615 lines
71 KiB
XML
615 lines
71 KiB
XML
<document id="FCAE613742F2E2A9BC8213A822A8A7A6" ID-CLB-Dataset="26963" ID-DOI="10.11646/zootaxa.4590.2.3" ID-GBIF-Dataset="833c2bc0-2a29-4efa-9058-04bcf3e7beed" ID-ISSN="1175-5326" ID-Zenodo-Dep="2651878" ID-ZooBank="3302E889-CDCF-4C0C-AFBE-BD287AFBD15E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1556256752834" checkinUser="plazi" docAuthor="Ziegler, Thomas, Nguyen, Truong Quang, Pham, Cuong The, Nguyen, Tao Thien, Pham, Anh Van, Schingen, Mona Van, Nguyen, Tham Thi & Le, Minh Duc" docDate="2019" docId="314D4322155E6352FF28D2AEFDABF827" docLanguage="en" docName="zootaxa.4590.2.3.pdf" docOrigin="Zootaxa 4590 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Achalinus juliani Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le 2019, sp. nov." docType="treatment" docVersion="10" lastPageNumber="255" masterDocId="CD743B5A155A6354FFBFD350FF9BFFD4" masterDocTitle="Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)" masterLastPageNumber="269" masterPageNumber="249" pageNumber="253" updateTime="1698723983114" updateUser="ExternalLinkService">
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<mods:titleInfo id="3136AE922C2C0A672E36B6241D42060B">
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<mods:title id="505F32109E1D358ACB73ACE2793A022F">Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)</mods:title>
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<mods:namePart id="0B0508333B84D4C97F810DA12411F3F9">Ziegler, Thomas</mods:namePart>
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<mods:namePart id="8510053729E57D015E460213340D39EA">Nguyen, Truong Quang</mods:namePart>
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<mods:namePart id="2CED052C728B4B95B2F7FA07E8DA34FC">Pham, Cuong The</mods:namePart>
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<mods:namePart id="31565E71633B2A30070271F9C641C99A">Nguyen, Tao Thien</mods:namePart>
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<mods:namePart id="9677A098E067ED751FD77E188FFA776D">Pham, Anh Van</mods:namePart>
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<mods:namePart id="281A6C475C70B4B3A9B5FE423A8BC234">Schingen, Mona Van</mods:namePart>
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<mods:namePart id="C8D4E418F3DB0A940F72FB0D51216B92">Nguyen, Tham Thi</mods:namePart>
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<mods:namePart id="9E2936812E8C46D2FF44ABC25069FDE0">Le, Minh Duc</mods:namePart>
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<mods:title id="99BCC0AFE5FFB0E81C8FA56576692801">Zootaxa</mods:title>
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<mods:part id="EE14E9D2B71A506E43873A6B783847C0">
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<mods:date id="82BAC8FF2E1B7DEDE8D21AA263CC0174">2019</mods:date>
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<mods:number id="DF724A6CD088E3C35EABB2B3875F6A33">2019-04-26</mods:number>
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<mods:number id="6462BB27FE9AE3791532171FD742CC43">4590</mods:number>
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<mods:identifier id="51EB434D480D2859D0C10E8DFEEFC2B6" type="DOI">10.11646/zootaxa.4590.2.3</mods:identifier>
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<mods:identifier id="C0A62134542C2EB9592B2BF4E82C0726" type="GBIF-Dataset">833c2bc0-2a29-4efa-9058-04bcf3e7beed</mods:identifier>
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<treatment id="314D4322155E6352FF28D2AEFDABF827" ID-DOI="http://doi.org/10.5281/zenodo.5586666" ID-GBIF-Taxon="156863566" ID-Zenodo-Dep="5586666" LSID="urn:lsid:plazi:treatment:314D4322155E6352FF28D2AEFDABF827" httpUri="http://treatment.plazi.org/id/314D4322155E6352FF28D2AEFDABF827" lastPageId="6" lastPageNumber="255" pageId="4" pageNumber="253">
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<subSubSection id="F1FEA1BF155E6350FF28D2AEFE54FDCC" box="[151,463,510,536]" pageId="4" pageNumber="253" type="nomenclature">
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<heading id="E2134558155E6350FF28D2AEFE54FDCC" bold="true" box="[151,463,510,536]" fontSize="11" level="1" pageId="4" pageNumber="253" reason="1">
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<emphasis id="8B902E26155E6350FF28D2AEFE54FDCC" bold="true" box="[151,463,510,536]" pageId="4" pageNumber="253">
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<taxonomicName id="7EE489B7155E6350FF28D2AEFEF1FDCC" authority="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le, 2019" authorityName="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le" authorityYear="2019" box="[151,362,510,536]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="253" phylum="Chordata" rank="species" species="juliani" status="sp. nov.">
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<emphasis id="8B902E26155E6350FF28D2AEFEF1FDCC" bold="true" box="[151,362,510,536]" italics="true" pageId="4" pageNumber="253">Achalinus juliani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="90A3935D155E6350FECED2AEFE54FDCC" box="[369,463,510,536]" pageId="4" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="F1FEA1BF155E6350FF28D171FE8AFDEE" box="[151,273,545,570]" pageId="4" pageNumber="253" type="description">
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<paragraph id="B95BF234155E6350FF28D171FE8AFDEE" blockId="4.[151,463,510,570]" box="[151,273,545,570]" pageId="4" pageNumber="253">
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(
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<figureCitation id="21DFEEB1155E6350FF20D171FE91FDEE" box="[159,266,545,570]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="5.[151,250,1977,1999]" captionStartId-1="7.[151,250,1998,2020]" captionStartId-2="8.[151,250,1179,1201]" captionStartId-3="8.[151,250,1977,1999]" captionTargetBox-0="[151,1437,1052,1960]" captionTargetBox-1="[165,1432,195,1987]" captionTargetBox-2="[150,1436,193,1158]" captionTargetBox-3="[151,1428,1282,1963]" captionTargetId-0="figure@5.[151,1437,1052,1960]" captionTargetId-1="figure@7.[165,1432,195,1987]" captionTargetId-2="figure@8.[150,1436,193,1158]" captionTargetId-3="figure@8.[151,1436,1282,1963]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="8" captionText-0="FIGURE 2. Head views of the preserved holotype of Achalinus juliani sp. nov. (IEBR A.2018.8) from Ha Lang District, Cao Bang Province, northern Vietnam. Phot. T. Ziegler" captionText-1="FIGURE 3. Overall view of the preserved holotype of Achalinus juliani sp. nov. (IEBR A.2018.8). Phot. T. Ziegler" captionText-2="FIGURE 4. Lateral view of the midbody region of the preserved holotype of Achalinus juliani sp. nov. (IEBR A.2018.8). Phot. T. Ziegler" captionText-3="FIGURE 5. Preserved paratypes of Achalinus juliani sp. nov. (IEBR A.2018.9, left, and VNMN 06924, right), both from Nguyen Binh District, Cao Bang Province, northern Vietnam. Phot. T. Ziegler and T. T. Nguyen" httpUri-0="https://zenodo.org/record/2651882/files/figure.png" httpUri-1="https://zenodo.org/record/2651884/files/figure.png" httpUri-2="https://zenodo.org/record/2651886/files/figure.png" httpUri-3="https://zenodo.org/record/2651888/files/figure.png" pageId="4" pageNumber="253">Figs. 2–5</figureCitation>
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)
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</paragraph>
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<subSubSection id="F1FEA1BF155E6350FF28D139FD7EFC76" pageId="4" pageNumber="253" type="materials_examined">
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<materialsCitation id="098CF869155E6350FF28D139FCADFD1E" ID-GBIF-Occurrence="2236983381" collectingDate="2014-06-14" collectionCode="IEBR" collectorName="C. T. Pham" country="Vietnam" elevation="477" latitude="22.718067" location="Ha Lang District" longLatPrecision="1" longitude="106.66088" pageId="4" pageNumber="253" specimenCode="CB 2014.18" specimenCount="1" stateProvince="Cao Bang" typeStatus="holotype">
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<typeStatus id="665F4C96155E6350FF28D139FE9DFD55" box="[151,262,617,641]" pageId="4" pageNumber="253" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="DFF56AF1155E6350FEA5D13AFEC1FD55" box="[282,346,618,641]" pageId="4" pageNumber="253">IEBR</collectionCode>
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A.2018.8 (Field no.
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<specimenCode id="E9425A4F155E6350FDDAD13AFD62FD56" box="[613,761,618,642]" collectionCode="CB" country="Australia" name="The CB Rhizobium Collection" pageId="4" pageNumber="253">CB 2014.18</specimenCode>
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), an adult male, from forest of Duc Quang Commune (
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,
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, at an elevation of
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above sea level
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</elevation>
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),
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,
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<collectingRegion id="7B203CD6155E6350FB09D1DEFA02FD72" box="[1206,1433,653,678]" country="Vietnam" name="Cao Bang" pageId="4" pageNumber="253">Cao Bang Province</collectingRegion>
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, northern
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<collectingCountry id="C1F3B2A4155E6350FF43D1E1FEC5FD1E" box="[252,350,689,714]" name="Vietnam" pageId="4" pageNumber="253">Vietnam</collectingCountry>
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, collected by
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<collectorName id="141197E2155E6350FE47D1E2FDF4FD1E" box="[504,623,689,714]" pageId="4" pageNumber="253">C.T. Pham</collectorName>
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on
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<collectingDate id="DD1E2D1C155E6350FD26D1E2FCA9FD1E" box="[665,818,690,714]" pageId="4" pageNumber="253" value="2014-06-14">
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<date id="CD5AD4F4155E6350FD26D1E2FCA9FD1E" box="[665,818,690,714]" pageId="4" pageNumber="253" value="2014-06-14">14 June 2014</date>
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</collectingDate>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="B95BF234155E6350FF78D185FD7EFC76" blockId="4.[151,1437,617,2010]" pageId="4" pageNumber="253">
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<materialsCitation id="098CF869155E6350FF78D185FC04FCE2" ID-GBIF-Occurrence="2236983380" collectingDate="2014-05-24" collectionCode="IEBR" collectorName="T. H. Pham" country="Vietnam" elevation="1461" latitude="22.60245" location="Phia Oac-Phia Den National Park" longLatPrecision="1" longitude="105.87877" municipality="Nguyen Binh District" pageId="4" pageNumber="253" specimenCode="CB 2014.19" specimenCount="1" stateProvince="Cao Bang" typeStatus="paratype">
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<typeStatus id="665F4C96155E6350FF78D185FEDAFD39" box="[199,321,725,749]" pageId="4" pageNumber="253" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<collectionCode id="DFF56AF1155E6350FEEBD186FE0FFD39" box="[340,404,726,749]" pageId="4" pageNumber="253">IEBR</collectionCode>
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A.2018.9 (Field no.
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<specimenCode id="E9425A4F155E6350FD25D186FCB7FD3A" box="[666,812,726,750]" collectionCode="CB" country="Australia" name="The CB Rhizobium Collection" pageId="4" pageNumber="253">CB 2014.19</specimenCode>
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), a female, from
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<location id="BC3BA4EF155E6350FBB8D185FA07FD3A" LSID="urn:lsid:plazi:treatment:314D4322155E6352FF28D2AEFDABF827:BC3BA4EF155E6350FBB8D185FA07FD3A" box="[1031,1436,725,750]" country="Vietnam" latitude="22.60245" longLatPrecision="1" longitude="105.87877" municipality="Nguyen Binh District" name="Phia Oac-Phia Den National Park" pageId="4" pageNumber="253" stateProvince="Cao Bang">Phia Oac-Phia Den National Park</location>
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(
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,
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, at an elevation of
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above sea level
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</elevation>
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),
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<collectingMunicipality id="593F684E155E6350FBAAD1AAFA8DFCC6" box="[1045,1302,761,786]" pageId="4" pageNumber="253">Nguyen Binh District</collectingMunicipality>
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,
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<collectingRegion id="7B203CD6155E6350FA98D1AAFF65FCE2" country="Vietnam" name="Cao Bang" pageId="4" pageNumber="253">Cao Bang Province</collectingRegion>
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, northern
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<collectingCountry id="C1F3B2A4155E6350FED2D04DFE54FCE2" box="[365,463,797,822]" name="Vietnam" pageId="4" pageNumber="253">Vietnam</collectingCountry>
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, collected by
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<collectorName id="141197E2155E6350FDD8D04EFD44FCE2" box="[615,735,797,822]" pageId="4" pageNumber="253">T.H. Pham</collectorName>
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on
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<collectingDate id="DD1E2D1C155E6350FCB7D04EFC04FCE2" box="[776,927,798,822]" pageId="4" pageNumber="253" value="2014-05-24">
|
||
<date id="CD5AD4F4155E6350FCB7D04EFC04FCE2" box="[776,927,798,822]" pageId="4" pageNumber="253" value="2014-05-24">24 May 2014</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="098CF869155E6350FC15D04EFD79FC76" ID-GBIF-Occurrence="2236983385" collectingDate="2010-05-10" collectionCode="VNMN" collectorName="T. T. Nguyen" country="Vietnam" county="Nguyen Binh District" elevation="1590" location="Thanh Cong Commune" municipality="Phia Oac-Phia Den National Park" pageId="4" pageNumber="253" specimenCode="TAO 766" specimenCount="1" stateProvince="Cao Bang" typeStatus="paratype">
|
||
<collectionCode id="DFF56AF1155E6350FC15D04EFC64FCE1" box="[938,1023,798,821]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="4" pageNumber="253">VNMN</collectionCode>
|
||
0 6924 (Field no.
|
||
<specimenCode id="E9425A4F155E6350FB7AD04EFAAAFCE2" box="[1221,1329,797,822]" collectionCode="TAO" pageId="4" pageNumber="253">TAO 766</specimenCode>
|
||
), an adult male, from forest of
|
||
<location id="BC3BA4EF155E6350FE10D011FD54FC8E" LSID="urn:lsid:plazi:treatment:314D4322155E6352FF28D2AEFDABF827:BC3BA4EF155E6350FE10D011FD54FC8E" box="[431,719,833,858]" country="Vietnam" county="Nguyen Binh District" municipality="Phia Oac-Phia Den National Park" name="Thanh Cong Commune" pageId="4" pageNumber="253" stateProvince="Cao Bang">Thanh Cong Commune</location>
|
||
within
|
||
<collectingMunicipality id="593F684E155E6350FC82D011FB7DFC8E" box="[829,1254,833,858]" pageId="4" pageNumber="253">Phia Oac-Phia Den National Park</collectingMunicipality>
|
||
(22°60.556’N, 105°87.530’E, at an elevation of
|
||
<elevation id="32C91507155E6350FDA7D035FCB4FCAA" box="[536,815,869,894]" metricMagnitude="3" metricUnit="m" metricValue="1.59" pageId="4" pageNumber="253" unit="m" value="1590.0">
|
||
<quantity id="7E1C5FD1155E6350FDA7D035FDEBFCA9" box="[536,624,869,894]" metricMagnitude="3" metricUnit="m" metricValue="1.59" pageId="4" pageNumber="253" unit="m" value="1590.0">1590 m</quantity>
|
||
above sea level
|
||
</elevation>
|
||
),
|
||
<collectingCounty id="503A8AB8155E6350FCFAD036FBA4FCAA" box="[837,1087,869,894]" pageId="4" pageNumber="253">Nguyen Binh District</collectingCounty>
|
||
,
|
||
<collectingRegion id="7B203CD6155E6350FBF2D036FAABFCAA" box="[1101,1328,869,894]" country="Vietnam" name="Cao Bang" pageId="4" pageNumber="253">Cao Bang Province</collectingRegion>
|
||
, northern
|
||
<collectingCountry id="C1F3B2A4155E6350FF28D0D9FF62FC76" box="[151,249,905,930]" name="Vietnam" pageId="4" pageNumber="253">Vietnam</collectingCountry>
|
||
, collected by
|
||
<collectorName id="141197E2155E6350FE2CD0DAFD84FC76" box="[403,543,906,930]" pageId="4" pageNumber="253">T.T. Nguyen</collectorName>
|
||
on
|
||
<collectingDate id="DD1E2D1C155E6350FDF6D0DAFD79FC76" box="[585,738,906,930]" pageId="4" pageNumber="253" value="2010-05-10">
|
||
<date id="CD5AD4F4155E6350FDF6D0DAFD79FC76" box="[585,738,906,930]" pageId="4" pageNumber="253" value="2010-05-10">10 May 2010</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F1FEA1BF155E6350FF78D0FCFD8EFB16" pageId="4" pageNumber="253" type="diagnosis">
|
||
<paragraph id="B95BF234155E6350FF78D0FCFD8EFB16" blockId="4.[151,1437,617,2010]" pageId="4" pageNumber="253">
|
||
<emphasis id="8B902E26155E6350FF78D0FCFEDAFC11" bold="true" box="[199,321,940,965]" pageId="4" pageNumber="253">Diagnosis.</emphasis>
|
||
A species of the genus
|
||
<taxonomicName id="7EE489B7155E6350FDEAD0FDFD5EFC11" authorityName="Peters" authorityYear="1869" box="[597,709,941,965]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="253" phylum="Chordata" rank="genus">
|
||
<emphasis id="8B902E26155E6350FDEAD0FDFD5EFC11" box="[597,709,941,965]" italics="true" pageId="4" pageNumber="253">Achalinus</emphasis>
|
||
</taxonomicName>
|
||
, characterized by a combination of the following characters: 1) maxillary teeth 28; 2) suture between internasals distinctly longer than that between the prefrontals; 3) internasal not fused to prefrontal; 4) loreal not fused with prefrontal; 5) infralabials 6; 6) mental separated from anterior chin shields; 7) two elongated anterior temporals, in contact with eye, and two posterior temporals; 8) dorsal scales in 25–23–23 rows, keeled; 9) ventrals 173–179; 10) subcaudals 77–91, unpaired; 11) a total length of at least
|
||
<quantity id="7E1C5FD1155E6350FAFFD76EFA06FB81" box="[1344,1437,1086,1110]" metricMagnitude="-1" metricUnit="m" metricValue="4.13" pageId="4" pageNumber="253" unit="mm" value="413.0">413 mm</quantity>
|
||
(with a maximum tail length of
|
||
<quantity id="7E1C5FD1155E6350FDB2D732FDF5FBAD" box="[525,622,1122,1146]" metricMagnitude="-1" metricUnit="m" metricValue="1.09" pageId="4" pageNumber="253" unit="mm" value="109.0">109 mm</quantity>
|
||
, and a tail/total length ratio of 0.22–0.37); 12) dorsum in preservative reddish to greyish brown above, the lower and posterior head sides paler; venter greyish cream, with the underside of the tail being somewhat darker.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F1FEA1BF155E6352FF78D79CFF7AF979" lastPageId="6" lastPageNumber="255" pageId="4" pageNumber="253" type="description">
|
||
<paragraph id="B95BF234155E6350FF78D79CFD32FA86" blockId="4.[151,1437,617,2010]" pageId="4" pageNumber="253">
|
||
<emphasis id="8B902E26155E6350FF78D79CFE7FFB31" bold="true" box="[199,484,1228,1253]" pageId="4" pageNumber="253">
|
||
Description of
|
||
<typeStatus id="665F4C96155E6350FEC9D79DFE44FB31" box="[374,479,1229,1253]" pageId="4" pageNumber="253" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Total length
|
||
<quantity id="7E1C5FD1155E6350FDC7D79DFD4EFB31" box="[632,725,1229,1254]" metricMagnitude="-1" metricUnit="m" metricValue="3.55" pageId="4" pageNumber="253" unit="mm" value="355.0">355 mm</quantity>
|
||
(SVL
|
||
<quantity id="7E1C5FD1155E6350FC9FD79DFCE6FB31" box="[800,893,1229,1254]" metricMagnitude="-1" metricUnit="m" metricValue="2.6" pageId="4" pageNumber="253" unit="mm" value="260.0">260 mm</quantity>
|
||
and TaL
|
||
<quantity id="7E1C5FD1155E6350FC58D79DFBA1FB32" box="[999,1082,1229,1254]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="4" pageNumber="253" unit="mm" value="95.0">95 mm</quantity>
|
||
); tail long, tail/total length ratio 0.365; body slender, cylindrical; head length
|
||
<quantity id="7E1C5FD1155E6350FD16D7A2FC98FADD" box="[681,771,1266,1290]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="4" pageNumber="253" unit="mm" value="8.9">8.9 mm</quantity>
|
||
(from tip of snout to posterior margin of parietal); head slightly distinct from neck, dorsally covered with large shields; eye small, with vertically subelliptic pupil; left maxilla with 28 equally sized and curved teeth.
|
||
</paragraph>
|
||
<paragraph id="B95BF234155E6350FF78D60DFDC4F96E" blockId="4.[151,1437,617,2010]" pageId="4" pageNumber="253">
|
||
Rostral small, triangular, slightly visible from above; suture between the internasals (
|
||
<quantity id="7E1C5FD1155E6350FB3BD60EFB47FAA2" box="[1156,1244,1374,1398]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="253" unit="mm" value="1.4">1.4 mm</quantity>
|
||
) longer than that between the prefrontals (
|
||
<quantity id="7E1C5FD1155E6350FE11D6D2FD91FA4E" box="[430,522,1410,1434]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="253" unit="mm" value="0.9">0.9 mm</quantity>
|
||
); nostril in the anterior part of the nasal; frontal pentagonal, slightly broader than long, pointed backwards, much shorter than the parietals; each parietal bordered by an elongated nuchal; nuchals separated from each other behind parietals by two small scales; second pair of nuchals about half the size than first pair; one loreal, distinctly wider than high, extending from the nasal to the eye; one supraocular; two anterior temporals, elongated, upper one smaller, both in contact with eye, and two elongated posterior temporals; supralabials six, the first smallest, fourth and fifth in contact with the eye, sixth longest; third and fourth in contact with the loreal; one mental, followed by six infralabials; first pair of infralabials in contact with each other; first four infralabials in contact with the first pair of chin shields; posterior pair of chin shields smaller, laterally in contact with fourth and fifth infralabials.
|
||
</paragraph>
|
||
<paragraph id="B95BF234155E6350FF78D595FD46F89E" blockId="4.[151,1437,617,2010]" pageId="4" pageNumber="253">Dorsal scales elliptical, keeled from the neck region onwards, 23 scale rows at midbody, those of the outer row enlarged, 25 scales round the anterior part of the body (one head length behind head), and 23 dorsal scale rows at posterior body (one head length before vent); ventrals 169 (plus one preventral), distinctly rounded laterally; subcaudals 91, unpaired, plus tail tip; cloacal entire.</paragraph>
|
||
<paragraph id="B95BF234155E6350FF78D405FB38F862" blockId="4.[151,1437,617,2010]" pageId="4" pageNumber="253">
|
||
The ethanol-preserved
|
||
<typeStatus id="665F4C96155E6350FE72D405FDB4F8BA" box="[461,559,1877,1902]" pageId="4" pageNumber="253" type="holotype">holotype</typeStatus>
|
||
greyish brown above, the lower and posterior head sides being paler; venter greyish cream, with the underside of the tail being somewhat darker and the chin region somewhat paler; infralabial and chin shields light greyish brown. Ventrals and subcaudals anteriorly and laterally darker.
|
||
</paragraph>
|
||
<paragraph id="B95BF234155E6351FF78D491FCCBFEC9" blockId="4.[151,1437,617,2010]" lastBlockId="5.[151,1437,151,1005]" lastPageId="5" lastPageNumber="254" pageId="4" pageNumber="253">
|
||
<emphasis id="8B902E26155E6350FF78D491FEA4F80E" bold="true" box="[199,319,1985,2010]" pageId="4" pageNumber="253">Variation.</emphasis>
|
||
For scalation details of the
|
||
<typeStatus id="665F4C96155E6350FD3ED492FD77F80E" box="[641,748,1986,2010]" pageId="4" pageNumber="253" type="paratype">paratypes</typeStatus>
|
||
see
|
||
<tableCitation id="F466C78F155E6350FC9ED491FCE3F80E" box="[801,888,1985,2010]" captionStart="TABLE 3" captionStartId="6.[151,239,152,175]" captionTargetPageId="6" captionText="TABLE 3. Sex, measurements, ratios, scalation and colour pattern of the type series of Achalinus juliani sp. nov.; * = in addition to the two elongated anterior temporals, a very small third anterior temporal is discernible in the upper back corner of the eye on the right side." httpUri="http://table.plazi.org/id/ED9BA2BC155C6352FF28D3C8FD9EFF3B" pageId="4" pageNumber="253" tableUuid="ED9BA2BC155C6352FF28D3C8FD9EFF3B">Table 3</tableCitation>
|
||
. In addition, the nuchals of the female
|
||
<typeStatus id="665F4C96155E6350FA83D492FA07F80E" box="[1340,1436,1986,2010]" pageId="4" pageNumber="253" type="paratype">paratype</typeStatus>
|
||
IEBR
|
||
<materialsCitation id="098CF869155F6351FF5ED3C8FECEFF00" ID-GBIF-Occurrence="2236983383" collectionCode="VNMN" pageId="5" pageNumber="254" specimenCount="1" typeStatus="paratype">
|
||
A.2018.9 are separated from each other behind the parietals by four small scales and those of the male
|
||
<typeStatus id="665F4C96155F6351FF28D3EDFF6CFF01" box="[151,247,189,213]" pageId="5" pageNumber="254" type="paratype">paratype</typeStatus>
|
||
<collectionCode id="DFF56AF1155F6351FEBFD3EDFECEFF00" box="[256,341,189,212]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="5" pageNumber="254">VNMN</collectionCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="098CF869155F6351FEE1D3ECFA07FF00" ID-GBIF-Occurrence="2236983382" box="[350,1436,188,213]" collectionCode="VNMN" pageId="5" pageNumber="254" specimenCount="1" typeStatus="paratype">
|
||
0 6924 by two large, two medium sized and a small scale. Furthermore, in the
|
||
<typeStatus id="665F4C96155F6351FB62D3EDFAA5FF01" box="[1245,1342,189,213]" pageId="5" pageNumber="254" type="paratype">paratype</typeStatus>
|
||
<collectionCode id="DFF56AF1155F6351FAF8D3EDFA07FF00" box="[1351,1436,189,212]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="5" pageNumber="254">VNMN</collectionCode>
|
||
</materialsCitation>
|
||
0 6924 the posterior pair of chin shields on the left side is in contact with third and fourth infralabials laterally. The hemipenes are basally everted in the specimen VNMN 0 6924.
|
||
</paragraph>
|
||
<paragraph id="B95BF234155F6351FF78D277FCAAFE5C" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="254">
|
||
<emphasis id="8B902E26155F6351FF78D277FEF6FE94" bold="true" box="[199,365,295,320]" pageId="5" pageNumber="254">Comparisons.</emphasis>
|
||
<taxonomicName id="7EE489B7155F6351FECAD278FDACFE94" authority="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le, 2019" authorityName="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le" authorityYear="2019" box="[373,567,296,320]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="juliani" status="sp. nov.">
|
||
<emphasis id="8B902E26155F6351FECAD278FDACFE94" box="[373,567,296,320]" italics="true" pageId="5" pageNumber="254">Achalinus juliani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90A3935D155F6351FDFFD278FD07FE94" box="[576,668,296,320]" pageId="5" pageNumber="254" rank="species">
|
||
<emphasis id="8B902E26155F6351FDFFD278FD07FE94" bold="true" box="[576,668,296,320]" pageId="5" pageNumber="254">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be differentiated from the remaining
|
||
<taxonomicName id="7EE489B7155F6351FBC4D278FB70FE94" authorityName="Peters" authorityYear="1869" box="[1147,1259,296,320]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="254" phylum="Chordata" rank="genus">
|
||
<emphasis id="8B902E26155F6351FBC4D278FB70FE94" box="[1147,1259,296,320]" italics="true" pageId="5" pageNumber="254">Achalinus</emphasis>
|
||
</taxonomicName>
|
||
representatives from
|
||
<collectingCountry id="C1F3B2A4155F6351FF6BD21CFEA8FEB1" box="[212,307,332,357]" name="Vietnam" pageId="5" pageNumber="254">Vietnam</collectingCountry>
|
||
by having more subcaudals, more dorsal scale rows on anterior part of the body, more ventrals, more infralabials, and a longer internasal suture (see
|
||
<tableCitation id="F466C78F155F6351FD18D23FFC9CFE5C" box="[679,775,367,392]" captionStart="TABLE 3" captionStartId="6.[151,239,152,175]" captionTargetPageId="6" captionText="TABLE 3. Sex, measurements, ratios, scalation and colour pattern of the type series of Achalinus juliani sp. nov.; * = in addition to the two elongated anterior temporals, a very small third anterior temporal is discernible in the upper back corner of the eye on the right side." httpUri="http://table.plazi.org/id/ED9BA2BC155C6352FF28D3C8FD9EFF3B" pageId="5" pageNumber="254" tableUuid="ED9BA2BC155C6352FF28D3C8FD9EFF3B">Tables 3</tableCitation>
|
||
,
|
||
<tableCitation id="F466C78F155F6351FCACD23FFCB8FE5C" box="[787,803,367,392]" captionStart="TABLE 5" captionStartId="18.[147,220,543,562]" captionTargetBox="[147,1872,667,1680]" captionText="TABLE 5. Morphometry, dentition and scalation of members of the genus Achalinus (species in bold are occurring in Vietnam) according to Van Denburgh (1912), Pope (1935), Bourret (1935, 1937), Smith (1943), Koshikawa (1982), Karsen et al. (1986), Ota & Toyama (1989), Inger et al. (1990), Ziegler (2002), Goris & Maeda (2004), Zhao (2006), and Das (2010); values given in brackets indicate infrequent condition: </= almost as long as or shorter; = equals; Int. fus. = internasal fused to prefrontal; Intern. sut. = internasal suture; prefr. sut. = prefrontal suture; Ment. sep. = Mental separated from anterior postmental / chin shields; * = by a suture; ** = according to Jian Wang (pers. comm.), but given as smooth in Ota & Toyama (1989); m = male; f = female; s+k = smooth on anterior, keeled on posterior body; s+k+k = keeled on mid and posterior body." httpUri="http://table.plazi.org/id/ED9BA2BC15486346FF2CD14FFD09FD41" pageId="5" pageNumber="254" tableUuid="ED9BA2BC15486346FF2CD14FFD09FD41">5</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B95BF234155F6351FF78D2C4FCCEFE04" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="254">
|
||
<taxonomicName id="7EE489B7155F6351FF78D2C4FE1CFE78" authority="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le, 2019" authorityName="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le" authorityYear="2019" box="[199,391,404,428]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="juliani" status="sp. nov.">
|
||
<emphasis id="8B902E26155F6351FF78D2C4FE1CFE78" box="[199,391,404,428]" italics="true" pageId="5" pageNumber="254">Achalinus juliani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90A3935D155F6351FE31D2C4FE7CFE78" box="[398,487,404,428]" pageId="5" pageNumber="254" rank="species">
|
||
<emphasis id="8B902E26155F6351FE31D2C4FE7CFE78" bold="true" box="[398,487,404,428]" pageId="5" pageNumber="254">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="7EE489B7155F6351FDC4D2C4FD5CFE78" authorityName="Bourret" authorityYear="1937" box="[635,711,404,428]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="ater">
|
||
<emphasis id="8B902E26155F6351FDC4D2C4FD5CFE78" box="[635,711,404,428]" italics="true" pageId="5" pageNumber="254">A. ater</emphasis>
|
||
</taxonomicName>
|
||
in having more dorsal scale rows in the anterior part of the body (25 versus 21–23) and more subcaudals (77–91 versus 47–70).
|
||
</paragraph>
|
||
<paragraph id="B95BF234155F6351FF78D28CFB2DFDCC" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="254">
|
||
<taxonomicName id="7EE489B7155F6351FF78D28CFE12FE20" authority="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le, 2019" authorityName="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le" authorityYear="2019" box="[199,393,476,500]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="juliani" status="sp. nov.">
|
||
<emphasis id="8B902E26155F6351FF78D28CFE12FE20" box="[199,393,476,500]" italics="true" pageId="5" pageNumber="254">Achalinus juliani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90A3935D155F6351FE2CD28CFE74FE20" box="[403,495,476,500]" pageId="5" pageNumber="254" rank="species">
|
||
<emphasis id="8B902E26155F6351FE2CD28CFE74FE20" bold="true" box="[403,495,476,500]" pageId="5" pageNumber="254">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="7EE489B7155F6351FD34D28CFC8CFE20" authorityName="Boulenger" authorityYear="1888" box="[651,791,476,500]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="rufescens">
|
||
<emphasis id="8B902E26155F6351FD34D28CFC8CFE20" box="[651,791,476,500]" italics="true" pageId="5" pageNumber="254">A. rufescens</emphasis>
|
||
</taxonomicName>
|
||
by having more infralabials (6 versus 5), more ventrals (163–179 versus 131–158), more subcaudals (77–91 versus 54–82), and more maxillary teeth.
|
||
</paragraph>
|
||
<paragraph id="B95BF234155F6351FF78D174FEDCFD7C" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="254">
|
||
<taxonomicName id="7EE489B7155F6351FF78D174FE11FDE8" authority="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le, 2019" authorityName="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le" authorityYear="2019" box="[199,394,548,572]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="juliani" status="sp. nov.">
|
||
<emphasis id="8B902E26155F6351FF78D174FE11FDE8" box="[199,394,548,572]" italics="true" pageId="5" pageNumber="254">Achalinus juliani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90A3935D155F6351FE2BD174FE6AFDE8" box="[404,497,548,572]" pageId="5" pageNumber="254" rank="species">
|
||
<emphasis id="8B902E26155F6351FE2BD174FE6AFDE8" bold="true" box="[404,497,548,572]" pageId="5" pageNumber="254">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="7EE489B7155F6351FD30D174FC93FDE8" authorityName="Peters" authorityYear="1869" box="[655,776,548,572]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="spinalis">
|
||
<emphasis id="8B902E26155F6351FD30D174FC93FDE8" box="[655,776,548,572]" italics="true" pageId="5" pageNumber="254">A. spinalis</emphasis>
|
||
</taxonomicName>
|
||
by the internasal suture being distinctly longer than that between the prefrontals (versus as long as or shorter), more dorsal scale rows in the anterior part of the body (25 versus 21–24), more subcaudals (77–91 versus 39–67), by lacking a black mid-dorsal line, and by having more maxillary teeth.
|
||
</paragraph>
|
||
<paragraph id="B95BF234155F6351FF78D1E4FE09FC54" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="254">
|
||
<taxonomicName id="7EE489B7155F6351FF78D1E4FE11FD18" authority="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le, 2019" authorityName="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le" authorityYear="2019" box="[199,394,692,716]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="juliani" status="sp. nov.">
|
||
<emphasis id="8B902E26155F6351FF78D1E4FE11FD18" box="[199,394,692,716]" italics="true" pageId="5" pageNumber="254">Achalinus juliani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90A3935D155F6351FE2BD1E4FE6BFD18" box="[404,496,692,716]" pageId="5" pageNumber="254" rank="species">
|
||
<emphasis id="8B902E26155F6351FE2BD1E4FE6BFD18" bold="true" box="[404,496,692,716]" pageId="5" pageNumber="254">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="7EE489B7155F6351FD33D1E4FCA9FD18" authorityName="Boulenger" authorityYear="1908" box="[652,818,692,716]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="formosanus">
|
||
<emphasis id="8B902E26155F6351FD33D1E4FCA9FD18" box="[652,818,692,716]" italics="true" pageId="5" pageNumber="254">A. formosanus</emphasis>
|
||
</taxonomicName>
|
||
by having the internasal suture distinctly longer than prefrontal suture (versus internasal suture almost as long as prefrontal suture), by the loreal not being fused with prefrontal, fewer dorsal scale rows (25–23–23 versus 25–29 – 25–27–25), more maxillary teeth (28 versus 14–17), and by lacking a black mid-dorsal line; in addition,
|
||
<taxonomicName id="7EE489B7155F6351FD44D070FC24FCEC" authority="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le, 2019" authorityName="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le" authorityYear="2019" box="[763,959,800,824]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="juliani" status="sp. nov.">
|
||
<emphasis id="8B902E26155F6351FD44D070FC24FCEC" box="[763,959,800,824]" italics="true" pageId="5" pageNumber="254">Achalinus juliani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90A3935D155F6351FC75D070FBBCFCEC" box="[970,1063,800,824]" pageId="5" pageNumber="254" rank="species">
|
||
<emphasis id="8B902E26155F6351FC75D070FBBCFCEC" bold="true" box="[970,1063,800,824]" pageId="5" pageNumber="254">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be distinguished from
|
||
<emphasis id="8B902E26155F6351FAD5D070FE81FC88" italics="true" pageId="5" pageNumber="254">
|
||
A. f.
|
||
<taxonomicName id="7EE489B7155F6351FF28D014FE81FC88" authorityName="Boulenger" authorityYear="1908" box="[151,282,836,860]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="formosanus">formosanus</taxonomicName>
|
||
</emphasis>
|
||
by having more subcaudals (77–91 versus 61–83), and from
|
||
<emphasis id="8B902E26155F6351FC72D014FBC7FC88" box="[973,1116,836,860]" italics="true" pageId="5" pageNumber="254">A. f. chigirai</emphasis>
|
||
by having fewer subcaudals (77–91 versus 96–97).
|
||
</paragraph>
|
||
<paragraph id="B95BF234155F6351FF78D0DCFEA7FC39" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="254">
|
||
<taxonomicName id="7EE489B7155F6351FF78D0DCFE10FC70" authority="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le, 2019" authorityName="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le" authorityYear="2019" box="[199,395,908,932]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="juliani" status="sp. nov.">
|
||
<emphasis id="8B902E26155F6351FF78D0DCFE10FC70" box="[199,395,908,932]" italics="true" pageId="5" pageNumber="254">Achalinus juliani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90A3935D155F6351FE29D0DCFE6EFC70" box="[406,501,908,932]" pageId="5" pageNumber="254" rank="species">
|
||
<emphasis id="8B902E26155F6351FE29D0DCFE6EFC70" bold="true" box="[406,501,908,932]" pageId="5" pageNumber="254">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="7EE489B7155F6351FD2AD0DCFC84FC70" authorityName="Huang" authorityYear="1975" box="[661,799,908,932]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="hainanus">
|
||
<emphasis id="8B902E26155F6351FD2AD0DCFC84FC70" box="[661,799,908,932]" italics="true" pageId="5" pageNumber="254">A. hainanus</emphasis>
|
||
</taxonomicName>
|
||
by having 6 versus 5 infralabials, 2 versus 1 anterior temporal, more dorsal scale rows in the anterior part of the body (25 versus 23), and more subcaudals (77–91 versus 67–69).
|
||
</paragraph>
|
||
<caption id="ED9BA2BC155F6351FF28D4E9FD0DF83A" httpUri="https://zenodo.org/record/2651882/files/figure.png" pageId="5" pageNumber="254" startId="5.[151,250,1977,1999]" targetBox="[151,1437,1052,1960]" targetPageId="5">
|
||
<paragraph id="B95BF234155F6351FF28D4E9FD0DF83A" blockId="5.[151,1436,1977,2030]" pageId="5" pageNumber="254">
|
||
<emphasis id="8B902E26155F6351FF28D4E9FE88F81A" bold="true" box="[151,275,1977,1999]" pageId="5" pageNumber="254">FIGURE 2.</emphasis>
|
||
Head views of the preserved holotype of
|
||
<taxonomicName id="7EE489B7155F6351FD00D4E9FCF6F81B" authority="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le, 2019" authorityName="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le" authorityYear="2019" box="[703,877,1977,1999]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="juliani" status="sp. nov.">
|
||
<emphasis id="8B902E26155F6351FD00D4E9FCF6F81B" box="[703,877,1977,1999]" italics="true" pageId="5" pageNumber="254">Achalinus juliani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90A3935D155F6351FCCBD4E9FC5EF81A" box="[884,965,1977,1998]" pageId="5" pageNumber="254" rank="species">
|
||
<emphasis id="8B902E26155F6351FCCBD4E9FC5EF81A" bold="true" box="[884,965,1977,1998]" pageId="5" pageNumber="254">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(IEBR A.2018.8) from Ha Lang District, Cao Bang Province, northern Vietnam. Phot. T. Ziegler
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="ED9BA2BC155C6352FF28D3C8FD9EFF3B" ID-Table-UUID="ED9BA2BC155C6352FF28D3C8FD9EFF3B" httpUri="http://table.plazi.org/id/ED9BA2BC155C6352FF28D3C8FD9EFF3B" pageId="6" pageNumber="255" targetBox="[159,1335,266,1259]" targetIsTable="true" targetPageId="6">
|
||
<paragraph id="B95BF234155C6352FF28D3C8FD9EFF3B" blockId="6.[151,1436,151,239]" pageId="6" pageNumber="255">
|
||
<emphasis id="8B902E26155C6352FF28D3C8FE92FF7B" bold="true" box="[151,265,152,175]" pageId="6" pageNumber="255">TABLE 3.</emphasis>
|
||
Sex, measurements, ratios, scalation and colour pattern of the type series of
|
||
<taxonomicName id="7EE489B7155C6352FB83D3C7FB68FF7B" authority="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le, 2019" authorityName="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le" authorityYear="2019" box="[1084,1267,151,175]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="juliani" status="sp. nov.">
|
||
<emphasis id="8B902E26155C6352FB83D3C7FB68FF7B" box="[1084,1267,151,175]" italics="true" pageId="6" pageNumber="255">Achalinus juliani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90A3935D155C6352FB45D3C8FAD4FF7B" box="[1274,1359,152,175]" pageId="6" pageNumber="255" rank="species">
|
||
<emphasis id="8B902E26155C6352FB45D3C8FAD4FF7B" bold="true" box="[1274,1359,152,175]" pageId="6" pageNumber="255">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; * = in addition to the two elongated anterior temporals, a very small third anterior temporal is discernible in the upper back corner of the eye on the right side.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B95BF234155C6352FF20D25AFAADFB3F" pageId="6" pageNumber="255">
|
||
<table id="CBE40094155C9CABFF20D25AFAACFB3F" box="[159,1335,266,1259]" gridcols="4" gridrows="23" pageId="6" pageNumber="255">
|
||
<tr id="07D4F076155C9CABFF20D25AFAACFE8A" box="[159,1335,266,350]" gridrow="0" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D25AFDAFFE8A" box="[159,564,266,350]" gridcol="0" gridrow="0" pageId="6" pageNumber="255">Character</th>
|
||
<th id="4405990A155C9CABFD2AD25AFCA8FE8A" box="[661,819,266,350]" gridcol="1" gridrow="0" pageId="6" pageNumber="255">IEBR A.2018.8 (CB 2014.18) holotype</th>
|
||
<th id="4405990A155C9CABFC2CD25AFBAAFE8A" box="[915,1073,266,350]" gridcol="2" gridrow="0" pageId="6" pageNumber="255">IEBR A.2018.9 (CB 2014.19) paratype</th>
|
||
<th id="4405990A155C9CABFB1BD25AFAACFE8A" box="[1188,1335,266,350]" gridcol="3" gridrow="0" pageId="6" pageNumber="255">VNMN 0 6924 (TAO 766) paratype</th>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D221FAACFE53" box="[159,1335,369,391]" gridrow="1" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D221FDAFFE53" box="[159,564,369,391]" gridcol="0" gridrow="1" pageId="6" pageNumber="255">Sex</th>
|
||
<td id="4405990A155C9CABFD2AD221FCA8FE53" box="[661,819,369,391]" gridcol="1" gridrow="1" pageId="6" pageNumber="255">male</td>
|
||
<td id="4405990A155C9CABFC2CD221FBAAFE53" box="[915,1073,369,391]" gridcol="2" gridrow="1" pageId="6" pageNumber="255">female</td>
|
||
<td id="4405990A155C9CABFB1BD221FAACFE53" box="[1188,1335,369,391]" gridcol="3" gridrow="1" pageId="6" pageNumber="255">male</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D2CAFAACFE64" box="[159,1335,410,432]" gridrow="2" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D2CAFDAFFE64" box="[159,564,410,432]" gridcol="0" gridrow="2" pageId="6" pageNumber="255">TL</th>
|
||
<td id="4405990A155C9CABFD2AD2CAFCA8FE64" box="[661,819,410,432]" gridcol="1" gridrow="2" pageId="6" pageNumber="255">355</td>
|
||
<td id="4405990A155C9CABFC2CD2CAFBAAFE64" box="[915,1073,410,432]" gridcol="2" gridrow="2" pageId="6" pageNumber="255">303</td>
|
||
<td id="4405990A155C9CABFB1BD2CAFAACFE64" box="[1188,1335,410,432]" gridcol="3" gridrow="2" pageId="6" pageNumber="255">413</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D294FAACFE0E" box="[159,1335,452,474]" gridrow="3" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D294FDAFFE0E" box="[159,564,452,474]" gridcol="0" gridrow="3" pageId="6" pageNumber="255">SVL</th>
|
||
<td id="4405990A155C9CABFD2AD294FCA8FE0E" box="[661,819,452,474]" gridcol="1" gridrow="3" pageId="6" pageNumber="255">260</td>
|
||
<td id="4405990A155C9CABFC2CD294FBAAFE0E" box="[915,1073,452,474]" gridcol="2" gridrow="3" pageId="6" pageNumber="255">235</td>
|
||
<td id="4405990A155C9CABFB1BD294FAACFE0E" box="[1188,1335,452,474]" gridcol="3" gridrow="3" pageId="6" pageNumber="255">304</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D2BDFAACFDD7" box="[159,1335,493,515]" gridrow="4" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D2BDFDAFFDD7" box="[159,564,493,515]" gridcol="0" gridrow="4" pageId="6" pageNumber="255">TaL</th>
|
||
<td id="4405990A155C9CABFD2AD2BDFCA8FDD7" box="[661,819,493,515]" gridcol="1" gridrow="4" pageId="6" pageNumber="255">95</td>
|
||
<td id="4405990A155C9CABFC2CD2BDFBAAFDD7" box="[915,1073,493,515]" gridcol="2" gridrow="4" pageId="6" pageNumber="255">68</td>
|
||
<td id="4405990A155C9CABFB1BD2BDFAACFDD7" box="[1188,1335,493,515]" gridcol="3" gridrow="4" pageId="6" pageNumber="255">109</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D146FAACFDF8" box="[159,1335,534,556]" gridrow="5" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D146FDAFFDF8" box="[159,564,534,556]" gridcol="0" gridrow="5" pageId="6" pageNumber="255">TaL/TL</th>
|
||
<td id="4405990A155C9CABFD2AD146FCA8FDF8" box="[661,819,534,556]" gridcol="1" gridrow="5" pageId="6" pageNumber="255">0.365</td>
|
||
<td id="4405990A155C9CABFC2CD146FBAAFDF8" box="[915,1073,534,556]" gridcol="2" gridrow="5" pageId="6" pageNumber="255">0.224</td>
|
||
<td id="4405990A155C9CABFB1BD146FAACFDF8" box="[1188,1335,534,556]" gridcol="3" gridrow="5" pageId="6" pageNumber="255">0.264</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D110FAACFD82" box="[159,1335,576,598]" gridrow="6" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D110FDAFFD82" box="[159,564,576,598]" gridcol="0" gridrow="6" pageId="6" pageNumber="255">SL</th>
|
||
<td id="4405990A155C9CABFD2AD110FCA8FD82" box="[661,819,576,598]" gridcol="1" gridrow="6" pageId="6" pageNumber="255">6</td>
|
||
<td id="4405990A155C9CABFC2CD110FBAAFD82" box="[915,1073,576,598]" gridcol="2" gridrow="6" pageId="6" pageNumber="255">6/7</td>
|
||
<td id="4405990A155C9CABFB1BD110FAACFD82" box="[1188,1335,576,598]" gridcol="3" gridrow="6" pageId="6" pageNumber="255">6</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D139FAACFDAB" box="[159,1335,617,639]" gridrow="7" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D139FDAFFDAB" box="[159,564,617,639]" gridcol="0" gridrow="7" pageId="6" pageNumber="255">SL/orbit</th>
|
||
<td id="4405990A155C9CABFD2AD139FCA8FDAB" box="[661,819,617,639]" gridcol="1" gridrow="7" pageId="6" pageNumber="255">4–5</td>
|
||
<td id="4405990A155C9CABFC2CD139FBAAFDAB" box="[915,1073,617,639]" gridcol="2" gridrow="7" pageId="6" pageNumber="255">4–5/5–6</td>
|
||
<td id="4405990A155C9CABFB1BD139FAACFDAB" box="[1188,1335,617,639]" gridcol="3" gridrow="7" pageId="6" pageNumber="255">4–5</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D1C2FAACFD7C" box="[159,1335,658,680]" gridrow="8" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D1C2FDAFFD7C" box="[159,564,658,680]" gridcol="0" gridrow="8" pageId="6" pageNumber="255">SL/loreal</th>
|
||
<td id="4405990A155C9CABFD2AD1C2FCA8FD7C" box="[661,819,658,680]" gridcol="1" gridrow="8" pageId="6" pageNumber="255">3–4</td>
|
||
<td id="4405990A155C9CABFC2CD1C2FBAAFD7C" box="[915,1073,658,680]" gridcol="2" gridrow="8" pageId="6" pageNumber="255">4–5/5–6</td>
|
||
<td id="4405990A155C9CABFB1BD1C2FAACFD7C" box="[1188,1335,658,680]" gridcol="3" gridrow="8" pageId="6" pageNumber="255">3–4</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D1ECFAACFD06" box="[159,1335,700,722]" gridrow="9" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D1ECFDAFFD06" box="[159,564,700,722]" gridcol="0" gridrow="9" pageId="6" pageNumber="255">Longest SL</th>
|
||
<td id="4405990A155C9CABFD2AD1ECFCA8FD06" box="[661,819,700,722]" gridcol="1" gridrow="9" pageId="6" pageNumber="255">6</td>
|
||
<td id="4405990A155C9CABFC2CD1ECFBAAFD06" box="[915,1073,700,722]" gridcol="2" gridrow="9" pageId="6" pageNumber="255">6/7</td>
|
||
<td id="4405990A155C9CABFB1BD1ECFAACFD06" box="[1188,1335,700,722]" gridcol="3" gridrow="9" pageId="6" pageNumber="255">6</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D1B5FAACFD2F" box="[159,1335,741,763]" gridrow="10" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D1B5FDAFFD2F" box="[159,564,741,763]" gridcol="0" gridrow="10" pageId="6" pageNumber="255">IL</th>
|
||
<td id="4405990A155C9CABFD2AD1B5FCA8FD2F" box="[661,819,741,763]" gridcol="1" gridrow="10" pageId="6" pageNumber="255">6</td>
|
||
<td id="4405990A155C9CABFC2CD1B5FBAAFD2F" box="[915,1073,741,763]" gridcol="2" gridrow="10" pageId="6" pageNumber="255">6</td>
|
||
<td id="4405990A155C9CABFB1BD1B5FAACFD2F" box="[1188,1335,741,763]" gridcol="3" gridrow="10" pageId="6" pageNumber="255">6</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D05EFAACFCF0" box="[159,1335,782,804]" gridrow="11" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D05EFDAFFCF0" box="[159,564,782,804]" gridcol="0" gridrow="11" pageId="6" pageNumber="255">IL in contact with 1. pair of chin shields</th>
|
||
<td id="4405990A155C9CABFD2AD05EFCA8FCF0" box="[661,819,782,804]" gridcol="1" gridrow="11" pageId="6" pageNumber="255">1–4</td>
|
||
<td id="4405990A155C9CABFC2CD05EFBAAFCF0" box="[915,1073,782,804]" gridcol="2" gridrow="11" pageId="6" pageNumber="255">1–4/1–3</td>
|
||
<td id="4405990A155C9CABFB1BD05EFAACFCF0" box="[1188,1335,782,804]" gridcol="3" gridrow="11" pageId="6" pageNumber="255">1–3</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D068FAACFC9A" box="[159,1335,824,846]" gridrow="12" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D068FDAFFC9A" box="[159,564,824,846]" gridcol="0" gridrow="12" pageId="6" pageNumber="255">Loreal</th>
|
||
<td id="4405990A155C9CABFD2AD068FCA8FC9A" box="[661,819,824,846]" gridcol="1" gridrow="12" pageId="6" pageNumber="255">1</td>
|
||
<td id="4405990A155C9CABFC2CD068FBAAFC9A" box="[915,1073,824,846]" gridcol="2" gridrow="12" pageId="6" pageNumber="255">1</td>
|
||
<td id="4405990A155C9CABFB1BD068FAACFC9A" box="[1188,1335,824,846]" gridcol="3" gridrow="12" pageId="6" pageNumber="255">1</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D031FAACFCA3" box="[159,1335,865,887]" gridrow="13" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D031FDAFFCA3" box="[159,564,865,887]" gridcol="0" gridrow="13" pageId="6" pageNumber="255">Supraocular</th>
|
||
<td id="4405990A155C9CABFD2AD031FCA8FCA3" box="[661,819,865,887]" gridcol="1" gridrow="13" pageId="6" pageNumber="255">1</td>
|
||
<td id="4405990A155C9CABFC2CD031FBAAFCA3" box="[915,1073,865,887]" gridcol="2" gridrow="13" pageId="6" pageNumber="255">1</td>
|
||
<td id="4405990A155C9CABFB1BD031FAACFCA3" box="[1188,1335,865,887]" gridcol="3" gridrow="13" pageId="6" pageNumber="255">1</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D0DAFAACFC74" box="[159,1335,906,928]" gridrow="14" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D0DAFDAFFC74" box="[159,564,906,928]" gridcol="0" gridrow="14" pageId="6" pageNumber="255">Elongated ATem</th>
|
||
<td id="4405990A155C9CABFD2AD0DAFCA8FC74" box="[661,819,906,928]" gridcol="1" gridrow="14" pageId="6" pageNumber="255">2</td>
|
||
<td id="4405990A155C9CABFC2CD0DAFBAAFC74" box="[915,1073,906,928]" gridcol="2" gridrow="14" pageId="6" pageNumber="255">2</td>
|
||
<td id="4405990A155C9CABFB1BD0DAFAACFC74" box="[1188,1335,906,928]" gridcol="3" gridrow="14" pageId="6" pageNumber="255">2*</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D0E4FAACFC1E" box="[159,1335,948,970]" gridrow="15" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D0E4FDAFFC1E" box="[159,564,948,970]" gridcol="0" gridrow="15" pageId="6" pageNumber="255">Elongated PTem</th>
|
||
<td id="4405990A155C9CABFD2AD0E4FCA8FC1E" box="[661,819,948,970]" gridcol="1" gridrow="15" pageId="6" pageNumber="255">2</td>
|
||
<td id="4405990A155C9CABFC2CD0E4FBAAFC1E" box="[915,1073,948,970]" gridcol="2" gridrow="15" pageId="6" pageNumber="255">2</td>
|
||
<td id="4405990A155C9CABFB1BD0E4FAACFC1E" box="[1188,1335,948,970]" gridcol="3" gridrow="15" pageId="6" pageNumber="255">2</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D08DFAACFC27" box="[159,1335,989,1011]" gridrow="16" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D08DFDAFFC27" box="[159,564,989,1011]" gridcol="0" gridrow="16" pageId="6" pageNumber="255">Ant DSR</th>
|
||
<td id="4405990A155C9CABFD2AD08DFCA8FC27" box="[661,819,989,1011]" gridcol="1" gridrow="16" pageId="6" pageNumber="255">25</td>
|
||
<td id="4405990A155C9CABFC2CD08DFBAAFC27" box="[915,1073,989,1011]" gridcol="2" gridrow="16" pageId="6" pageNumber="255">25</td>
|
||
<td id="4405990A155C9CABFB1BD08DFAACFC27" box="[1188,1335,989,1011]" gridcol="3" gridrow="16" pageId="6" pageNumber="255">25</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D756FAACFBC8" box="[159,1335,1030,1052]" gridrow="17" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D756FDAFFBC8" box="[159,564,1030,1052]" gridcol="0" gridrow="17" pageId="6" pageNumber="255">Mid DSR</th>
|
||
<td id="4405990A155C9CABFD2AD756FCA8FBC8" box="[661,819,1030,1052]" gridcol="1" gridrow="17" pageId="6" pageNumber="255">23</td>
|
||
<td id="4405990A155C9CABFC2CD756FBAAFBC8" box="[915,1073,1030,1052]" gridcol="2" gridrow="17" pageId="6" pageNumber="255">23</td>
|
||
<td id="4405990A155C9CABFB1BD756FAACFBC8" box="[1188,1335,1030,1052]" gridcol="3" gridrow="17" pageId="6" pageNumber="255">23</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D760FAACFB92" box="[159,1335,1072,1094]" gridrow="18" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D760FDAFFB92" box="[159,564,1072,1094]" gridcol="0" gridrow="18" pageId="6" pageNumber="255">Post DSR</th>
|
||
<td id="4405990A155C9CABFD2AD760FCA8FB92" box="[661,819,1072,1094]" gridcol="1" gridrow="18" pageId="6" pageNumber="255">23</td>
|
||
<td id="4405990A155C9CABFC2CD760FBAAFB92" box="[915,1073,1072,1094]" gridcol="2" gridrow="18" pageId="6" pageNumber="255">23</td>
|
||
<td id="4405990A155C9CABFB1BD760FAACFB92" box="[1188,1335,1072,1094]" gridcol="3" gridrow="18" pageId="6" pageNumber="255">23</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D709FAACFBBB" box="[159,1335,1113,1135]" gridrow="19" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D709FDAFFBBB" box="[159,564,1113,1135]" gridcol="0" gridrow="19" pageId="6" pageNumber="255">Ven</th>
|
||
<td id="4405990A155C9CABFD2AD709FCA8FBBB" box="[661,819,1113,1135]" gridcol="1" gridrow="19" pageId="6" pageNumber="255">169</td>
|
||
<td id="4405990A155C9CABFC2CD709FBAAFBBB" box="[915,1073,1113,1135]" gridcol="2" gridrow="19" pageId="6" pageNumber="255">179</td>
|
||
<td id="4405990A155C9CABFB1BD709FAACFBBB" box="[1188,1335,1113,1135]" gridcol="3" gridrow="19" pageId="6" pageNumber="255">163</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D7D2FAACFB4C" box="[159,1335,1154,1176]" gridrow="20" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D7D2FDAFFB4C" box="[159,564,1154,1176]" gridcol="0" gridrow="20" pageId="6" pageNumber="255">Prec</th>
|
||
<td id="4405990A155C9CABFD2AD7D2FCA8FB4C" box="[661,819,1154,1176]" gridcol="1" gridrow="20" pageId="6" pageNumber="255">single</td>
|
||
<td id="4405990A155C9CABFC2CD7D2FBAAFB4C" box="[915,1073,1154,1176]" gridcol="2" gridrow="20" pageId="6" pageNumber="255">single</td>
|
||
<td id="4405990A155C9CABFB1BD7D2FAACFB4C" box="[1188,1335,1154,1176]" gridcol="3" gridrow="20" pageId="6" pageNumber="255">single</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D7FCFAACFB16" box="[159,1335,1196,1218]" gridrow="21" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D7FCFDAFFB16" box="[159,564,1196,1218]" gridcol="0" gridrow="21" pageId="6" pageNumber="255">Subc</th>
|
||
<td id="4405990A155C9CABFD2AD7FCFCA8FB16" box="[661,819,1196,1218]" gridcol="1" gridrow="21" pageId="6" pageNumber="255">91</td>
|
||
<td id="4405990A155C9CABFC2CD7FCFBAAFB16" box="[915,1073,1196,1218]" gridcol="2" gridrow="21" pageId="6" pageNumber="255">77</td>
|
||
<td id="4405990A155C9CABFB1BD7FCFAACFB16" box="[1188,1335,1196,1218]" gridcol="3" gridrow="21" pageId="6" pageNumber="255">81+</td>
|
||
</tr>
|
||
<tr id="07D4F076155C9CABFF20D785FAACFB3F" box="[159,1335,1237,1259]" gridrow="22" pageId="6" pageNumber="255">
|
||
<th id="4405990A155C9CABFF20D785FDAFFB3F" box="[159,564,1237,1259]" gridcol="0" gridrow="22" pageId="6" pageNumber="255">Dorsal coloration in preservative</th>
|
||
<td id="4405990A155C9CABFD2AD785FCA8FB3F" box="[661,819,1237,1259]" gridcol="1" gridrow="22" pageId="6" pageNumber="255">greyish brown</td>
|
||
<td id="4405990A155C9CABFC2CD785FBAAFB3F" box="[915,1073,1237,1259]" gridcol="2" gridrow="22" pageId="6" pageNumber="255">reddish brown</td>
|
||
<td id="4405990A155C9CABFB1BD785FAACFB3F" box="[1188,1335,1237,1259]" gridcol="3" gridrow="22" pageId="6" pageNumber="255">greyish brown</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="B95BF234155C6352FF78D67CFA8DFABC" blockId="6.[151,1437,1324,2035]" pageId="6" pageNumber="255">
|
||
<taxonomicName id="7EE489B7155C6352FF78D67CFE1CFA90" authority="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le, 2019" authorityName="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le" authorityYear="2019" box="[199,391,1324,1348]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="juliani" status="sp. nov.">
|
||
<emphasis id="8B902E26155C6352FF78D67CFE1CFA90" box="[199,391,1324,1348]" italics="true" pageId="6" pageNumber="255">Achalinus juliani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90A3935D155C6352FE31D67CFE7CFA90" box="[398,487,1324,1348]" pageId="6" pageNumber="255" rank="species">
|
||
<emphasis id="8B902E26155C6352FE31D67CFE7CFA90" bold="true" box="[398,487,1324,1348]" pageId="6" pageNumber="255">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="7EE489B7155C6352FDC4D67CFCADFA90" baseAuthorityName="Zong & Ma" baseAuthorityYear="1983" box="[635,822,1324,1348]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="jinggangensis">
|
||
<emphasis id="8B902E26155C6352FDC4D67CFCADFA90" box="[635,822,1324,1348]" italics="true" pageId="6" pageNumber="255">A. jinggangensis</emphasis>
|
||
</taxonomicName>
|
||
by loreal not being fused with prefrontal, having more ventrals (163–179 versus 156–164), more subcaudals (77–91 versus 51–64), and more maxillary teeth.
|
||
</paragraph>
|
||
<paragraph id="B95BF234155C6352FF78D624FBDEFA01" blockId="6.[151,1437,1324,2035]" pageId="6" pageNumber="255">
|
||
<taxonomicName id="7EE489B7155C6352FF78D624FE11FA58" authority="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le, 2019" authorityName="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le" authorityYear="2019" box="[199,394,1396,1420]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="juliani" status="sp. nov.">
|
||
<emphasis id="8B902E26155C6352FF78D624FE11FA58" box="[199,394,1396,1420]" italics="true" pageId="6" pageNumber="255">Achalinus juliani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90A3935D155C6352FE2BD624FE6BFA58" box="[404,496,1396,1420]" pageId="6" pageNumber="255" rank="species">
|
||
<emphasis id="8B902E26155C6352FE2BD624FE6BFA58" bold="true" box="[404,496,1396,1420]" pageId="6" pageNumber="255">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="7EE489B7155C6352FD33D624FCB1FA58" box="[652,810,1396,1420]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="meiguensis">
|
||
<emphasis id="8B902E26155C6352FD33D624FCB1FA58" box="[652,810,1396,1420]" italics="true" pageId="6" pageNumber="255">A. meiguensis</emphasis>
|
||
</taxonomicName>
|
||
by mental being separated from anterior chin shields, internasal not fused to prefrontal, by different dorsal scale row counts (25–23–23 versus 21–23 – 19–23 – 19), by having more subcaudals (77–91 versus 39–62), and by having more maxillary teeth.
|
||
</paragraph>
|
||
<paragraph id="B95BF234155C6352FF78D6B0FB6BF994" blockId="6.[151,1437,1324,2035]" pageId="6" pageNumber="255">
|
||
<taxonomicName id="7EE489B7155C6352FF78D6B0FE17FA2C" authority="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le, 2019" authorityName="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le" authorityYear="2019" box="[199,396,1504,1528]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="juliani" status="sp. nov.">
|
||
<emphasis id="8B902E26155C6352FF78D6B0FE17FA2C" box="[199,396,1504,1528]" italics="true" pageId="6" pageNumber="255">Achalinus juliani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90A3935D155C6352FE27D6B0FE6DFA2C" box="[408,502,1504,1528]" pageId="6" pageNumber="255" rank="species">
|
||
<emphasis id="8B902E26155C6352FE27D6B0FE6DFA2C" bold="true" box="[408,502,1504,1528]" pageId="6" pageNumber="255">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="7EE489B7155C6352FD26D6B0FD63FA23" authorityName="Bourret" authorityYear="1935" box="[665,760,1504,1527]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="niger">
|
||
<emphasis id="8B902E26155C6352FD26D6B0FD63FA23" box="[665,760,1504,1527]" italics="true" pageId="6" pageNumber="255">A. niger</emphasis>
|
||
</taxonomicName>
|
||
by the internasal suture being distinctly longer than that between the prefrontals (versus almost as long as or shorter), fewer midbody dorsal scale rows (23 versus 25), more subcaudals (77–91 versus 52–72), and keeled body scales (versus smooth on anterior part of body).
|
||
</paragraph>
|
||
<paragraph id="B95BF234155C6352FF78D51CFF7AF979" blockId="6.[151,1437,1324,2035]" pageId="6" pageNumber="255">
|
||
<taxonomicName id="7EE489B7155C6352FF78D51CFE11F9B0" authority="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le, 2019" authorityName="Ziegler & Nguyen & Pham & Nguyen & Pham & Schingen & Nguyen & Le" authorityYear="2019" box="[199,394,1612,1636]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="juliani" status="sp. nov.">
|
||
<emphasis id="8B902E26155C6352FF78D51CFE11F9B0" box="[199,394,1612,1636]" italics="true" pageId="6" pageNumber="255">Achalinus juliani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90A3935D155C6352FE2BD51CFE6AF9B0" box="[404,497,1612,1636]" pageId="6" pageNumber="255" rank="species">
|
||
<emphasis id="8B902E26155C6352FE2BD51CFE6AF9B0" bold="true" box="[404,497,1612,1636]" pageId="6" pageNumber="255">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="7EE489B7155C6352FD30D51CFC93F9B0" authorityName="Van Denburgh" authorityYear="1912" box="[655,776,1612,1636]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="werneri">
|
||
<emphasis id="8B902E26155C6352FD30D51CFC93F9B0" box="[655,776,1612,1636]" italics="true" pageId="6" pageNumber="255">A. werneri</emphasis>
|
||
</taxonomicName>
|
||
by the internasal suture being distinctly longer than that between the prefrontals (versus almost as long as), and by lacking a black mid-dorsal line and a dark subcaudal streak.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F1FEA1BF155C6352FF78D5E8FF78F921" pageId="6" pageNumber="255" type="etymology">
|
||
<paragraph id="B95BF234155C6352FF78D5E8FF78F921" blockId="6.[151,1437,1324,2035]" pageId="6" pageNumber="255">
|
||
<emphasis id="8B902E26155C6352FF78D5E8FED5F904" bold="true" box="[199,334,1720,1744]" pageId="6" pageNumber="255">Etymology.</emphasis>
|
||
We name this species after Julian L. Ziegler. As common name we propose Julian’s Burrowing Snake.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F1FEA1BF155C6352FF78D5AFFDABF827" pageId="6" pageNumber="255" type="distribution">
|
||
<paragraph id="B95BF234155C6352FF78D5AFFE6CF8E8" blockId="6.[151,1437,1324,2035]" pageId="6" pageNumber="255">
|
||
<emphasis id="8B902E26155C6352FF78D5AFFEFBF8CC" bold="true" box="[199,352,1791,1816]" pageId="6" pageNumber="255">Distribution.</emphasis>
|
||
The new species currently is only known from two localities within
|
||
<collectingRegion id="7B203CD6155C6352FB10D450FA0CF8CC" box="[1199,1431,1791,1817]" country="Vietnam" name="Cao Bang" pageId="6" pageNumber="255">Cao Bang Province</collectingRegion>
|
||
, northeastern
|
||
<collectingCountry id="C1F3B2A4155C6352FE97D474FE1CF8E9" box="[296,391,1828,1853]" name="Vietnam" pageId="6" pageNumber="255">Vietnam</collectingCountry>
|
||
(
|
||
<figureCitation id="21DFEEB1155C6352FE29D474FE71F8E8" box="[406,490,1828,1853]" captionStart="FIGURE 12" captionStartId="17.[151,250,1739,1761]" captionTargetBox="[347,1236,332,1718]" captionTargetId="figure@17.[347,1239,332,1721]" captionTargetPageId="17" captionText="FIGURE 12. Map showing the currently known distribution of Achalinus juliani sp. nov. in Cao Bang Province (1 = Ha Lang, type locality, 2 = Nguyen Binh), of Achalinus timi sp. nov. in Son La Province (3 = Copia Nature Reserve), and of Achalinus emilyae sp. nov. in Quang Ninh and Bac Giang provinces (4 = Dong Son-Ky Thuong Nature Reserve, type locality, 5 = Suoi Tuyen)." httpUri="https://zenodo.org/record/2651902/files/figure.png" pageId="6" pageNumber="255">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B95BF234155C6352FF78D418FDABF827" blockId="6.[151,1437,1324,2035]" pageId="6" pageNumber="255">
|
||
<emphasis id="8B902E26155C6352FF78D418FE1DF8B4" bold="true" box="[199,390,1863,1888]" pageId="6" pageNumber="255">Natural history.</emphasis>
|
||
Specimens were found between 20:00 and 22:00 h on forest paths. The surrounding habitat was secondary forest of large, medium and small hardwoods mixed with shrubs and vines. Air temperature at the sites were 25.4–32.6
|
||
<superScript id="4E915F7C155C6352FE39D4DFFE16F849" attach="right" box="[390,397,1935,1949]" fontSize="6" pageId="6" pageNumber="255">o</superScript>
|
||
C and relative humidity 65–80%. The new species was found in different habitat
|
||
<typeStatus id="665F4C96155C6352FAE6D4C3FA03F87F" box="[1369,1432,1939,1963]" pageId="6" pageNumber="255">types</typeStatus>
|
||
: lowland forest in Phia Oac-Phia Den National Park and limestone karst forest in Ha Lang District, at elevations from
|
||
<quantity id="7E1C5FD1155C6352FF6BD48BFEECF826" box="[212,375,2010,2035]" metricMagnitude="2" metricUnit="m" metricValue="9.65" metricValueMax="14.6" metricValueMin="4.7" pageId="6" pageNumber="255" unit="m" value="965.0" valueMax="1460.0" valueMin="470.0">470 to 1460 m</quantity>
|
||
above sea level.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |