText="TABLE 2. Morphological and meristic data for Metriaclima gallireyae from Gallireya Reef, Lake Malaŵi, PSU 13370, 13371, n=20. Ranges include holotype." pageId="7" pageNumber="426">Table 2</tableCitation>
. Small mbuna with ovoid body (mean BD 38.6% SL) with greatest depth between the 3 
<superScript id="5AF09BCF8B372C08FDB1F9AFB931391A" attach="left" box="[518,532,1575,1589]" fontSize="6" pageId="7" pageNumber="426">rd</superScript>
and 4 
<superScript id="5AF09BCF8B372C08FDE3F9AFB944391A" attach="left" box="[596,609,1575,1589]" fontSize="6" pageId="7" pageNumber="426">th</superScript>
dorsal spine. Dorsal body profile with gradual curve downward posteriorly (more pronounced than in 
<taxonomicName id="6A854D048B372C08FE74F9C4B91D394A" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[451,568,1612,1637]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B372C08FE74F9C4B91D394A" box="[451,568,1612,1637]" italics="true" pageId="7" pageNumber="426">M. ngarae</emphasis>
</taxonomicName>
) with highest point between the 4 
<superScript id="5AF09BCF8B372C08FC0EF9C3B8E33976" attach="left" box="[953,966,1611,1625]" fontSize="6" pageId="7" pageNumber="426">th</superScript>
and 5 
<superScript id="5AF09BCF8B372C08FBBBF9C3BF3C3976" attach="left" box="[1036,1049,1611,1625]" fontSize="6" pageId="7" pageNumber="426">th</superScript>
spine, less deep towards posterior end of dorsal fin and beginning of caudal peduncle; ventral body profile flat between pelvic fins and origin of anal fin with slight upward angle to caudal fin. Dorsal head profile round with continuous curve between interorbital and dorsal-fin origin; short snout with isognathous to slightly retrognathous jaws; teeth in upper and lower jaws in 3–4 rows (mode 3); teeth in outer row bicuspid, inner rows unicuspid or tricuspid.
</paragraph>
<paragraph id="AD3A36878B372C07FF70F888BA093FFA" blockId="7.[151,1437,1539,2034]" lastBlockId="8.[151,1437,150,465]" lastPageId="8" lastPageNumber="427" pageId="7" pageNumber="426">
Dorsal fin with XVI or XVIII (mode XVII) spines and 7–9 (mode 8) rays. Anal fin with III spines and 7–8 rays. First 5–6 dorsal-fin spines gradually longer posteriorly with sixth spine about twice as long as first spine; posterior 13 spines slightly longer posteriorly with last spine longest; soft dorsal fin with subacuminate tip, 2nd or 3rd ray longest, extending to approximately 1/3–1/2 length of caudal fin. Anal-fin spines progressively longer posteriorly; 3 
<superScript id="5AF09BCF8B372C08FF12F807BB9638B2" attach="left" box="[165,179,1935,1949]" fontSize="6" pageId="7" pageNumber="426">rd</superScript>
or 4 
<superScript id="5AF09BCF8B372C08FF5DF807BBD238B2" attach="left" box="[234,247,1935,1949]" fontSize="6" pageId="7" pageNumber="426">th</superScript>
ray longest, extending to halfway caudal fin, equal to or slightly further posterior than tip of dorsal fin. Caudal fin subtruncate to emarginate. Pelvic fin reaching to first anal-fin spine. Pectoral fin rounded, paddle-shaped, short, reaching vertical through base of 10th or 11th dorsal-fin spine. Flank scales ctenoid with small scales on breast and belly; 29–32 (mode 30) lateral-line scales, cheek with 4–5 rows of small scales. Small scales over proximal ¼ of caudal fin.
</paragraph>
<paragraph id="AD3A36878B382C07FF70FF57BA9B3EFE" blockId="8.[151,1437,150,465]" pageId="8" pageNumber="427">
Head of male brown with dark brown interorbital bar; cheek and preopercle with blue highlights; opercle with blue and green highlights and black spot; throat brown (
<figureCitation id="35BE2A028B382C07FC96FE8BB84D3E32" box="[801,872,259,285]" captionStart="FIGURE 7" captionStartId="10.[151,250,1571,1595]" captionTargetBox="[176,1412,633,1548]" captionTargetId="figure-225@10.[176,1412,633,1548]" captionTargetPageId="10" captionText="FIGURE 7. A male Metriaclima gallireyae at Gallireya Reef (10.500153 S, 34.233362 E), Lake Malaŵi." figureDoi="http://doi.org/10.5281/zenodo.5572232" httpUri="https://zenodo.org/record/5572232/files/figure.png" pageId="8" pageNumber="427">Fig. 7</figureCitation>
). Flank dark brown to blue (in territorial males), center of scales brown with blue outline; 4–5 brown bars, obscured in fully mature males; breast and belly black. Dorsal fin brown with black submarginal bar; orange and brown lappets, with orange pigmentation in membranes of rayed portion. Caudal fin with black dorsal and ventral margin; brown rays; blue membranes. Anal fin with proximal ¼ gray, distal ¾ black; one yellow ocellus. Pelvic fin black with white leading edge. Pectoral fin with clear membranes and gray rays.
</paragraph>
</subSubSection>
<caption id="F9FA660F8B382C07FF20FA17B8583AF4" ID-DOI="http://doi.org/10.5281/zenodo.5572230" ID-Zenodo-Dep="5572230" httpUri="https://zenodo.org/record/5572230/files/figure.png" pageId="8" pageNumber="427" startId="8.[151,250,1439,1463]" targetBox="[196,1392,489,1415]" targetPageId="8">
<paragraph id="AD3A36878B382C07FF20FA17B8583AF4" blockId="8.[151,1437,1439,1499]" pageId="8" pageNumber="427">
<emphasis id="9FF1EA958B382C07FF20FA17BA373A98" bold="true" box="[151,274,1439,1463]" pageId="8" pageNumber="427">FIGURE 6.</emphasis>
CT scan of the cranium of 
<taxonomicName id="6A854D048B382C07FD93FA17B8283A98" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[548,781,1439,1463]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B382C07FD93FA17B8283A98" box="[548,781,1439,1463]" italics="true" pageId="8" pageNumber="427">Metriaclima gallireyae</emphasis>
</taxonomicName>
. Holotype, PSU 13370, adult male, 54.4 mm SL, Gallireya Reef, Lake Malaŵi; angle of ethmovomerine block is 49.6° with parasphenoid.
</paragraph>
</caption>
<subSubSection id="E59F650C8B382C05FF70F981BA603FD6" lastPageId="10" lastPageNumber="429" pageId="8" pageNumber="427" type="distribution">
<paragraph id="AD3A36878B382C07FF70F981B8E03968" blockId="8.[151,1437,1545,2039]" pageId="8" pageNumber="427">
<emphasis id="9FF1EA958B382C07FF70F981BA44390C" bold="true" box="[199,353,1545,1571]" pageId="8" pageNumber="427">Distribution.</emphasis>
<taxonomicName id="6A854D048B382C07FED0F981B94D390C" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[359,616,1545,1571]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B382C07FED0F981B94D390C" box="[359,616,1545,1571]" italics="true" pageId="8" pageNumber="427">Metriaclima gallireyae</emphasis>
</taxonomicName>
is found on Gallireya Reef (
<geoCoordinate id="C8B150408B382C07FC13F981BF10390C" box="[932,1077,1545,1572]" degrees="10.500153" direction="south" orientation="latitude" pageId="8" pageNumber="427" precision="1" value="-10.500153">10.500153 S</geoCoordinate>
, 
<geoCoordinate id="C8B150408B382C07FBF6F981BFF7390B" box="[1089,1234,1545,1572]" degrees="34.233362" direction="east" orientation="longitude" pageId="8" pageNumber="427" precision="1" value="34.23336">34.233362 E</geoCoordinate>
), Youngs Bay and in Chitimba Bay (
<geoCoordinate id="C8B150408B382C07FEE8F9A5BACA3968" box="[351,495,1581,1608]" degrees="10.581122" direction="south" orientation="latitude" pageId="8" pageNumber="427" precision="1" value="-10.581122">10.581122 S</geoCoordinate>
, 34.23692 Lake 
<collectingCountry id="D59276178B382C07FD1DF9A5B8273968" box="[682,770,1581,1607]" name="Malawi" pageId="8" pageNumber="427">Malaŵi</collectingCountry>
, 
<collectingCountry id="D59276178B382C07FCB9F9A5B8413968" box="[782,868,1581,1607]" name="Malawi" pageId="8" pageNumber="427">Malaŵi</collectingCountry>
(
<figureCitation id="35BE2A028B382C07FCC4F9A5B8913968" box="[883,948,1581,1607]" captionStart="FIGURE 1" captionStartId="2.[151,250,1776,1800]" captionTargetBox="[161,1425,463,1748]" captionTargetId="figure-160@2.[151,1436,453,1753]" captionTargetPageId="2" captionText="FIGURE 1. Map of Lake Malaŵi with the localities marked that are mentioned in the text." figureDoi="http://doi.org/10.5281/zenodo.5572220" httpUri="https://zenodo.org/record/5572220/files/figure.png" pageId="8" pageNumber="427">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="AD3A36878B382C07FF70F9D9BE69386C" blockId="8.[151,1437,1545,2039]" pageId="8" pageNumber="427">
<emphasis id="9FF1EA958B382C07FF70F9D9BA8F3944" bold="true" box="[199,426,1617,1643]" pageId="8" pageNumber="427">Field observations.</emphasis>
The population density of 
<taxonomicName id="6A854D048B382C07FD68F9D9B8C43944" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[735,993,1617,1643]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B382C07FD68F9D9B8C43944" box="[735,993,1617,1643]" italics="true" pageId="8" pageNumber="427">Metriaclima gallireyae</emphasis>
</taxonomicName>
is much higher than that of 
<taxonomicName id="6A854D048B382C07FA91F9DABEB93944" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[1318,1436,1618,1643]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B382C07FA91F9DABEB93944" box="[1318,1436,1618,1643]" italics="true" pageId="8" pageNumber="427">M. ngarae</emphasis>
</taxonomicName>
and is often seen in large groups of up to 20 individuals in the vicinity of empty shells of 
<taxonomicName id="6A854D048B382C07FB3BF9FEBE3B39A0" authorityName="Dohrn" authorityYear="1865" box="[1164,1310,1654,1679]" class="Gastropoda" family="Ampullariidae" genus="Lanistes" kingdom="Animalia" order="Architaenioglossa" pageId="8" pageNumber="427" phylum="Mollusca" rank="species" species="nyassanus">
<emphasis id="9FF1EA958B382C07FB3BF9FEBE3B39A0" box="[1164,1310,1654,1679]" italics="true" pageId="8" pageNumber="427">L. nyassanus</emphasis>
</taxonomicName>
. Both sites where this small species was observed contained either many football-sized (lava?) rocks (Gallireya Reef) or large slabs of sandstone (Chitimba Bay) that may provide additional shelter to individuals that were not able to secure an empty shell for protection. The depth range at Gallireya Reef is about 
<quantity id="6A7D9B628B382C07FC47F969BF6D39D3" box="[1008,1096,1761,1788]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="8" pageNumber="427" unit="m" value="10.0" valueMax="12.0" valueMin="8.0">8–12 m</quantity>
while the reef (almost solely consisting of sandstone slabs) in Chitimba Bay is at a depth of about 
<quantity id="6A7D9B628B382C07FC19F88DBF30380F" box="[942,1045,1797,1824]" metricMagnitude="1" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="8" pageNumber="427" unit="m" value="22.5" valueMax="25.0" valueMin="20.0">
20– 
<specimenCount id="BB83FD0E8B382C07FC6EF88DBF30380F" box="[985,1045,1797,1824]" pageId="8" pageNumber="427" type="generic">25 m</specimenCount>
</quantity>
. Pure sandy habitats in the area of these two reefs were not visited other than near shore (shallower than 
<quantity id="6A7D9B628B382C07FC10F8A1B8F6386B" box="[935,979,1833,1860]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="427" unit="m" value="5.0">5 m</quantity>
) where 
<taxonomicName id="6A854D048B382C07FB9AF8A2BFE6386C" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[1069,1219,1833,1859]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B382C07FB9AF8A2BFE6386C" box="[1069,1219,1833,1859]" italics="true" pageId="8" pageNumber="427">M. gallireyae</emphasis>
</taxonomicName>
was absent.
</paragraph>
<paragraph id="AD3A36878B382C07FF70F8C6BA1238D8" blockId="8.[151,1437,1545,2039]" pageId="8" pageNumber="427">
No stomach analyses were performed but individuals were observed picking from the rocks and slabs, particularly from patches that showed less sediment cover. Repeated application of the fully opened mouth to a sediment-free spot on a rock suggests that it combs the algal matrix for loose strands of (blue-green) algae. Females and immatures were often seen in small foraging groups mainly feeding from plankton in the water column close (&lt;
<quantity id="6A7D9B628B382C07FABCF831BE6938FC" box="[1291,1356,1977,2003]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="427" unit="cm" value="20.0">20cm</quantity>
) to the sandy bottom.
</paragraph>
<caption id="F9FA660F8B392C06FF20FF1FB91E3FFC" pageId="9" pageNumber="428" startId="9.[151,239,151,175]" targetBox="[166,1350,228,2023]" targetIsTable="true" targetPageId="9">
<paragraph id="AD3A36878B392C06FF20FF1FB91E3FFC" blockId="9.[151,1436,151,212]" pageId="9" pageNumber="428">
<emphasis id="9FF1EA958B392C06FF20FF1FBA2F3F80" bold="true" box="[151,266,151,175]" pageId="9" pageNumber="428">TABLE 2.</emphasis>
Morphological and meristic data for 
<taxonomicName id="6A854D048B392C06FD2BFF1FB8B73F80" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[668,914,151,175]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B392C06FD2BFF1FB8B73F80" box="[668,914,151,175]" italics="true" pageId="9" pageNumber="428">Metriaclima gallireyae</emphasis>
</taxonomicName>
from Gallireya Reef, Lake Malaŵi, PSU 13370, 13371, n=20. Ranges include holotype.
</paragraph>
</caption>
<paragraph id="AD3A36878B392C06FF11FF6CBFDF38C8" pageId="9" pageNumber="428">
<table id="DF85C4278B39D3F0FF11FF6CBE6338C8" box="[166,1350,228,2023]" colsContinueFrom="5.[166,1351,228,2023]" gridcols="5" gridrows="45" pageId="9" pageNumber="428">
<tr id="13B534C58B39D3F0FF11FF6CBE633FD3" box="[166,1350,228,252]" gridrow="0" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FF6CB9293FD3" box="[166,524,228,252]" gridcol="0" gridrow="0" pageId="9" pageNumber="428">Variable</th>
<th id="50645DB98B39D3F0FD0EFF6CB8323FD3" box="[697,791,228,252]" gridcol="1" gridrow="0" pageId="9" pageNumber="428">Holotype</th>
<th id="50645DB98B39D3F0FCE9FF6CB8BC3FD3" box="[862,921,228,252]" gridcol="2" gridrow="0" pageId="9" pageNumber="428">Mean</th>
<th id="50645DB98B39D3F0FC57FF6CBF523FD3" box="[992,1143,228,252]" gridcol="3" gridrow="0" pageId="9" pageNumber="428">Std Dev</th>
<th id="50645DB98B39D3F0FB63FF6CBE633FD3" box="[1236,1350,228,252]" gridcol="4" gridrow="0" pageId="9" pageNumber="428">Range</th>
</tr>
<tr id="13B534C58B39D3F0FF11FE84BE633E0B" box="[166,1350,268,292]" gridrow="1" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FE84B9293E0B" box="[166,524,268,292]" gridcol="0" gridrow="1" pageId="9" pageNumber="428">Standard length (mm)</th>
<td id="50645DB98B39D3F0FD0EFE84B8323E0B" box="[697,791,268,292]" gridcol="1" gridrow="1" pageId="9" pageNumber="428">54.4</td>
<td id="50645DB98B39D3F0FCE9FE84B8BC3E0B" box="[862,921,268,292]" gridcol="2" gridrow="1" pageId="9" pageNumber="428">45.8</td>
<td id="50645DB98B39D3F0FC57FE84BF523E0B" box="[992,1143,268,292]" gridcol="3" gridrow="1" pageId="9" pageNumber="428">5.0</td>
<td id="50645DB98B39D3F0FB63FE84BE633E0B" box="[1236,1350,268,292]" gridcol="4" gridrow="1" pageId="9" pageNumber="428">36.4–54.4</td>
</tr>
<tr id="13B534C58B39D3F0FF11FEBCBE633E63" box="[166,1350,308,332]" gridrow="2" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FEBCB9293E63" box="[166,524,308,332]" gridcol="0" gridrow="2" pageId="9" pageNumber="428">Head length (mm)</th>
<td id="50645DB98B39D3F0FD0EFEBCB8323E63" box="[697,791,308,332]" gridcol="1" gridrow="2" pageId="9" pageNumber="428">17.7</td>
<td id="50645DB98B39D3F0FCE9FEBCB8BC3E63" box="[862,921,308,332]" gridcol="2" gridrow="2" pageId="9" pageNumber="428">15.1</td>
<td id="50645DB98B39D3F0FC57FEBCBF523E63" box="[992,1143,308,332]" gridcol="3" gridrow="2" pageId="9" pageNumber="428">1.4</td>
<td id="50645DB98B39D3F0FB63FEBCBE633E63" box="[1236,1350,308,332]" gridcol="4" gridrow="2" pageId="9" pageNumber="428">12.4–17.7</td>
</tr>
<tr id="13B534C58B39D3F0FF11FED4BE633E5A" box="[166,1350,348,373]" gridrow="3" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FED4BE633E5A" box="[166,1350,348,373]" colspan="5" colspanRight="4" gridcol="0" gridrow="3" pageId="9" pageNumber="428">Percent head length (%)</th>
</tr>
<tr id="13B534C58B39D3F0FF11FE0DBE633EB2" box="[166,1350,389,413]" gridrow="4" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FE0DB9293EB2" box="[166,524,389,413]" gridcol="0" gridrow="4" pageId="9" pageNumber="428">Snout length</th>
<td id="50645DB98B39D3F0FD0EFE0DB8323EB2" box="[697,791,389,413]" gridcol="1" gridrow="4" pageId="9" pageNumber="428">28.3</td>
<td id="50645DB98B39D3F0FCE9FE0DB8BC3EB2" box="[862,921,389,413]" gridcol="2" gridrow="4" pageId="9" pageNumber="428">31.0</td>
<td id="50645DB98B39D3F0FC57FE0DBF523EB2" box="[992,1143,389,413]" gridcol="3" gridrow="4" pageId="9" pageNumber="428">2.3</td>
<td id="50645DB98B39D3F0FB63FE0DBE633EB2" box="[1236,1350,389,413]" gridcol="4" gridrow="4" pageId="9" pageNumber="428">26.2–35.0</td>
</tr>
<tr id="13B534C58B39D3F0FF11FE25BE633EEA" box="[166,1350,429,453]" gridrow="5" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FE25B9293EEA" box="[166,524,429,453]" gridcol="0" gridrow="5" pageId="9" pageNumber="428">Postorbital head length</th>
<td id="50645DB98B39D3F0FD0EFE25B8323EEA" box="[697,791,429,453]" gridcol="1" gridrow="5" pageId="9" pageNumber="428">41.9</td>
<td id="50645DB98B39D3F0FCE9FE25B8BC3EEA" box="[862,921,429,453]" gridcol="2" gridrow="5" pageId="9" pageNumber="428">42.7</td>
<td id="50645DB98B39D3F0FC57FE25BF523EEA" box="[992,1143,429,453]" gridcol="3" gridrow="5" pageId="9" pageNumber="428">1.8</td>
<td id="50645DB98B39D3F0FB63FE25BE633EEA" box="[1236,1350,429,453]" gridcol="4" gridrow="5" pageId="9" pageNumber="428">39.0–46.0</td>
</tr>
<tr id="13B534C58B39D3F0FF11FE5DBE633EC1" box="[166,1350,469,494]" gridrow="6" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FE5DB9293EC1" box="[166,524,469,494]" gridcol="0" gridrow="6" pageId="9" pageNumber="428">Horizontal eye diameter</th>
<td id="50645DB98B39D3F0FD0EFE5DB8323EC1" box="[697,791,469,494]" gridcol="1" gridrow="6" pageId="9" pageNumber="428">32.3</td>
<td id="50645DB98B39D3F0FCE9FE5DB8BC3EC1" box="[862,921,469,494]" gridcol="2" gridrow="6" pageId="9" pageNumber="428">31.1</td>
<td id="50645DB98B39D3F0FC57FE5DBF523EC1" box="[992,1143,469,494]" gridcol="3" gridrow="6" pageId="9" pageNumber="428">1.6</td>
<td id="50645DB98B39D3F0FB63FE5DBE633EC1" box="[1236,1350,469,494]" gridcol="4" gridrow="6" pageId="9" pageNumber="428">28.5–34.2</td>
</tr>
<tr id="13B534C58B39D3F0FF11FE75BE633D39" box="[166,1350,509,534]" gridrow="7" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FE75B9293D39" box="[166,524,509,534]" gridcol="0" gridrow="7" pageId="9" pageNumber="428">Vertical eye diameter</th>
<td id="50645DB98B39D3F0FD0EFE75B8323D39" box="[697,791,509,534]" gridcol="1" gridrow="7" pageId="9" pageNumber="428">30.8</td>
<td id="50645DB98B39D3F0FCE9FE75B8BC3D39" box="[862,921,509,534]" gridcol="2" gridrow="7" pageId="9" pageNumber="428">29.0</td>
<td id="50645DB98B39D3F0FC57FE75BF523D39" box="[992,1143,509,534]" gridcol="3" gridrow="7" pageId="9" pageNumber="428">1.8</td>
<td id="50645DB98B39D3F0FB63FE75BE633D39" box="[1236,1350,509,534]" gridcol="4" gridrow="7" pageId="9" pageNumber="428">25.1–32.4</td>
</tr>
<tr id="13B534C58B39D3F0FF11FDAEBE633D11" box="[166,1350,550,574]" gridrow="8" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FDAEB9293D11" box="[166,524,550,574]" gridcol="0" gridrow="8" pageId="9" pageNumber="428">Preorbital depth</th>
<td id="50645DB98B39D3F0FD0EFDAEB8323D11" box="[697,791,550,574]" gridcol="1" gridrow="8" pageId="9" pageNumber="428">17.9</td>
<td id="50645DB98B39D3F0FCE9FDAEB8BC3D11" box="[862,921,550,574]" gridcol="2" gridrow="8" pageId="9" pageNumber="428">16.3</td>
<td id="50645DB98B39D3F0FC57FDAEBF523D11" box="[992,1143,550,574]" gridcol="3" gridrow="8" pageId="9" pageNumber="428">1.5</td>
<td id="50645DB98B39D3F0FB63FDAEBE633D11" box="[1236,1350,550,574]" gridcol="4" gridrow="8" pageId="9" pageNumber="428">14.1–18.9</td>
</tr>
<tr id="13B534C58B39D3F0FF11FDC6BE633D49" box="[166,1350,590,614]" gridrow="9" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FDC6B9293D49" box="[166,524,590,614]" gridcol="0" gridrow="9" pageId="9" pageNumber="428">Cheek depth</th>
<td id="50645DB98B39D3F0FD0EFDC6B8323D49" box="[697,791,590,614]" gridcol="1" gridrow="9" pageId="9" pageNumber="428">31.1</td>
<td id="50645DB98B39D3F0FCE9FDC6B8BC3D49" box="[862,921,590,614]" gridcol="2" gridrow="9" pageId="9" pageNumber="428">26.3</td>
<td id="50645DB98B39D3F0FC57FDC6BF523D49" box="[992,1143,590,614]" gridcol="3" gridrow="9" pageId="9" pageNumber="428">3.6</td>
<td id="50645DB98B39D3F0FB63FDC6BE633D49" box="[1236,1350,590,614]" gridcol="4" gridrow="9" pageId="9" pageNumber="428">21.1–33.5</td>
</tr>
<tr id="13B534C58B39D3F0FF11FDFEBE633DA1" box="[166,1350,630,654]" gridrow="10" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FDFEB9293DA1" box="[166,524,630,654]" gridcol="0" gridrow="10" pageId="9" pageNumber="428">Lower jaw length</th>
<td id="50645DB98B39D3F0FD0EFDFEB8323DA1" box="[697,791,630,654]" gridcol="1" gridrow="10" pageId="9" pageNumber="428">26.9</td>
<td id="50645DB98B39D3F0FCE9FDFEB8BC3DA1" box="[862,921,630,654]" gridcol="2" gridrow="10" pageId="9" pageNumber="428">26.0</td>
<td id="50645DB98B39D3F0FC57FDFEBF523DA1" box="[992,1143,630,654]" gridcol="3" gridrow="10" pageId="9" pageNumber="428">3.0</td>
<td id="50645DB98B39D3F0FB63FDFEBE633DA1" box="[1236,1350,630,654]" gridcol="4" gridrow="10" pageId="9" pageNumber="428">21.6–33.5</td>
</tr>
<tr id="13B534C58B39D3F0FF11FD16BE633D98" box="[166,1350,670,695]" gridrow="11" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FD16B9293D98" box="[166,524,670,695]" gridcol="0" gridrow="11" pageId="9" pageNumber="428">Head depth</th>
<td id="50645DB98B39D3F0FD0EFD16B8323D98" box="[697,791,670,695]" gridcol="1" gridrow="11" pageId="9" pageNumber="428">104.8</td>
<td id="50645DB98B39D3F0FCE9FD16B8BC3D98" box="[862,921,670,695]" gridcol="2" gridrow="11" pageId="9" pageNumber="428">100.1</td>
<td id="50645DB98B39D3F0FC57FD16BF523D98" box="[992,1143,670,695]" gridcol="3" gridrow="11" pageId="9" pageNumber="428">5.4</td>
<td id="50645DB98B39D3F0FB63FD16BE633D98" box="[1236,1350,670,695]" gridcol="4" gridrow="11" pageId="9" pageNumber="428">91.3–109.7</td>
</tr>
<tr id="13B534C58B39D3F0FF11FD4FBE633DF0" box="[166,1350,711,735]" gridrow="12" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FD4FB9293DF0" box="[166,524,711,735]" gridcol="0" gridrow="12" pageId="9" pageNumber="428">Inter-orbital width</th>
<td id="50645DB98B39D3F0FD0EFD4FB8323DF0" box="[697,791,711,735]" gridcol="1" gridrow="12" pageId="9" pageNumber="428">26.2</td>
<td id="50645DB98B39D3F0FCE9FD4FB8BC3DF0" box="[862,921,711,735]" gridcol="2" gridrow="12" pageId="9" pageNumber="428">27.3</td>
<td id="50645DB98B39D3F0FC57FD4FBF523DF0" box="[992,1143,711,735]" gridcol="3" gridrow="12" pageId="9" pageNumber="428">1.5</td>
<td id="50645DB98B39D3F0FB63FD4FBE633DF0" box="[1236,1350,711,735]" gridcol="4" gridrow="12" pageId="9" pageNumber="428">23.6–29.7</td>
</tr>
<tr id="13B534C58B39D3F0FF11FD67BE633C28" box="[166,1350,751,775]" gridrow="13" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FD67B9293C28" box="[166,524,751,775]" gridcol="0" gridrow="13" pageId="9" pageNumber="428">Premaxillary pedicel length</th>
<td id="50645DB98B39D3F0FD0EFD67B8323C28" box="[697,791,751,775]" gridcol="1" gridrow="13" pageId="9" pageNumber="428">15.7</td>
<td id="50645DB98B39D3F0FCE9FD67B8BC3C28" box="[862,921,751,775]" gridcol="2" gridrow="13" pageId="9" pageNumber="428">13.7</td>
<td id="50645DB98B39D3F0FC57FD67BF523C28" box="[992,1143,751,775]" gridcol="3" gridrow="13" pageId="9" pageNumber="428">1.7</td>
<td id="50645DB98B39D3F0FB63FD67BE633C28" box="[1236,1350,751,775]" gridcol="4" gridrow="13" pageId="9" pageNumber="428">10.9–16.4</td>
</tr>
<tr id="13B534C58B39D3F0FF11FC9FBE633C00" box="[166,1350,791,815]" gridrow="14" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FC9FBE633C00" box="[166,1350,791,815]" colspan="5" colspanRight="4" gridcol="0" gridrow="14" pageId="9" pageNumber="428">Percent standard length (%)</th>
</tr>
<tr id="13B534C58B39D3F0FF11FCB7BE633C77" box="[166,1350,831,856]" gridrow="15" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FCB7B9293C77" box="[166,524,831,856]" gridcol="0" gridrow="15" pageId="9" pageNumber="428">Head length</th>
<td id="50645DB98B39D3F0FD0EFCB7B8323C77" box="[697,791,831,856]" gridcol="1" gridrow="15" pageId="9" pageNumber="428">32.7</td>
<td id="50645DB98B39D3F0FCE9FCB7B8BC3C77" box="[862,921,831,856]" gridcol="2" gridrow="15" pageId="9" pageNumber="428">33.0</td>
<td id="50645DB98B39D3F0FC57FCB7BF523C77" box="[992,1143,831,856]" gridcol="3" gridrow="15" pageId="9" pageNumber="428">0.8</td>
<td id="50645DB98B39D3F0FB63FCB7BE633C77" box="[1236,1350,831,856]" gridcol="4" gridrow="15" pageId="9" pageNumber="428">31.5–34.4</td>
</tr>
<tr id="13B534C58B39D3F0FF11FCE0BE633CAF" box="[166,1350,872,896]" gridrow="16" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FCE0B9293CAF" box="[166,524,872,896]" gridcol="0" gridrow="16" pageId="9" pageNumber="428">Body depth</th>
<td id="50645DB98B39D3F0FD0EFCE0B8323CAF" box="[697,791,872,896]" gridcol="1" gridrow="16" pageId="9" pageNumber="428">43.3</td>
<td id="50645DB98B39D3F0FCE9FCE0B8BC3CAF" box="[862,921,872,896]" gridcol="2" gridrow="16" pageId="9" pageNumber="428">38.6</td>
<td id="50645DB98B39D3F0FC57FCE0BF523CAF" box="[992,1143,872,896]" gridcol="3" gridrow="16" pageId="9" pageNumber="428">1.8</td>
<td id="50645DB98B39D3F0FB63FCE0BE633CAF" box="[1236,1350,872,896]" gridcol="4" gridrow="16" pageId="9" pageNumber="428">34.9–43.3</td>
</tr>
<tr id="13B534C58B39D3F0FF11FC18BE633C87" box="[166,1350,912,936]" gridrow="17" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FC18B9293C87" box="[166,524,912,936]" gridcol="0" gridrow="17" pageId="9" pageNumber="428">Snout to dorsal fin origin</th>
<td id="50645DB98B39D3F0FD0EFC18B8323C87" box="[697,791,912,936]" gridcol="1" gridrow="17" pageId="9" pageNumber="428">39.5</td>
<td id="50645DB98B39D3F0FCE9FC18B8BC3C87" box="[862,921,912,936]" gridcol="2" gridrow="17" pageId="9" pageNumber="428">35.7</td>
<td id="50645DB98B39D3F0FC57FC18BF523C87" box="[992,1143,912,936]" gridcol="3" gridrow="17" pageId="9" pageNumber="428">1.3</td>
<td id="50645DB98B39D3F0FB63FC18BE633C87" box="[1236,1350,912,936]" gridcol="4" gridrow="17" pageId="9" pageNumber="428">32.6–39.5</td>
</tr>
<tr id="13B534C58B39D3F0FF11FC30BE633CFF" box="[166,1350,952,976]" gridrow="18" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FC30B9293CFF" box="[166,524,952,976]" gridcol="0" gridrow="18" pageId="9" pageNumber="428">Snout to pelvic fin origin</th>
<td id="50645DB98B39D3F0FD0EFC30B8323CFF" box="[697,791,952,976]" gridcol="1" gridrow="18" pageId="9" pageNumber="428">38.9</td>
<td id="50645DB98B39D3F0FCE9FC30B8BC3CFF" box="[862,921,952,976]" gridcol="2" gridrow="18" pageId="9" pageNumber="428">40.6</td>
<td id="50645DB98B39D3F0FC57FC30BF523CFF" box="[992,1143,952,976]" gridcol="3" gridrow="18" pageId="9" pageNumber="428">1.5</td>
<td id="50645DB98B39D3F0FB63FC30BE633CFF" box="[1236,1350,952,976]" gridcol="4" gridrow="18" pageId="9" pageNumber="428">37.4–42.5</td>
</tr>
<tr id="13B534C58B39D3F0FF11FC68BE633CD7" box="[166,1350,992,1016]" gridrow="19" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FC68B9293CD7" box="[166,524,992,1016]" gridcol="0" gridrow="19" pageId="9" pageNumber="428">Dorsal fin base length</th>
<td id="50645DB98B39D3F0FD0EFC68B8323CD7" box="[697,791,992,1016]" gridcol="1" gridrow="19" pageId="9" pageNumber="428">60.2</td>
<td id="50645DB98B39D3F0FCE9FC68B8BC3CD7" box="[862,921,992,1016]" gridcol="2" gridrow="19" pageId="9" pageNumber="428">57.2</td>
<td id="50645DB98B39D3F0FC57FC68BF523CD7" box="[992,1143,992,1016]" gridcol="3" gridrow="19" pageId="9" pageNumber="428">2.2</td>
<td id="50645DB98B39D3F0FB63FC68BE633CD7" box="[1236,1350,992,1016]" gridcol="4" gridrow="19" pageId="9" pageNumber="428">52.4–60.2</td>
</tr>
<tr id="13B534C58B39D3F0FF11FB80BE633B0E" box="[166,1350,1032,1057]" gridrow="20" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FB80B9293B0E" box="[166,524,1032,1057]" gridcol="0" gridrow="20" pageId="9" pageNumber="428">Anterior dorsal to anterior anal</th>
<td id="50645DB98B39D3F0FD0EFB80B8323B0E" box="[697,791,1032,1057]" gridcol="1" gridrow="20" pageId="9" pageNumber="428">55.3</td>
<td id="50645DB98B39D3F0FCE9FB80B8BC3B0E" box="[862,921,1032,1057]" gridcol="2" gridrow="20" pageId="9" pageNumber="428">51.8</td>
<td id="50645DB98B39D3F0FC57FB80BF523B0E" box="[992,1143,1032,1057]" gridcol="3" gridrow="20" pageId="9" pageNumber="428">1.7</td>
<td id="50645DB98B39D3F0FB63FB80BE633B0E" box="[1236,1350,1032,1057]" gridcol="4" gridrow="20" pageId="9" pageNumber="428">48.6–55.3</td>
</tr>
<tr id="13B534C58B39D3F0FF11FBB9BE633B66" box="[166,1350,1073,1097]" gridrow="21" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FBB9B9293B66" box="[166,524,1073,1097]" gridcol="0" gridrow="21" pageId="9" pageNumber="428">Anterior dorsal to posterior anal</th>
<td id="50645DB98B39D3F0FD0EFBB9B8323B66" box="[697,791,1073,1097]" gridcol="1" gridrow="21" pageId="9" pageNumber="428">65.3</td>
<td id="50645DB98B39D3F0FCE9FBB9B8BC3B66" box="[862,921,1073,1097]" gridcol="2" gridrow="21" pageId="9" pageNumber="428">62.5</td>
<td id="50645DB98B39D3F0FC57FBB9BF523B66" box="[992,1143,1073,1097]" gridcol="3" gridrow="21" pageId="9" pageNumber="428">1.8</td>
<td id="50645DB98B39D3F0FB63FBB9BE633B66" box="[1236,1350,1073,1097]" gridcol="4" gridrow="21" pageId="9" pageNumber="428">58.0–65.3</td>
</tr>
<tr id="13B534C58B39D3F0FF11FBD1BE633B5E" box="[166,1350,1113,1137]" gridrow="22" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FBD1B9293B5E" box="[166,524,1113,1137]" gridcol="0" gridrow="22" pageId="9" pageNumber="428">Posterior dorsal to anterior anal</th>
<td id="50645DB98B39D3F0FD0EFBD1B8323B5E" box="[697,791,1113,1137]" gridcol="1" gridrow="22" pageId="9" pageNumber="428">34.9</td>
<td id="50645DB98B39D3F0FCE9FBD1B8BC3B5E" box="[862,921,1113,1137]" gridcol="2" gridrow="22" pageId="9" pageNumber="428">33.0</td>
<td id="50645DB98B39D3F0FC57FBD1BF523B5E" box="[992,1143,1113,1137]" gridcol="3" gridrow="22" pageId="9" pageNumber="428">1.4</td>
<td id="50645DB98B39D3F0FB63FBD1BE633B5E" box="[1236,1350,1113,1137]" gridcol="4" gridrow="22" pageId="9" pageNumber="428">30.2–35.0</td>
</tr>
<tr id="13B534C58B39D3F0FF11FB09BE633BB6" box="[166,1350,1153,1177]" gridrow="23" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FB09B9293BB6" box="[166,524,1153,1177]" gridcol="0" gridrow="23" pageId="9" pageNumber="428">Posterior dorsal to posterior anal</th>
<td id="50645DB98B39D3F0FD0EFB09B8323BB6" box="[697,791,1153,1177]" gridcol="1" gridrow="23" pageId="9" pageNumber="428">19.9</td>
<td id="50645DB98B39D3F0FCE9FB09B8BC3BB6" box="[862,921,1153,1177]" gridcol="2" gridrow="23" pageId="9" pageNumber="428">18.6</td>
<td id="50645DB98B39D3F0FC57FB09BF523BB6" box="[992,1143,1153,1177]" gridcol="3" gridrow="23" pageId="9" pageNumber="428">1.0</td>
<td id="50645DB98B39D3F0FB63FB09BE633BB6" box="[1236,1350,1153,1177]" gridcol="4" gridrow="23" pageId="9" pageNumber="428">17.0–19.9</td>
</tr>
<tr id="13B534C58B39D3F0FF11FB21BE633BED" box="[166,1350,1193,1218]" gridrow="24" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FB21B9293BED" box="[166,524,1193,1218]" gridcol="0" gridrow="24" pageId="9" pageNumber="428">Posterior dorsal to ventral caudal</th>
<td id="50645DB98B39D3F0FD0EFB21B8323BED" box="[697,791,1193,1218]" gridcol="1" gridrow="24" pageId="9" pageNumber="428">20.9</td>
<td id="50645DB98B39D3F0FCE9FB21B8BC3BED" box="[862,921,1193,1218]" gridcol="2" gridrow="24" pageId="9" pageNumber="428">19.3</td>
<td id="50645DB98B39D3F0FC57FB21BF523BED" box="[992,1143,1193,1218]" gridcol="3" gridrow="24" pageId="9" pageNumber="428">0.9</td>
<td id="50645DB98B39D3F0FB63FB21BE633BED" box="[1236,1350,1193,1218]" gridcol="4" gridrow="24" pageId="9" pageNumber="428">17.5–20.9</td>
</tr>
<tr id="13B534C58B39D3F0FF11FB5ABE633BC5" box="[166,1350,1234,1258]" gridrow="25" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FB5AB9293BC5" box="[166,524,1234,1258]" gridcol="0" gridrow="25" pageId="9" pageNumber="428">Posterior anal to dorsal caudal</th>
<td id="50645DB98B39D3F0FD0EFB5AB8323BC5" box="[697,791,1234,1258]" gridcol="1" gridrow="25" pageId="9" pageNumber="428">23.2</td>
<td id="50645DB98B39D3F0FCE9FB5AB8BC3BC5" box="[862,921,1234,1258]" gridcol="2" gridrow="25" pageId="9" pageNumber="428">22.0</td>
<td id="50645DB98B39D3F0FC57FB5ABF523BC5" box="[992,1143,1234,1258]" gridcol="3" gridrow="25" pageId="9" pageNumber="428">0.9</td>
<td id="50645DB98B39D3F0FB63FB5ABE633BC5" box="[1236,1350,1234,1258]" gridcol="4" gridrow="25" pageId="9" pageNumber="428">21.0–23.8</td>
</tr>
<tr id="13B534C58B39D3F0FF11FB72BE633A3D" box="[166,1350,1274,1298]" gridrow="26" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FB72B9293A3D" box="[166,524,1274,1298]" gridcol="0" gridrow="26" pageId="9" pageNumber="428">Anterior dorsal to pelvic-fin origin</th>
<td id="50645DB98B39D3F0FD0EFB72B8323A3D" box="[697,791,1274,1298]" gridcol="1" gridrow="26" pageId="9" pageNumber="428">42.7</td>
<td id="50645DB98B39D3F0FCE9FB72B8BC3A3D" box="[862,921,1274,1298]" gridcol="2" gridrow="26" pageId="9" pageNumber="428">38.5</td>
<td id="50645DB98B39D3F0FC57FB72BF523A3D" box="[992,1143,1274,1298]" gridcol="3" gridrow="26" pageId="9" pageNumber="428">1.8</td>
<td id="50645DB98B39D3F0FB63FB72BE633A3D" box="[1236,1350,1274,1298]" gridcol="4" gridrow="26" pageId="9" pageNumber="428">35.0–42.7</td>
</tr>
<tr id="13B534C58B39D3F0FF11FAAABE633A15" box="[166,1350,1314,1338]" gridrow="27" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FAAAB9293A15" box="[166,524,1314,1338]" gridcol="0" gridrow="27" pageId="9" pageNumber="428">Posterior dorsal to pelvic-fin origin</th>
<td id="50645DB98B39D3F0FD0EFAAAB8323A15" box="[697,791,1314,1338]" gridcol="1" gridrow="27" pageId="9" pageNumber="428">61.1</td>
<td id="50645DB98B39D3F0FCE9FAAAB8BC3A15" box="[862,921,1314,1338]" gridcol="2" gridrow="27" pageId="9" pageNumber="428">57.5</td>
<td id="50645DB98B39D3F0FC57FAAABF523A15" box="[992,1143,1314,1338]" gridcol="3" gridrow="27" pageId="9" pageNumber="428">1.7</td>
<td id="50645DB98B39D3F0FB63FAAABE633A15" box="[1236,1350,1314,1338]" gridcol="4" gridrow="27" pageId="9" pageNumber="428">54.7–61.1</td>
</tr>
<tr id="13B534C58B39D3F0FF11FAC2BE633A4C" box="[166,1350,1354,1379]" gridrow="28" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FAC2B9293A4C" box="[166,524,1354,1379]" gridcol="0" gridrow="28" pageId="9" pageNumber="428">Caudal peduncle length</th>
<td id="50645DB98B39D3F0FD0EFAC2B8323A4C" box="[697,791,1354,1379]" gridcol="1" gridrow="28" pageId="9" pageNumber="428">13.6</td>
<td id="50645DB98B39D3F0FCE9FAC2B8BC3A4C" box="[862,921,1354,1379]" gridcol="2" gridrow="28" pageId="9" pageNumber="428">13.3</td>
<td id="50645DB98B39D3F0FC57FAC2BF523A4C" box="[992,1143,1354,1379]" gridcol="3" gridrow="28" pageId="9" pageNumber="428">1.0</td>
<td id="50645DB98B39D3F0FB63FAC2BE633A4C" box="[1236,1350,1354,1379]" gridcol="4" gridrow="28" pageId="9" pageNumber="428">11.5–14.8</td>
</tr>
<tr id="13B534C58B39D3F0FF11FAFBBE633AA4" box="[166,1350,1395,1419]" gridrow="29" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FAFBB9293AA4" box="[166,524,1395,1419]" gridcol="0" gridrow="29" pageId="9" pageNumber="428">Least caudal peduncle depth</th>
<td id="50645DB98B39D3F0FD0EFAFBB8323AA4" box="[697,791,1395,1419]" gridcol="1" gridrow="29" pageId="9" pageNumber="428">15.8</td>
<td id="50645DB98B39D3F0FCE9FAFBB8BC3AA4" box="[862,921,1395,1419]" gridcol="2" gridrow="29" pageId="9" pageNumber="428">15.6</td>
<td id="50645DB98B39D3F0FC57FAFBBF523AA4" box="[992,1143,1395,1419]" gridcol="3" gridrow="29" pageId="9" pageNumber="428">0.7</td>
<td id="50645DB98B39D3F0FB63FAFBBE633AA4" box="[1236,1350,1395,1419]" gridcol="4" gridrow="29" pageId="9" pageNumber="428">14.4–16.3</td>
</tr>
<tr id="13B534C58B39D3F0FF11FA13BE633A9B" box="[166,1350,1435,1460]" gridrow="30" pageId="9" pageNumber="428" rowspan-1="1">
<th id="50645DB98B39D3F0FF11FA13B9293A9B" box="[166,524,1435,1460]" gridcol="0" gridrow="30" pageId="9" pageNumber="428">Meristics</th>
<td id="50645DB98B39D3F0FCE9FA13B8BC3A9B" box="[862,921,1435,1460]" gridcol="2" gridrow="30" pageId="9" pageNumber="428">Mode</td>
<td id="50645DB98B39D3F0FC57FA13BF523A9B" box="[992,1143,1435,1460]" gridcol="3" gridrow="30" pageId="9" pageNumber="428">Frequency (%)</td>
<td id="50645DB98B39D3F0FB63FA13BE633A9B" box="[1236,1350,1435,1460]" gridcol="4" gridrow="30" pageId="9" pageNumber="428">Range</td>
</tr>
<tr id="13B534C58B39D3F0FF11FA4BBE633AF4" box="[166,1350,1475,1499]" gridrow="31" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FA4BB9293AF4" box="[166,524,1475,1499]" gridcol="0" gridrow="31" pageId="9" pageNumber="428">Dorsal-fin spines</th>
<td id="50645DB98B39D3F0FD0EFA4BB8323AF4" box="[697,791,1475,1499]" gridcol="1" gridrow="31" pageId="9" pageNumber="428">17</td>
<td id="50645DB98B39D3F0FCE9FA4BB8BC3AF4" box="[862,921,1475,1499]" gridcol="2" gridrow="31" pageId="9" pageNumber="428">17</td>
<td id="50645DB98B39D3F0FC57FA4BBF523AF4" box="[992,1143,1475,1499]" gridcol="3" gridrow="31" pageId="9" pageNumber="428">80.0</td>
<td id="50645DB98B39D3F0FB63FA4BBE633AF4" box="[1236,1350,1475,1499]" gridcol="4" gridrow="31" pageId="9" pageNumber="428">16–18</td>
</tr>
<tr id="13B534C58B39D3F0FF11FA63BE63392B" box="[166,1350,1515,1540]" gridrow="32" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11FA63B929392B" box="[166,524,1515,1540]" gridcol="0" gridrow="32" pageId="9" pageNumber="428">Dorsal-fin rays</th>
<td id="50645DB98B39D3F0FD0EFA63B832392B" box="[697,791,1515,1540]" gridcol="1" gridrow="32" pageId="9" pageNumber="428">8</td>
<td id="50645DB98B39D3F0FCE9FA63B8BC392B" box="[862,921,1515,1540]" gridcol="2" gridrow="32" pageId="9" pageNumber="428">8</td>
<td id="50645DB98B39D3F0FC57FA63BF52392B" box="[992,1143,1515,1540]" gridcol="3" gridrow="32" pageId="9" pageNumber="428">80.0</td>
<td id="50645DB98B39D3F0FB63FA63BE63392B" box="[1236,1350,1515,1540]" gridcol="4" gridrow="32" pageId="9" pageNumber="428">7–9</td>
</tr>
<tr id="13B534C58B39D3F0FF11F99CBE633903" box="[166,1350,1556,1580]" gridrow="33" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11F99CB9293903" box="[166,524,1556,1580]" gridcol="0" gridrow="33" pageId="9" pageNumber="428">Anal-fin spines</th>
<td id="50645DB98B39D3F0FD0EF99CB8323903" box="[697,791,1556,1580]" gridcol="1" gridrow="33" pageId="9" pageNumber="428">3</td>
<td id="50645DB98B39D3F0FCE9F99CB8BC3903" box="[862,921,1556,1580]" gridcol="2" gridrow="33" pageId="9" pageNumber="428">3</td>
<td id="50645DB98B39D3F0FC57F99CBF523903" box="[992,1143,1556,1580]" gridcol="3" gridrow="33" pageId="9" pageNumber="428">100.0</td>
<td id="50645DB98B39D3F0FB63F99CBE633903" box="[1236,1350,1556,1580]" gridcol="4" gridrow="33" pageId="9" pageNumber="428">3–3</td>
</tr>
<tr id="13B534C58B39D3F0FF11F9B4BE63397A" box="[166,1350,1596,1621]" gridrow="34" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11F9B4B929397A" box="[166,524,1596,1621]" gridcol="0" gridrow="34" pageId="9" pageNumber="428">Anal-fin rays</th>
<td id="50645DB98B39D3F0FD0EF9B4B832397A" box="[697,791,1596,1621]" gridcol="1" gridrow="34" pageId="9" pageNumber="428">7</td>
<td id="50645DB98B39D3F0FCE9F9B4B8BC397A" box="[862,921,1596,1621]" gridcol="2" gridrow="34" pageId="9" pageNumber="428">8</td>
<td id="50645DB98B39D3F0FC57F9B4BF52397A" box="[992,1143,1596,1621]" gridcol="3" gridrow="34" pageId="9" pageNumber="428">75.0</td>
<td id="50645DB98B39D3F0FB63F9B4BE63397A" box="[1236,1350,1596,1621]" gridcol="4" gridrow="34" pageId="9" pageNumber="428">7–8</td>
</tr>
<tr id="13B534C58B39D3F0FF11F9ECBE633952" box="[166,1350,1636,1661]" gridrow="35" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11F9ECB9293952" box="[166,524,1636,1661]" gridcol="0" gridrow="35" pageId="9" pageNumber="428">Pectoral-fin rays</th>
<td id="50645DB98B39D3F0FD0EF9ECB8323952" box="[697,791,1636,1661]" gridcol="1" gridrow="35" pageId="9" pageNumber="428">14</td>
<td id="50645DB98B39D3F0FCE9F9ECB8BC3952" box="[862,921,1636,1661]" gridcol="2" gridrow="35" pageId="9" pageNumber="428">14</td>
<td id="50645DB98B39D3F0FC57F9ECBF523952" box="[992,1143,1636,1661]" gridcol="3" gridrow="35" pageId="9" pageNumber="428">45.0</td>
<td id="50645DB98B39D3F0FB63F9ECBE633952" box="[1236,1350,1636,1661]" gridcol="4" gridrow="35" pageId="9" pageNumber="428">12–15</td>
</tr>
<tr id="13B534C58B39D3F0FF11F904BE63398A" box="[166,1350,1676,1701]" gridrow="36" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11F904B929398A" box="[166,524,1676,1701]" gridcol="0" gridrow="36" pageId="9" pageNumber="428">Pelvic-fin rays</th>
<td id="50645DB98B39D3F0FD0EF904B832398A" box="[697,791,1676,1701]" gridcol="1" gridrow="36" pageId="9" pageNumber="428">6</td>
<td id="50645DB98B39D3F0FCE9F904B8BC398A" box="[862,921,1676,1701]" gridcol="2" gridrow="36" pageId="9" pageNumber="428">6</td>
<td id="50645DB98B39D3F0FC57F904BF52398A" box="[992,1143,1676,1701]" gridcol="3" gridrow="36" pageId="9" pageNumber="428">90.0</td>
<td id="50645DB98B39D3F0FB63F904BE63398A" box="[1236,1350,1676,1701]" gridcol="4" gridrow="36" pageId="9" pageNumber="428">6–7</td>
</tr>
<tr id="13B534C58B39D3F0FF11F93DBE6339E2" box="[166,1350,1717,1741]" gridrow="37" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11F93DB92939E2" box="[166,524,1717,1741]" gridcol="0" gridrow="37" pageId="9" pageNumber="428">Lateral line scales</th>
<td id="50645DB98B39D3F0FD0EF93DB83239E2" box="[697,791,1717,1741]" gridcol="1" gridrow="37" pageId="9" pageNumber="428">30</td>
<td id="50645DB98B39D3F0FCE9F93DB8BC39E2" box="[862,921,1717,1741]" gridcol="2" gridrow="37" pageId="9" pageNumber="428">30</td>
<td id="50645DB98B39D3F0FC57F93DBF5239E2" box="[992,1143,1717,1741]" gridcol="3" gridrow="37" pageId="9" pageNumber="428">40.0</td>
<td id="50645DB98B39D3F0FB63F93DBE6339E2" box="[1236,1350,1717,1741]" gridcol="4" gridrow="37" pageId="9" pageNumber="428">29–32</td>
</tr>
<tr id="13B534C58B39D3F0FF11F955BE6339DA" box="[166,1350,1757,1781]" gridrow="38" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11F955B92939DA" box="[166,524,1757,1781]" gridcol="0" gridrow="38" pageId="9" pageNumber="428">Pored scales caudal</th>
<td id="50645DB98B39D3F0FD0EF955B83239DA" box="[697,791,1757,1781]" gridcol="1" gridrow="38" pageId="9" pageNumber="428">2</td>
<td id="50645DB98B39D3F0FCE9F955B8BC39DA" box="[862,921,1757,1781]" gridcol="2" gridrow="38" pageId="9" pageNumber="428">1</td>
<td id="50645DB98B39D3F0FC57F955BF5239DA" box="[992,1143,1757,1781]" gridcol="3" gridrow="38" pageId="9" pageNumber="428">40.0</td>
<td id="50645DB98B39D3F0FB63F955BE6339DA" box="[1236,1350,1757,1781]" gridcol="4" gridrow="38" pageId="9" pageNumber="428">0–3</td>
</tr>
<tr id="13B534C58B39D3F0FF11F88DBE633832" box="[166,1350,1797,1821]" gridrow="39" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11F88DB9293832" box="[166,524,1797,1821]" gridcol="0" gridrow="39" pageId="9" pageNumber="428">Cheek scale rows</th>
<td id="50645DB98B39D3F0FD0EF88DB8323832" box="[697,791,1797,1821]" gridcol="1" gridrow="39" pageId="9" pageNumber="428">4</td>
<td id="50645DB98B39D3F0FCE9F88DB8BC3832" box="[862,921,1797,1821]" gridcol="2" gridrow="39" pageId="9" pageNumber="428">4</td>
<td id="50645DB98B39D3F0FC57F88DBF523832" box="[992,1143,1797,1821]" gridcol="3" gridrow="39" pageId="9" pageNumber="428">85.0</td>
<td id="50645DB98B39D3F0FB63F88DBE633832" box="[1236,1350,1797,1821]" gridcol="4" gridrow="39" pageId="9" pageNumber="428">4–5</td>
</tr>
<tr id="13B534C58B39D3F0FF11F8A5BE633869" box="[166,1350,1837,1862]" gridrow="40" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11F8A5B9293869" box="[166,524,1837,1862]" gridcol="0" gridrow="40" pageId="9" pageNumber="428">Gill rakers 1st ceratobranchial</th>
<td id="50645DB98B39D3F0FD0EF8A5B8323869" box="[697,791,1837,1862]" gridcol="1" gridrow="40" pageId="9" pageNumber="428">10</td>
<td id="50645DB98B39D3F0FCE9F8A5B8BC3869" box="[862,921,1837,1862]" gridcol="2" gridrow="40" pageId="9" pageNumber="428">10</td>
<td id="50645DB98B39D3F0FC57F8A5BF523869" box="[992,1143,1837,1862]" gridcol="3" gridrow="40" pageId="9" pageNumber="428">70.0</td>
<td id="50645DB98B39D3F0FB63F8A5BE633869" box="[1236,1350,1837,1862]" gridcol="4" gridrow="40" pageId="9" pageNumber="428">9–11</td>
</tr>
<tr id="13B534C58B39D3F0FF11F8DEBE633841" box="[166,1350,1878,1902]" gridrow="41" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11F8DEB9293841" box="[166,524,1878,1902]" gridcol="0" gridrow="41" pageId="9" pageNumber="428">Gill rakers 1st epibranchial</th>
<td id="50645DB98B39D3F0FD0EF8DEB8323841" box="[697,791,1878,1902]" gridcol="1" gridrow="41" pageId="9" pageNumber="428">5</td>
<td id="50645DB98B39D3F0FCE9F8DEB8BC3841" box="[862,921,1878,1902]" gridcol="2" gridrow="41" pageId="9" pageNumber="428">4</td>
<td id="50645DB98B39D3F0FC57F8DEBF523841" box="[992,1143,1878,1902]" gridcol="3" gridrow="41" pageId="9" pageNumber="428">85.0</td>
<td id="50645DB98B39D3F0FB63F8DEBE633841" box="[1236,1350,1878,1902]" gridcol="4" gridrow="41" pageId="9" pageNumber="428">3–5</td>
</tr>
<tr id="13B534C58B39D3F0FF11F8F6BE6338B9" box="[166,1350,1918,1942]" gridrow="42" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11F8F6B92938B9" box="[166,524,1918,1942]" gridcol="0" gridrow="42" pageId="9" pageNumber="428">Teeth outer left lower jaw</th>
<td id="50645DB98B39D3F0FD0EF8F6B83238B9" box="[697,791,1918,1942]" gridcol="1" gridrow="42" pageId="9" pageNumber="428">12</td>
<td id="50645DB98B39D3F0FCE9F8F6B8BC38B9" box="[862,921,1918,1942]" gridcol="2" gridrow="42" pageId="9" pageNumber="428">10</td>
<td id="50645DB98B39D3F0FC57F8F6BF5238B9" box="[992,1143,1918,1942]" gridcol="3" gridrow="42" pageId="9" pageNumber="428">25.0</td>
<td id="50645DB98B39D3F0FB63F8F6BE6338B9" box="[1236,1350,1918,1942]" gridcol="4" gridrow="42" pageId="9" pageNumber="428">6–17</td>
</tr>
<tr id="13B534C58B39D3F0FF11F82EBE633891" box="[166,1350,1958,1982]" gridrow="43" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11F82EB9293891" box="[166,524,1958,1982]" gridcol="0" gridrow="43" pageId="9" pageNumber="428">Tooth rows upper jaw</th>
<td id="50645DB98B39D3F0FD0EF82EB8323891" box="[697,791,1958,1982]" gridcol="1" gridrow="43" pageId="9" pageNumber="428">4</td>
<td id="50645DB98B39D3F0FCE9F82EB8BC3891" box="[862,921,1958,1982]" gridcol="2" gridrow="43" pageId="9" pageNumber="428">3</td>
<td id="50645DB98B39D3F0FC57F82EBF523891" box="[992,1143,1958,1982]" gridcol="3" gridrow="43" pageId="9" pageNumber="428">70.0</td>
<td id="50645DB98B39D3F0FB63F82EBE633891" box="[1236,1350,1958,1982]" gridcol="4" gridrow="43" pageId="9" pageNumber="428">3–4</td>
</tr>
<tr id="13B534C58B39D3F0FF11F846BE6338C8" box="[166,1350,1998,2023]" gridrow="44" pageId="9" pageNumber="428">
<th id="50645DB98B39D3F0FF11F846B92938C8" box="[166,524,1998,2023]" gridcol="0" gridrow="44" pageId="9" pageNumber="428">Tooth rows lower jaw</th>
<td id="50645DB98B39D3F0FD0EF846B83238C8" box="[697,791,1998,2023]" gridcol="1" gridrow="44" pageId="9" pageNumber="428">4</td>
<td id="50645DB98B39D3F0FCE9F846B8BC38C8" box="[862,921,1998,2023]" gridcol="2" gridrow="44" pageId="9" pageNumber="428">3</td>
<td id="50645DB98B39D3F0FC57F846BF5238C8" box="[992,1143,1998,2023]" gridcol="3" gridrow="44" pageId="9" pageNumber="428">90.0.</td>
<td id="50645DB98B39D3F0FB63F846BE6338C8" box="[1236,1350,1998,2023]" gridcol="4" gridrow="44" pageId="9" pageNumber="428">3–4</td>
</tr>
</table>
</paragraph>
<paragraph id="AD3A36878B3A2C05FF70FF1FBA603FD6" blockId="10.[151,1437,151,609]" pageId="10" pageNumber="429">Mature males protect an empty shell as their territory but mouth-brooding females at Gallireya Reef were frequently encountered among the rocks of the habitat. Mouth-brooding females in Chitimba Bay were mostly seen near their shell.</paragraph>
</subSubSection>
<subSubSection id="E59F650C8B3A2C05FF70FE8BBA483E6E" pageId="10" pageNumber="429" type="etymology">
<paragraph id="AD3A36878B3A2C05FF70FE8BBA483E6E" blockId="10.[151,1437,151,609]" pageId="10" pageNumber="429">
<emphasis id="9FF1EA958B3A2C05FF70FE8BBA6B3E32" bold="true" box="[199,334,259,285]" pageId="10" pageNumber="429">Etymology.</emphasis>
The specific epithet 
<taxonomicName id="6A854D048B3A2C05FD8DFE8BB98E3E32" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[570,683,259,285]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="429" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B3A2C05FD8DFE8BB98E3E32" box="[570,683,259,285]" italics="true" pageId="10" pageNumber="429">gallireyae</emphasis>
</taxonomicName>
is a genitive of Gallireya Reef, the local name for a reef in Youngs Bay, Lake 
<collectingCountry id="D59276178B3A2C05FEB8FEAFBA4D3E6E" box="[271,360,295,321]" name="Malawi" pageId="10" pageNumber="429">Malaŵi</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="E59F650C8B3A2C05FF70FEC3BFCB3D4E" pageId="10" pageNumber="429" type="discussion">
<paragraph id="AD3A36878B3A2C05FF70FEC3BFCB3D4E" blockId="10.[151,1437,151,609]" pageId="10" pageNumber="429">
<emphasis id="9FF1EA958B3A2C05FF70FEC3BA113E4A" bold="true" box="[199,308,331,357]" pageId="10" pageNumber="429">Remarks</emphasis>
. There was complete separation of the minimum polygon clusters when the sheared second principal components (SPCA2) of the morphometric data were plotted against the first principal components (PC1) of the meristic data for 
<taxonomicName id="6A854D048B3A2C05FE3CFE1BB9263E83" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[395,515,403,428]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="429" phylum="Chordata" rank="species" species="ngarae">
<emphasis id="9FF1EA958B3A2C05FE3CFE1BB9263E83" box="[395,515,403,428]" italics="true" pageId="10" pageNumber="429">M. ngarae</emphasis>
</taxonomicName>
and 
<taxonomicName id="6A854D048B3A2C05FDF7FE1BB9FD3E82" authorityName="Miller &amp; Konings &amp; Stauffer" authorityYear="2021" box="[576,728,403,429]" class="Actinopterygii" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="429" phylum="Chordata" rank="species" species="gallireyae">
<emphasis id="9FF1EA958B3A2C05FDF7FE1BB9FD3E82" box="[576,728,403,429]" italics="true" pageId="10" pageNumber="429">M. gallireyae</emphasis>
</taxonomicName>
(
<figureCitation id="35BE2A028B3A2C05FD5CFE1BB8163E82" box="[747,819,403,429]" captionStart="FIGURE 8" captionStartId="11.[151,250,1779,1803]" captionTargetBox="[153,1419,568,1749]" captionTargetId="figure-223@11.[151,1436,561,1755]" captionTargetPageId="11" captionText="FIGURE 8. Sheared second principal components (morphometric data) plotted against the first principal components (meristic data) of 14 specimens of Metriaclima ngarae from Ngara (x) and 20 specimens of Metriaclima gallireyae from Gallireya Reef (+)." figureDoi="http://doi.org/10.5281/zenodo.5572234" httpUri="https://zenodo.org/record/5572234/files/figure.png" pageId="10" pageNumber="429">Fig. 8</figureCitation>
). The first principal component (size variable) of the morphometric data explained 87.8% of the observed variance and SPCA2 explained 5.4%. Variables that had the highest loadings on SPCA2 were the premaxillary pedicel length (0.89), snout length (-0.24), and cheek depth (- 0.21). The first principal component of the meristic data explained 24.7% of the variance. Variables with the highest loadings on the first principal components of the meristic data were pored lateral-line scales posterior to the hypural plate (0.50), number of tooth rows on the upper jaw (0.44), and tooth rows on the lower jaw (0.28).
</paragraph>
</subSubSection>
</treatment>
</document>