191 lines
18 KiB
XML
191 lines
18 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4762191" ID-GBIF-Dataset="df79feb7-c7e6-4267-b324-91597bc182fd" ID-GBIF-Taxon="182070955" ID-ISSN="1854-0392" ID-Zenodo-Dep="4762191" checkinTime="1620963448601" checkinUser="felipe" docAuthor="Sivec, Ignac & Stark, Bill P." docDate="2011" docId="ED2FE866E4697D6EFE84E4512C878D51" docLanguage="en" docName="Illiesia.7.24.264-279.pdf" docOrigin="Illiesia 7 (24)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Neoperla andreas Sivec & Stark 2011, sp. n." docType="treatment" docVersion="6" lastPageId="3" lastPageNumber="267" masterDocId="1116901EE46B7D6DFFDAE1242D7E8B32" masterDocTitle="New Species Of Neoperla Needham And Phanoperla Banks (Plecoptera: Perlidae) From The Philippine Islands" masterLastPageNumber="279" masterPageNumber="264" pageId="2" pageNumber="266" updateTime="1632023202857" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New Species Of Neoperla Needham And Phanoperla Banks (Plecoptera: Perlidae) From The Philippine Islands</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sivec, Ignac</mods:namePart>
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<mods:affiliation>Slovenian Museum of Natural History, Prešernova 20, P. O. Box 290, SLO- 1001 Ljubljana, Slovenia E-mail: isivec @ pms-lj. si</mods:affiliation>
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<mods:nameIdentifier type="email">isivec@pms-lj.si</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Stark, Bill P.</mods:namePart>
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<mods:affiliation>Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu</mods:affiliation>
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<mods:nameIdentifier type="email">stark@mc.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Illiesia</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>7</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>24</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>264</mods:start>
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<mods:end>279</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4760560</mods:identifier>
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<mods:identifier type="GBIF-Dataset">df79feb7-c7e6-4267-b324-91597bc182fd</mods:identifier>
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<mods:identifier type="ISSN">1854-0392</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4760560</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4762191" ID-GBIF-Taxon="182070955" ID-Zenodo-Dep="4762191" LSID="urn:lsid:plazi:treatment:ED2FE866E4697D6EFE84E4512C878D51" httpUri="http://treatment.plazi.org/id/ED2FE866E4697D6EFE84E4512C878D51" lastPageId="3" lastPageNumber="267" pageId="2" pageNumber="266">
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<subSubSection box="[350,636,1397,1421]" pageId="2" pageNumber="266" type="nomenclature">
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<paragraph blockId="2.[350,636,1397,1421]" box="[350,636,1397,1421]" pageId="2" pageNumber="266">
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<heading bold="true" box="[350,636,1397,1421]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="266" reason="2">
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<emphasis bold="true" box="[350,636,1397,1421]" pageId="2" pageNumber="266">
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<taxonomicName authority="Sivec & Stark, 2011" authorityName="Sivec & Stark" authorityYear="2011" box="[350,558,1397,1421]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="266" phylum="Arthropoda" rank="species" species="andreas" status="sp. n.">
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<emphasis bold="true" box="[350,558,1397,1421]" italics="true" pageId="2" pageNumber="266">Neoperla andreas</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[571,636,1397,1421]" pageId="2" pageNumber="266" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[429,558,1433,1457]" pageId="2" pageNumber="266" type="description">
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<paragraph blockId="2.[429,558,1433,1457]" box="[429,558,1433,1457]" pageId="2" pageNumber="266">
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(
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<figureCitation box="[437,549,1433,1457]" captionStart="Figs" captionStartId="3.[189,241,971,995]" captionTargetBox="[249,1383,290,912]" captionTargetId="figure-381@3.[249,1383,252,849]" captionTargetPageId="3" captionText="Figs. 9-11. Neoperla andreas. 9. Head and pronotum. 10. Male terminalia. 11. Penis tube and everted sac, lateral aspect, and ventral aspect inset." figureDoi="http://doi.org/10.5281/zenodo.4760566" httpUri="https://zenodo.org/record/4760566/files/figure.png" pageId="2" pageNumber="266">Figs. 9-11</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="266" type="materials_examined">
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<paragraph blockId="2.[189,798,1504,1920]" pageId="2" pageNumber="266">
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<emphasis bold="true" box="[189,426,1504,1528]" pageId="2" pageNumber="266">Material examined.</emphasis>
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<materialsCitation ID-GBIF-Occurrence="3111458329" collectingDate="1996-03-19" collectingDateMax="1996-03-19" collectingDateMin="1995-03-03" country="Philippines" county="Salakot Falls Road" elevation="300" location="Salakot Falls Road" municipality="Salakot Falls" pageId="2" pageNumber="266" specimenCount="2" specimenCount-male="2" stateProvince="Palawan" typeStatus="holotype, paratype">
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<typeStatus box="[434,546,1504,1528]" pageId="2" pageNumber="266">Holotype</typeStatus>
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<specimenCount box="[554,574,1504,1528]" pageId="2" pageNumber="266" type="male">♂</specimenCount>
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and 1♂ paratypes,
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<emphasis bold="true" box="[189,487,1539,1564]" pageId="2" pageNumber="266">
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<collectingCountry box="[189,487,1539,1564]" name="Philippines" pageId="2" pageNumber="266">PHILIPPINE ISLANDS</collectingCountry>
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</emphasis>
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,
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<collectingRegion box="[512,615,1540,1564]" country="Philippines" name="Palawan" pageId="2" pageNumber="266">Palawan</collectingRegion>
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,
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<collectingCounty pageId="2" pageNumber="266">Salakot Falls Road</collectingCounty>
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,
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<quantity box="[265,338,1576,1600]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="266" unit="m" value="300.0">
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<elevation box="[265,338,1576,1600]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="266" unit="m" value="300.0">300 m</elevation>
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</quantity>
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,
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<date box="[351,526,1576,1600]" pageId="2" pageNumber="266" value="1996-03-19">
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<collectingDate box="[351,526,1576,1600]" pageId="2" pageNumber="266" value="1996-03-19">19 March 1996</collectingDate>
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</date>
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.
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</materialsCitation>
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<materialsCitation ID-GBIF-Occurrence="3111458334" collectingDate="1995-03-03" collectingDateMax="1996-03-19" collectingDateMin="1995-03-03" country="Philippines" county="Salakot Falls Road" elevation="300" location="Estrella Falls" municipality="Salakot Falls" pageId="2" pageNumber="266" specimenCount="3" specimenCount-male="3" stateProvince="Palawan" typeStatus="holotype">
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Additional
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<typeStatus box="[673,792,1576,1600]" pageId="2" pageNumber="266">paratypes</typeStatus>
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:
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<collectingRegion box="[189,292,1611,1635]" country="Philippines" name="Palawan" pageId="2" pageNumber="266">Palawan</collectingRegion>
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,
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<location LSID="urn:lsid:plazi:treatment:ED2FE866E4697D6EFE84E4512C878D51:60590FABE4697D6FFEE1E76F2CA18D51" box="[315,479,1611,1635]" country="Philippines" county="Salakot Falls Road" municipality="Salakot Falls" name="Estrella Falls" pageId="2" pageNumber="266" stateProvince="Palawan">Estrella Falls</location>
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,
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<date box="[502,684,1611,1635]" pageId="2" pageNumber="266" value="1995-03-03">
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<collectingDate box="[502,684,1611,1635]" pageId="2" pageNumber="266" value="1995-03-03">3 March 1995</collectingDate>
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</date>
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,
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<specimenCount box="[707,741,1611,1635]" pageId="2" pageNumber="266" type="male">1♂</specimenCount>
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(all specimens deposited in the
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<collectionCode box="[515,549,1647,1670]" pageId="2" pageNumber="266">PZ</collectionCode>
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collection).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="267" pageId="2" pageNumber="266" type="description">
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<paragraph blockId="2.[189,798,1504,1920]" lastBlockId="3.[189,798,1113,1635]" lastPageId="3" lastPageNumber="267" pageId="2" pageNumber="266">
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<emphasis bold="true" box="[189,366,1682,1706]" pageId="2" pageNumber="266">Adult habitus.</emphasis>
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General color pale brown patterned with yellow. Head pale brown over most of frons, but with a dark spot forward of M-line and a pair of brown spots forward of callosities (
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<figureCitation box="[597,662,1789,1813]" captionStart="Figs" captionStartId="3.[189,241,971,995]" captionTargetBox="[249,1383,290,912]" captionTargetId="figure-381@3.[249,1383,252,849]" captionTargetPageId="3" captionText="Figs. 9-11. Neoperla andreas. 9. Head and pronotum. 10. Male terminalia. 11. Penis tube and everted sac, lateral aspect, and ventral aspect inset." figureDoi="http://doi.org/10.5281/zenodo.4760566" httpUri="https://zenodo.org/record/4760566/files/figure.png" pageId="2" pageNumber="266">Fig. 9</figureCitation>
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). Pronotum with narrow median and sublateral dark bands. Legs, antennae and palpi pale; wings pale brown with darker venation. Ocelli large, about 1 diameter apart.
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<emphasis bold="true" box="[835,904,1397,1421]" pageId="2" pageNumber="266">Male.</emphasis>
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Forewing length
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<quantity box="[1115,1199,1397,1421]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="266" unit="mm" value="13.0">13 mm</quantity>
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. Tergum 7 bearing a small posteromedian triangular lobe and a median sclerotized area (
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<figureCitation box="[1043,1129,1468,1493]" captionStart="Figs" captionStartId="3.[189,241,971,995]" captionTargetBox="[249,1383,290,912]" captionTargetId="figure-381@3.[249,1383,252,849]" captionTargetPageId="3" captionText="Figs. 9-11. Neoperla andreas. 9. Head and pronotum. 10. Male terminalia. 11. Penis tube and everted sac, lateral aspect, and ventral aspect inset." figureDoi="http://doi.org/10.5281/zenodo.4760566" httpUri="https://zenodo.org/record/4760566/files/figure.png" pageId="2" pageNumber="266">Fig. 10</figureCitation>
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). Tergum 8 with a small, median, recurved, tongue-shaped process. Tergum 9 with a small median patch of sensilla. Hemitergal lobes slender, conspicuously narrowed to apex, scarcely bent and directed mesad. Penis tube weakly sclerotized along ventral margin, bare, and subequal to sac in length (
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<figureCitation box="[1021,1100,1682,1706]" captionStart="Figs" captionStartId="3.[189,241,971,995]" captionTargetBox="[249,1383,290,912]" captionTargetId="figure-381@3.[249,1383,252,849]" captionTargetPageId="3" captionText="Figs. 9-11. Neoperla andreas. 9. Head and pronotum. 10. Male terminalia. 11. Penis tube and everted sac, lateral aspect, and ventral aspect inset." figureDoi="http://doi.org/10.5281/zenodo.4760566" httpUri="https://zenodo.org/record/4760566/files/figure.png" pageId="2" pageNumber="266">Fig. 11</figureCitation>
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). Sac curved dorsad near apex and armed over much of surface with variably sized spines, largest spines form a moderately wide penultimate band; subapex of sac bearing a narrow ring of fine spines; venter of sac bearing near midlength, a large, basally directed, weakly bilobed process, mostly bare but weakly armed on tips; dorsolateral of ventral lobe, a pair of spherical, spiny lobes are located.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.4760566" ID-Zenodo-Dep="4760566" httpUri="https://zenodo.org/record/4760566/files/figure.png" pageId="3" pageNumber="267" startId="3.[189,241,971,995]" targetBox="[249,1383,290,912]" targetPageId="3">
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<paragraph blockId="3.[189,1442,971,1030]" pageId="3" pageNumber="267">
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Figs. 9-11.
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<taxonomicName authorityName="Sivec & Stark" authorityYear="2011" box="[311,495,971,995]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="267" phylum="Arthropoda" rank="species" species="andreas">
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<emphasis box="[311,495,971,995]" italics="true" pageId="3" pageNumber="267">Neoperla andreas</emphasis>
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</taxonomicName>
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. 9. Head and pronotum. 10. Male terminalia. 11. Penis tube and everted sac, lateral aspect, and ventral aspect inset.
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</paragraph>
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</caption>
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<paragraph blockId="3.[189,798,1113,1635]" box="[189,416,1184,1208]" pageId="3" pageNumber="267">
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<emphasis bold="true" box="[189,283,1184,1208]" pageId="3" pageNumber="267">Female.</emphasis>
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Unknown.
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</paragraph>
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<paragraph blockId="3.[189,798,1113,1635]" box="[189,396,1220,1244]" pageId="3" pageNumber="267">
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<emphasis bold="true" box="[189,264,1220,1244]" pageId="3" pageNumber="267">Larva.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="267" type="etymology">
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<paragraph blockId="3.[189,798,1113,1635]" pageId="3" pageNumber="267">
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<emphasis bold="true" box="[189,327,1256,1280]" pageId="3" pageNumber="267">Etymology.</emphasis>
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The species name honors Dr. Andreas Zwick, who graciously forwarded this material for our study.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="267" type="diagnosis">
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<paragraph blockId="3.[189,798,1113,1635]" pageId="3" pageNumber="267">
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<emphasis bold="true" box="[189,319,1362,1386]" pageId="3" pageNumber="267">Diagnosis.</emphasis>
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<taxonomicName authorityName="Sivec & Stark" authorityYear="2011" box="[329,519,1362,1386]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="267" phylum="Arthropoda" rank="species" species="andreas">
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<emphasis box="[329,519,1362,1386]" italics="true" pageId="3" pageNumber="267">Neoperla andreas</emphasis>
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</taxonomicName>
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is a member of the
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<taxonomicName authority="Banks, 1913" authorityName="Banks" authorityYear="1913" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="267" phylum="Arthropoda" rank="species" species="recta">
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<emphasis italics="true" pageId="3" pageNumber="267">N. recta</emphasis>
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<bibRefCitation author="Banks, N." box="[246,383,1398,1422]" pageId="3" pageNumber="267" pagination="170 - 180" refId="ref5968" refString="Banks, N. 1913. On a collection of neuropteroid insects from the Philippine Islands. Proceedings of the Entomological Society of Washington, 15: 170 - 180." type="journal article" year="1913">Banks, 1913</bibRefCitation>
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</taxonomicName>
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complex of the
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<taxonomicName authorityName="P.Zwick" authorityYear="1977" box="[564,711,1399,1422]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="267" phylum="Arthropoda" rank="species" species="montivaga">
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<emphasis box="[564,711,1399,1422]" italics="true" pageId="3" pageNumber="267">N. montivaga</emphasis>
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</taxonomicName>
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species group (
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<bibRefCitation author="Sivec, I." box="[287,425,1434,1458]" pageId="3" pageNumber="267" pagination="1 - 44" refId="ref6167" refString="Sivec, I. 1984. Study of genus Neoperla (Plecoptera: Perlidae) from the Philippines. Scopolia, 7: 1 - 44." type="journal article" year="1984">Sivec 1984</bibRefCitation>
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;
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<bibRefCitation author="Zwick, P." box="[447,598,1434,1458]" pageId="3" pageNumber="267" pagination="1 - 53" refId="ref6252" refString="Zwick, P. 1986. The Bornean species of the stonefly genus Neoperla (Plecoptera: Perlidae). Aquatic Insects, 8: 1 - 53." type="journal article" year="1986">Zwick 1986</bibRefCitation>
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). This complex currently includes four Philippine species,
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<taxonomicName authority="Sivec, 1984" authorityName="Sivec" authorityYear="1984" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="267" phylum="Arthropoda" rank="species" species="nishidai">
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<emphasis box="[682,798,1469,1493]" italics="true" pageId="3" pageNumber="267">N. nishidai</emphasis>
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<bibRefCitation author="Sivec, I." box="[189,318,1505,1529]" pageId="3" pageNumber="267" pagination="1 - 44" refId="ref6167" refString="Sivec, I. 1984. Study of genus Neoperla (Plecoptera: Perlidae) from the Philippines. Scopolia, 7: 1 - 44." type="journal article" year="1984">Sivec, 1984</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName authority="Sivec, 1984" authorityName="Sivec" authorityYear="1984" box="[328,621,1505,1529]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="267" phylum="Arthropoda" rank="species" species="pseudorecta">
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<emphasis box="[328,487,1505,1529]" italics="true" pageId="3" pageNumber="267">N. pseudorecta</emphasis>
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<bibRefCitation author="Sivec, I." box="[492,621,1505,1529]" pageId="3" pageNumber="267" pagination="1 - 44" refId="ref6167" refString="Sivec, I. 1984. Study of genus Neoperla (Plecoptera: Perlidae) from the Philippines. Scopolia, 7: 1 - 44." type="journal article" year="1984">Sivec, 1984</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName authorityName="Banks" authorityYear="1913" box="[631,715,1506,1529]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="267" phylum="Arthropoda" rank="species" species="recta">
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<emphasis box="[631,715,1506,1529]" italics="true" pageId="3" pageNumber="267">N. recta</emphasis>
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</taxonomicName>
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and
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<taxonomicName authority="Sivec, 1984" authorityName="Sivec" authorityYear="1984" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="267" phylum="Arthropoda" rank="species" species="zwicki">
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<emphasis italics="true" pageId="3" pageNumber="267">N. zwicki</emphasis>
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<bibRefCitation author="Sivec, I." box="[263,394,1540,1565]" pageId="3" pageNumber="267" pagination="1 - 44" refId="ref6167" refString="Sivec, I. 1984. Study of genus Neoperla (Plecoptera: Perlidae) from the Philippines. Scopolia, 7: 1 - 44." type="journal article" year="1984">Sivec, 1984</bibRefCitation>
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</taxonomicName>
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. The new species is the only one of this group in which the ventral penial lobe is bilobed and at least weakly armed.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |