270 lines
33 KiB
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270 lines
33 KiB
XML
<document id="E8286E5450AED154343E881EE95812BC" ID-DOI="10.37828/em.2020.35.10" ID-ISSN="2336-9744" ID-Zenodo-Dep="13232490" ID-ZooBank="urn:lsid:zoobank.org:pub:F63DAA57-8D7A-4EC7-9FA0-602943FED1FB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722913961471" checkinUser="felipe" docAuthor="Georgiev, Dilian & Glöer, Peter" docDate="2020" docId="C97AB10B4B35FFF662D2710EFDC96F96" docLanguage="en" docName="EcolMontenegrina.35.129-137.pdf" docOrigin="Ecologica Montenegrina 35" docSource="http://dx.doi.org/10.37828/em.2020.35.10" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Grossuana thasia Georgiev & Glöer, 2020, n. sp." docType="treatment" docUuid="019E5D1D-E033-4913-896D-4C779ACA9198" docUuidSource="ZooBank" docVersion="2" lastPageNumber="133" masterDocId="3543C9734B31FFF262457045FFEE693E" masterDocTitle="Three new hydrobiid species from Thassos Island (Greece) with a re-description of Amnicola charpentieri Roth, 1855" masterLastPageNumber="137" masterPageNumber="129" pageNumber="133" updateTime="1722916995419" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="86762631DAD7A088AAEF0F3FBBACE5C6">
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<mods:title id="13E1A82AC3B88CDF931C18CA394CF627">Three new hydrobiid species from Thassos Island (Greece) with a re-description of Amnicola charpentieri Roth, 1855</mods:title>
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<mods:name id="1E9E458079D0F82AB3F63402D650DB38" type="personal">
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<mods:role id="4E56E26698ED09420148A7CC4BEEDFB7">
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<mods:roleTerm id="0EA680DD9D03FC3F63B7F0C9C130F297">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="17BB246AB2A82F5D59BD8061876489F0">Georgiev, Dilian</mods:namePart>
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<mods:affiliation id="254A1A6FEB1E6F596F8C52C7A9944A4B">Department of Ecology and Nature Conservation, Faculty of Biology, Paisii Hilendarski University of Plovdiv, 24 Tsar Assen Str., 4000 Plovdiv, Bulgaria</mods:affiliation>
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</mods:name>
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<mods:name id="18D8AC9C5C91B6409D450537FD49336B" type="personal">
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<mods:roleTerm id="C5F32775CF12340A3D3C8EBDA561D0E6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="511F3ED2410544374829877D9838B1E7">Glöer, Peter</mods:namePart>
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<mods:affiliation id="815668B406789314C91802B08131532A">Biodiversity Research Laboratory, Schulstrasse 3, D- 25491 Hetlingen, Germany Corresponding author. E-mail: diliangeorgiev @ abv. bg</mods:affiliation>
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<mods:typeOfResource id="0C758736E1B007B4A832CB25604B9671">text</mods:typeOfResource>
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<mods:title id="7507E27D5A4F5378617554DD8A0C5713">Ecologica Montenegrina</mods:title>
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</mods:titleInfo>
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<mods:part id="FF432CB953517BE9BDF2AE7B1CECDA4E">
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<mods:date id="7FEF73E1F3B040D0AFD1D627E104ED36">2020</mods:date>
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<mods:number id="E06D0D9E34D2439BD176D4BD09EF2F79">2020-10-22</mods:number>
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<mods:number id="0E8550AFF4485E4EBBD7A42E72AB8721">35</mods:number>
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<mods:start id="AB0C08D139B36F6023871D952EBC1336">129</mods:start>
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<mods:end id="42C8C3F3E2ADEE448EFC4A4B672E4D36">137</mods:end>
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<mods:url id="DBABE4A95BDCFE4923FB797B708284F1">http://dx.doi.org/10.37828/em.2020.35.10</mods:url>
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<mods:classification id="49623660DF83381E06E4D8FD18A1EE22">journal article</mods:classification>
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<mods:identifier id="1C4874D5AE3018752F83CBBC441E9849" type="DOI">10.37828/em.2020.35.10</mods:identifier>
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<treatment id="C97AB10B4B35FFF662D2710EFDC96F96" ID-DOI="http://doi.org/10.5281/zenodo.13232609" ID-Zenodo-Dep="13232609" LSID="urn:lsid:zoobank.org:act:019E5D1D-E033-4913-896D-4C779ACA9198" httpUri="http://treatment.plazi.org/id/C97AB10B4B35FFF662D2710EFDC96F96" lastPageNumber="133" pageId="4" pageNumber="133">
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<subSubSection id="09C953964B35FFF662D2710EFB9D68BD" pageId="4" pageNumber="133" type="nomenclature">
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<paragraph id="416C001D4B35FFF662D2710EFD6D685B" blockId="4.[151,1139,330,387]" box="[151,643,330,357]" pageId="4" pageNumber="133">
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<heading id="1A24B7714B35FFF662D2710EFD6D685B" box="[151,643,330,357]" fontSize="11" level="2" pageId="4" pageNumber="133" reason="2">
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<emphasis id="73A7DC0F4B35FFF662D2710EFE58685B" bold="true" box="[151,438,331,357]" pageId="4" pageNumber="133">
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<taxonomicName id="86D37B9E4B35FFF662D2710EFE83685B" ID-CoL="7RG9Y" authorityName="Georgiev & Glöer" authorityYear="2020" box="[151,365,331,357]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="thasia" status="sp. nov.">
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<emphasis id="73A7DC0F4B35FFF662D2710EFE83685B" bold="true" box="[151,365,331,357]" italics="true" pageId="4" pageNumber="133">Grossuana thasia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="689461744B35FFF66336710EFE58685B" box="[371,438,331,357]" pageId="4" pageNumber="133" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="D9E81C984B35FFF66382710FFDF6685B" box="[455,536,330,357]" captionStart="Figure 2" captionStartId="2.[151,226,729,753]" captionTargetBox="[151,1437,223,711]" captionTargetId="figure-21@2.[151,1437,223,711]" captionTargetPageId="2" captionText="Figure 2. The sampling sites of the species under discussion on Thassos. 1: Rachoni village: type locality of Bythinella rachonica n. sp., localiy of Grossuana beroni n. sp.; 2: Drakotrype cave, Panagia village: localiy of Grossuana beroni n. sp.; 3: type localiy of Grossuana beroni; 4: type locality of Grossuana thasia n. sp., 5: Pernes Mountain, type locality of Bythinella reischuetzi n. sp., 6: Hymettos, type locality of Bythinella charpentieri (Roth, 1855)." figureDoi="http://doi.org/10.5281/zenodo.13232496" httpUri="https://zenodo.org/record/13232496/files/figure.png" pageId="4" pageNumber="133">Figs. 2</figureCitation>
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,
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<figureCitation id="D9E81C984B35FFF66060710FFDD8685B" box="[549,566,330,357]" captionStart="Figures 3-5. 3" captionStartId="3.[151,236,1449,1473]" captionTargetBox="[151,1439,404,1431]" captionTargetId="figure-76@3.[151,1439,404,1431]" captionTargetPageId="3" captionText="Figures 3-5. 3: Type locality of Grossuana beroni n. sp., water source south of Skala Potamia near the round road of the island. 4: Type locality of Grossuana thasia n. sp., a spring south of Kinira near the round road of the island. 5: Type locality of Bythinella rachonica n. sp." figureDoi="http://doi.org/10.5281/zenodo.13232498" httpUri="https://zenodo.org/record/13232498/files/figure.png" pageId="4" pageNumber="133">4</figureCitation>
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,
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<figureCitation id="D9E81C984B35FFF66007710FFD97685B" box="[578,633,330,357]" captionStart="Figures 6-10" captionStartId="3.[151,236,1972,1996]" captionTargetBox="[151,1444,1565,1953]" captionTargetId="figure-136@3.[151,1444,1565,1953]" captionTargetPageId="3" captionText="Figures 6-10. The Grossuana spp. 6-7: G. beroni n. sp. (paratypes), 8: penis of G. beroni n. sp. 9: G. thasia n. sp (holotype), 10: penis of G. thasia n. sp." figureDoi="http://doi.org/10.5281/zenodo.13232500" httpUri="https://zenodo.org/record/13232500/files/figure.png" pageId="4" pageNumber="133">9-10</figureCitation>
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)
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</heading>
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</paragraph>
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<paragraph id="416C001D4B35FFF662D2712EFB9D68BD" blockId="4.[151,1139,330,387]" box="[151,1139,363,387]" pageId="4" pageNumber="133">
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https://zoobank.org/
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<uri id="35420C1F4B35FFF66328712EFB9D68BD" box="[365,1139,363,387]" pageId="4" pageNumber="133">
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urn:lsid:zoobank.org:act:
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<uuid id="35753AC84B35FFF66033712EFB9D68BD" box="[630,1139,363,387]" pageId="4" pageNumber="133">019E5D1D-E033-4913-896D-4C779ACA9198</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="09C953964B35FFF662D271E8FC786BAC" pageId="4" pageNumber="133" type="materials_examined">
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<paragraph id="416C001D4B35FFF662D271E8FE9E68F9" blockId="4.[151,1427,429,557]" box="[151,368,429,455]" pageId="4" pageNumber="133">
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<emphasis id="73A7DC0F4B35FFF662D271E8FE9E68F9" bold="true" box="[151,368,429,455]" pageId="4" pageNumber="133">Material studied:</emphasis>
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</paragraph>
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<paragraph id="416C001D4B35FFF662B0718AFB5668D7" blockId="4.[151,1427,429,557]" box="[245,1208,462,489]" pageId="4" pageNumber="133">
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<materialsCitation id="F1BB0A404B35FFF662B0718AFB5668D7" box="[245,1208,462,489]" collectionCode="ZMH" pageId="4" pageNumber="133" specimenCode="ZMH 140793" specimenCount="1" typeStatus="holotype">
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<emphasis id="73A7DC0F4B35FFF662B0718AFE9F68D7" bold="true" box="[245,369,463,489]" pageId="4" pageNumber="133">
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<typeStatus id="9E68BEBF4B35FFF662B0718AFE8268D7" box="[245,364,463,489]" pageId="4" pageNumber="133" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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shell height
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<quantity id="862BADF84B35FFF6604D718BFD8A68D7" box="[520,612,462,489]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="133" unit="mm" value="1.7">1.7 mm</quantity>
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, width
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<quantity id="862BADF84B35FFF660FF718BFCF968D7" box="[698,791,462,489]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="133" unit="mm" value="1.1">1.1 mm</quantity>
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, from type locality (
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<specimenCode id="1175A8664B35FFF66646718BFB4568D7" box="[1027,1195,462,489]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="4" pageNumber="133" type="Museum">ZMH 140793</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="416C001D4B35FFF662B071B4FEE26B13" blockId="4.[151,1427,429,557]" pageId="4" pageNumber="133">
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<materialsCitation id="F1BB0A404B35FFF662B071B4FCA06B35" box="[245,846,496,523]" collectionCode="ZMH" pageId="4" pageNumber="133" specimenCode="ZMH 140794" specimenCount="5" typeStatus="paratype">
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<emphasis id="73A7DC0F4B35FFF662B071B4FE926B35" bold="true" box="[245,380,497,523]" pageId="4" pageNumber="133">
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<typeStatus id="9E68BEBF4B35FFF662B071B4FE996B35" box="[245,375,497,523]" pageId="4" pageNumber="133" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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<specimenCount id="57D5CB944B35FFF663C171B5FDFD6B35" box="[388,531,496,523]" count="5" pageId="4" pageNumber="133" type="generic">5 specimens</specimenCount>
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in ethanol (
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<specimenCode id="1175A8664B35FFF660DA71B5FCA86B35" box="[671,838,496,523]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="4" pageNumber="133" type="Museum">ZMH 140794</specimenCode>
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)
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</materialsCitation>
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,
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<specimenCount id="57D5CB944B35FFF6611F71B5FC076B35" box="[858,1001,496,523]" count="5" pageId="4" pageNumber="133" type="generic">5 specimens</specimenCount>
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in coll. Glöer,
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<specimenCount id="57D5CB944B35FFF666DF71B5FAD56B35" box="[1178,1339,496,523]" count="11" pageId="4" pageNumber="133" type="generic">11 specimens</specimenCount>
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in coll. Georgiev.
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</paragraph>
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<paragraph id="416C001D4B35FFF662D27213FC786BAC" blockId="4.[151,1435,597,658]" pageId="4" pageNumber="133">
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<emphasis id="73A7DC0F4B35FFF662D27213FEAD6B4E" bold="true" box="[151,323,598,624]" pageId="4" pageNumber="133">Type locality:</emphasis>
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<materialsCitation id="F1BB0A404B35FFF6630E7210FC7C6BAC" country="Greece" elevation="144" latitude="40.635418" location="Kinira" longLatPrecision="1" longitude="24.7645" municipality="Thassos Island" pageId="4" pageNumber="133" specimenCount="1" stateProvince="Voreio Aigaio">
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<collectingCountry id="39C4408D4B35FFF6630E7210FE4F6B4E" box="[331,417,597,624]" name="Greece" pageId="4" pageNumber="133">Greece</collectingCountry>
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,
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<collectingMunicipality id="A1089A674B35FFF663E87210FDB06B4E" box="[429,606,597,624]" pageId="4" pageNumber="133">Thassos Island</collectingMunicipality>
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,
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<collectingRegion id="8317CEFF4B35FFF6602F7210FCFD6B4E" box="[618,787,597,624]" country="Greece" name="Voreio Aigaio" pageId="4" pageNumber="133">North Aegean</collectingRegion>
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, a spring (water source) south of
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<location id="440C56C64B35FFF666DB7210FB066B4E" LSID="urn:lsid:plazi:treatment:C97AB10B4B35FFF662D2710EFDC96F96:440C56C64B35FFF666DB7210FB066B4E" box="[1182,1256,597,624]" country="Greece" latitude="40.635418" longLatPrecision="1" longitude="24.7645" municipality="Thassos Island" name="Kinira" pageId="4" pageNumber="133" stateProvince="Voreio Aigaio">Kinira</location>
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near the round road of the island,
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<geoCoordinate id="24E766DA4B35FFF663377232FDEA6BAC" box="[370,516,631,658]" degrees="40" direction="north" minutes="38" orientation="latitude" pageId="4" pageNumber="133" precision="1" seconds="07.5" value="40.635418">N40 38 07.5</geoCoordinate>
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<geoCoordinate id="24E766DA4B35FFF6604E7232FD726BAC" box="[523,668,631,658]" degrees="24" direction="east" minutes="45" orientation="longitude" pageId="4" pageNumber="133" precision="1" seconds="52.2" value="24.7645">E24 45 52.2</geoCoordinate>
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,
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<elevation id="CAFEE72E4B35FFF660ED7232FCDE6BAC" box="[680,816,631,658]" metricMagnitude="2" metricUnit="m" metricValue="1.44" pageId="4" pageNumber="133" unit="m" value="144.0">
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<quantity id="862BADF84B35FFF660ED7232FD1C6BAC" box="[680,754,631,658]" metricMagnitude="2" metricUnit="m" metricValue="1.44" pageId="4" pageNumber="133" unit="m" value="144.0">144 m</quantity>
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a.s.l.
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</elevation>
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(
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<figureCitation id="D9E81C984B35FFF661057232FC686BAC" box="[832,902,631,658]" captionStart="Figures 3-5. 3" captionStartId="3.[151,236,1449,1473]" captionTargetBox="[151,1439,404,1431]" captionTargetId="figure-76@3.[151,1439,404,1431]" captionTargetPageId="3" captionText="Figures 3-5. 3: Type locality of Grossuana beroni n. sp., water source south of Skala Potamia near the round road of the island. 4: Type locality of Grossuana thasia n. sp., a spring south of Kinira near the round road of the island. 5: Type locality of Bythinella rachonica n. sp." figureDoi="http://doi.org/10.5281/zenodo.13232498" httpUri="https://zenodo.org/record/13232498/files/figure.png" pageId="4" pageNumber="133">Fig. 3</figureCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="09C953964B35FFF662D272FEFC596BEB" box="[151,951,698,725]" pageId="4" pageNumber="133" type="etymology">
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<paragraph id="416C001D4B35FFF662D272FEFC596BEB" blockId="4.[151,951,698,725]" box="[151,951,698,725]" pageId="4" pageNumber="133">
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<emphasis id="73A7DC0F4B35FFF662D272FEFEC66BEB" bold="true" box="[151,296,699,725]" pageId="4" pageNumber="133">Etymology:</emphasis>
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Named after Thassos Island on which the species lives.
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</paragraph>
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</subSubSection>
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<subSubSection id="09C953964B35FFF662D272BBFBC36CC1" pageId="4" pageNumber="133" type="description">
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<paragraph id="416C001D4B35FFF662D272BBFEEB6ADD" blockId="4.[151,1436,766,995]" pageId="4" pageNumber="133">
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<emphasis id="73A7DC0F4B35FFF662D272BBFEDC6A26" bold="true" box="[151,306,766,792]" pageId="4" pageNumber="133">Description:</emphasis>
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Shell: The corneous shell is elongate ovate with a prolonged body whorl (
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<figureCitation id="D9E81C984B35FFF666EA72BBFB146A27" box="[1199,1274,766,793]" captionStart="Figure 2" captionStartId="2.[151,226,729,753]" captionTargetBox="[151,1437,223,711]" captionTargetId="figure-21@2.[151,1437,223,711]" captionTargetPageId="2" captionText="Figure 2. The sampling sites of the species under discussion on Thassos. 1: Rachoni village: type locality of Bythinella rachonica n. sp., localiy of Grossuana beroni n. sp.; 2: Drakotrype cave, Panagia village: localiy of Grossuana beroni n. sp.; 3: type localiy of Grossuana beroni; 4: type locality of Grossuana thasia n. sp., 5: Pernes Mountain, type locality of Bythinella reischuetzi n. sp., 6: Hymettos, type locality of Bythinella charpentieri (Roth, 1855)." figureDoi="http://doi.org/10.5281/zenodo.13232496" httpUri="https://zenodo.org/record/13232496/files/figure.png" pageId="4" pageNumber="133">Fig. 2</figureCitation>
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). The apex is rounded and often eroded. The 4-4.5 whorls are slightly convex with a weak suture, with rough irregular growth lines. The aperture is ovate slightly angled at the top with a sharp periostome thickened at the columella. The umbilicus is closed. The operculum is dark red. Animal: The animal is mostly black colored. The mantle is black with a thin white border. The snout and tentacles are black with white or grayish tips. Penis: The penis is nearly regularly broad with a long tapered penis-tip. There is a dark spot at the near the penis-tip.
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</paragraph>
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<paragraph id="416C001D4B35FFF662D27449FEB16C2D" blockId="4.[151,1436,1036,1535]" pageId="4" pageNumber="133">
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<emphasis id="73A7DC0F4B35FFF662D27449FE246D18" bold="true" box="[151,458,1036,1062]" pageId="4" pageNumber="133">Differentiating features:</emphasis>
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The new species is most similar to
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<taxonomicName id="86D37B9E4B35FFF661C87448FF3C6D77" authority="Gloer & Georgiev, 2009" authorityName="Gloer & Georgiev" authorityYear="2009" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="angeltsekovi">
|
|
<emphasis id="73A7DC0F4B35FFF661C87448FB426D19" box="[909,1196,1036,1063]" italics="true" pageId="4" pageNumber="133">Grossuana angeltsekovi</emphasis>
|
|
<bibRefCitation id="25427DEC4B35FFF666F27449FF3C6D77" author="Gloer, P. & Georgiev, D." pageId="4" pageNumber="133" pagination="123 - 136" refId="ref4427" refString="Gloer, P., & Georgiev, D. (2009) New Rissooidea from Bulgaria (Gastropoda: Rissooidea). Mollusca, 27 (2), 123 - 136." type="journal article" year="2009">Glöer & Georgiev, 2009</bibRefCitation>
|
|
</taxonomicName>
|
|
which is known to be widely distributed in
|
|
<collectingCountry id="39C4408D4B35FFF66092746BFCD26D77" box="[727,828,1070,1097]" name="Bulgaria" pageId="4" pageNumber="133">Bulgaria</collectingCountry>
|
|
and North
|
|
<collectingCountry id="39C4408D4B35FFF66187746BFBFB6D77" box="[962,1045,1070,1097]" name="Greece" pageId="4" pageNumber="133">Greece</collectingCountry>
|
|
(
|
|
<bibRefCitation id="25427DEC4B35FFF66661746BFB376D77" author="Georgiev, D." box="[1060,1241,1070,1097]" pageId="4" pageNumber="133" pagination="1 - 4" refId="ref4338" refString="Georgiev, D. (2013) First record of Grossuana angeltsekovi Gloer & Georgiev, 2009 (Gastropoda: Risooidea) from Greece. ZooNotes, 37, 1 - 4." type="journal article" year="2013">Georgiev 2013</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="25427DEC4B35FFF666A0746BFF3B6D54" author="Georgiev, D. & Gloer, P. & Dedov, I. & Irikov, A." pageId="4" pageNumber="133" pagination="159 - 166" refId="ref4371" refString="Georgiev, D., Gloer, P., Dedov, I. & Irikov, A. (2015) Review of the Genus Grossuana Radoman, 1973 (Gastropoda: Truncatelloidea) from Bulgaria, with a description of a new species. Acta Zoologica Bulgarica, 67, 159 - 166." type="journal article" year="2015">Georgiev et al., 2015</bibRefCitation>
|
|
, Falniowski et al. 2015,
|
|
<bibRefCitation id="25427DEC4B35FFF66041740AFD0E6D54" author="Gloer, P. & Reuselaars, R. & Papavasileiou, K." box="[516,736,1103,1130]" pageId="4" pageNumber="133" pagination="14 - 19" refId="ref4461" refString="Gloer, P., Reuselaars, R. & Papavasileiou, K. (2018) Grossuana Radoman, 1973 from Macedonia (Greece) (Gastropoda: Truncatelloidea) with the description of three new species. Ecologica Montenegrina, 17, 14 - 19. https: // doi. org / 10.37828 / em. 2018.17.2" type="journal article" year="2018">Glöer et al., 2018</bibRefCitation>
|
|
). From this species
|
|
<taxonomicName id="86D37B9E4B35FFF661937415FB476D54" authorityName="Georgiev & Glöer" authorityYear="2020" box="[982,1193,1103,1130]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="thasia" status="sp. nov.">
|
|
<emphasis id="73A7DC0F4B35FFF661937415FB476D54" box="[982,1193,1103,1130]" italics="true" pageId="4" pageNumber="133">Grossuana thasia</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="689461744B35FFF666F07415FB196D54" box="[1205,1271,1104,1130]" pageId="4" pageNumber="133" rank="species">n. sp.</taxonomicNameLabel>
|
|
differs by its more elongated and slender shell with flatter whorls, which have rough surface with irregular growth lines, and by its more rounded, often eroded apex. The whorls of the shell of the new species are 4-4.5 while in
|
|
<taxonomicName id="86D37B9E4B35FFF667C574D1FEC76DF1" authorityName="Gloer & Georgiev" authorityYear="2009" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="angeltsekovi">
|
|
<emphasis id="73A7DC0F4B35FFF667C574D1FEC76DF1" italics="true" pageId="4" pageNumber="133">G. angeltsekovi</emphasis>
|
|
</taxonomicName>
|
|
they are 5-5.5 and the later is larger (height of the
|
|
<typeStatus id="9E68BEBF4B35FFF661C574F1FC086DF1" box="[896,998,1204,1231]" pageId="4" pageNumber="133" type="holotype">holotype</typeStatus>
|
|
=
|
|
<quantity id="862BADF84B35FFF6664274F1FB886DF1" box="[1031,1126,1204,1231]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="133" unit="mm" value="2.1">2.1 mm</quantity>
|
|
). In addition
|
|
<taxonomicName id="86D37B9E4B35FFF6674474F0FA806DF1" authorityName="Georgiev & Glöer" authorityYear="2020" box="[1281,1390,1204,1231]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="thasia">
|
|
<emphasis id="73A7DC0F4B35FFF6674474F0FA806DF1" box="[1281,1390,1204,1231]" italics="true" pageId="4" pageNumber="133">G. thasia</emphasis>
|
|
</taxonomicName>
|
|
has a well visible bifurcated penis lobe, while in
|
|
<taxonomicName id="86D37B9E4B35FFF660877492FC956DCF" authorityName="Gloer & Georgiev" authorityYear="2009" box="[706,891,1238,1265]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="angeltsekovi">
|
|
<emphasis id="73A7DC0F4B35FFF660877492FC956DCF" box="[706,891,1238,1265]" italics="true" pageId="4" pageNumber="133">G. angeltsekovi</emphasis>
|
|
</taxonomicName>
|
|
it is very small and hardly visible (
|
|
<bibRefCitation id="25427DEC4B35FFF667707493FEBD6C2D" author="Gloer, P. & Georgiev, D." pageId="4" pageNumber="133" pagination="123 - 136" refId="ref4427" refString="Gloer, P., & Georgiev, D. (2009) New Rissooidea from Bulgaria (Gastropoda: Rissooidea). Mollusca, 27 (2), 123 - 136." type="journal article" year="2009">Glöer & Georgiev, 2009</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="416C001D4B35FFF662B0755EFCED6C46" blockId="4.[151,1436,1036,1535]" pageId="4" pageNumber="133">
|
|
<taxonomicName id="86D37B9E4B35FFF662B0755EFE286C0B" authorityName="Georgiev & Glöer" authorityYear="2020" box="[245,454,1306,1333]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="thasia" status="sp. nov.">
|
|
<emphasis id="73A7DC0F4B35FFF662B0755EFE286C0B" box="[245,454,1306,1333]" italics="true" pageId="4" pageNumber="133">Grossuana thasia</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="689461744B35FFF66388755EFDE26C0B" box="[461,524,1307,1333]" pageId="4" pageNumber="133" rank="species">n. sp.</taxonomicNameLabel>
|
|
is similar by its shell morphology to
|
|
<taxonomicName id="86D37B9E4B35FFF66181755EFE4C6C68" authority="Gloer, Reuselaars & Papavasileiou, 2018" authorityName="Gloer, Reuselaars & Papavasileiou" authorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="stenaensis">
|
|
<emphasis id="73A7DC0F4B35FFF66181755EFB2A6C0A" box="[964,1220,1307,1333]" italics="true" pageId="4" pageNumber="133">Grossuana stenaensis</emphasis>
|
|
<bibRefCitation id="25427DEC4B35FFF66689755FFE4C6C68" author="Gloer, P. & Reuselaars, R. & Papavasileiou, K." pageId="4" pageNumber="133" pagination="14 - 19" refId="ref4461" refString="Gloer, P., Reuselaars, R. & Papavasileiou, K. (2018) Grossuana Radoman, 1973 from Macedonia (Greece) (Gastropoda: Truncatelloidea) with the description of three new species. Ecologica Montenegrina, 17, 14 - 19. https: // doi. org / 10.37828 / em. 2018.17.2" type="journal article" year="2018">Glöer, Reuselaars & Papavasileiou, 2018</bibRefCitation>
|
|
</taxonomicName>
|
|
described from the nearby mainland of
|
|
<collectingCountry id="39C4408D4B35FFF6613D757EFC246C68" box="[888,970,1339,1366]" name="Greece" pageId="4" pageNumber="133">Greece</collectingCountry>
|
|
(
|
|
<bibRefCitation id="25427DEC4B35FFF6619C757EFB446C68" author="Gloer, P. & Reuselaars, R. & Papavasileiou, K." box="[985,1194,1339,1366]" pageId="4" pageNumber="133" pagination="14 - 19" refId="ref4461" refString="Gloer, P., Reuselaars, R. & Papavasileiou, K. (2018) Grossuana Radoman, 1973 from Macedonia (Greece) (Gastropoda: Truncatelloidea) with the description of three new species. Ecologica Montenegrina, 17, 14 - 19. https: // doi. org / 10.37828 / em. 2018.17.2" type="journal article" year="2018">Glöer et al., 2018</bibRefCitation>
|
|
) but this species has a penis with broader base and shorter and thicker tip.
|
|
</paragraph>
|
|
<paragraph id="416C001D4B35FFF662B0753AFBC36CC1" blockId="4.[151,1436,1036,1535]" pageId="4" pageNumber="133">
|
|
Previously all
|
|
<taxonomicName id="86D37B9E4B35FFF663E275C5FDC76CA4" authorityName="Radoman" authorityYear="1983" box="[423,553,1408,1434]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="genus">
|
|
<emphasis id="73A7DC0F4B35FFF663E275C5FDC76CA4" box="[423,553,1408,1434]" italics="true" pageId="4" pageNumber="133">Grossuana</emphasis>
|
|
</taxonomicName>
|
|
of Thassos were assigned to
|
|
<taxonomicName id="86D37B9E4B35FFF661DD75C5FF316C82" authority="(Kuster, 1853) (Reischutz, 1983)" authorityName="Reischutz" authorityYear="1983" baseAuthorityName="Kuster" baseAuthorityYear="1853" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="hohenackeri">
|
|
<emphasis id="73A7DC0F4B35FFF661DD75C5FBBF6CA4" box="[920,1105,1407,1434]" italics="true" pageId="4" pageNumber="133">G. hohenackeri</emphasis>
|
|
(Küster, 1853) (
|
|
<bibRefCitation id="25427DEC4B35FFF66765753AFF386C82" author="Reischutz, P." pageId="4" pageNumber="133" pagination="133 - 146" refId="ref4865" refString="Reischutz, P. (1983) Ein Beitrag zur Molluskenfauna der Insel Thasos (Griechenland). Annalen des Naturhistorischen Museums in Wien, 85 B, 133 - 146." type="journal article" year="1983">Reischütz, 1983</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
. It was later re-described and split into two species:
|
|
<taxonomicName id="86D37B9E4B35FFF6611B75E4FA896C82" authority="(Bourguignat, 1852)" baseAuthorityName="Bourguignat" baseAuthorityYear="1852" box="[862,1383,1441,1468]" class="Gastropoda" family="Hydrobiidae" genus="Radomaniola" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="tritonum">
|
|
<emphasis id="73A7DC0F4B35FFF6611B75E4FB826C85" box="[862,1132,1441,1468]" italics="true" pageId="4" pageNumber="133">Radomaniola tritonum</emphasis>
|
|
(Bourguignat, 1852)
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="86D37B9E4B35FFF662D27586FCF26CE3" authority="Boeters, Gloer & Falniowski (2018)" authorityName="Boeters, Gloer & Falniowski" authorityYear="2018" box="[151,796,1474,1501]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="tembii">
|
|
<emphasis id="73A7DC0F4B35FFF662D27586FE876CE3" box="[151,361,1474,1501]" italics="true" pageId="4" pageNumber="133">Grossuana tembii</emphasis>
|
|
<bibRefCitation id="25427DEC4B35FFF663347586FCF26CE3" author="Boeters, H. D. & Gloer, P. & Falniowski, A." box="[369,796,1474,1501]" pageId="4" pageNumber="133" pagination="83 - 88" refId="ref3918" refString="Boeters, H. D., Gloer, P. & Falniowski, A. (2018) A new species of Grossuana (Caenogastropoda: Truncatelloidea: Hydrobiidae). Folia Malacologica, 26 (2), 83 - 88. https: // doi. org / 10.12657 / folmal. 026.010" type="journal article" year="2018">Boeters, Glöer & Falniowski (2018)</bibRefCitation>
|
|
</taxonomicName>
|
|
(
|
|
<bibRefCitation id="25427DEC4B35FFF661697587FBF66CE3" author="Boeters, H. D. & Gloer, P. & Falniowski, A." box="[812,1048,1474,1501]" pageId="4" pageNumber="133" pagination="83 - 88" refId="ref3918" refString="Boeters, H. D., Gloer, P. & Falniowski, A. (2018) A new species of Grossuana (Caenogastropoda: Truncatelloidea: Hydrobiidae). Folia Malacologica, 26 (2), 83 - 88. https: // doi. org / 10.12657 / folmal. 026.010" type="journal article" year="2018">Boeters et al., 2018</bibRefCitation>
|
|
).
|
|
<taxonomicName id="86D37B9E4B35FFF6666B7586FB746CE3" authorityName="Georgiev & Glöer" authorityYear="2020" box="[1070,1178,1474,1501]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="thasia" status="n. sp.">
|
|
<emphasis id="73A7DC0F4B35FFF6666B7586FB746CE3" box="[1070,1178,1474,1501]" italics="true" pageId="4" pageNumber="133">G. thasia</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="689461744B35FFF666E67586FB0C6CE3" box="[1187,1250,1475,1501]" pageId="4" pageNumber="133" rank="species">n. sp.</taxonomicNameLabel>
|
|
differs from
|
|
<taxonomicName id="86D37B9E4B35FFF667C57586FF0E6CC1" authorityName="Boeters, Gloer & Falniowski" authorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="tembii">
|
|
<emphasis id="73A7DC0F4B35FFF667C57586FF0E6CC1" italics="true" pageId="4" pageNumber="133">G. tembii</emphasis>
|
|
</taxonomicName>
|
|
by its higher whorl number (
|
|
<quantity id="862BADF84B35FFF6607375A1FD566CC1" box="[566,696,1508,1535]" metricMagnitude="-2" metricUnit="m" metricValue="8.5725" metricValueMax="8.89" metricValueMin="8.255" pageId="4" pageNumber="133" unit="in" value="3.375" valueMax="3.5" valueMin="3.25">3.25-3.5 in</quantity>
|
|
<taxonomicName id="86D37B9E4B35FFF6608575A0FCC36CC1" authorityName="Boeters, Gloer & Falniowski" authorityYear="2018" box="[704,813,1508,1535]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="133" phylum="Mollusca" rank="species" species="tembii">
|
|
<emphasis id="73A7DC0F4B35FFF6608575A0FCC36CC1" box="[704,813,1508,1535]" italics="true" pageId="4" pageNumber="133">G. tembii</emphasis>
|
|
</taxonomicName>
|
|
) and longer penis tip.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="09C953964B35FFF662D2766DFDC96F96" pageId="4" pageNumber="133" type="distribution">
|
|
<paragraph id="416C001D4B35FFF662D2766DFD316F7D" blockId="4.[151,735,1576,1603]" box="[151,735,1576,1603]" pageId="4" pageNumber="133">
|
|
<emphasis id="73A7DC0F4B35FFF662D2766DFED46F7C" bold="true" box="[151,314,1576,1602]" pageId="4" pageNumber="133">Distribution:</emphasis>
|
|
Known only from the
|
|
<typeStatus id="9E68BEBF4B35FFF66003766CFD966F7D" box="[582,632,1577,1603]" pageId="4" pageNumber="133">type</typeStatus>
|
|
locality.
|
|
</paragraph>
|
|
<paragraph id="416C001D4B35FFF662D2762EFDC96F96" blockId="4.[151,1417,1643,1704]" pageId="4" pageNumber="133">
|
|
<emphasis id="73A7DC0F4B35FFF662D2762EFEEF6FBB" bold="true" box="[151,257,1643,1669]" pageId="4" pageNumber="133">Habitat:</emphasis>
|
|
Karst spring in a mixed broad leaf forest (mainly
|
|
<taxonomicName id="86D37B9E4B35FFF66108762EFBC36FB8" box="[845,1069,1643,1670]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="4" pageNumber="133" phylum="Tracheophyta" rank="species" species="orientalis">
|
|
<emphasis id="73A7DC0F4B35FFF66108762EFBC36FB8" box="[845,1069,1643,1670]" italics="true" pageId="4" pageNumber="133">Platanus orientalis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="86D37B9E4B35FFF666237629FAEB6FB8" box="[1126,1285,1643,1670]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="133" phylum="Tracheophyta" rank="species" species="undetermined">
|
|
<emphasis id="73A7DC0F4B35FFF666237629FB276FB8" box="[1126,1225,1644,1670]" italics="true" pageId="4" pageNumber="133">Quercus</emphasis>
|
|
spp.
|
|
</taxonomicName>
|
|
). Found on moss, dry leaves, sand and stones.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |