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<document id="F01EF21F62F8028D822F415BCC938C23" ID-CLB-Dataset="22296" ID-DOI="10.5852/ejt.2018.456" ID-GBIF-Dataset="0566a11e-d6a3-4283-89c7-c7feea96fdbf" ID-Zenodo-Dep="3817823" ID-ZooBank="DE1B1DEE-9871-4803-9F67-025F2B439560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588965964057" checkinUser="valdenar" docAuthor="Sørensen, Martin V., Rohal, Melissa &amp; Thistle, David" docDate="2018" docId="F64287A2507BFFAD1451FED5FD5F0B8F" docLanguage="en" docName="ejt-456_sorensen_rohal_thistle.pdf.imf" docOrigin="European Journal of Taxonomy 456" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Echinoderes lupherorum Sørensen &amp; Rohal &amp; Thistle 2018, sp. nov." docType="treatment" docUuid="B3E96033-83C3-4924-8FFE-3592E427620C" docUuidSource="ZooBank" docVersion="8" lastPageNumber="52" masterDocId="0A7BFFDA5056FF9E1600FFDEFFDE0C64" masterDocTitle="Deep-sea Echinoderidae (Kinorhyncha: Cyclorhagida) from the Northwest Pacific" masterLastPageNumber="75" masterPageNumber="1" pageNumber="46" updateTime="1698839057356" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="49CC4BD2581F3313ADC09FA362B98B80">Deep-sea Echinoderidae (Kinorhyncha: Cyclorhagida) from the Northwest Pacific</mods:title>
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<mods:namePart id="C5F63DACF785CE1850F965D0EC0EF0FF">Sørensen, Martin V.</mods:namePart>
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<mods:affiliation id="6F267289203B434F8CFE9CFA2E47B728">Section for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, DK- 1350 Copenhagen, Denmark. Harte Research Institute, Texas A &amp; M University - Corpus Christi, Corpus Christi, TX 78412, USA. Department of Earth, Ocean and Atmospheric Sciences, Florida State University, Tallahassee, FL 32306 - 4520, USA.</mods:affiliation>
<mods:nameIdentifier id="E77054816C6541CC05C79EE377C648C5" type="email">mvsorensen@snm.ku.dk</mods:nameIdentifier>
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<mods:namePart id="740FC82CB5AE50914D07F892E0B8A15B">Rohal, Melissa</mods:namePart>
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<mods:nameIdentifier id="7BE209A95314105E9901423025014D56" type="email">mrohal@islander.tamucc.edu</mods:nameIdentifier>
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<mods:namePart id="7A19C95FD5238BC592E4E8AEB40DACA6">Thistle, David</mods:namePart>
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<mods:nameIdentifier id="E2072892DDF35C5EE2432A29170624CD" type="email">dthistle@fsu.edu</mods:nameIdentifier>
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<mods:title id="A4F37CA967A31FD54B88879E3742C7C3">European Journal of Taxonomy</mods:title>
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<mods:date id="F099570E2C46D0AB66A9A37BEB5445B6">2018</mods:date>
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<treatment id="F64287A2507BFFAD1451FED5FD5F0B8F" ID-DOI="http://doi.org/10.5281/zenodo.3818840" ID-GBIF-Taxon="163993845" ID-Zenodo-Dep="3818840" LSID="urn:lsid:zoobank.org:act:B3E96033-83C3-4924-8FFE-3592E427620C" httpUri="http://treatment.plazi.org/id/F64287A2507BFFAD1451FED5FD5F0B8F" lastPageId="51" lastPageNumber="52" pageId="45" pageNumber="46">
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<paragraph id="7E5436B4507BFFB31451FED5FC3D0D42" blockId="45.[379,1208,267,365]" box="[593,995,267,294]" pageId="45" pageNumber="46">
<heading id="251C81D8507BFFB31451FED5FC3D0D42" box="[593,995,267,294]" centered="true" fontSize="11" level="2" pageId="45" pageNumber="46" reason="2">
<taxonomicName id="B9EB4D37507BFFB31451FED5FC5F0D42" authority="Sørensen &amp; Rohal &amp; Thistle, 2018" authorityName="Sørensen &amp; Rohal &amp; Thistle" authorityYear="2018" box="[593,897,267,294]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="45" pageNumber="46" phylum="Cephalorhyncha" rank="species" species="lupherorum" status="sp. nov.">
<emphasis id="4C9FEAA6507BFFB31451FED5FC5F0D42" bold="true" box="[593,897,267,294]" italics="true" pageId="45" pageNumber="46">Echinoderes lupherorum</emphasis>
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urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="36F1653F507BFFB31483FE8DFC6F0D09" box="[643,945,339,365]" pageId="45" pageNumber="46" type="description">
<paragraph id="7E5436B4507BFFB31483FE8DFC6F0D09" blockId="45.[379,1208,267,365]" box="[643,945,339,365]" pageId="45" pageNumber="46">
<figureCitation id="E6D02A31507BFFB31483FE8DFCD70D09" box="[643,777,339,365]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="46.[189,232,1754,1780]" captionStartId-1="47.[189,232,1647,1673]" captionStartId-2="48.[189,232,1647,1673]" captionTargetBox-0="[278,1310,269,1745]" captionTargetBox-1="[233,1360,265,1627]" captionTargetBox-2="[191,1396,382,1597]" captionTargetId-0="figure@46.[277,1310,265,1745]" captionTargetId-1="figure@47.[233,1360,265,1627]" captionTargetId-2="figure@48.[191,1396,382,1597]" captionTargetPageId-0="46" captionTargetPageId-1="47" captionTargetPageId-2="48" captionText-0="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." captionText-1="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." captionText-2="Fig. 20. Scanning electron micrographs showing overviews and details of Echinoderes lupherorum sp. nov. A. Lateroventral overview. B. Laterodorsal overview. C. Introvert sectors 10 and 9. D. Segments 1 to 3, ventral view. E. Segments 1 to 2, dorsal view. F. Detail showing middorsal (right) to laterodorsal (left) parts of segments 1 and 2. Inset shows close-up of subdorsal glandular cell outlet type 2; scale = 1 µm. G. Detail showing middorsal (right) to laterodorsal (left) parts of 8. H. Close-up showing laterodorsal sensory spot and glandular cell outlet type 2 on segment 8. I. Segments 9 to 10, lateral view. J. ♀, segments 10 to 11, dorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvt = lateroventral tube; mds = middorsal spine; mdss = middorsal sensory spot; mlt = midlateral tube; pdss = paradorsal sensory spot; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid." figureDoi-0="http://doi.org/10.5281/zenodo.3817853" figureDoi-1="http://doi.org/10.5281/zenodo.3817855" figureDoi-2="http://doi.org/10.5281/zenodo.3817857" httpUri-0="https://zenodo.org/record/3817853/files/figure.png" httpUri-1="https://zenodo.org/record/3817855/files/figure.png" httpUri-2="https://zenodo.org/record/3817857/files/figure.png" pageId="45" pageNumber="46">Figs 1820</figureCitation>
,
<tableCitation id="3369030F507BFFB31514FE8DFC6F0D09" box="[788,945,339,365]" captionStart-0="Table 12" captionStart-1="Table 13" captionStartId-0="49.[189,258,268,294]" captionStartId-1="50.[189,258,268,294]" captionTargetBox-0="[355,1197,515,1620]" captionTargetPageId-0="49" captionText-0="Table 12. Measurements from light microscopy of Echinoderes lupherorum sp. nov. (in µm), including number of measured specimens (n) and standard deviation (SD). Abbreviations: (ac) = acicular spine; LTAS = lateral terminal accessory spine; LTS = lateral terminal spine; LV = lateroventral; MD = middorsal; ML = midlateral; MSW-8 = maximum sternal width, measured on segment 8 in this species; S = segment lengths; SW-10 = standard width, always measured on segment 10; TE = tergal extension; TL = trunk length; (tu) = tube." captionText-1="Table 13. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes lupherorum sp. nov. Abbreviations: LA = lateral accessory; LD = laterodorsal; LV = lateroventral; MD = middorsal; ML = midlateral; PD = paradorsal; PV = paraventral; SD = subdorsal; SL = sublateral; VL = ventrolateral; VM = ventromedial; ac = acicular spine; gco1/2 = glandular cell outlet type 1/2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; pa = papilla; pe = penile spines; si = sieve plate; ss = sensory spot; tu = tube; (♀) = female and (♂) = male conditions of sexually dimorphic characters. * = indicates that the structure is not present in all specimens." pageId="45" pageNumber="46">Tables 1213</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="36F1653F507BFFB316BDFE4BFDB10ED3" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="7E5436B4507BFFB316BDFE4BFEE80DCB" blockId="45.[189,310,405,431]" box="[189,310,405,431]" pageId="45" pageNumber="46">
<heading id="251C81D8507BFFB316BDFE4BFEE80DCB" bold="true" box="[189,310,405,431]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="4C9FEAA6507BFFB316BDFE4BFEE80DCB" bold="true" box="[189,310,405,431]" pageId="45" pageNumber="46">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="7E5436B4507BFFB316BDFE1BFDB10ED3" blockId="45.[189,1399,453,696]" pageId="45" pageNumber="46">
<taxonomicName id="B9EB4D37507BFFB316BDFE1BFE910DBB" authorityName="Claparede" authorityYear="1863" box="[189,335,453,479]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="45" pageNumber="46" phylum="Cephalorhyncha" rank="genus">
<emphasis id="4C9FEAA6507BFFB316BDFE1BFE910DBB" box="[189,335,453,479]" italics="true" pageId="45" pageNumber="46">Echinoderes</emphasis>
</taxonomicName>
with middorsal spines on segments 4 to 8, and spines in lateroventral positions on segments 6 to 9. Tubes present in lateroventral positions on segments 2 and 5, and in midlateral positions of segment 10. Very minute glandular cell outlets
<typeStatus id="A1508816507BFFB314EDFDD0FCFE0E4C" box="[749,800,526,552]" pageId="45" pageNumber="46">type</typeStatus>
2 present in subdorsal positions on segment 2 and in laterodorsal positions on segments 8 and 9. Tergal plate of segment 11 terminates in pointed, acicular tergal extensions, constituting 6 to 8% of trunk length. Males with three pairs of penile spines; females with long lateral terminal accessory spines, and papillae in ventrolateral positions of segment 7 and ventromedial positions of segment 8.
</paragraph>
</subSubSection>
<subSubSection id="36F1653F507BFFB316BDFD01FE8C0F29" pageId="45" pageNumber="46" type="etymology">
<paragraph id="7E5436B4507BFFB316BDFD01FE9A0E9D" blockId="45.[189,324,735,761]" box="[189,324,735,761]" pageId="45" pageNumber="46">
<heading id="251C81D8507BFFB316BDFD01FE9A0E9D" bold="true" box="[189,324,735,761]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="4C9FEAA6507BFFB316BDFD01FE9A0E9D" bold="true" box="[189,324,735,761]" pageId="45" pageNumber="46">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="7E5436B4507BFFB316BDFCD1FE8C0F29" blockId="45.[189,1398,783,845]" pageId="45" pageNumber="46">The second author (MR) dedicates this species to her fiancé Brach Lupher and the Lupher family her family to be.</paragraph>
</subSubSection>
<subSubSection id="36F1653F507BFFB316BDFCABFDC60907" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="7E5436B4507BFFB316BDFCABFE740FEB" blockId="45.[189,426,885,911]" box="[189,426,885,911]" pageId="45" pageNumber="46">
<heading id="251C81D8507BFFB316BDFCABFE740FEB" bold="true" box="[189,426,885,911]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="4C9FEAA6507BFFB316BDFCABFE740FEB" bold="true" box="[189,426,885,911]" pageId="45" pageNumber="46">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="CE833CE9507BFFB316EAFC7AFB0D084E" ID-GBIF-Occurrence="2611490307" collectingDate="2008-09-17" collectionCode="US" collectorName="St. &amp; Fluoromount G &amp; See" country="United States of America" latitude="42.56389" location="West Coast" longLatPrecision="19" longitude="-131.99667" pageId="45" pageNumber="46" specimenCount="1" specimenCount-female="1" stateProvince="Oregon" typeStatus="holotype">
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<typeStatus id="A1508816507BFFB316EAFC7AFE820FDA" box="[234,348,932,958]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7E5436B4507BFFB316BDFC17FB0D084E" blockId="45.[189,1399,932,1067]" pageId="45" pageNumber="46">
<collectingCountry id="06FC7624507BFFB316BDFC17FDB50F86" box="[189,619,968,995]" name="United States of America" pageId="45" pageNumber="46">UNITED STATES OF AMERICA</collectingCountry>
: adult
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,
<collectionCode id="18FAAE71507BFFB314F1FC16FCC80F86" box="[753,790,968,994]" country="South Africa" name="University of Stellenbosch" pageId="45" pageNumber="46">US</collectionCode>
<location id="7B34606F507BFFB31524FC17FC6C0F86" LSID="urn:lsid:plazi:treatment:F64287A2507BFFAD1451FED5FD5F0B8F:7B34606F507BFFB31524FC17FC6C0F86" box="[804,946,968,995]" country="United States of America" latitude="42.56389" longLatPrecision="19" longitude="-131.99667" name="West Coast" pageId="45" pageNumber="46" stateProvince="Oregon">West Coast</location>
, off southern
<collectingRegion id="BC2FF856507BFFB31268FC16FB1D0F86" box="[1128,1219,968,994]" country="United States of America" name="Oregon" pageId="45" pageNumber="46">Oregon</collectingRegion>
,
<geoCoordinate id="1BDF5073507BFFB312D5FC18FAAC0F87" box="[1237,1394,966,995]" degrees="42" direction="north" minutes="33" orientation="latitude" pageId="45" pageNumber="46" precision="15" seconds="50" value="42.56389">42°3350″ N</geoCoordinate>
,
<geoCoordinate id="1BDF5073507BFFB316BDFC34FEBA0863" box="[189,356,1002,1031]" degrees="131" direction="west" minutes="59" orientation="longitude" pageId="45" pageNumber="46" precision="15" seconds="48" value="-131.99667">131°5948″ W</geoCoordinate>
,
<collectorName id="D31E5362507BFFB3176FFC32FE510862" box="[367,399,1004,1030]" pageId="45" pageNumber="46">St.</collectorName>
2, 3601 m deep, collected from mud,
<date id="0A551074507BFFB31549FC32FC3B0862" box="[841,997,1004,1030]" pageId="45" pageNumber="46" value="2008-09-17">
<collectingDate id="1A11E99C507BFFB31549FC32FC3B0862" box="[841,997,1004,1030]" pageId="45" pageNumber="46" value="2008-09-17">17 Sep. 2008</collectingDate>
</date>
, mounted in
<collectorName id="D31E5362507BFFB3127BFC32FAEE0862" box="[1147,1328,1004,1030]" pageId="45" pageNumber="46">Fluoromount G</collectorName>
on an H-S slide (NHMD-225220).
<collectorName id="D31E5362507BFFB31410FBCEFDE4084E" box="[528,570,1040,1066]" pageId="45" pageNumber="46">See</collectorName>
<figureCitation id="E6D02A31507BFFB31442FBCEFD58084E" box="[578,646,1040,1066]" captionStart="Fig" captionStartId="2.[189,232,2000,2026]" captionTargetBox="[241,1347,1019,1957]" captionTargetId="figure@2.[241,1347,1019,1958]" captionTargetPageId="2" captionText="Fig. 1. Map showing the eight sampling stations off the US West Coast. See Table 1 for further details." figureDoi="http://doi.org/10.5281/zenodo.3817825" httpUri="https://zenodo.org/record/3817825/files/figure.png" pageId="45" pageNumber="46">Fig. 1</figureCitation>
for localities and
<tableCitation id="3369030F507BFFB31559FBCEFC6E084E" box="[857,944,1040,1066]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionTargetBox="[202,1399,283,441]" captionTargetPageId="4" captionText="Table 1 (continued on next page). Summary of data on stations, species identities, and catalogue numbers." httpUri="http://table.plazi.org/id/2A94663C5055FF9D16BDFED2FAA90D41" pageId="45" pageNumber="46" tableUuid="2A94663C5055FF9D16BDFED2FAA90D41">Table 1</tableCitation>
for detailed station data.
</paragraph>
</materialsCitation>
<materialsCitation id="CE833CE9507BFFB316EAFB8DFED908BC" ID-GBIF-Occurrence="2611490309" pageId="45" pageNumber="46" typeStatus="paratype">
<paragraph id="7E5436B4507BFFB316EAFB8DFEB90809" blockId="45.[189,1399,1107,1240]" box="[234,359,1107,1133]" pageId="45" pageNumber="46">
<emphasis id="4C9FEAA6507BFFB316EAFB8DFEB90809" bold="true" box="[234,359,1107,1133]" pageId="45" pageNumber="46">
<typeStatus id="A1508816507BFFB316EAFB8DFEB90809" box="[234,359,1107,1133]" pageId="45" pageNumber="46" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="7E5436B4507BFFB316BDFBA9FED908BC" blockId="45.[189,1399,1107,1240]" pageId="45" pageNumber="46">
<collectingCountry id="06FC7624507BFFB316BDFBA9FD8D08F5" box="[189,595,1143,1169]" name="United States of America" pageId="45" pageNumber="46">UNITED STATES OF AMERICA</collectingCountry>
:
<specimenCount id="68EDFD3D507BFFB3145DFBA9FD4108F5" box="[605,671,1143,1169]" pageId="45" pageNumber="46" type="female">4 ♀♀</specimenCount>
, 1 ♂, same collecting data as for
<typeStatus id="A1508816507BFFB31220FBA8FB5408F4" box="[1056,1162,1142,1168]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
;
<specimenCount id="68EDFD3D507BFFB31295FBA9FB1E08F5" box="[1173,1216,1143,1169]" pageId="45" pageNumber="46" type="female">1 ♀</specimenCount>
, St. 3 (NHMD- 225221225226). All
<typeStatus id="A1508816507BFFB317C1FB45FDEF08D1" box="[449,561,1179,1205]" pageId="45" pageNumber="46" type="paratype">paratypes</typeStatus>
are mounted in Fluoromount G, three on H-S slides and three on glass slides.
</paragraph>
</materialsCitation>
<paragraph id="7E5436B4507BFFB316EAFADFFD89097F" blockId="45.[189,1399,1281,1379]" box="[234,599,1281,1307]" pageId="45" pageNumber="46">
<emphasis id="4C9FEAA6507BFFB316EAFADFFD89097F" bold="true" box="[234,599,1281,1307]" pageId="45" pageNumber="46">Additional non-type material</emphasis>
</paragraph>
<paragraph id="7E5436B4507BFFB316BDFAFBFDC60907" blockId="45.[189,1399,1281,1379]" pageId="45" pageNumber="46">
<collectingCountry id="06FC7624507BFFB316BDFAFBFD8A095B" box="[189,596,1317,1343]" name="United States of America" pageId="45" pageNumber="46">UNITED STATES OF AMERICA</collectingCountry>
:
<specimenCount id="68EDFD3D507BFFB3145EFAFBFD7F095B" box="[606,673,1317,1343]" pageId="45" pageNumber="46" type="female">3 ♀♀</specimenCount>
, 4 ♂♂, St. 6, mounted for SEM and stored in the first authors personal reference collection.
</paragraph>
</subSubSection>
<subSubSection id="36F1653F507BFFAD16BDFA55FAC70B31" lastPageId="51" lastPageNumber="52" pageId="45" pageNumber="46" type="description">
<paragraph id="7E5436B4507BFFB316BDFA55FE9009C1" blockId="45.[189,334,1419,1445]" box="[189,334,1419,1445]" pageId="45" pageNumber="46">
<heading id="251C81D8507BFFB316BDFA55FE9009C1" bold="true" box="[189,334,1419,1445]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="4C9FEAA6507BFFB316BDFA55FE9009C1" bold="true" box="[189,334,1419,1445]" pageId="45" pageNumber="46">Description</emphasis>
</heading>
</paragraph>
<paragraph id="7E5436B4507BFFB316BDFA64FB750AEC" blockId="45.[189,1399,1466,1673]" pageId="45" pageNumber="46">
Adults with head, neck and eleven trunk segments (
<figureCitation id="E6D02A31507BFFB31520FA64FCA709B0" box="[800,889,1466,1492]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="45" pageNumber="46">Figs 18</figureCitation>
AB, 19A, 20AB). The trunk is relatively large, appears slender, tapered in both ends, and broadest from segment 5 to 8. Lateral terminal spines are long and slender (
<figureCitation id="E6D02A31507BFFB3178CF9DCFE2B0A78" box="[396,501,1538,1564]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="45" pageNumber="46">Fig. 18C</figureCitation>
), from 80% and up to same length as the trunk, and lateral terminal accessory spines in females are shorter (
<figureCitation id="E6D02A31507BFFB3142EF9F8FD570A24" box="[558,649,1574,1600]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="45" pageNumber="46">Figs 18</figureCitation>
AB), around 20% of the lateral terminal spine length. For a complete overview of measurements and dimensions, see
<tableCitation id="3369030F507BFFB31566F994FC100A00" box="[870,974,1610,1636]" captionStart="Table 12" captionStartId="49.[189,258,268,294]" captionTargetBox="[355,1197,515,1620]" captionTargetPageId="49" captionText="Table 12. Measurements from light microscopy of Echinoderes lupherorum sp. nov. (in µm), including number of measured specimens (n) and standard deviation (SD). Abbreviations: (ac) = acicular spine; LTAS = lateral terminal accessory spine; LTS = lateral terminal spine; LV = lateroventral; MD = middorsal; ML = midlateral; MSW-8 = maximum sternal width, measured on segment 8 in this species; S = segment lengths; SW-10 = standard width, always measured on segment 10; TE = tergal extension; TL = trunk length; (tu) = tube." pageId="45" pageNumber="46">Table 12</tableCitation>
. Distribution of cuticular structures, i.e., sensory spots, glandular cell outlets, spines and tubes, is summarized in
<tableCitation id="3369030F507BFFB3123EF9B0FB780AEC" box="[1086,1190,1646,1672]" captionStart="Table 13" captionStartId="50.[189,258,268,294]" captionText="Table 13. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes lupherorum sp. nov. Abbreviations: LA = lateral accessory; LD = laterodorsal; LV = lateroventral; MD = middorsal; ML = midlateral; PD = paradorsal; PV = paraventral; SD = subdorsal; SL = sublateral; VL = ventrolateral; VM = ventromedial; ac = acicular spine; gco1/2 = glandular cell outlet type 1/2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; pa = papilla; pe = penile spines; si = sieve plate; ss = sensory spot; tu = tube; (♀) = female and (♂) = male conditions of sexually dimorphic characters. * = indicates that the structure is not present in all specimens." pageId="45" pageNumber="46">Table 13</tableCitation>
.
</paragraph>
<paragraph id="7E5436B4507BFFAF16BDF96FFEB00ABF" blockId="45.[189,1399,1713,2027]" lastBlockId="49.[189,1398,1693,1755]" lastPageId="49" lastPageNumber="50" pageId="45" pageNumber="46">
The head consists of a retractable mouth cone and an introvert. Inner oral styles are present, but their exact number and arrangement could not be determined. The external mouth cone armature consists of nine outer oral styles, each with four or five basal fringe tips. The introvert sectors are defined by ten primary spinoscalids in Ring 01. Each primary spinoscalid consists of a basal sheath and a distal end piece with a blunt tip (
<figureCitation id="E6D02A31507BFFB31400F89FFDB00B3F" box="[512,622,1857,1883]" captionStart="Fig" captionStartId="48.[189,232,1647,1673]" captionTargetBox="[191,1396,382,1597]" captionTargetId="figure@48.[191,1396,382,1597]" captionTargetPageId="48" captionText="Fig. 20. Scanning electron micrographs showing overviews and details of Echinoderes lupherorum sp. nov. A. Lateroventral overview. B. Laterodorsal overview. C. Introvert sectors 10 and 9. D. Segments 1 to 3, ventral view. E. Segments 1 to 2, dorsal view. F. Detail showing middorsal (right) to laterodorsal (left) parts of segments 1 and 2. Inset shows close-up of subdorsal glandular cell outlet type 2; scale = 1 µm. G. Detail showing middorsal (right) to laterodorsal (left) parts of 8. H. Close-up showing laterodorsal sensory spot and glandular cell outlet type 2 on segment 8. I. Segments 9 to 10, lateral view. J. ♀, segments 10 to 11, dorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvt = lateroventral tube; mds = middorsal spine; mdss = middorsal sensory spot; mlt = midlateral tube; pdss = paradorsal sensory spot; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid." figureDoi="http://doi.org/10.5281/zenodo.3817857" httpUri="https://zenodo.org/record/3817857/files/figure.png" pageId="45" pageNumber="46">Fig. 20C</figureCitation>
). The basal sheaths have marginal extensions forming two layers of transverse fringes. End pieces are smooth. The arrangement of scalids follows the same pattern as in
<taxonomicName id="B9EB4D37507BFFB316DEF857FE2D0BC7" authority="Sørensen &amp; Rohal &amp; Thistle, 2018" authorityName="Sørensen &amp; Rohal &amp; Thistle" authorityYear="2018" box="[222,499,1929,1955]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="45" pageNumber="46" phylum="Cephalorhyncha" rank="species" species="hviidarum" status="sp. nov.">
<emphasis id="4C9FEAA6507BFFB316DEF857FE2D0BC7" box="[222,499,1929,1955]" italics="true" pageId="45" pageNumber="46">Echinoderes hviidarum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57AC57DD507BFFB317FDF857FD840BC7" box="[509,602,1929,1955]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
described above; hence, see
<figureCitation id="E6D02A31507BFFB315BDF857FBCC0BC7" box="[957,1042,1929,1955]" captionStart="Fig" captionStartId="30.[189,232,1930,1956]" captionTargetBox="[201,1388,269,1911]" captionTargetId="figure@30.[200,1389,265,1912]" captionTargetPageId="30" captionText="Fig. 11. Diagram of mouth cone (grey area), introvert and placids in Echinoderes hviidarum sp. nov., showing distribution of outer oral styles, scalids and trichoscalids. Table shows the scalid arrangement by sector; single-lined boxes mark quincunxes, double-lined boxes mark “double diamonds”." figureDoi="http://doi.org/10.5281/zenodo.3817845" httpUri="https://zenodo.org/record/3817845/files/figure.png" pageId="45" pageNumber="46">Fig. 11</figureCitation>
for a complete overview. All spinoscalids consist of a basal sheath and a pointed end piece. The basal sheaths terminate in fringed margins in spinoscalids of Rings 02 and 04, whereas the sheaths of Ring 03 have a median spike only. Spinoscalids of Rings 05 to 07 are composed as those in the preceding ring, but with shorter end pieces (
<figureCitation id="E6D02A315067FFAF16C5F91FFEC10ABF" box="[197,287,1729,1755]" captionStart="Fig" captionStartId="30.[189,232,1930,1956]" captionTargetBox="[201,1388,269,1911]" captionTargetId="figure@30.[200,1389,265,1912]" captionTargetPageId="30" captionText="Fig. 11. Diagram of mouth cone (grey area), introvert and placids in Echinoderes hviidarum sp. nov., showing distribution of outer oral styles, scalids and trichoscalids. Table shows the scalid arrangement by sector; single-lined boxes mark quincunxes, double-lined boxes mark “double diamonds”." figureDoi="http://doi.org/10.5281/zenodo.3817845" httpUri="https://zenodo.org/record/3817845/files/figure.png" pageId="49" pageNumber="50">Figs 11</figureCitation>
,
<figureCitation id="E6D02A315067FFAF172CF91FFEBF0ABF" box="[300,353,1729,1755]" captionStart="Fig" captionStartId="48.[189,232,1647,1673]" captionTargetBox="[191,1396,382,1597]" captionTargetId="figure@48.[191,1396,382,1597]" captionTargetPageId="48" captionText="Fig. 20. Scanning electron micrographs showing overviews and details of Echinoderes lupherorum sp. nov. A. Lateroventral overview. B. Laterodorsal overview. C. Introvert sectors 10 and 9. D. Segments 1 to 3, ventral view. E. Segments 1 to 2, dorsal view. F. Detail showing middorsal (right) to laterodorsal (left) parts of segments 1 and 2. Inset shows close-up of subdorsal glandular cell outlet type 2; scale = 1 µm. G. Detail showing middorsal (right) to laterodorsal (left) parts of 8. H. Close-up showing laterodorsal sensory spot and glandular cell outlet type 2 on segment 8. I. Segments 9 to 10, lateral view. J. ♀, segments 10 to 11, dorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvt = lateroventral tube; mds = middorsal spine; mdss = middorsal sensory spot; mlt = midlateral tube; pdss = paradorsal sensory spot; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid." figureDoi="http://doi.org/10.5281/zenodo.3817857" httpUri="https://zenodo.org/record/3817857/files/figure.png" pageId="49" pageNumber="50">20C</figureCitation>
).
</paragraph>
<caption id="2A94663C5078FFB016BDF904FCC60B8E" ID-DOI="http://doi.org/10.5281/zenodo.3817853" ID-Zenodo-Dep="3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="46" pageNumber="47" startId="46.[189,232,1754,1780]" targetBox="[278,1310,269,1745]" targetPageId="46">
<paragraph id="7E5436B45078FFB016BDF904FCC60B8E" blockId="46.[189,1399,1754,2027]" pageId="46" pageNumber="47">
<emphasis id="4C9FEAA65078FFB016BDF904FEC70A90" bold="true" box="[189,281,1754,1780]" pageId="46" pageNumber="47">Fig. 18.</emphasis>
Line art illustrations of
<taxonomicName id="B9EB4D375078FFB01433F904FC8B0A90" authority="Sørensen &amp; Rohal &amp; Thistle, 2018" authorityName="Sørensen &amp; Rohal &amp; Thistle" authorityYear="2018" box="[563,853,1754,1780]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="46" pageNumber="47" phylum="Cephalorhyncha" rank="species" species="lupherorum" status="sp. nov.">
<emphasis id="4C9FEAA65078FFB01433F904FC8B0A90" box="[563,853,1754,1780]" italics="true" pageId="46" pageNumber="47">Echinoderes lupherorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57AC57DD5078FFB0155CF904FC690A90" box="[860,951,1754,1780]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="4C9FEAA65078FFB015BDF904FC0C0A90" bold="true" box="[957,978,1754,1780]" pageId="46" pageNumber="47">A</emphasis>
. ♀, dorsal view.
<emphasis id="4C9FEAA65078FFB01299F904FB730A90" bold="true" box="[1177,1197,1754,1780]" pageId="46" pageNumber="47">B</emphasis>
. ♀, ventral view.
<emphasis id="4C9FEAA65078FFB016BDF923FF0C0B73" bold="true" box="[189,210,1789,1815]" pageId="46" pageNumber="47">C</emphasis>
. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view.
<emphasis id="4C9FEAA65078FFB012C0F923FB0B0B73" bold="true" box="[1216,1237,1789,1815]" pageId="46" pageNumber="47">D</emphasis>
. ♂, segments 10 to 11, dorsal view.
<emphasis id="4C9FEAA65078FFB017D1F8FFFE3B0B5F" bold="true" box="[465,485,1825,1851]" pageId="46" pageNumber="47">E</emphasis>
. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension.
</paragraph>
</caption>
<caption id="2A94663C5079FFB116BDF9B1FF280B8D" ID-DOI="http://doi.org/10.5281/zenodo.3817855" ID-Zenodo-Dep="3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="47" pageNumber="48" startId="47.[189,232,1647,1673]" targetBox="[233,1360,265,1627]" targetPageId="47">
<paragraph id="7E5436B45079FFB116BDF9B1FF280B8D" blockId="47.[189,1399,1647,2025]" pageId="47" pageNumber="48">
<emphasis id="4C9FEAA65079FFB116BDF9B1FEFF0AED" bold="true" box="[189,289,1647,1673]" pageId="47" pageNumber="48">Fig. 19.</emphasis>
Light micrographs showing overviews and details of
<taxonomicName id="B9EB4D375079FFB115D5F9B1FB200AED" authority="Sørensen &amp; Rohal &amp; Thistle, 2018" authorityName="Sørensen &amp; Rohal &amp; Thistle" authorityYear="2018" box="[981,1278,1647,1673]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="47" pageNumber="48" phylum="Cephalorhyncha" rank="species" species="lupherorum" status="sp. nov.">
<emphasis id="4C9FEAA65079FFB115D5F9B1FB200AED" box="[981,1278,1647,1673]" italics="true" pageId="47" pageNumber="48">Echinoderes lupherorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57AC57DD5079FFB1130DF9B1FAAC0AED" box="[1293,1394,1647,1673]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225).
<emphasis id="4C9FEAA65079FFB1179DF96BFE6C0AAB" bold="true" box="[413,434,1717,1743]" pageId="47" pageNumber="48">A</emphasis>
. Ventral overview.
<emphasis id="4C9FEAA65079FFB1149CF96BFD6E0AAB" bold="true" box="[668,688,1717,1743]" pageId="47" pageNumber="48">B</emphasis>
. Segments 1 to 3, dorsal view.
<emphasis id="4C9FEAA65079FFB1122CF96BFB9F0AAB" bold="true" box="[1068,1089,1717,1743]" pageId="47" pageNumber="48">C</emphasis>
. Segments 1 to 3, ventral view.
<emphasis id="4C9FEAA65079FFB11702F907FEC90A97" bold="true" box="[258,279,1753,1779]" pageId="47" pageNumber="48">D</emphasis>
. Segments 3 to 6, dorsal view.
<emphasis id="4C9FEAA65079FFB1147EF907FD4C0A97" bold="true" box="[638,658,1753,1779]" pageId="47" pageNumber="48">E</emphasis>
. Segments 3 to 6, ventral view.
<emphasis id="4C9FEAA65079FFB11203F907FBCB0A97" bold="true" box="[1027,1045,1753,1779]" pageId="47" pageNumber="48">F</emphasis>
. Segments 8 to 9, dorsal view.
<emphasis id="4C9FEAA65079FFB116BDF925FF0A0B71" bold="true" box="[189,212,1787,1813]" pageId="47" pageNumber="48">G</emphasis>
. Segments 7 to 8, ventral view.
<emphasis id="4C9FEAA65079FFB11449F922FDBE0B72" bold="true" box="[585,608,1788,1814]" pageId="47" pageNumber="48">H</emphasis>
. ♀, segments 10 to 11, ventral view.
<emphasis id="4C9FEAA65079FFB11210F922FBC50B72" bold="true" box="[1040,1051,1788,1814]" pageId="47" pageNumber="48">I</emphasis>
. ♂, segments 10 to 11, dorsal view.
<emphasis id="4C9FEAA65079FFB11702F8C1FECF0B5D" bold="true" box="[258,273,1823,1849]" pageId="47" pageNumber="48">J</emphasis>
. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot.
</paragraph>
</caption>
<caption id="2A94663C5066FFAE16BDF9B1FB6A0B8D" ID-DOI="http://doi.org/10.5281/zenodo.3817857" ID-Zenodo-Dep="3817857" httpUri="https://zenodo.org/record/3817857/files/figure.png" pageId="48" pageNumber="49" startId="48.[189,232,1647,1673]" targetBox="[191,1396,382,1597]" targetPageId="48">
<paragraph id="7E5436B45066FFAE16BDF9B1FB6A0B8D" blockId="48.[189,1399,1647,2025]" pageId="48" pageNumber="49">
<emphasis id="4C9FEAA65066FFAE16BDF9B1FEC30AED" bold="true" box="[189,285,1647,1673]" pageId="48" pageNumber="49">Fig. 20.</emphasis>
Scanning electron micrographs showing overviews and details of
<taxonomicName id="B9EB4D375066FFAE1250F9B1FAA80AED" authority="Sørensen &amp; Rohal &amp; Thistle, 2018" authorityName="Sørensen &amp; Rohal &amp; Thistle" authorityYear="2018" box="[1104,1398,1647,1673]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="48" pageNumber="49" phylum="Cephalorhyncha" rank="species" species="lupherorum" status="sp. nov.">
<emphasis id="4C9FEAA65066FFAE1250F9B1FAA80AED" box="[1104,1398,1647,1673]" italics="true" pageId="48" pageNumber="49">Echinoderes lupherorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57AC57DD5066FFAE16BDF94DFEC80AC9" box="[189,278,1683,1709]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="4C9FEAA65066FFAE171BF94CFEEE0AC8" bold="true" box="[283,304,1682,1708]" pageId="48" pageNumber="49">A</emphasis>
. Lateroventral overview.
<emphasis id="4C9FEAA65066FFAE145AF94CFDB00AC8" bold="true" box="[602,622,1682,1708]" pageId="48" pageNumber="49">B</emphasis>
. Laterodorsal overview.
<emphasis id="4C9FEAA65066FFAE158DF94CFC7C0AC8" bold="true" box="[909,930,1682,1708]" pageId="48" pageNumber="49">C</emphasis>
. Introvert sectors 10 and 9.
<emphasis id="4C9FEAA65066FFAE12E2F94CFB290AC8" bold="true" box="[1250,1271,1682,1708]" pageId="48" pageNumber="49">D</emphasis>
. Segments 1 to 3, ventral view.
<emphasis id="4C9FEAA65066FFAE17AAF96BFE600AAB" bold="true" box="[426,446,1717,1743]" pageId="48" pageNumber="49">E</emphasis>
. Segments 1 to 2, dorsal view.
<emphasis id="4C9FEAA65066FFAE1528F96BFCE40AAB" bold="true" box="[808,826,1717,1743]" pageId="48" pageNumber="49">F</emphasis>
. Detail showing middorsal (right) to laterodorsal (left) parts of segments 1 and 2. Inset shows close-up of subdorsal glandular cell outlet type 2; scale = 1 µm.
<emphasis id="4C9FEAA65066FFAE172DF925FE9A0B71" bold="true" box="[301,324,1787,1813]" pageId="48" pageNumber="49">G</emphasis>
. Detail showing middorsal (right) to laterodorsal (left) parts of 8.
<emphasis id="4C9FEAA65066FFAE1275F922FB520B72" bold="true" box="[1141,1164,1788,1814]" pageId="48" pageNumber="49">H</emphasis>
. Close-up showing laterodorsal sensory spot and glandular cell outlet type 2 on segment 8.
<emphasis id="4C9FEAA65066FFAE15FAF8C1FBDB0B5D" bold="true" box="[1018,1029,1823,1849]" pageId="48" pageNumber="49">I</emphasis>
. Segments 9 to 10, lateral view.
<emphasis id="4C9FEAA65066FFAE16BDF89CFF120B38" bold="true" box="[189,204,1858,1884]" pageId="48" pageNumber="49">J</emphasis>
. ♀, segments 10 to 11, dorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvt = lateroventral tube; mds = middorsal spine; mdss = middorsal sensory spot; mlt = midlateral tube; pdss = paradorsal sensory spot; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid.
</paragraph>
</caption>
<caption id="2A94663C5067FFAF16BDFED2FDFF0DB1" pageId="49" pageNumber="50" targetBox="[355,1197,515,1620]" targetIsTable="true" targetPageId="49">
<paragraph id="7E5436B45067FFAF16BDFED2FDFF0DB1" blockId="49.[189,1399,267,470]" pageId="49" pageNumber="50">
<emphasis id="4C9FEAA65067FFAF16BDFED2FEF00D41" bold="true" box="[189,302,267,294]" pageId="49" pageNumber="50">Table 12.</emphasis>
Measurements from light microscopy of
<taxonomicName id="B9EB4D375067FFAF1518FED5FBE70D41" authority="Sørensen &amp; Rohal &amp; Thistle, 2018" authorityName="Sørensen &amp; Rohal &amp; Thistle" authorityYear="2018" box="[792,1081,267,293]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="49" pageNumber="50" phylum="Cephalorhyncha" rank="species" species="lupherorum" status="sp. nov.">
<emphasis id="4C9FEAA65067FFAF1518FED5FBE70D41" box="[792,1081,267,293]" italics="true" pageId="49" pageNumber="50">Echinoderes lupherorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57AC57DD5067FFAF1240FED2FB440D42" box="[1088,1178,268,294]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
(in µm), including number of measured specimens (n) and standard deviation (SD). Abbreviations: (ac) = acicular spine; LTAS = lateral terminal accessory spine; LTS = lateral terminal spine; LV = lateroventral; MD = middorsal; ML = midlateral; MSW-8 = maximum sternal width, measured on segment 8 in this species; S = segment lengths; SW-10 = standard width, always measured on segment 10; TE = tergal extension; TL = trunk length; (tu) = tube.
</paragraph>
</caption>
<paragraph id="7E5436B45067FFAF1766FDDDFB7F0A30" pageId="49" pageNumber="50">
<table id="0CEBC414506700611763FDDDFB730A30" box="[355,1197,515,1620]" colsContinueFrom="41.[358,1195,473,1431]" colsContinueIn="55.[358,1195,484,1531]" gridcols="5" gridrows="32" pageId="49" pageNumber="50">
<tr id="C0DB34F6506700611763FDDDFB730E7A" box="[355,1197,515,542]" gridrow="0" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FDDDFE380E7A" box="[355,486,515,542]" gridcol="0" gridrow="0" pageId="49" pageNumber="50">
<emphasis id="4C9FEAA65067FFAF1766FDDDFE3F0E7A" bold="true" box="[358,481,515,542]" pageId="49" pageNumber="50">Character</emphasis>
</th>
<th id="830A5D8A50670061143AFDDDFD940E7A" box="[570,586,515,542]" gridcol="1" gridrow="0" pageId="49" pageNumber="50">
<emphasis id="4C9FEAA65067FFAF143AFDDAFD940E7A" bold="true" box="[570,586,516,542]" pageId="49" pageNumber="50">n</emphasis>
</th>
<th id="830A5D8A506700611496FDDDFCF50E7A" box="[662,811,515,542]" gridcol="2" gridrow="0" pageId="49" pageNumber="50">
<emphasis id="4C9FEAA65067FFAF14BBFDDAFCD90E7A" bold="true" box="[699,775,516,542]" pageId="49" pageNumber="50">Range</emphasis>
</th>
<th id="830A5D8A506700611590FDDDFC070E7A" box="[912,985,515,542]" gridcol="3" gridrow="0" pageId="49" pageNumber="50">
<emphasis id="4C9FEAA65067FFAF1592FDDAFC080E7A" bold="true" box="[914,982,516,542]" pageId="49" pageNumber="50">Mean</emphasis>
</th>
<th id="830A5D8A506700611264FDDDFB730E7A" box="[1124,1197,515,542]" gridcol="4" gridrow="0" pageId="49" pageNumber="50">
<emphasis id="4C9FEAA65067FFAF1276FDDDFB440E7A" bold="true" box="[1142,1178,515,542]" pageId="49" pageNumber="50">SD</emphasis>
</th>
</tr>
<tr id="C0DB34F6506700611763FDF2FB730E2C" box="[355,1197,556,584]" gridrow="1" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FDF2FE380E2C" box="[355,486,556,584]" gridcol="0" gridrow="1" pageId="49" pageNumber="50">TL</th>
<td id="830A5D8A50670061143AFDF2FD940E2C" box="[570,586,556,584]" gridcol="1" gridrow="1" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FDF2FCF50E2C" box="[662,811,556,584]" gridcol="2" gridrow="1" pageId="49" pageNumber="50">331415</td>
<td id="830A5D8A506700611590FDF2FC070E2C" box="[912,985,556,584]" gridcol="3" gridrow="1" pageId="49" pageNumber="50">371</td>
<td id="830A5D8A506700611264FDF2FB730E2C" box="[1124,1197,556,584]" gridcol="4" gridrow="1" pageId="49" pageNumber="50">36.42</td>
</tr>
<tr id="C0DB34F6506700611763FD90FB730E0D" box="[355,1197,590,617]" gridrow="2" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FD90FE380E0D" box="[355,486,590,617]" gridcol="0" gridrow="2" pageId="49" pageNumber="50">MSW-8</th>
<td id="830A5D8A50670061143AFD90FD940E0D" box="[570,586,590,617]" gridcol="1" gridrow="2" pageId="49" pageNumber="50">6</td>
<td id="830A5D8A506700611496FD90FCF50E0D" box="[662,811,590,617]" gridcol="2" gridrow="2" pageId="49" pageNumber="50">7581</td>
<td id="830A5D8A506700611590FD90FC070E0D" box="[912,985,590,617]" gridcol="3" gridrow="2" pageId="49" pageNumber="50">78</td>
<td id="830A5D8A506700611264FD90FB730E0D" box="[1124,1197,590,617]" gridcol="4" gridrow="2" pageId="49" pageNumber="50">2.16</td>
</tr>
<tr id="C0DB34F6506700611763FDAEFB730EEF" box="[355,1197,624,651]" gridrow="3" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FDAEFE380EEF" box="[355,486,624,651]" gridcol="0" gridrow="3" pageId="49" pageNumber="50">MSW-8/TL</th>
<td id="830A5D8A50670061143AFDAEFD940EEF" box="[570,586,624,651]" gridcol="1" gridrow="3" pageId="49" pageNumber="50">6</td>
<td id="830A5D8A506700611496FDAEFCF50EEF" box="[662,811,624,651]" gridcol="2" gridrow="3" pageId="49" pageNumber="50">19.323.9%</td>
<td id="830A5D8A506700611590FDAEFC070EEF" box="[912,985,624,651]" gridcol="3" gridrow="3" pageId="49" pageNumber="50">21.0%</td>
<td id="830A5D8A506700611264FDAEFB730EEF" box="[1124,1197,624,651]" gridcol="4" gridrow="3" pageId="49" pageNumber="50">1.83%</td>
</tr>
<tr id="C0DB34F6506700611763FD4FFB730EC8" box="[355,1197,657,684]" gridrow="4" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FD4FFE380EC8" box="[355,486,657,684]" gridcol="0" gridrow="4" pageId="49" pageNumber="50">SW-10</th>
<td id="830A5D8A50670061143AFD4FFD940EC8" box="[570,586,657,684]" gridcol="1" gridrow="4" pageId="49" pageNumber="50">6</td>
<td id="830A5D8A506700611496FD4FFCF50EC8" box="[662,811,657,684]" gridcol="2" gridrow="4" pageId="49" pageNumber="50">6571</td>
<td id="830A5D8A506700611590FD4FFC070EC8" box="[912,985,657,684]" gridcol="3" gridrow="4" pageId="49" pageNumber="50">67</td>
<td id="830A5D8A506700611264FD4FFB730EC8" box="[1124,1197,657,684]" gridcol="4" gridrow="4" pageId="49" pageNumber="50">2.58</td>
</tr>
<tr id="C0DB34F6506700611763FD6DFB730EAA" box="[355,1197,691,718]" gridrow="5" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FD6DFE380EAA" box="[355,486,691,718]" gridcol="0" gridrow="5" pageId="49" pageNumber="50">SW-10/TL</th>
<td id="830A5D8A50670061143AFD6DFD940EAA" box="[570,586,691,718]" gridcol="1" gridrow="5" pageId="49" pageNumber="50">6</td>
<td id="830A5D8A506700611496FD6DFCF50EAA" box="[662,811,691,718]" gridcol="2" gridrow="5" pageId="49" pageNumber="50">15.720.2%</td>
<td id="830A5D8A506700611590FD6DFC070EAA" box="[912,985,691,718]" gridcol="3" gridrow="5" pageId="49" pageNumber="50">18.1%</td>
<td id="830A5D8A506700611264FD6DFB730EAA" box="[1124,1197,691,718]" gridcol="4" gridrow="5" pageId="49" pageNumber="50">1.77%</td>
</tr>
<tr id="C0DB34F6506700611763FD0AFB730E94" box="[355,1197,724,752]" gridrow="6" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FD0AFE380E94" box="[355,486,724,752]" gridcol="0" gridrow="6" pageId="49" pageNumber="50">S1</th>
<td id="830A5D8A50670061143AFD0AFD940E94" box="[570,586,724,752]" gridcol="1" gridrow="6" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FD0AFCF50E94" box="[662,811,724,752]" gridcol="2" gridrow="6" pageId="49" pageNumber="50">3542</td>
<td id="830A5D8A506700611590FD0AFC070E94" box="[912,985,724,752]" gridcol="3" gridrow="6" pageId="49" pageNumber="50">39</td>
<td id="830A5D8A506700611264FD0AFB730E94" box="[1124,1197,724,752]" gridcol="4" gridrow="6" pageId="49" pageNumber="50">2.54</td>
</tr>
<tr id="C0DB34F6506700611763FD28FB730F75" box="[355,1197,758,785]" gridrow="7" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FD28FE380F75" box="[355,486,758,785]" gridcol="0" gridrow="7" pageId="49" pageNumber="50">S2</th>
<td id="830A5D8A50670061143AFD28FD940F75" box="[570,586,758,785]" gridcol="1" gridrow="7" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FD28FCF50F75" box="[662,811,758,785]" gridcol="2" gridrow="7" pageId="49" pageNumber="50">3337</td>
<td id="830A5D8A506700611590FD28FC070F75" box="[912,985,758,785]" gridcol="3" gridrow="7" pageId="49" pageNumber="50">35</td>
<td id="830A5D8A506700611264FD28FB730F75" box="[1124,1197,758,785]" gridcol="4" gridrow="7" pageId="49" pageNumber="50">1.46</td>
</tr>
<tr id="C0DB34F6506700611763FCC6FB730F57" box="[355,1197,792,819]" gridrow="8" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FCC6FE380F57" box="[355,486,792,819]" gridcol="0" gridrow="8" pageId="49" pageNumber="50">S3</th>
<td id="830A5D8A50670061143AFCC6FD940F57" box="[570,586,792,819]" gridcol="1" gridrow="8" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FCC6FCF50F57" box="[662,811,792,819]" gridcol="2" gridrow="8" pageId="49" pageNumber="50">3943</td>
<td id="830A5D8A506700611590FCC6FC070F57" box="[912,985,792,819]" gridcol="3" gridrow="8" pageId="49" pageNumber="50">41</td>
<td id="830A5D8A506700611264FCC6FB730F57" box="[1124,1197,792,819]" gridcol="4" gridrow="8" pageId="49" pageNumber="50">1.50</td>
</tr>
<tr id="C0DB34F6506700611763FCE7FB730F31" box="[355,1197,825,853]" gridrow="9" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FCE7FE380F31" box="[355,486,825,853]" gridcol="0" gridrow="9" pageId="49" pageNumber="50">S4</th>
<td id="830A5D8A50670061143AFCE7FD940F31" box="[570,586,825,853]" gridcol="1" gridrow="9" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FCE7FCF50F31" box="[662,811,825,853]" gridcol="2" gridrow="9" pageId="49" pageNumber="50">4247</td>
<td id="830A5D8A506700611590FCE7FC070F31" box="[912,985,825,853]" gridcol="3" gridrow="9" pageId="49" pageNumber="50">45</td>
<td id="830A5D8A506700611264FCE7FB730F31" box="[1124,1197,825,853]" gridcol="4" gridrow="9" pageId="49" pageNumber="50">1.90</td>
</tr>
<tr id="C0DB34F6506700611763FC85FB730F12" box="[355,1197,859,886]" gridrow="10" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FC85FE380F12" box="[355,486,859,886]" gridcol="0" gridrow="10" pageId="49" pageNumber="50">S5</th>
<td id="830A5D8A50670061143AFC85FD940F12" box="[570,586,859,886]" gridcol="1" gridrow="10" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FC85FCF50F12" box="[662,811,859,886]" gridcol="2" gridrow="10" pageId="49" pageNumber="50">4252</td>
<td id="830A5D8A506700611590FC85FC070F12" box="[912,985,859,886]" gridcol="3" gridrow="10" pageId="49" pageNumber="50">48</td>
<td id="830A5D8A506700611264FC85FB730F12" box="[1124,1197,859,886]" gridcol="4" gridrow="10" pageId="49" pageNumber="50">3.78</td>
</tr>
<tr id="C0DB34F6506700611763FCA2FB730FFC" box="[355,1197,892,920]" gridrow="11" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FCA2FE380FFC" box="[355,486,892,920]" gridcol="0" gridrow="11" pageId="49" pageNumber="50">S6</th>
<td id="830A5D8A50670061143AFCA2FD940FFC" box="[570,586,892,920]" gridcol="1" gridrow="11" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FCA2FCF50FFC" box="[662,811,892,920]" gridcol="2" gridrow="11" pageId="49" pageNumber="50">5057</td>
<td id="830A5D8A506700611590FCA2FC070FFC" box="[912,985,892,920]" gridcol="3" gridrow="11" pageId="49" pageNumber="50">54</td>
<td id="830A5D8A506700611264FCA2FB730FFC" box="[1124,1197,892,920]" gridcol="4" gridrow="11" pageId="49" pageNumber="50">2.70</td>
</tr>
<tr id="C0DB34F6506700611763FC40FB730FDD" box="[355,1197,926,953]" gridrow="12" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FC40FE380FDD" box="[355,486,926,953]" gridcol="0" gridrow="12" pageId="49" pageNumber="50">S7</th>
<td id="830A5D8A50670061143AFC40FD940FDD" box="[570,586,926,953]" gridcol="1" gridrow="12" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FC40FCF50FDD" box="[662,811,926,953]" gridcol="2" gridrow="12" pageId="49" pageNumber="50">5460</td>
<td id="830A5D8A506700611590FC40FC070FDD" box="[912,985,926,953]" gridcol="3" gridrow="12" pageId="49" pageNumber="50">57</td>
<td id="830A5D8A506700611264FC40FB730FDD" box="[1124,1197,926,953]" gridcol="4" gridrow="12" pageId="49" pageNumber="50">2.34</td>
</tr>
<tr id="C0DB34F6506700611763FC1EFB730FBF" box="[355,1197,960,987]" gridrow="13" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FC1EFE380FBF" box="[355,486,960,987]" gridcol="0" gridrow="13" pageId="49" pageNumber="50">S8</th>
<td id="830A5D8A50670061143AFC1EFD940FBF" box="[570,586,960,987]" gridcol="1" gridrow="13" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FC1EFCF50FBF" box="[662,811,960,987]" gridcol="2" gridrow="13" pageId="49" pageNumber="50">5462</td>
<td id="830A5D8A506700611590FC1EFC070FBF" box="[912,985,960,987]" gridcol="3" gridrow="13" pageId="49" pageNumber="50">59</td>
<td id="830A5D8A506700611264FC1EFB730FBF" box="[1124,1197,960,987]" gridcol="4" gridrow="13" pageId="49" pageNumber="50">2.82</td>
</tr>
<tr id="C0DB34F6506700611763FC3FFB730F99" box="[355,1197,993,1021]" gridrow="14" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FC3FFE380F99" box="[355,486,993,1021]" gridcol="0" gridrow="14" pageId="49" pageNumber="50">S9</th>
<td id="830A5D8A50670061143AFC3FFD940F99" box="[570,586,993,1021]" gridcol="1" gridrow="14" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FC3FFCF50F99" box="[662,811,993,1021]" gridcol="2" gridrow="14" pageId="49" pageNumber="50">5962</td>
<td id="830A5D8A506700611590FC3FFC070F99" box="[912,985,993,1021]" gridcol="3" gridrow="14" pageId="49" pageNumber="50">61</td>
<td id="830A5D8A506700611264FC3FFB730F99" box="[1124,1197,993,1021]" gridcol="4" gridrow="14" pageId="49" pageNumber="50">1.38</td>
</tr>
<tr id="C0DB34F6506700611763FBDDFB73087A" box="[355,1197,1027,1054]" gridrow="15" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FBDDFE38087A" box="[355,486,1027,1054]" gridcol="0" gridrow="15" pageId="49" pageNumber="50">S10</th>
<td id="830A5D8A50670061143AFBDDFD94087A" box="[570,586,1027,1054]" gridcol="1" gridrow="15" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FBDDFCF5087A" box="[662,811,1027,1054]" gridcol="2" gridrow="15" pageId="49" pageNumber="50">4451</td>
<td id="830A5D8A506700611590FBDDFC07087A" box="[912,985,1027,1054]" gridcol="3" gridrow="15" pageId="49" pageNumber="50">49</td>
<td id="830A5D8A506700611264FBDDFB73087A" box="[1124,1197,1027,1054]" gridcol="4" gridrow="15" pageId="49" pageNumber="50">2.51</td>
</tr>
<tr id="C0DB34F6506700611763FBFAFB730824" box="[355,1197,1060,1088]" gridrow="16" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FBFAFE380824" box="[355,486,1060,1088]" gridcol="0" gridrow="16" pageId="49" pageNumber="50">S11</th>
<td id="830A5D8A50670061143AFBFAFD940824" box="[570,586,1060,1088]" gridcol="1" gridrow="16" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FBFAFCF50824" box="[662,811,1060,1088]" gridcol="2" gridrow="16" pageId="49" pageNumber="50">5358</td>
<td id="830A5D8A506700611590FBFAFC070824" box="[912,985,1060,1088]" gridcol="3" gridrow="16" pageId="49" pageNumber="50">56</td>
<td id="830A5D8A506700611264FBFAFB730824" box="[1124,1197,1060,1088]" gridcol="4" gridrow="16" pageId="49" pageNumber="50">1.91</td>
</tr>
<tr id="C0DB34F6506700611763FB8EFB73080F" box="[355,1197,1104,1131]" gridrow="17" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FB8EFE38080F" box="[355,486,1104,1131]" gridcol="0" gridrow="17" pageId="49" pageNumber="50">MD4 (ac)</th>
<td id="830A5D8A50670061143AFB8EFD94080F" box="[570,586,1104,1131]" gridcol="1" gridrow="17" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FB8EFCF5080F" box="[662,811,1104,1131]" gridcol="2" gridrow="17" pageId="49" pageNumber="50">3136</td>
<td id="830A5D8A506700611590FB8EFC07080F" box="[912,985,1104,1131]" gridcol="3" gridrow="17" pageId="49" pageNumber="50">33</td>
<td id="830A5D8A506700611264FB8EFB73080F" box="[1124,1197,1104,1131]" gridcol="4" gridrow="17" pageId="49" pageNumber="50">1.95</td>
</tr>
<tr id="C0DB34F6506700611763FBAFFB7308E8" box="[355,1197,1137,1164]" gridrow="18" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FBAFFE3808E8" box="[355,486,1137,1164]" gridcol="0" gridrow="18" pageId="49" pageNumber="50">MD5 (ac)</th>
<td id="830A5D8A50670061143AFBAFFD9408E8" box="[570,586,1137,1164]" gridcol="1" gridrow="18" pageId="49" pageNumber="50">6</td>
<td id="830A5D8A506700611496FBAFFCF508E8" box="[662,811,1137,1164]" gridcol="2" gridrow="18" pageId="49" pageNumber="50">4048</td>
<td id="830A5D8A506700611590FBAFFC0708E8" box="[912,985,1137,1164]" gridcol="3" gridrow="18" pageId="49" pageNumber="50">44</td>
<td id="830A5D8A506700611264FBAFFB7308E8" box="[1124,1197,1137,1164]" gridcol="4" gridrow="18" pageId="49" pageNumber="50">3.27</td>
</tr>
<tr id="C0DB34F6506700611763FB4DFB7308CA" box="[355,1197,1171,1198]" gridrow="19" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FB4DFE3808CA" box="[355,486,1171,1198]" gridcol="0" gridrow="19" pageId="49" pageNumber="50">MD6 (ac)</th>
<td id="830A5D8A50670061143AFB4DFD9408CA" box="[570,586,1171,1198]" gridcol="1" gridrow="19" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FB4DFCF508CA" box="[662,811,1171,1198]" gridcol="2" gridrow="19" pageId="49" pageNumber="50">5067</td>
<td id="830A5D8A506700611590FB4DFC0708CA" box="[912,985,1171,1198]" gridcol="3" gridrow="19" pageId="49" pageNumber="50">54</td>
<td id="830A5D8A506700611264FB4DFB7308CA" box="[1124,1197,1171,1198]" gridcol="4" gridrow="19" pageId="49" pageNumber="50">6.00</td>
</tr>
<tr id="C0DB34F6506700611763FB6AFB7308B4" box="[355,1197,1204,1232]" gridrow="20" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FB6AFE3808B4" box="[355,486,1204,1232]" gridcol="0" gridrow="20" pageId="49" pageNumber="50">MD7 (ac)</th>
<td id="830A5D8A50670061143AFB6AFD9408B4" box="[570,586,1204,1232]" gridcol="1" gridrow="20" pageId="49" pageNumber="50">6</td>
<td id="830A5D8A506700611496FB6AFCF508B4" box="[662,811,1204,1232]" gridcol="2" gridrow="20" pageId="49" pageNumber="50">5483</td>
<td id="830A5D8A506700611590FB6AFC0708B4" box="[912,985,1204,1232]" gridcol="3" gridrow="20" pageId="49" pageNumber="50">70</td>
<td id="830A5D8A506700611264FB6AFB7308B4" box="[1124,1197,1204,1232]" gridcol="4" gridrow="20" pageId="49" pageNumber="50">11.15</td>
</tr>
<tr id="C0DB34F6506700611763FB09FB730895" box="[355,1197,1239,1265]" gridrow="21" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FB09FE380895" box="[355,486,1239,1265]" gridcol="0" gridrow="21" pageId="49" pageNumber="50">MD8 (ac)</th>
<td id="830A5D8A50670061143AFB09FD940895" box="[570,586,1239,1265]" gridcol="1" gridrow="21" pageId="49" pageNumber="50">6</td>
<td id="830A5D8A506700611496FB09FCF50895" box="[662,811,1239,1265]" gridcol="2" gridrow="21" pageId="49" pageNumber="50">90128</td>
<td id="830A5D8A506700611590FB09FC070895" box="[912,985,1239,1265]" gridcol="3" gridrow="21" pageId="49" pageNumber="50">112</td>
<td id="830A5D8A506700611264FB09FB730895" box="[1124,1197,1239,1265]" gridcol="4" gridrow="21" pageId="49" pageNumber="50">16.97</td>
</tr>
<tr id="C0DB34F6506700611763FADCFB730978" box="[355,1197,1282,1308]" gridrow="22" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FADCFE380978" box="[355,486,1282,1308]" gridcol="0" gridrow="22" pageId="49" pageNumber="50">LV6 (ac)</th>
<td id="830A5D8A50670061143AFADCFD940978" box="[570,586,1282,1308]" gridcol="1" gridrow="22" pageId="49" pageNumber="50">6</td>
<td id="830A5D8A506700611496FADCFCF50978" box="[662,811,1282,1308]" gridcol="2" gridrow="22" pageId="49" pageNumber="50">3243</td>
<td id="830A5D8A506700611590FADCFC070978" box="[912,985,1282,1308]" gridcol="3" gridrow="22" pageId="49" pageNumber="50">38</td>
<td id="830A5D8A506700611264FADCFB730978" box="[1124,1197,1282,1308]" gridcol="4" gridrow="22" pageId="49" pageNumber="50">4.32</td>
</tr>
<tr id="C0DB34F6506700611763FAFDFB73095A" box="[355,1197,1315,1342]" gridrow="23" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FAFDFE38095A" box="[355,486,1315,1342]" gridcol="0" gridrow="23" pageId="49" pageNumber="50">LV7 (ac)</th>
<td id="830A5D8A50670061143AFAFDFD94095A" box="[570,586,1315,1342]" gridcol="1" gridrow="23" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FAFDFCF5095A" box="[662,811,1315,1342]" gridcol="2" gridrow="23" pageId="49" pageNumber="50">4553</td>
<td id="830A5D8A506700611590FAFDFC07095A" box="[912,985,1315,1342]" gridcol="3" gridrow="23" pageId="49" pageNumber="50">48</td>
<td id="830A5D8A506700611264FAFDFB73095A" box="[1124,1197,1315,1342]" gridcol="4" gridrow="23" pageId="49" pageNumber="50">3.86</td>
</tr>
<tr id="C0DB34F6506700611763FA9AFB730904" box="[355,1197,1348,1376]" gridrow="24" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FA9AFE380904" box="[355,486,1348,1376]" gridcol="0" gridrow="24" pageId="49" pageNumber="50">LV8 (ac)</th>
<td id="830A5D8A50670061143AFA9AFD940904" box="[570,586,1348,1376]" gridcol="1" gridrow="24" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FA9AFCF50904" box="[662,811,1348,1376]" gridcol="2" gridrow="24" pageId="49" pageNumber="50">5368</td>
<td id="830A5D8A506700611590FA9AFC070904" box="[912,985,1348,1376]" gridcol="3" gridrow="24" pageId="49" pageNumber="50">60</td>
<td id="830A5D8A506700611264FA9AFB730904" box="[1124,1197,1348,1376]" gridcol="4" gridrow="24" pageId="49" pageNumber="50">5.77</td>
</tr>
<tr id="C0DB34F6506700611763FAB8FB7309E5" box="[355,1197,1382,1409]" gridrow="25" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FAB8FE3809E5" box="[355,486,1382,1409]" gridcol="0" gridrow="25" pageId="49" pageNumber="50">LV9 (ac)</th>
<td id="830A5D8A50670061143AFAB8FD9409E5" box="[570,586,1382,1409]" gridcol="1" gridrow="25" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FAB8FCF509E5" box="[662,811,1382,1409]" gridcol="2" gridrow="25" pageId="49" pageNumber="50">5474</td>
<td id="830A5D8A506700611590FAB8FC0709E5" box="[912,985,1382,1409]" gridcol="3" gridrow="25" pageId="49" pageNumber="50">64</td>
<td id="830A5D8A506700611264FAB8FB7309E5" box="[1124,1197,1382,1409]" gridcol="4" gridrow="25" pageId="49" pageNumber="50">7.63</td>
</tr>
<tr id="C0DB34F6506700611763FA56FB7309C7" box="[355,1197,1416,1443]" gridrow="26" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FA56FE3809C7" box="[355,486,1416,1443]" gridcol="0" gridrow="26" pageId="49" pageNumber="50">ML10 (tu)</th>
<td id="830A5D8A50670061143AFA56FD9409C7" box="[570,586,1416,1443]" gridcol="1" gridrow="26" pageId="49" pageNumber="50">6</td>
<td id="830A5D8A506700611496FA56FCF509C7" box="[662,811,1416,1443]" gridcol="2" gridrow="26" pageId="49" pageNumber="50">2934</td>
<td id="830A5D8A506700611590FA56FC0709C7" box="[912,985,1416,1443]" gridcol="3" gridrow="26" pageId="49" pageNumber="50">31</td>
<td id="830A5D8A506700611264FA56FB7309C7" box="[1124,1197,1416,1443]" gridcol="4" gridrow="26" pageId="49" pageNumber="50">1.75</td>
</tr>
<tr id="C0DB34F6506700611763FA6AFB7309AA" box="[355,1197,1460,1486]" gridrow="27" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FA6AFE3809AA" box="[355,486,1460,1486]" gridcol="0" gridrow="27" pageId="49" pageNumber="50">TE</th>
<td id="830A5D8A50670061143AFA6AFD9409AA" box="[570,586,1460,1486]" gridcol="1" gridrow="27" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FA6AFCF509AA" box="[662,811,1460,1486]" gridcol="2" gridrow="27" pageId="49" pageNumber="50">2428</td>
<td id="830A5D8A506700611590FA6AFC0709AA" box="[912,985,1460,1486]" gridcol="3" gridrow="27" pageId="49" pageNumber="50">26</td>
<td id="830A5D8A506700611264FA6AFB7309AA" box="[1124,1197,1460,1486]" gridcol="4" gridrow="27" pageId="49" pageNumber="50">1.40</td>
</tr>
<tr id="C0DB34F6506700611763FA0BFB730994" box="[355,1197,1493,1520]" gridrow="28" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FA0BFE380994" box="[355,486,1493,1520]" gridcol="0" gridrow="28" pageId="49" pageNumber="50">TE/TL</th>
<td id="830A5D8A50670061143AFA0BFD940994" box="[570,586,1493,1520]" gridcol="1" gridrow="28" pageId="49" pageNumber="50">7</td>
<td id="830A5D8A506700611496FA0BFCF50994" box="[662,811,1493,1520]" gridcol="2" gridrow="28" pageId="49" pageNumber="50">6.08.2%</td>
<td id="830A5D8A506700611590FA0BFC070994" box="[912,985,1493,1520]" gridcol="3" gridrow="28" pageId="49" pageNumber="50">7.2%</td>
<td id="830A5D8A506700611264FA0BFB730994" box="[1124,1197,1493,1520]" gridcol="4" gridrow="28" pageId="49" pageNumber="50">0.94%</td>
</tr>
<tr id="C0DB34F6506700611763FA28FB730A75" box="[355,1197,1526,1553]" gridrow="29" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763FA28FE380A75" box="[355,486,1526,1553]" gridcol="0" gridrow="29" pageId="49" pageNumber="50">LTS</th>
<td id="830A5D8A50670061143AFA28FD940A75" box="[570,586,1526,1553]" gridcol="1" gridrow="29" pageId="49" pageNumber="50">6</td>
<td id="830A5D8A506700611496FA28FCF50A75" box="[662,811,1526,1553]" gridcol="2" gridrow="29" pageId="49" pageNumber="50">324355</td>
<td id="830A5D8A506700611590FA28FC070A75" box="[912,985,1526,1553]" gridcol="3" gridrow="29" pageId="49" pageNumber="50">341</td>
<td id="830A5D8A506700611264FA28FB730A75" box="[1124,1197,1526,1553]" gridcol="4" gridrow="29" pageId="49" pageNumber="50">12.45</td>
</tr>
<tr id="C0DB34F6506700611763F9C6FB730A57" box="[355,1197,1560,1587]" gridrow="30" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763F9C6FE380A57" box="[355,486,1560,1587]" gridcol="0" gridrow="30" pageId="49" pageNumber="50">LTS/TL</th>
<td id="830A5D8A50670061143AF9C6FD940A57" box="[570,586,1560,1587]" gridcol="1" gridrow="30" pageId="49" pageNumber="50">6</td>
<td id="830A5D8A506700611496F9C6FCF50A57" box="[662,811,1560,1587]" gridcol="2" gridrow="30" pageId="49" pageNumber="50">80.5100.9%</td>
<td id="830A5D8A506700611590F9C6FC070A57" box="[912,985,1560,1587]" gridcol="3" gridrow="30" pageId="49" pageNumber="50">91.5%</td>
<td id="830A5D8A506700611264F9C6FB730A57" box="[1124,1197,1560,1587]" gridcol="4" gridrow="30" pageId="49" pageNumber="50">7.59%</td>
</tr>
<tr id="C0DB34F6506700611763F9E7FB730A30" box="[355,1197,1593,1620]" gridrow="31" pageId="49" pageNumber="50">
<th id="830A5D8A506700611763F9E7FE380A30" box="[355,486,1593,1620]" gridcol="0" gridrow="31" pageId="49" pageNumber="50">LTAS</th>
<td id="830A5D8A50670061143AF9E7FD940A30" box="[570,586,1593,1620]" gridcol="1" gridrow="31" pageId="49" pageNumber="50">6</td>
<td id="830A5D8A506700611496F9E7FCF50A30" box="[662,811,1593,1620]" gridcol="2" gridrow="31" pageId="49" pageNumber="50">6283</td>
<td id="830A5D8A506700611590F9E7FC070A30" box="[912,985,1593,1620]" gridcol="3" gridrow="31" pageId="49" pageNumber="50">75</td>
<td id="830A5D8A506700611264F9E7FB730A30" box="[1124,1197,1593,1620]" gridcol="4" gridrow="31" pageId="49" pageNumber="50">7.40</td>
</tr>
</table>
</paragraph>
<paragraph id="7E5436B45067FFAF16BDF8DCFB680BE2" blockId="49.[189,1399,1794,1926]" pageId="49" pageNumber="50">
The neck has 16 placids, measuring
<quantity id="B9139B515067FFAF1476F8DCFD1B0B79" box="[630,709,1794,1821]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="49" pageNumber="50" unit="mm" value="18.0">18 µm</quantity>
in length, except for the slightly longer midventral ones, measuring
<quantity id="B9139B515067FFAF173DF8FBFE570B24" box="[317,393,1829,1856]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="49" pageNumber="50" unit="mm" value="20.0">20 µm</quantity>
in length. The midventral placid is also broadest, measuring
<quantity id="B9139B515067FFAF1247F8FBFB4D0B24" box="[1095,1171,1829,1856]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="49" pageNumber="50" unit="mm" value="16.0">16 µm</quantity>
in width at its base, whereas all others are narrower, measuring
<quantity id="B9139B515067FFAF14C3F897FCCE0B07" box="[707,784,1865,1891]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="49" pageNumber="50" unit="mm" value="10.0">10 µm</quantity>
in width at their bases. Four dorsal and two ventral trichoscalid plates are present; trichoscalid plates are fairly small, and rounded to oval.
</paragraph>
<paragraph id="7E5436B45067FFAC16BDF873FDD7094B" blockId="49.[189,1399,1965,2027]" lastBlockId="50.[189,1399,1125,1327]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
Segment 1 consists of a complete cuticular ring. Sensory spots are located on the anterior half of the segment, but not at the anterior margin in subdorsal and laterodorsal positions (
<figureCitation id="E6D02A315067FFAF1274F80EFB380B8E" box="[1140,1254,2000,2026]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="49" pageNumber="50">Figs 18A</figureCitation>
,
<figureCitation id="E6D02A315067FFAF12F4F80EFAF90B8E" box="[1268,1319,2000,2026]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="49" pageNumber="50">19B</figureCitation>
,
<figureCitation id="E6D02A315067FFAF1335F80EFAB70B8E" box="[1333,1385,2000,2026]" captionStart="Fig" captionStartId="48.[189,232,1647,1673]" captionTargetBox="[191,1396,382,1597]" captionTargetId="figure@48.[191,1396,382,1597]" captionTargetPageId="48" captionText="Fig. 20. Scanning electron micrographs showing overviews and details of Echinoderes lupherorum sp. nov. A. Lateroventral overview. B. Laterodorsal overview. C. Introvert sectors 10 and 9. D. Segments 1 to 3, ventral view. E. Segments 1 to 2, dorsal view. F. Detail showing middorsal (right) to laterodorsal (left) parts of segments 1 and 2. Inset shows close-up of subdorsal glandular cell outlet type 2; scale = 1 µm. G. Detail showing middorsal (right) to laterodorsal (left) parts of 8. H. Close-up showing laterodorsal sensory spot and glandular cell outlet type 2 on segment 8. I. Segments 9 to 10, lateral view. J. ♀, segments 10 to 11, dorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvt = lateroventral tube; mds = middorsal spine; mdss = middorsal sensory spot; mlt = midlateral tube; pdss = paradorsal sensory spot; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid." figureDoi="http://doi.org/10.5281/zenodo.3817857" httpUri="https://zenodo.org/record/3817857/files/figure.png" pageId="49" pageNumber="50">20E</figureCitation>
); sensory spots are small and rounded, with a few papillae. Their minute size, combined with the dense hair covering on most segments, makes it very difficult to observe the sensory spots in SEM, and often they are more easily observed in LM. Glandular cell outlets
<typeStatus id="A15088165064FFAC154BFB72FCA008A2" box="[843,894,1196,1222]" pageId="50" pageNumber="51">type</typeStatus>
1 only observed in lateroventral positions. Dorsal and lateral sides have scattered cuticular hairs, whereas the ventral side is nearly devoid of hairs (
<figureCitation id="E6D02A315064FFAC16C6FB2CFEC30968" box="[198,285,1266,1292]" captionStart="Fig" captionStartId="48.[189,232,1647,1673]" captionTargetBox="[191,1396,382,1597]" captionTargetId="figure@48.[191,1396,382,1597]" captionTargetPageId="48" captionText="Fig. 20. Scanning electron micrographs showing overviews and details of Echinoderes lupherorum sp. nov. A. Lateroventral overview. B. Laterodorsal overview. C. Introvert sectors 10 and 9. D. Segments 1 to 3, ventral view. E. Segments 1 to 2, dorsal view. F. Detail showing middorsal (right) to laterodorsal (left) parts of segments 1 and 2. Inset shows close-up of subdorsal glandular cell outlet type 2; scale = 1 µm. G. Detail showing middorsal (right) to laterodorsal (left) parts of 8. H. Close-up showing laterodorsal sensory spot and glandular cell outlet type 2 on segment 8. I. Segments 9 to 10, lateral view. J. ♀, segments 10 to 11, dorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvt = lateroventral tube; mds = middorsal spine; mdss = middorsal sensory spot; mlt = midlateral tube; pdss = paradorsal sensory spot; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid." figureDoi="http://doi.org/10.5281/zenodo.3817857" httpUri="https://zenodo.org/record/3817857/files/figure.png" pageId="50" pageNumber="51">Fig. 20</figureCitation>
DE). The posterior segment margin is nearly straight, terminating in a pectinate fringe with long and flexible fringe tips.
</paragraph>
<caption id="2A94663C5064FFAC16BDFED2FEE10E78" pageId="50" pageNumber="51">
<paragraph id="7E5436B45064FFAC16BDFED2FEE10E78" blockId="50.[189,1399,267,540]" pageId="50" pageNumber="51">
<emphasis id="4C9FEAA65064FFAC16BDFED2FEEF0D41" bold="true" box="[189,305,267,294]" pageId="50" pageNumber="51">Table 13.</emphasis>
Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in
<taxonomicName id="B9EB4D375064FFAC17C4FEF1FD370D2D" authority="Sørensen &amp; Rohal &amp; Thistle, 2018" authorityName="Sørensen &amp; Rohal &amp; Thistle" authorityYear="2018" box="[452,745,303,329]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="50" pageNumber="51" phylum="Cephalorhyncha" rank="species" species="lupherorum" status="sp. nov.">
<emphasis id="4C9FEAA65064FFAC17C4FEF1FD370D2D" box="[452,745,303,329]" italics="true" pageId="50" pageNumber="51">Echinoderes lupherorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57AC57DD5064FFAC14F3FEF1FC8F0D2D" box="[755,849,303,329]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviations: LA = lateral accessory; LD = laterodorsal; LV = lateroventral; MD = middorsal; ML = midlateral; PD = paradorsal; PV = paraventral; SD = subdorsal; SL = sublateral; VL = ventrolateral; VM = ventromedial; ac = acicular spine; gco1/2 = glandular cell outlet type 1/2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; pa = papilla; pe = penile spines; si = sieve plate; ss = sensory spot; tu = tube; (♀) = female and (♂) = male conditions of sexually dimorphic characters. * = indicates that the structure is not present in all specimens.
</paragraph>
</caption>
<paragraph id="7E5436B45064FFAC14C9FD9AFBFA0866" pageId="50" pageNumber="51">
<table id="0CEBC4145064006116C1FD9AFAB10866" box="[193,1391,580,1026]" gridcols="11" gridrows="13" pageId="50" pageNumber="51">
<tr id="C0DB34F65064006116C1FD9AFAB10E38" box="[193,1391,580,604]" gridrow="0" pageId="50" pageNumber="51" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="830A5D8A50640061147BFD9AFC8D0E38" box="[635,851,580,604]" gridcol="4" gridrow="0" pageId="50" pageNumber="51">
<emphasis id="4C9FEAA65064FFAC14C9FD9AFCFB0E38" bold="true" box="[713,805,580,604]" pageId="50" pageNumber="51">Position</emphasis>
</th>
</tr>
<tr id="C0DB34F65064006116C1FDACFAB10EEE" box="[193,1391,626,650]" gridrow="1" pageId="50" pageNumber="51">
<th id="830A5D8A5064006116C1FDACFE400EEE" box="[193,414,626,650]" colspan="2" colspanRight="1" gridcol="0" gridrow="1" pageId="50" pageNumber="51">
<emphasis id="4C9FEAA65064FFAC16C1FDACFEA70EEE" bold="true" box="[193,377,626,650]" pageId="50" pageNumber="51">Segment MD</emphasis>
</th>
<td id="830A5D8A5064006117B1FDACFE0B0EEE" box="[433,469,626,650]" gridcol="2" gridrow="1" pageId="50" pageNumber="51">
<emphasis id="4C9FEAA65064FFAC17B1FDACFE0B0EEE" bold="true" box="[433,469,626,650]" pageId="50" pageNumber="51">PD</emphasis>
</td>
<td id="830A5D8A5064006117F7FDACFD840EEE" box="[503,602,626,650]" gridcol="3" gridrow="1" pageId="50" pageNumber="51">
<emphasis id="4C9FEAA65064FFAC1417FDACFDE70EEE" bold="true" box="[535,569,626,650]" pageId="50" pageNumber="51">SD</emphasis>
</td>
<td id="830A5D8A50640061147BFDACFC8D0EEE" box="[635,851,626,650]" gridcol="4" gridrow="1" pageId="50" pageNumber="51">
<emphasis id="4C9FEAA65064FFAC1493FDACFCEB0EEE" bold="true" box="[659,821,626,650]" pageId="50" pageNumber="51">LD ML</emphasis>
</td>
<td id="830A5D8A50640061156DFDACFC500EEE" box="[877,910,626,650]" gridcol="5" gridrow="1" pageId="50" pageNumber="51">
<emphasis id="4C9FEAA65064FFAC156DFDACFC500EEE" bold="true" box="[877,910,626,650]" pageId="50" pageNumber="51">SL</emphasis>
</td>
<td id="830A5D8A5064006115A2FDACFC300EEE" box="[930,1006,626,650]" gridcol="6" gridrow="1" pageId="50" pageNumber="51">
<emphasis id="4C9FEAA65064FFAC15B6FDACFC050EEE" bold="true" box="[950,987,626,650]" pageId="50" pageNumber="51">LA</emphasis>
</td>
<td id="830A5D8A5064006115FEFDACFBEC0EEE" box="[1022,1074,626,650]" gridcol="7" gridrow="1" pageId="50" pageNumber="51">
<emphasis id="4C9FEAA65064FFAC1207FDACFBF90EEE" bold="true" box="[1031,1063,626,650]" pageId="50" pageNumber="51">LV</emphasis>
</td>
<td id="830A5D8A50640061124FFDACFB500EEE" box="[1103,1166,626,650]" gridcol="8" gridrow="1" pageId="50" pageNumber="51">
<emphasis id="4C9FEAA65064FFAC125CFDACFB5F0EEE" bold="true" box="[1116,1153,626,650]" pageId="50" pageNumber="51">VL</emphasis>
</td>
<td id="830A5D8A5064006112A9FDACFAF40EEE" box="[1193,1322,626,650]" gridcol="9" gridrow="1" pageId="50" pageNumber="51">
<emphasis id="4C9FEAA65064FFAC12D3FDACFB210EEE" bold="true" box="[1235,1279,626,650]" pageId="50" pageNumber="51">VM</emphasis>
</td>
<td id="830A5D8A50640061133BFDACFAB10EEE" box="[1339,1391,626,650]" gridcol="10" gridrow="1" pageId="50" pageNumber="51">
<emphasis id="4C9FEAA65064FFAC1344FDACFAB60EEE" bold="true" box="[1348,1384,626,650]" pageId="50" pageNumber="51">PV</emphasis>
</td>
</tr>
<tr id="C0DB34F65064006116C1FD44FAB10ED6" box="[193,1391,666,690]" gridrow="2" pageId="50" pageNumber="51" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-8="1" rowspan-9="1">
<th id="830A5D8A5064006116C1FD44FF200ED6" box="[193,254,666,690]" gridcol="0" gridrow="2" pageId="50" pageNumber="51">1</th>
<td id="830A5D8A5064006117F7FD44FD840ED6" box="[503,602,666,690]" gridcol="3" gridrow="2" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A50640061147BFD44FC8D0ED6" box="[635,851,666,690]" gridcol="4" gridrow="2" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A5064006115FEFD44FBEC0ED6" box="[1022,1074,666,690]" gridcol="7" gridrow="2" pageId="50" pageNumber="51">gco1</td>
</tr>
<tr id="C0DB34F65064006116C1FD65FAB10EB7" box="[193,1391,699,723]" gridrow="3" pageId="50" pageNumber="51" rowspan-10="1" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-8="1">
<th id="830A5D8A5064006116C1FD65FF200EB7" box="[193,254,699,723]" gridcol="0" gridrow="3" pageId="50" pageNumber="51">2</th>
<td id="830A5D8A506400611757FD65FE400EB7" box="[343,414,699,723]" gridcol="1" gridrow="3" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A5064006117F7FD65FD840EB7" box="[503,602,699,723]" gridcol="3" gridrow="3" pageId="50" pageNumber="51">gco2, ss</td>
<td id="830A5D8A50640061147BFD65FC8D0EB7" box="[635,851,699,723]" gridcol="4" gridrow="3" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A5064006115FEFD65FBEC0EB7" box="[1022,1074,699,723]" gridcol="7" gridrow="3" pageId="50" pageNumber="51">tu</td>
<td id="830A5D8A5064006112A9FD65FAF40EB7" box="[1193,1322,699,723]" gridcol="9" gridrow="3" pageId="50" pageNumber="51">ss</td>
</tr>
<tr id="C0DB34F65064006116C1FD03FAB10E91" box="[193,1391,733,757]" gridrow="4" pageId="50" pageNumber="51" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="830A5D8A5064006116C1FD03FF200E91" box="[193,254,733,757]" gridcol="0" gridrow="4" pageId="50" pageNumber="51">3</th>
<td id="830A5D8A5064006117F7FD03FD840E91" box="[503,602,733,757]" gridcol="3" gridrow="4" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A50640061156DFD03FC500E91" box="[877,910,733,757]" gridcol="5" gridrow="4" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A5064006112A9FD03FAF40E91" box="[1193,1322,733,757]" gridcol="9" gridrow="4" pageId="50" pageNumber="51">gco1</td>
</tr>
<tr id="C0DB34F65064006116C1FD20FAB10F72" box="[193,1391,766,790]" gridrow="5" pageId="50" pageNumber="51" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="830A5D8A5064006116C1FD20FF200F72" box="[193,254,766,790]" gridcol="0" gridrow="5" pageId="50" pageNumber="51">4</th>
<td id="830A5D8A506400611757FD20FE400F72" box="[343,414,766,790]" gridcol="1" gridrow="5" pageId="50" pageNumber="51">ac</td>
<td id="830A5D8A5064006112A9FD20FAF40F72" box="[1193,1322,766,790]" gridcol="9" gridrow="5" pageId="50" pageNumber="51">ss, gco1</td>
</tr>
<tr id="C0DB34F65064006116C1FCFEFAB10F5C" box="[193,1391,800,824]" gridrow="6" pageId="50" pageNumber="51" rowspan-10="1" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-8="1">
<th id="830A5D8A5064006116C1FCFEFF200F5C" box="[193,254,800,824]" gridcol="0" gridrow="6" pageId="50" pageNumber="51">5</th>
<td id="830A5D8A506400611757FCFEFE400F5C" box="[343,414,800,824]" gridcol="1" gridrow="6" pageId="50" pageNumber="51">ac</td>
<td id="830A5D8A5064006117F7FCFEFD840F5C" box="[503,602,800,824]" gridcol="3" gridrow="6" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A50640061147BFCFEFC8D0F5C" box="[635,851,800,824]" gridcol="4" gridrow="6" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A5064006115FEFCFEFBEC0F5C" box="[1022,1074,800,824]" gridcol="7" gridrow="6" pageId="50" pageNumber="51">tu</td>
<td id="830A5D8A5064006112A9FCFEFAF40F5C" box="[1193,1322,800,824]" gridcol="9" gridrow="6" pageId="50" pageNumber="51">gco1</td>
</tr>
<tr id="C0DB34F65064006116C1FC9FFAB10F3E" box="[193,1391,833,858]" gridrow="7" pageId="50" pageNumber="51" rowspan-10="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-8="1">
<th id="830A5D8A5064006116C1FC9FFF200F3E" box="[193,254,833,858]" gridcol="0" gridrow="7" pageId="50" pageNumber="51">6</th>
<td id="830A5D8A506400611757FC9FFE400F3E" box="[343,414,833,858]" gridcol="1" gridrow="7" pageId="50" pageNumber="51">ac</td>
<td id="830A5D8A5064006117B1FC9FFE0B0F3E" box="[433,469,833,858]" gridcol="2" gridrow="7" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A50640061147BFC9FFC8D0F3E" box="[635,851,833,858]" gridcol="4" gridrow="7" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A5064006115FEFC9FFBEC0F3E" box="[1022,1074,833,858]" gridcol="7" gridrow="7" pageId="50" pageNumber="51">ac</td>
<td id="830A5D8A5064006112A9FC9FFAF40F3E" box="[1193,1322,833,858]" gridcol="9" gridrow="7" pageId="50" pageNumber="51">ss, gco1</td>
</tr>
<tr id="C0DB34F65064006116C1FCBCFAB10F18" box="[193,1391,866,892]" gridrow="8" pageId="50" pageNumber="51" rowspan-10="1" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<th id="830A5D8A5064006116C1FCBCFF200F18" box="[193,254,866,892]" gridcol="0" gridrow="8" pageId="50" pageNumber="51">7</th>
<td id="830A5D8A506400611757FCBCFE400F18" box="[343,414,866,892]" gridcol="1" gridrow="8" pageId="50" pageNumber="51">ac</td>
<td id="830A5D8A5064006117B1FCBCFE0B0F18" box="[433,469,866,892]" gridcol="2" gridrow="8" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A50640061147BFCBCFC8D0F18" box="[635,851,866,892]" gridcol="4" gridrow="8" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A5064006115FEFCBCFBEC0F18" box="[1022,1074,866,892]" gridcol="7" gridrow="8" pageId="50" pageNumber="51">ac</td>
<td id="830A5D8A50640061124FFCBCFB500F18" box="[1103,1166,866,892]" gridcol="8" gridrow="8" pageId="50" pageNumber="51">pa(♀)</td>
<td id="830A5D8A5064006112A9FCBCFAF40F18" box="[1193,1322,866,892]" gridcol="9" gridrow="8" pageId="50" pageNumber="51">ss(♂), gco1</td>
</tr>
<tr id="C0DB34F65064006116C1FC5BFAB10FF9" box="[193,1391,901,925]" gridrow="9" pageId="50" pageNumber="51" rowspan-10="1" rowspan-5="1" rowspan-6="1" rowspan-8="1">
<th id="830A5D8A5064006116C1FC5BFF200FF9" box="[193,254,901,925]" gridcol="0" gridrow="9" pageId="50" pageNumber="51">8</th>
<td id="830A5D8A506400611757FC5BFE400FF9" box="[343,414,901,925]" gridcol="1" gridrow="9" pageId="50" pageNumber="51">ac</td>
<td id="830A5D8A5064006117B1FC5BFE0B0FF9" box="[433,469,901,925]" gridcol="2" gridrow="9" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A5064006117F7FC5BFD840FF9" box="[503,602,901,925]" gridcol="3" gridrow="9" pageId="50" pageNumber="51">gco1, ss*</td>
<td id="830A5D8A50640061147BFC5BFC8D0FF9" box="[635,851,901,925]" gridcol="4" gridrow="9" pageId="50" pageNumber="51">gco2, ss</td>
<td id="830A5D8A5064006115FEFC5BFBEC0FF9" box="[1022,1074,901,925]" gridcol="7" gridrow="9" pageId="50" pageNumber="51">ac</td>
<td id="830A5D8A5064006112A9FC5BFAF40FF9" box="[1193,1322,901,925]" gridcol="9" gridrow="9" pageId="50" pageNumber="51">pa(♀), gco1</td>
</tr>
<tr id="C0DB34F65064006116C1FC78FAB10FDB" box="[193,1391,934,959]" gridrow="10" pageId="50" pageNumber="51" rowspan-1="1" rowspan-10="1" rowspan-5="1">
<th id="830A5D8A5064006116C1FC78FF200FDB" box="[193,254,934,959]" gridcol="0" gridrow="10" pageId="50" pageNumber="51">9</th>
<td id="830A5D8A5064006117B1FC78FE0B0FDB" box="[433,469,934,959]" gridcol="2" gridrow="10" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A5064006117F7FC78FD840FDB" box="[503,602,934,959]" gridcol="3" gridrow="10" pageId="50" pageNumber="51">gco1, ss</td>
<td id="830A5D8A50640061147BFC78FC8D0FDB" box="[635,851,934,959]" gridcol="4" gridrow="10" pageId="50" pageNumber="51">gco2, ss</td>
<td id="830A5D8A5064006115A2FC78FC300FDB" box="[930,1006,934,959]" gridcol="6" gridrow="10" pageId="50" pageNumber="51">si</td>
<td id="830A5D8A5064006115FEFC78FBEC0FDB" box="[1022,1074,934,959]" gridcol="7" gridrow="10" pageId="50" pageNumber="51">ac</td>
<td id="830A5D8A50640061124FFC78FB500FDB" box="[1103,1166,934,959]" gridcol="8" gridrow="10" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A5064006112A9FC78FAF40FDB" box="[1193,1322,934,959]" gridcol="9" gridrow="10" pageId="50" pageNumber="51">gco1</td>
</tr>
<tr id="C0DB34F65064006116C1FC16FAB10F84" box="[193,1391,968,992]" gridrow="11" pageId="50" pageNumber="51" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="830A5D8A5064006116C1FC16FE400F84" box="[193,414,968,992]" colspan="2" colspanRight="1" gridcol="0" gridrow="11" pageId="50" pageNumber="51">10 gco1, gco1</th>
<td id="830A5D8A5064006117B1FC16FE0B0F84" box="[433,469,968,992]" gridcol="2" gridrow="11" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A50640061147BFC16FC8D0F84" box="[635,851,968,992]" gridcol="4" gridrow="11" pageId="50" pageNumber="51">tu</td>
<td id="830A5D8A50640061124FFC16FB500F84" box="[1103,1166,968,992]" gridcol="8" gridrow="11" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A50640061133BFC16FAB10F84" box="[1339,1391,968,992]" gridcol="10" gridrow="11" pageId="50" pageNumber="51">gco1</td>
</tr>
<tr id="C0DB34F65064006116C1FC36FAB10866" box="[193,1391,1000,1026]" gridrow="12" pageId="50" pageNumber="51" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-5="1" rowspan-8="1" rowspan-9="1">
<th id="830A5D8A5064006116C1FC36FF200866" box="[193,254,1000,1026]" gridcol="0" gridrow="12" pageId="50" pageNumber="51">11</th>
<td id="830A5D8A5064006117F7FC36FD840866" box="[503,602,1000,1026]" gridcol="3" gridrow="12" pageId="50" pageNumber="51">ss</td>
<td id="830A5D8A50640061147BFC36FC8D0866" box="[635,851,1000,1026]" gridcol="4" gridrow="12" pageId="50" pageNumber="51">3 x pe(♂)</td>
<td id="830A5D8A5064006115A2FC36FC300866" box="[930,1006,1000,1026]" gridcol="6" gridrow="12" pageId="50" pageNumber="51">ltas(♀)</td>
<td id="830A5D8A5064006115FEFC36FBEC0866" box="[1022,1074,1000,1026]" gridcol="7" gridrow="12" pageId="50" pageNumber="51">lts</td>
</tr>
</table>
</paragraph>
<paragraph id="7E5436B45064FFAC16BDFA89FB480A0C" blockId="50.[189,1399,1367,1640]" pageId="50" pageNumber="51">
Segment 2 consists of a complete cuticular ring. The pachycyclus of the anterior segment margin is relatively thin and not interrupted. Very minute glandular cell outlets
<typeStatus id="A15088165064FFAC158EFAA4FC6109F0" box="[910,959,1402,1428]" pageId="50" pageNumber="51">type</typeStatus>
2 are present in subdorsal positions (
<figureCitation id="E6D02A315064FFAC16C5FA43FEE709D3" box="[197,313,1437,1463]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="50" pageNumber="51">Figs 18A</figureCitation>
,
<figureCitation id="E6D02A315064FFAC1749FA43FEA509D3" box="[329,379,1437,1463]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="50" pageNumber="51">19B</figureCitation>
,
<figureCitation id="E6D02A315064FFAC178DFA43FE7509D3" box="[397,427,1437,1463]" captionStart="Fig" captionStartId="48.[189,232,1647,1673]" captionTargetBox="[191,1396,382,1597]" captionTargetId="figure@48.[191,1396,382,1597]" captionTargetPageId="48" captionText="Fig. 20. Scanning electron micrographs showing overviews and details of Echinoderes lupherorum sp. nov. A. Lateroventral overview. B. Laterodorsal overview. C. Introvert sectors 10 and 9. D. Segments 1 to 3, ventral view. E. Segments 1 to 2, dorsal view. F. Detail showing middorsal (right) to laterodorsal (left) parts of segments 1 and 2. Inset shows close-up of subdorsal glandular cell outlet type 2; scale = 1 µm. G. Detail showing middorsal (right) to laterodorsal (left) parts of 8. H. Close-up showing laterodorsal sensory spot and glandular cell outlet type 2 on segment 8. I. Segments 9 to 10, lateral view. J. ♀, segments 10 to 11, dorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvt = lateroventral tube; mds = middorsal spine; mdss = middorsal sensory spot; mlt = midlateral tube; pdss = paradorsal sensory spot; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid." figureDoi="http://doi.org/10.5281/zenodo.3817857" httpUri="https://zenodo.org/record/3817857/files/figure.png" pageId="50" pageNumber="51">20</figureCitation>
EF), and well-developed tubes in lateroventral positions (
<figureCitation id="E6D02A315064FFAC126EFA43FB0109D3" box="[1134,1247,1437,1463]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="50" pageNumber="51">Figs 18B</figureCitation>
,
<figureCitation id="E6D02A315064FFAC12F1FA43FAFD09D3" box="[1265,1315,1437,1463]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="50" pageNumber="51">19C</figureCitation>
,
<figureCitation id="E6D02A315064FFAC1334FA43FAB409D3" box="[1332,1386,1437,1463]" captionStart="Fig" captionStartId="48.[189,232,1647,1673]" captionTargetBox="[191,1396,382,1597]" captionTargetId="figure@48.[191,1396,382,1597]" captionTargetPageId="48" captionText="Fig. 20. Scanning electron micrographs showing overviews and details of Echinoderes lupherorum sp. nov. A. Lateroventral overview. B. Laterodorsal overview. C. Introvert sectors 10 and 9. D. Segments 1 to 3, ventral view. E. Segments 1 to 2, dorsal view. F. Detail showing middorsal (right) to laterodorsal (left) parts of segments 1 and 2. Inset shows close-up of subdorsal glandular cell outlet type 2; scale = 1 µm. G. Detail showing middorsal (right) to laterodorsal (left) parts of 8. H. Close-up showing laterodorsal sensory spot and glandular cell outlet type 2 on segment 8. I. Segments 9 to 10, lateral view. J. ♀, segments 10 to 11, dorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvt = lateroventral tube; mds = middorsal spine; mdss = middorsal sensory spot; mlt = midlateral tube; pdss = paradorsal sensory spot; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid." figureDoi="http://doi.org/10.5281/zenodo.3817857" httpUri="https://zenodo.org/record/3817857/files/figure.png" pageId="50" pageNumber="51">20D</figureCitation>
). Sensory spots are located in middorsal (
<figureCitation id="E6D02A315064FFAC1489FA1EFD0309BE" box="[649,733,1472,1498]" captionStart="Fig" captionStartId="48.[189,232,1647,1673]" captionTargetBox="[191,1396,382,1597]" captionTargetId="figure@48.[191,1396,382,1597]" captionTargetPageId="48" captionText="Fig. 20. Scanning electron micrographs showing overviews and details of Echinoderes lupherorum sp. nov. A. Lateroventral overview. B. Laterodorsal overview. C. Introvert sectors 10 and 9. D. Segments 1 to 3, ventral view. E. Segments 1 to 2, dorsal view. F. Detail showing middorsal (right) to laterodorsal (left) parts of segments 1 and 2. Inset shows close-up of subdorsal glandular cell outlet type 2; scale = 1 µm. G. Detail showing middorsal (right) to laterodorsal (left) parts of 8. H. Close-up showing laterodorsal sensory spot and glandular cell outlet type 2 on segment 8. I. Segments 9 to 10, lateral view. J. ♀, segments 10 to 11, dorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvt = lateroventral tube; mds = middorsal spine; mdss = middorsal sensory spot; mlt = midlateral tube; pdss = paradorsal sensory spot; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid." figureDoi="http://doi.org/10.5281/zenodo.3817857" httpUri="https://zenodo.org/record/3817857/files/figure.png" pageId="50" pageNumber="51">Fig. 20</figureCitation>
EF), subdorsal, laterodorsal (
<figureCitation id="E6D02A315064FFAC1237FA1EFB4009BE" box="[1079,1182,1472,1498]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="50" pageNumber="51">Fig. 19B</figureCitation>
) and ventromedial positions. Glandular cell outlets
<typeStatus id="A15088165064FFAC1434FA3AFDBB099A" box="[564,613,1508,1534]" pageId="50" pageNumber="51">type</typeStatus>
1 are not observed. A secondary pectinate fringe is present near the anterior segment margin of this and the following segments, but it is usually covered by the preceding segment. On this and the following eight segments, the cuticular hairs are bracteate and densely cover the dorsal and lateral areas. The posterior segment margin is as on the preceding segment.
</paragraph>
<paragraph id="7E5436B45064FFAC16BDF951FCEC0B3D" blockId="50.[189,1399,1678,1881]" pageId="50" pageNumber="51">
Segment 3, and the remaining segments, consist of one tergal and two sternal plates (
<figureCitation id="E6D02A315064FFAC12BEF950FAF00ACD" box="[1214,1326,1678,1705]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="50" pageNumber="51">Figs 18B</figureCitation>
,
<figureCitation id="E6D02A315064FFAC133DF951FEFF0AA8" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="50" pageNumber="51">19A, C, E, G</figureCitation>
). The pachycyclus of the anterior segment margin is well-developed and interrupted at the tergosternal junctions and middorsally. Sensory spots are present in subdorsal (
<figureCitation id="E6D02A315064FFAC1259F90BFB1D0A8B" box="[1113,1219,1749,1775]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="50" pageNumber="51">Fig. 19D</figureCitation>
) and sublateral positions, as well as glandular cell outlets
<typeStatus id="A15088165064FFAC14C6F927FD270B77" box="[710,761,1785,1811]" pageId="50" pageNumber="51">type</typeStatus>
<quantity id="B9139B515064FFAC1503F926FCEA0B76" box="[771,820,1784,1810]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="50" pageNumber="51" unit="in" value="1.0">1 in</quantity>
ventromedial positions. Cuticular hairs densely cover the tergal plate and the lateral halves of the sternal plates. The posterior segment margin is nearly straight, with fringe tips as on the preceding segment.
</paragraph>
<paragraph id="7E5436B45064FFAC16BDF85EFCFE0B85" blockId="50.[189,1398,1920,2017]" pageId="50" pageNumber="51">
Segment 4 with a middorsal acicular spine (
<figureCitation id="E6D02A315064FFAC14C0F85EFCEF0BFE" box="[704,817,1920,1946]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="50" pageNumber="51">Figs 18A</figureCitation>
,
<figureCitation id="E6D02A315064FFAC153DF85EFCAA0BFE" box="[829,884,1920,1946]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="50" pageNumber="51">19D</figureCitation>
). There are sensory spots and glandular cell outlets
<typeStatus id="A15088165064FFAC1716F87AFE970BDA" box="[278,329,1956,1982]" pageId="50" pageNumber="51">type</typeStatus>
1 present in ventromedial positions (
<figureCitation id="E6D02A315064FFAC150FF87DFCA20BD9" box="[783,892,1955,1981]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="50" pageNumber="51">Fig. 19E</figureCitation>
). Pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="7E5436B45065FFAD16BDFED5FBBD0DF6" blockId="51.[189,1399,267,402]" pageId="51" pageNumber="52">
Segment 5 with a middorsal acicular spine and well-developed tubes in lateroventral positions (
<figureCitation id="E6D02A315065FFAD16C5FEF1FEC20D2D" box="[197,284,303,329]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="51" pageNumber="52">Figs 18</figureCitation>
AB, 19DE); tubes have weakly developed, yet visible bases and lateral wings. Sensory spots are present in subdorsal (
<figureCitation id="E6D02A315065FFAD17E4FE8DFD910D09" box="[484,591,339,365]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="51" pageNumber="52">Fig. 19D</figureCitation>
) and midlateral positions. Cuticular hairs, glandular cell outlets
<typeStatus id="A15088165065FFAD1344FE8AFAA90D0A" box="[1348,1399,340,366]" pageId="51" pageNumber="52">type</typeStatus>
1, pachycycli and pectinate fringe of posterior margin as on preceding segment.
</paragraph>
<paragraph id="7E5436B45065FFAD16BDFE64FAC70E78" blockId="51.[189,1399,441,540]" pageId="51" pageNumber="52">
Segment 6 with acicular spines in middorsal and lateroventral positions (
<figureCitation id="E6D02A315065FFAD120FFE67FBB80DB0" box="[1039,1126,441,468]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="51" pageNumber="52">Figs 18</figureCitation>
AB, 19DE). Sensory spots present in paradorsal (
<figureCitation id="E6D02A315065FFAD1400FE03FDB50D93" box="[512,619,477,503]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="51" pageNumber="52">Fig. 19D</figureCitation>
), midlateral and ventromedial positions. Glandular cell outlets
<typeStatus id="A15088165065FFAD1344FE00FAA90D9C" box="[1348,1399,478,504]" pageId="51" pageNumber="52">type</typeStatus>
1, pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="7E5436B45065FFAD16BDFD9AFAC70E8A" blockId="51.[189,1399,579,750]" pageId="51" pageNumber="52">
Segment 7 with acicular spines in middorsal and lateroventral positions (
<figureCitation id="E6D02A315065FFAD1210FD9DFBB90E3A" box="[1040,1127,579,606]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="51" pageNumber="52">Figs 18</figureCitation>
AB, 19FG). Sensory spots present in paradorsal and midlateral positions, and males in addition with sensory spots in ventromedial positions. Females with papillary structures in ventrolateral positions (
<figureCitation id="E6D02A315065FFAD128FFD55FB230EC2" box="[1167,1277,651,678]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="51" pageNumber="52">Figs 18B</figureCitation>
,
<figureCitation id="E6D02A315065FFAD1308FD52FAE00EC2" box="[1288,1342,652,678]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="51" pageNumber="52">19G</figureCitation>
); the structures are quite prominent in LM (
<figureCitation id="E6D02A315065FFAD147FFD71FD320EAE" box="[639,748,687,714]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="51" pageNumber="52">Fig. 19G</figureCitation>
), but hardly visible in SEM. Glandular cell outlets
<typeStatus id="A15088165065FFAD1344FD6EFAA90EAE" box="[1348,1399,688,714]" pageId="51" pageNumber="52">type</typeStatus>
1, pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="7E5436B45065FFAD16BDFCC8FBB0086C" blockId="51.[189,1399,789,1032]" pageId="51" pageNumber="52">
Segment 8 with acicular spines in middorsal and lateroventral positions (
<figureCitation id="E6D02A315065FFAD122EFCCBFB570F54" box="[1070,1161,789,816]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="51" pageNumber="52">Figs 18</figureCitation>
AB, 19FG). Very minute glandular cell outlets
<typeStatus id="A15088165065FFAD1416FCE4FD970F30" box="[534,585,826,852]" pageId="51" pageNumber="52">type</typeStatus>
2 present in laterodorsal positions (
<figureCitation id="E6D02A315065FFAD15F0FCE7FBBF0F30" box="[1008,1121,825,852]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="51" pageNumber="52">Figs 18A</figureCitation>
,
<figureCitation id="E6D02A315065FFAD126EFCE4FB420F30" box="[1134,1180,826,852]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="51" pageNumber="52">19F</figureCitation>
,
<figureCitation id="E6D02A315065FFAD12A8FCE4FB190F30" box="[1192,1223,826,852]" captionStart="Fig" captionStartId="48.[189,232,1647,1673]" captionTargetBox="[191,1396,382,1597]" captionTargetId="figure@48.[191,1396,382,1597]" captionTargetPageId="48" captionText="Fig. 20. Scanning electron micrographs showing overviews and details of Echinoderes lupherorum sp. nov. A. Lateroventral overview. B. Laterodorsal overview. C. Introvert sectors 10 and 9. D. Segments 1 to 3, ventral view. E. Segments 1 to 2, dorsal view. F. Detail showing middorsal (right) to laterodorsal (left) parts of segments 1 and 2. Inset shows close-up of subdorsal glandular cell outlet type 2; scale = 1 µm. G. Detail showing middorsal (right) to laterodorsal (left) parts of 8. H. Close-up showing laterodorsal sensory spot and glandular cell outlet type 2 on segment 8. I. Segments 9 to 10, lateral view. J. ♀, segments 10 to 11, dorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvt = lateroventral tube; mds = middorsal spine; mdss = middorsal sensory spot; mlt = midlateral tube; pdss = paradorsal sensory spot; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid." figureDoi="http://doi.org/10.5281/zenodo.3817857" httpUri="https://zenodo.org/record/3817857/files/figure.png" pageId="51" pageNumber="52">20</figureCitation>
GH). Sensory spots present in paradorsal (
<figureCitation id="E6D02A315065FFAD1410FC83FD5F0F1C" box="[528,641,861,888]" captionStart="Fig" captionStartId="48.[189,232,1647,1673]" captionTargetBox="[191,1396,382,1597]" captionTargetId="figure@48.[191,1396,382,1597]" captionTargetPageId="48" captionText="Fig. 20. Scanning electron micrographs showing overviews and details of Echinoderes lupherorum sp. nov. A. Lateroventral overview. B. Laterodorsal overview. C. Introvert sectors 10 and 9. D. Segments 1 to 3, ventral view. E. Segments 1 to 2, dorsal view. F. Detail showing middorsal (right) to laterodorsal (left) parts of segments 1 and 2. Inset shows close-up of subdorsal glandular cell outlet type 2; scale = 1 µm. G. Detail showing middorsal (right) to laterodorsal (left) parts of 8. H. Close-up showing laterodorsal sensory spot and glandular cell outlet type 2 on segment 8. I. Segments 9 to 10, lateral view. J. ♀, segments 10 to 11, dorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvt = lateroventral tube; mds = middorsal spine; mdss = middorsal sensory spot; mlt = midlateral tube; pdss = paradorsal sensory spot; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid." figureDoi="http://doi.org/10.5281/zenodo.3817857" httpUri="https://zenodo.org/record/3817857/files/figure.png" pageId="51" pageNumber="52">Fig. 20G</figureCitation>
), subdorsal (
<figureCitation id="E6D02A315065FFAD151BFC83FC5A0F1C" box="[795,900,861,888]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="51" pageNumber="52">Fig. 19F</figureCitation>
) and laterodorsal (
<figureCitation id="E6D02A315065FFAD1268FC83FB610F1C" box="[1128,1215,861,888]" captionStart="Fig" captionStartId="48.[189,232,1647,1673]" captionTargetBox="[191,1396,382,1597]" captionTargetId="figure@48.[191,1396,382,1597]" captionTargetPageId="48" captionText="Fig. 20. Scanning electron micrographs showing overviews and details of Echinoderes lupherorum sp. nov. A. Lateroventral overview. B. Laterodorsal overview. C. Introvert sectors 10 and 9. D. Segments 1 to 3, ventral view. E. Segments 1 to 2, dorsal view. F. Detail showing middorsal (right) to laterodorsal (left) parts of segments 1 and 2. Inset shows close-up of subdorsal glandular cell outlet type 2; scale = 1 µm. G. Detail showing middorsal (right) to laterodorsal (left) parts of 8. H. Close-up showing laterodorsal sensory spot and glandular cell outlet type 2 on segment 8. I. Segments 9 to 10, lateral view. J. ♀, segments 10 to 11, dorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvt = lateroventral tube; mds = middorsal spine; mdss = middorsal sensory spot; mlt = midlateral tube; pdss = paradorsal sensory spot; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid." figureDoi="http://doi.org/10.5281/zenodo.3817857" httpUri="https://zenodo.org/record/3817857/files/figure.png" pageId="51" pageNumber="52">Fig. 20</figureCitation>
GH) positions (one specimen without subdorsal sensory spots though (
<figureCitation id="E6D02A315065FFAD155DFC5FFC130FF8" box="[861,973,897,924]" captionStart="Fig" captionStartId="48.[189,232,1647,1673]" captionTargetBox="[191,1396,382,1597]" captionTargetId="figure@48.[191,1396,382,1597]" captionTargetPageId="48" captionText="Fig. 20. Scanning electron micrographs showing overviews and details of Echinoderes lupherorum sp. nov. A. Lateroventral overview. B. Laterodorsal overview. C. Introvert sectors 10 and 9. D. Segments 1 to 3, ventral view. E. Segments 1 to 2, dorsal view. F. Detail showing middorsal (right) to laterodorsal (left) parts of segments 1 and 2. Inset shows close-up of subdorsal glandular cell outlet type 2; scale = 1 µm. G. Detail showing middorsal (right) to laterodorsal (left) parts of 8. H. Close-up showing laterodorsal sensory spot and glandular cell outlet type 2 on segment 8. I. Segments 9 to 10, lateral view. J. ♀, segments 10 to 11, dorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvt = lateroventral tube; mds = middorsal spine; mdss = middorsal sensory spot; mlt = midlateral tube; pdss = paradorsal sensory spot; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid." figureDoi="http://doi.org/10.5281/zenodo.3817857" httpUri="https://zenodo.org/record/3817857/files/figure.png" pageId="51" pageNumber="52">Fig. 20G</figureCitation>
), and one too dirty to confirm their presence). Female specimens with papillary structures in ventromedial positions (
<figureCitation id="E6D02A315065FFAD12ACFC7BFAFF0FA4" box="[1196,1313,933,960]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="51" pageNumber="52">Figs 18B</figureCitation>
,
<figureCitation id="E6D02A315065FFAD1333FC78FAB70FA4" box="[1331,1385,934,960]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="51" pageNumber="52">19G</figureCitation>
). Glandular cell outlets
<typeStatus id="A15088165065FFAD17C8FC14FE250F80" box="[456,507,970,996]" pageId="51" pageNumber="52">type</typeStatus>
1 present in subdorsal (
<figureCitation id="E6D02A315065FFAD1520FC17FC560F87" box="[800,904,969,995]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="51" pageNumber="52">Fig. 19F</figureCitation>
) and ventromedial positions. Pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="7E5436B45065FFAD16BDFBEEFCC30946" blockId="51.[189,1399,1071,1314]" pageId="51" pageNumber="52">
Segment 9 with acicular spines in lateroventral positions (
<figureCitation id="E6D02A315065FFAD1565FBF1FC0A082E" box="[869,980,1071,1098]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="51" pageNumber="52">Figs 18B</figureCitation>
). Very minute glandular cell outlets
<typeStatus id="A15088165065FFAD16BDFB8AFF2E080A" box="[189,240,1108,1134]" pageId="51" pageNumber="52">type</typeStatus>
2 present in laterodorsal positions (
<figureCitation id="E6D02A315065FFAD14B5FB8DFCF5080A" box="[693,811,1107,1134]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="51" pageNumber="52">Figs 18A</figureCitation>
,
<figureCitation id="E6D02A315065FFAD153DFB8AFCB5080A" box="[829,875,1108,1134]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="51" pageNumber="52">19F</figureCitation>
,
<figureCitation id="E6D02A315065FFAD157CFB8AFC79080A" box="[892,935,1108,1134]" captionStart="Fig" captionStartId="48.[189,232,1647,1673]" captionTargetBox="[191,1396,382,1597]" captionTargetId="figure@48.[191,1396,382,1597]" captionTargetPageId="48" captionText="Fig. 20. Scanning electron micrographs showing overviews and details of Echinoderes lupherorum sp. nov. A. Lateroventral overview. B. Laterodorsal overview. C. Introvert sectors 10 and 9. D. Segments 1 to 3, ventral view. E. Segments 1 to 2, dorsal view. F. Detail showing middorsal (right) to laterodorsal (left) parts of segments 1 and 2. Inset shows close-up of subdorsal glandular cell outlet type 2; scale = 1 µm. G. Detail showing middorsal (right) to laterodorsal (left) parts of 8. H. Close-up showing laterodorsal sensory spot and glandular cell outlet type 2 on segment 8. I. Segments 9 to 10, lateral view. J. ♀, segments 10 to 11, dorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvt = lateroventral tube; mds = middorsal spine; mdss = middorsal sensory spot; mlt = midlateral tube; pdss = paradorsal sensory spot; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid." figureDoi="http://doi.org/10.5281/zenodo.3817857" httpUri="https://zenodo.org/record/3817857/files/figure.png" pageId="51" pageNumber="52">20I</figureCitation>
). Sensory spots present in paradorsal (
<figureCitation id="E6D02A315065FFAD16C6FBA9FEF308F6" box="[198,301,1143,1170]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="51" pageNumber="52">Fig. 19F</figureCitation>
), subdorsal (
<figureCitation id="E6D02A315065FFAD17C1FBA9FDF608F6" box="[449,552,1143,1170]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="51" pageNumber="52">Fig. 19F</figureCitation>
), laterodorsal (
<figureCitation id="E6D02A315065FFAD14D7FBA9FCE808F6" box="[727,822,1143,1170]" captionStart="Fig" captionStartId="48.[189,232,1647,1673]" captionTargetBox="[191,1396,382,1597]" captionTargetId="figure@48.[191,1396,382,1597]" captionTargetPageId="48" captionText="Fig. 20. Scanning electron micrographs showing overviews and details of Echinoderes lupherorum sp. nov. A. Lateroventral overview. B. Laterodorsal overview. C. Introvert sectors 10 and 9. D. Segments 1 to 3, ventral view. E. Segments 1 to 2, dorsal view. F. Detail showing middorsal (right) to laterodorsal (left) parts of segments 1 and 2. Inset shows close-up of subdorsal glandular cell outlet type 2; scale = 1 µm. G. Detail showing middorsal (right) to laterodorsal (left) parts of 8. H. Close-up showing laterodorsal sensory spot and glandular cell outlet type 2 on segment 8. I. Segments 9 to 10, lateral view. J. ♀, segments 10 to 11, dorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvt = lateroventral tube; mds = middorsal spine; mdss = middorsal sensory spot; mlt = midlateral tube; pdss = paradorsal sensory spot; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid." figureDoi="http://doi.org/10.5281/zenodo.3817857" httpUri="https://zenodo.org/record/3817857/files/figure.png" pageId="51" pageNumber="52">Fig. 20I</figureCitation>
) and ventrolateral positions. Small rounded sieve plates are present in lateral accessory positions (
<figureCitation id="E6D02A315065FFAD14FAFB45FCB808D2" box="[762,870,1179,1206]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="51" pageNumber="52">Fig. 18B</figureCitation>
). Glandular cell outlets
<typeStatus id="A15088165065FFAD1284FB42FB6908D2" box="[1156,1207,1180,1206]" pageId="51" pageNumber="52">type</typeStatus>
1 are present in subdorsal and ventromedial positions. Cuticular hair covering on tergal plate is less dense than on the preceding segments, and the subdorsal to middorsal areas have no hair at all. Pachycycli and pectinate fringe of posterior margin as on preceding segment.
</paragraph>
<paragraph id="7E5436B45065FFAD16BDFA94FAEE0A58" blockId="51.[189,1399,1353,1596]" pageId="51" pageNumber="52">
Segment 10 with tubes in midlateral positions (
<figureCitation id="E6D02A315065FFAD14E3FA97FCE00900" box="[739,830,1353,1380]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="51" pageNumber="52">Figs 18</figureCitation>
,
<figureCitation id="E6D02A315065FFAD154BFA94FC470900" box="[843,921,1354,1380]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="51" pageNumber="52">19H, J</figureCitation>
,
<figureCitation id="E6D02A315065FFAD15A7FA94FC1B0900" box="[935,965,1354,1380]" captionStart="Fig" captionStartId="48.[189,232,1647,1673]" captionTargetBox="[191,1396,382,1597]" captionTargetId="figure@48.[191,1396,382,1597]" captionTargetPageId="48" captionText="Fig. 20. Scanning electron micrographs showing overviews and details of Echinoderes lupherorum sp. nov. A. Lateroventral overview. B. Laterodorsal overview. C. Introvert sectors 10 and 9. D. Segments 1 to 3, ventral view. E. Segments 1 to 2, dorsal view. F. Detail showing middorsal (right) to laterodorsal (left) parts of segments 1 and 2. Inset shows close-up of subdorsal glandular cell outlet type 2; scale = 1 µm. G. Detail showing middorsal (right) to laterodorsal (left) parts of 8. H. Close-up showing laterodorsal sensory spot and glandular cell outlet type 2 on segment 8. I. Segments 9 to 10, lateral view. J. ♀, segments 10 to 11, dorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvt = lateroventral tube; mds = middorsal spine; mdss = middorsal sensory spot; mlt = midlateral tube; pdss = paradorsal sensory spot; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid." figureDoi="http://doi.org/10.5281/zenodo.3817857" httpUri="https://zenodo.org/record/3817857/files/figure.png" pageId="51" pageNumber="52">20</figureCitation>
IJ); tubes appear fairly stout and are usually much easier to observe in LM (
<figureCitation id="E6D02A315065FFAD1474FAB3FD0509EC" box="[628,731,1389,1416]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="51" pageNumber="52">Fig. 19H</figureCitation>
) than tubes on segment
<quantity id="B9139B515065FFAD15EBFAB0FBFD09E3" box="[1003,1059,1389,1416]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="51" pageNumber="52" unit="in" value="10.0">10 in</quantity>
other species of
<taxonomicName id="B9EB4D375065FFAD12E0FAB3FAB009E3" authorityName="Claparede" authorityYear="1863" box="[1248,1390,1389,1415]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="51" pageNumber="52" phylum="Cephalorhyncha" rank="genus">
<emphasis id="4C9FEAA65065FFAD12E0FAB3FAB009E3" box="[1248,1390,1389,1415]" italics="true" pageId="51" pageNumber="52">Echinoderes</emphasis>
</taxonomicName>
; differentiated bases and lateral wings apparently not present. Sensory spots are present in paradorsal and ventrolateral positions. Glandular cell outlets
<typeStatus id="A15088165065FFAD1500FA68FCEF09B4" box="[768,817,1462,1488]" pageId="51" pageNumber="52">type</typeStatus>
1 are present as two middorsal ones and a pair in paraventral positions. Cuticular hairs are scattered over the tergal plate and lateral parts of the sternal plates, but not as densely as on preceding segments. The posterior segment margin of the tergal plate is straight, whereas the margins of the sternal plates are broadly concave. Pachycycli as on preceding segment.
</paragraph>
<paragraph id="7E5436B45065FFAD16BDF9BAFAC70B31" blockId="51.[189,1399,1635,1878]" pageId="51" pageNumber="52">
Segment 11 with long lateral terminal spines (
<figureCitation id="E6D02A315065FFAD14FCF9BDFCB70A1A" box="[764,873,1635,1662]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="51" pageNumber="52">Fig. 18C</figureCitation>
). Males with three pairs of penile spines (
<figureCitation id="E6D02A315065FFAD16C5F959FEC10AC6" box="[197,287,1671,1698]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="51" pageNumber="52">Figs 18</figureCitation>
DE, 19J); dorsal and ventral penile spines are thin, and quite long flexible tubes, whereas the median one is thick, conical, and stout. Females with lateral terminal accessory spines (
<figureCitation id="E6D02A315065FFAD12F9F975FA8E0AA1" box="[1273,1360,1707,1733]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="51" pageNumber="52">Figs 18</figureCitation>
A B, 19H). Sensory spots present in subdorsal positions. The segment is completely devoid of cuticular hairs, besides the short hair-like extensions in the mid- to subdorsal areas. Tergal extensions are long and slender, extending into long but yet delicate spinous tips (
<figureCitation id="E6D02A315065FFAD15A5F8C9FBDF0B55" box="[933,1025,1815,1841]" captionStart="Fig" captionStartId="46.[189,232,1754,1780]" captionTargetBox="[278,1310,269,1745]" captionTargetId="figure@46.[277,1310,265,1745]" captionTargetPageId="46" captionText="Fig. 18. Line art illustrations of Echinoderes lupherorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlss = midlateral sensory spot; mlt = midlateral tube; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slss = sublateral sensory spot; te = tergal extension." figureDoi="http://doi.org/10.5281/zenodo.3817853" httpUri="https://zenodo.org/record/3817853/files/figure.png" pageId="51" pageNumber="52">Figs 18</figureCitation>
,
<figureCitation id="E6D02A315065FFAD1210F8C9FBF20B55" box="[1040,1068,1815,1841]" captionStart="Fig" captionStartId="47.[189,232,1647,1673]" captionTargetBox="[233,1360,265,1627]" captionTargetId="figure@47.[233,1360,265,1627]" captionTargetPageId="47" captionText="Fig. 19. Light micrographs showing overviews and details of Echinoderes lupherorum sp. nov., holotype, ♀ (AC, H) (NHMD-225220), paratype, ♀ (DG) (NHMD-225226) and paratype, ♂ (IJ) (NHMD-225225). A. Ventral overview. B. Segments 1 to 3, dorsal view. C. Segments 1 to 3, ventral view. D. Segments 3 to 6, dorsal view. E. Segments 3 to 6, ventral view. F. Segments 8 to 9, dorsal view. G. Segments 7 to 8, ventral view. H. ♀, segments 10 to 11, ventral view. I. ♂, segments 10 to 11, dorsal view. J. ♂, segments 10 to 11, ventral view. Abbreviations: fpa = female papillae; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlt = midlateral tube; pdss = paradorsal sensory spot; pe = penile spine; sdgco1/2 = subdorsal glandular cell outlet type 1 or 2; sdss = subdorsal sensory spot; te = tergal extensions; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817855" httpUri="https://zenodo.org/record/3817855/files/figure.png" pageId="51" pageNumber="52">19</figureCitation>
HI, 20J); tergal extensions measure
<quantity id="B9139B515065FFAD1726F8E5FE660B32" box="[294,440,1851,1878]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="2.8" metricValueMin="2.4" pageId="51" pageNumber="52" unit="mm" value="26.0" valueMax="28.0" valueMin="24.0">24 to 28 µm</quantity>
, equal to 6 to 8% of trunk length. Sternal extensions shorter and rounded.
</paragraph>
</subSubSection>
<subSubSection id="36F1653F5065FFAD16BDF8A0FD5F0B8F" pageId="51" pageNumber="52" type="discussion">
<paragraph id="7E5436B45065FFAD16BDF8A0FEF10BFC" blockId="51.[189,303,1918,1944]" box="[189,303,1918,1944]" pageId="51" pageNumber="52">
<heading id="251C81D85065FFAD16BDF8A0FEF10BFC" bold="true" box="[189,303,1918,1944]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="4C9FEAA65065FFAD16BDF8A0FEF10BFC" bold="true" box="[189,303,1918,1944]" pageId="51" pageNumber="52">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="7E5436B45065FFAD16BDF873FD5F0B8F" blockId="51.[189,1398,1965,2027]" pageId="51" pageNumber="52">
Due to their close resemblance, diagnostic remarks for
<taxonomicName id="B9EB4D375065FFAD1537F870FC010BA3" authority="Sørensen &amp; Rohal &amp; Thistle, 2018" authorityName="Sørensen &amp; Rohal &amp; Thistle" authorityYear="2018" box="[823,991,1965,1991]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="51" pageNumber="52" phylum="Cephalorhyncha" rank="species" species="lupherorum" status="sp. nov.">
<emphasis id="4C9FEAA65065FFAD1537F870FC010BA3" box="[823,991,1965,1991]" italics="true" pageId="51" pageNumber="52">E. lupherorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57AC57DD5065FFAD15E5F873FBE00BA3" box="[997,1086,1965,1991]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
will follow below together with remarks for
<taxonomicName id="B9EB4D375065FFAD1788F80CFDC10B8F" authority="Sørensen &amp; Rohal &amp; Thistle, 2018" authorityName="Sørensen &amp; Rohal &amp; Thistle" authorityYear="2018" box="[392,543,2001,2027]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="51" pageNumber="52" phylum="Cephalorhyncha" rank="species" species="yamasakii" status="sp. nov.">
<emphasis id="4C9FEAA65065FFAD1788F80CFDC10B8F" box="[392,543,2001,2027]" italics="true" pageId="51" pageNumber="52">E. yamasakii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57AC57DD5065FFAD1427F80FFD5F0B8F" box="[551,641,2001,2027]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>