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<document id="D62739DFF223756C29D3924225B6AABA" ID-CLB-Dataset="24911" ID-DOI="10.11646/phytotaxa.17.1" ID-GBIF-Dataset="65962ba7-eec1-40e7-aed6-cef94e99ca39" ID-ISSN="1179-3163" ID-Zenodo-Dep="3538362" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573511149647" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2011" docId="FA4887FA242EBD76FF37DA8C1EA4A720" docLanguage="en" docName="Phytotaxa.17.1.pdf.imd" docOrigin="Phytotaxa 17" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Geonoma concinna Burret 1930" docType="treatment" docVersion="7" lastPageNumber="50" masterDocId="0671FF82241EBD47FFA0DE421B3FA301" masterDocTitle="A revision of Geonoma (Arecaceae)" masterLastPageNumber="271" masterPageNumber="1" pageNumber="49" updateTime="1698765326825" updateUser="ExternalLinkService">
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<mods:title id="54FF82220C60B93FD2759B04911BB0E8">A revision of Geonoma (Arecaceae)</mods:title>
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<mods:namePart id="193F5FB685DF073420A67C284AEE8A78">Henderson, Andrew</mods:namePart>
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<mods:title id="2BD2A808FC9A0ED989318ACE6CCCEEAC">Phytotaxa</mods:title>
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<mods:date id="FAA4859FAEE58F37C0ABECD69DC18E40">2011</mods:date>
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<treatment id="FA4887FA242EBD76FF37DA8C1EA4A720" ID-DOI="http://doi.org/10.5281/zenodo.5608918" ID-GBIF-Taxon="160342495" ID-Zenodo-Dep="5608918" LSID="urn:lsid:plazi:treatment:FA4887FA242EBD76FF37DA8C1EA4A720" httpUri="http://treatment.plazi.org/id/FA4887FA242EBD76FF37DA8C1EA4A720" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<subSubSection id="3AFB6567242EBD77FF37DA8C1877A65C" pageId="48" pageNumber="49" type="nomenclature">
<paragraph id="725E36EC242EBD77FF37DA8C1877A65C" blockId="48.[151,1437,1230,1373]" pageId="48" pageNumber="49">
<emphasis id="4095EAFE242EBD77FF37DA8C1B83A7E9" bold="true" box="[151,188,1230,1256]" pageId="48" pageNumber="49">11.</emphasis>
<taxonomicName id="B5E14D6F242EBD77FF64DA8C1983A7E9" ID-CoL="3FQHP" authority="Burret (1930 a: 229)" authorityName="Burret" authorityYear="1930" box="[196,700,1230,1256]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="concinna">
<emphasis id="4095EAFE242EBD77FF64DA8C1A84A7E9" bold="true" box="[196,443,1230,1256]" italics="true" pageId="48" pageNumber="49">Geonoma concinna</emphasis>
<bibRefCitation id="16704B1D242EBD77FE63DA8C1983A7E9" author="Burret, M." box="[451,700,1230,1256]" journalOrPublisher="Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie" pageId="48" pageNumber="49" pagination="123 - 270" part="63" refId="ref121027" refString="Burret, M. (1930 a) Geonomeae americanae. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 63: 123 - 270." title="Geonomeae americanae" type="journal article" year="1930">Burret (1930a: 229)</bibRefCitation>
</taxonomicName>
. Type:
<collectingCountry id="0AF6767C242EBD77FCB6DA8C1884A7E9" box="[790,955,1230,1256]" name="Colombia" pageId="48" pageNumber="49">COLOMBIA</collectingCountry>
.
<collectingRegion id="B025F80E242EBD77FC68DA8C1F77A7E9" box="[968,1096,1230,1256]" country="Colombia" name="Antioquia" pageId="48" pageNumber="49">Antioquia</collectingRegion>
: Tabor,
<quantity id="B5199B09242EBD77FB09DA8C1E35A7E9" box="[1193,1290,1230,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="48" pageNumber="49" unit="m" value="1950.0">1950 m</quantity>
,
<date id="065F102C242EBD77FAB7DA8C1BE9A60E" pageId="48" pageNumber="49" value="1880-01-27">27 January 1880</date>
,
<emphasis id="4095EAFE242EBD77FF43DAB41AFAA60E" box="[227,453,1269,1295]" italics="true" pageId="48" pageNumber="49">W. Kalbreyer 1367</emphasis>
(
<typeStatus id="AD5A884E242EBD77FE75DAB7197FA60E" box="[469,576,1269,1295]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
B, destroyed).
<typeStatus id="AD5A884E242EBD77FD5BDAB7185FA60E" box="[763,864,1269,1295]" pageId="48" pageNumber="49" type="neotype">Neotype</typeStatus>
(selected by
<bibRefCitation id="16704B1D242EBD77FBA2DAB71FDFA60E" author="Bernal, R. &amp; Galeano-Garces, G. &amp; Henderson, A." box="[1026,1248,1269,1295]" journalOrPublisher="Taxon" pageId="48" pageNumber="49" pagination="98 - 107" part="38" refId="ref120832" refString="Bernal, R., Galeano-Garces, G. &amp; Henderson, A. (1989) Neotypification of Colombian palms collected by W. Kalbreyer. Taxon 38: 98 - 107." title="Neotypification of Colombian palms collected by W. Kalbreyer" type="journal article" year="1989">
Bernal
<emphasis id="4095EAFE242EBD77FBFADAB41FA8A60E" box="[1114,1175,1269,1295]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
1989
</bibRefCitation>
):
<collectingCountry id="0AF6767C242EBD77FB56DAB71EA6A60E" box="[1270,1433,1269,1295]" name="Colombia" pageId="48" pageNumber="49">COLOMBIA</collectingCountry>
.
<collectingRegion id="B025F80E242EBD77FF37DB5E1A2BA637" box="[151,276,1308,1334]" country="Colombia" name="Antioquia" pageId="48" pageNumber="49">Antioquia</collectingRegion>
: carretera Granada-San Luis,
<quantity id="B5199B09242EBD77FDDDDB5E19EEA637" box="[637,721,1308,1334]" metricMagnitude="3" metricUnit="m" metricValue="5.5" pageId="48" pageNumber="49" unit="km" value="5.5">5.5 km</quantity>
adelante de El Chocó,
<quantity id="B5199B09242EBD77FC4EDB5E1F73A637" box="[1006,1100,1308,1334]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="48" pageNumber="49" unit="m" value="1750.0">1750 m</quantity>
,
<date id="065F102C242EBD77FBFADB5E1E4AA637" box="[1114,1397,1308,1334]" pageId="48" pageNumber="49" value="1987-09-20" valueMax="1987-09-21" valueMin="1987-09-20">2021 September 1987</date>
,
<emphasis id="4095EAFE242EBD77FA22DB5F1A8BA65C" italics="true" pageId="48" pageNumber="49">R. Bernal &amp; L. Tobón 1358</emphasis>
(
<typeStatus id="AD5A884E242EBD77FE64DB01191DA65C" box="[452,546,1347,1373]" pageId="48" pageNumber="49" type="neotype">neotype</typeStatus>
COL!, isoneotype NY!).
</paragraph>
</subSubSection>
<subSubSection id="3AFB6567242EBD76FF37DBD2184FA06B" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="description">
<paragraph id="725E36EC242EBD76FF37DBD21FFCA2E6" blockId="48.[151,1437,1424,2030]" lastBlockId="49.[151,1437,152,874]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<emphasis id="4095EAFE242EBD77FF37DBD21BDAA6AB" box="[151,229,1424,1450]" italics="true" pageId="48" pageNumber="49">Plants</emphasis>
1.8(1.52.5) m tall; stems
<quantity id="B5199B09242EBD77FD92DBD21948A6AB" box="[562,631,1424,1450]" metricMagnitude="0" metricUnit="m" metricValue="3.5" pageId="48" pageNumber="49" unit="m" value="3.5">3.5 m</quantity>
tall, 1.0(0.91.2) cm in diameter, clustered, cane-like; internodes 3.5(2.54.5) cm long, yellowish and smooth.
<emphasis id="4095EAFE242EBD77FD0FDBFA183EA6D0" box="[687,769,1464,1489]" italics="true" pageId="48" pageNumber="49">Leaves</emphasis>
8 per stem, undivided or irregularly pinnate, not plicate, bases of blades running diagonally into the rachis; sheaths 14.0 cm long; petioles 9.0 cm long, drying green or yellowish; rachis 27.4(20.338.5) cm long, 3.4(3.13.8) mm in diameter; veins raised and rectangular in cross-section adaxially; pinnae 2(14) per side of rachis; basal pinna 30.5(24.037.0) cm long, 8.2(8.08.4) cm wide, forming an angle of 44(3555)° with the rachis; apical pinna 17.9(14.020.8) cm long, 10.0(8.0 12.0) cm wide, forming an angle of 38(3242)° with the rachis.
<emphasis id="4095EAFE242EBD77FC34D83A1F01A593" box="[916,1086,1656,1682]" italics="true" pageId="48" pageNumber="49">Inflorescences</emphasis>
branched 2 orders; prophylls and peduncular bracts not ribbed with elongate, unbranched fibers, flattened, deciduous; prophylls 10.8(9.0 12.5) cm long, not short and asymmetrically apiculate, the surfaces ridged with close, equal, parallel, nondividing ridges, scarcely tomentose between the ridges, without unequally wide ridges; peduncular bracts
<quantity id="B5199B09242EBD77FAD8D8AE1B83A42C" metricMagnitude="-2" metricUnit="m" metricValue="7.2" pageId="48" pageNumber="49" unit="cm" value="7.2">7.2 cm</quantity>
long, well-developed, inserted 0.4(0.20.6) cm above the prophyll; peduncles 11.1(8.213.2) cm long, 3.6(3.04.2) mm in diameter; rachillae 19(1321), 1.7(8.014.8) cm long, 1.7(1.22.7) mm in diameter, the surfaces without spiky, fibrous projections or ridges, drying brown, with faint to pronounced, short, transverse ridges, not filiform and not narrowed between the flower pits; flower pits spirally arranged, glabrous internally; proximal lips without a central notch before anthesis, not recurved after anthesis, hood-shaped at anthesis, sometimes splitting post-anthesis; proximal and distal lips drying the same color as the rachillae, not joined to form a raised cupule, the proximal lip margins overlapping the distal lip margins; distal lips welldeveloped; staminate and pistillate petals not emergent, not valvate throughout; staminate flowers deciduous after anthesis; stamens 6; thecae diverging at anthesis, inserted almost directly onto the filament apices, the connectives bifid but scarcely developed; anthers short and curled over at anthesis; non-fertilized pistillate flowers deciduous after anthesis; staminodial tubes crenulate or shallowly lobed at the apex, those of nonfertilized flowers not projecting and persistent after anthesis;
<emphasis id="4095EAFE242FBD76FCC5DF1B189CA272" box="[869,931,345,371]" italics="true" pageId="49" pageNumber="50">fruits</emphasis>
7.7(7.18.3) mm long, 6.7(6.17.3) mm in diameter, the bases without a prominent stipe, the apices not conical, the surfaces not splitting at maturity, without fibers emerging, bumpy from the numerous, subepidermal, tangential, short fibers present, these coming to a point at fruit apices; locular epidermis with operculum, smooth, without pores.
</paragraph>
<paragraph id="725E36EC242FBD76FF66DFB618CDA1A9" blockId="49.[151,1437,152,874]" pageId="49" pageNumber="50">
<emphasis id="4095EAFE242FBD76FF66DFB61A81A10F" bold="true" box="[198,446,500,526]" pageId="49" pageNumber="50">Taxonomic notes:—</emphasis>
<taxonomicName id="B5E14D6F242FBD76FE1EDFB6199DA10F" authorityName="Burret" authorityYear="1930" box="[446,674,500,526]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="concinna">
<emphasis id="4095EAFE242FBD76FE1EDFB6199DA10F" box="[446,674,500,526]" italics="true" pageId="49" pageNumber="50">Geonoma concinna</emphasis>
</taxonomicName>
is a member of the
<taxonomicName id="B5E14D6F242FBD76FC29DFB61F2FA10F" authorityName="Wendland ex Spruce" authorityYear="1871" box="[905,1040,500,526]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="congesta">
<emphasis id="4095EAFE242FBD76FC29DFB61F2FA10F" box="[905,1040,500,526]" italics="true" pageId="49" pageNumber="50">G. congesta</emphasis>
</taxonomicName>
clade, and most closely related to
<taxonomicName id="B5E14D6F242FBD76FF37DC581A5AA135" authorityName="Henderson" authorityYear="2011" box="[151,357,538,564]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="concinnoidea">
<emphasis id="4095EAFE242FBD76FF37DC581A5AA135" box="[151,357,538,564]" italics="true" pageId="49" pageNumber="50">G. concinnoidea</emphasis>
</taxonomicName>
, from which it differs by its fruits without fibers emerging.
<taxonomicName id="B5E14D6F242FBD76FBCDDC581E5CA134" authorityName="Burret" authorityYear="1930" box="[1133,1379,538,565]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="concinna">
<emphasis id="4095EAFE242FBD76FBCDDC581E5CA134" box="[1133,1379,538,565]" italics="true" pageId="49" pageNumber="50">Geonoma concinna</emphasis>
</taxonomicName>
was considered by
<bibRefCitation id="16704B1D242FBD76FEE3DC031965A15A" author="Henderson, A. &amp; Galeano, G. &amp; Bernal, R." box="[323,602,577,603]" journalOrPublisher="Princeton University Press, Princeton, New Jersey" pageId="49" pageNumber="50" pagination="1 - 352" refId="ref122861" refString="Henderson, A., Galeano, G. &amp; Bernal, R. (1995) A field guide to the palms of the Americas. Princeton University Press, Princeton, New Jersey, pp. 1 - 352." title="A field guide to the palms of the Americas" type="book chapter" year="1995">
Henderson
<emphasis id="4095EAFE242FBD76FE69DC00193BA15A" box="[457,516,577,603]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
(1995)
</bibRefCitation>
to be a poorly understood species. This is still true because of the lack of specimens (only four specimens known). In Henderson
<emphasis id="4095EAFE242FBD76FCE7DC2B18B6A183" box="[839,905,616,642]" italics="true" pageId="49" pageNumber="50">et al.,</emphasis>
specimens from
<collectingCountry id="0AF6767C242FBD76FBF3DC2A1F8FA183" box="[1107,1200,616,642]" name="Panama" pageId="49" pageNumber="50">Panama</collectingCountry>
were included in
<taxonomicName id="B5E14D6F242FBD76FA23DC2A1A3CA1A8" authorityName="Burret" authorityYear="1930" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="concinna">
<emphasis id="4095EAFE242FBD76FA23DC2A1A3CA1A8" italics="true" pageId="49" pageNumber="50">G. concinna</emphasis>
</taxonomicName>
; here they are recognized as a distinct species,
<taxonomicName id="B5E14D6F242FBD76FC8DDCCC18D4A1A9" authorityName="Henderson" authorityYear="2011" box="[813,1003,654,680]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="concinnoidea">
<emphasis id="4095EAFE242FBD76FC8DDCCC18D4A1A9" box="[813,1003,654,680]" italics="true" pageId="49" pageNumber="50">G. concinnoidea</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="725E36EC242FBD76FF66DCF7184FA06B" blockId="49.[151,1437,152,874]" pageId="49" pageNumber="50">
<emphasis id="4095EAFE242FBD76FF66DCF71AC1A1CE" bold="true" box="[198,510,693,719]" pageId="49" pageNumber="50">Subspecific variation:—</emphasis>
One trait (leaf division) varies within this species. There is geographic discontinuity and specimens occur in two separate areas in
<collectingCountry id="0AF6767C242FBD76FCDEDC9E18C3A1F7" box="[894,1020,732,758]" name="Colombia" pageId="49" pageNumber="50">Colombia</collectingCountry>
. There are too few specimens for analysis. Based on the trait distribution and geography, two subgroups can be recognized and these are treated as subspecies (subspp.
<emphasis id="4095EAFE242FBD76FE16DD6819AAA042" box="[438,661,809,835]" italics="true" pageId="49" pageNumber="50">
<taxonomicName id="B5E14D6F242FBD76FE16DD681915A042" authorityName="Burret" authorityYear="1930" box="[438,554,810,835]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="concinna">concinna</taxonomicName>
, simplex
</emphasis>
). Subspecies
<taxonomicName id="B5E14D6F242FBD76FCE2DD68188CA042" authorityName="Burret" authorityYear="1930" box="[834,947,810,835]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="concinna">
<emphasis id="4095EAFE242FBD76FCE2DD68188CA042" box="[834,947,810,835]" italics="true" pageId="49" pageNumber="50">concinna</emphasis>
</taxonomicName>
occurs at higher elevations than
<taxonomicName id="B5E14D6F242FBD76FAEEDD6B1BCEA06B" authorityName="Henderson" authorityYear="2011" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="subSpecies" species="concinna" subSpecies="simplex">
subsp.
<emphasis id="4095EAFE242FBD76FF37DD121BCEA06B" box="[151,241,848,874]" italics="true" pageId="49" pageNumber="50">simplex</emphasis>
</taxonomicName>
; 1475(12001750) versus 777(755800) m elevation.
</paragraph>
</subSubSection>
<subSubSection id="3AFB6567242FBD76FF37DDDF1EA4A720" pageId="49" pageNumber="50" type="key">
<paragraph id="725E36EC242FBD76FF37DDDF1967A0B7" blockId="49.[151,600,924,951]" box="[151,600,924,951]" pageId="49" pageNumber="50">
<heading id="29168180242FBD76FF37DDDF1967A0B7" bold="true" box="[151,600,924,951]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="2">
<emphasis id="4095EAFE242FBD76FF37DDDF1967A0B7" bold="true" box="[151,600,924,951]" pageId="49" pageNumber="50">
Key to the subspecies of
<taxonomicName id="B5E14D6F242FBD76FE68DDDE1967A0B7" ID-CoL="3FQHP" authorityName="Burret" authorityYear="1930" box="[456,600,924,950]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="concinna">
<emphasis id="4095EAFE242FBD76FE68DDDE1967A0B7" bold="true" box="[456,600,924,950]" italics="true" pageId="49" pageNumber="50">G. concinna</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="725E36EC242FBD76FF37DDA81EA4A720" blockId="49.[151,1436,1002,1057]" pageId="49" pageNumber="50">
1 Leaves pinnate; Central
<collectingRegion id="B025F80E242FBD76FE6CDDA81907A700" box="[460,568,1002,1025]" country="Paraguay" name="Cordillera" pageId="49" pageNumber="50">Cordillera</collectingRegion>
.......................................................................................................
<taxonomicName id="B5E14D6F242FBD76FB50DDA81EA3A702" ID-CoL="7K4G4" box="[1264,1436,1002,1027]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="subSpecies" species="concinna" subSpecies="concinna">
subsp.
<emphasis id="4095EAFE242FBD76FA9ADDAE1EA3A702" box="[1338,1436,1004,1027]" italics="true" pageId="49" pageNumber="50">concinna</emphasis>
</taxonomicName>
- Leaves undivided; Western
<collectingRegion id="B025F80E242FBD76FE50DA481963A720" box="[496,604,1034,1057]" country="Paraguay" name="Cordillera" pageId="49" pageNumber="50">Cordillera</collectingRegion>
.....................................................................................................
<taxonomicName id="B5E14D6F242FBD76FAA6DA481EA4A720" ID-CoL="5HFVF" authorityName="Henderson" authorityYear="2011" box="[1286,1435,1034,1057]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="subSpecies" species="concinna" subSpecies="simplex">
subsp.
<emphasis id="4095EAFE242FBD76FAEADA481EA4A720" box="[1354,1435,1034,1057]" italics="true" pageId="49" pageNumber="50">simplex</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>