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<document id="1CBF3EF2E24CACC05B5B62504AA258D3" ID-CLB-Dataset="21863" ID-DOI="10.5852/ejt.2016.241" ID-GBIF-Dataset="f694842b-3af3-4198-b01e-13d0b5466b5d" ID-ISSN="2118-9773" ID-Zenodo-Dep="3850299" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590443252253" checkinUser="valdenar" docAuthor="Frolov, Andrey V. &amp; Akhmetova, Lilia A." docDate="2016" docId="C4027A0914095554D361F9F0FB32B1C3" docLanguage="en" docName="ejt-2016-241.pdf.imf" docOrigin="European Journal of Taxonomy 241" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Orphnus (Phornus) parastrangulatus Frolov &amp; Akhmetova 2016, sp. nov." docType="treatment" docUuid="FDAC941D-8E3B-4496-A8BB-2A2E58BDBB2B" docUuidSource="ZooBank" docVersion="10" lastPageNumber="15" masterDocId="383B02711405555AD167FFB9FFF7B60D" masterDocTitle="Revision of the subgenus Orphnus (Phornus) (Coleoptera, Scarabaeidae, Orphninae)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="13" updateTime="1698841278433" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5CA8B3DE6FA1C0F109066007F82795E7">Revision of the subgenus Orphnus (Phornus) (Coleoptera, Scarabaeidae, Orphninae)</mods:title>
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<mods:namePart id="86909AC92CC27AECCDD9B25ED8972915">Frolov, Andrey V.</mods:namePart>
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<mods:affiliation id="789F2BA8895205D359D308F899EB0014">Laboratory of Insect Systematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, St. - Petersburg, 199034 Russia. Federal University of Mato Grosso, Av. Fernando Corrêa da Costa, 2367, Boa Esperança, 78060 - 900 Cuiabá, MT, Brazil.</mods:affiliation>
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<mods:namePart id="62A0AB9099A4AA81BFC37D105B24416B">Akhmetova, Lilia A.</mods:namePart>
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<mods:affiliation id="3A16FD45B626E83C6D398526CCA93176">Laboratory of Insect Systematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, St. - Petersburg, 199034 Russia. Federal University of Mato Grosso, Av. Fernando Corrêa da Costa, 2367, Boa Esperança, 78060 - 900 Cuiabá, MT, Brazil.</mods:affiliation>
<mods:nameIdentifier id="A3384C216BB04E473E154BB0DBC3DF25" type="email">akhmetova@scarabaeoidea.com</mods:nameIdentifier>
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<mods:title id="38A5CE0C652189956EC259B7D08785DF">European Journal of Taxonomy</mods:title>
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<mods:date id="F99A707FDF502A76B9D918C7C247A1F6">2016</mods:date>
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<treatment id="C4027A0914095554D361F9F0FB32B1C3" ID-DOI="http://doi.org/10.5281/zenodo.3854786" ID-GBIF-Taxon="164283136" ID-Zenodo-Dep="3854786" LSID="urn:lsid:zoobank.org:act:FDAC941D-8E3B-4496-A8BB-2A2E58BDBB2B" httpUri="http://treatment.plazi.org/id/C4027A0914095554D361F9F0FB32B1C3" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<subSubSection id="04B1989414095556D361F9F0FBD9B069" box="[518,1070,1609,1636]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="4C14CB1F14095556D361F9F0FBD9B069" blockId="12.[350,1237,1609,1706]" box="[518,1070,1609,1636]" pageId="12" pageNumber="13">
<heading id="175C7C7314095556D361F9F0FBD9B069" box="[518,1070,1609,1636]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName id="8BABB09C14095556D361F9F0FC3BB069" authority="Frolov &amp; Akhmetova, 2016" authorityName="Frolov &amp; Akhmetova" authorityYear="2016" box="[518,972,1609,1636]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="parastrangulatus" status="sp. nov." subGenus="Phornus">
<emphasis id="7EDF170D14095556D361F9F0FC3BB069" bold="true" box="[518,972,1609,1636]" pageId="12" pageNumber="13">
<emphasis id="7EDF170D14095556D361F9F0FD85B069" bold="true" box="[518,626,1609,1636]" italics="true" pageId="12" pageNumber="13">Orphnus</emphasis>
(
<emphasis id="7EDF170D14095556D3E4F9F0FD1CB069" bold="true" box="[643,747,1609,1636]" italics="true" pageId="12" pageNumber="13">Phornus</emphasis>
)
<emphasis id="7EDF170D14095556D39AF9F0FC3BB069" bold="true" box="[765,972,1609,1636]" italics="true" pageId="12" pageNumber="13">parastrangulatus</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65ECAA7614095556D2B4F9F3FBD9B069" box="[979,1070,1610,1636]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<uri id="383AC71D14095556D039F9D5FB22B08A" box="[350,1237,1644,1671]" pageId="12" pageNumber="13">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
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<subSubSection id="04B1989414095556D3A7F936FC83B0A7" box="[704,884,1679,1706]" pageId="12" pageNumber="13" type="description">
<paragraph id="4C14CB1F14095556D3A7F936FC83B0A7" blockId="12.[350,1237,1609,1706]" box="[704,884,1679,1706]" pageId="12" pageNumber="13">
<figureCitation id="D490D79A14095556D3A7F936FCE8B0A7" box="[704,799,1679,1706]" captionStart="Fig" captionStartId="4.[189,232,1860,1886]" captionTargetBox="[228,1350,266,1814]" captionTargetId="figure@4.[227,1357,265,1825]" captionTargetPageId="4" captionText="Fig. 1. — AM. Orphnus giganteus Paulian, 1948. AB. Habitus. CD. Head in dorsal and apical view. EG. Antennae. H. Aedeagus in lateral view. I. Parameres in dorsal view. J. Abdomen in dorsal view. KL. Stridulatory field. M. Habitus. — N. O. renaudi sp. nov., holotype, habitus. — O. O. ferrierei sp. nov., holotype, habitus. — P. O. valeriae sp. nov., holotype, habitus. — Q. O. parastrangulatus sp. nov., holotype, habitus. — R. O. strangulatus Paulian, 1948, habitus. MR = not to scale." figureDoi="http://doi.org/10.5281/zenodo.3850303" httpUri="https://zenodo.org/record/3850303/files/figure.png" pageId="12" pageNumber="13">Figs 1Q</figureCitation>
,
<figureCitation id="D490D79A14095556D24BF936FCCEB0A7" box="[812,825,1679,1706]" captionStart="Fig" captionStartId="13.[189,232,1862,1888]" captionTargetBox="[190,1383,262,1815]" captionTargetId="figure@13.[189,1394,261,1821]" captionTargetPageId="13" captionText="Fig. 5. — AH. O. parastrangulatus sp. nov. A. Habitus. BC. Head in dorsal and apical view. D. Aedeagus in lateral view. E. Parameres in dorsal view. F. Label of holotype. G. Abdomen in dorsal view. H. Stridulatory field. — IP. O. strangulatus Paulian, 1948. I. Habitus. JK. Head in dorsal and apical view. L. Aedeagus in lateral view. M. Parameres in dorsal view. N. Label of holotype. O. Abdomen in dorsal view. P. Stridulatory field. AH, I, N. Holotypes." figureDoi="http://doi.org/10.5281/zenodo.3850319" httpUri="https://zenodo.org/record/3850319/files/figure.png" pageId="12" pageNumber="13">5</figureCitation>
AH
</paragraph>
</subSubSection>
<subSubSection id="04B1989414095556D1DAF968FE6AB189" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="4C14CB1F14095556D1DAF968FEC1B0E6" blockId="12.[189,310,1745,1771]" box="[189,310,1745,1771]" pageId="12" pageNumber="13">
<heading id="175C7C7314095556D1DAF968FEC1B0E6" bold="true" box="[189,310,1745,1771]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="7EDF170D14095556D1DAF968FEC1B0E6" bold="true" box="[189,310,1745,1771]" pageId="12" pageNumber="13">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="4C14CB1F14095556D1DAF8B9FE6AB189" blockId="12.[189,1399,1791,1924]" pageId="12" pageNumber="13">
<taxonomicName id="8BABB09C14095556D1DAF8B9FE00B117" authority="Frolov &amp; Akhmetova, 2016" authorityName="Frolov &amp; Akhmetova" authorityYear="2016" box="[189,503,1792,1818]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="parastrangulatus" status="sp. nov.">
<emphasis id="7EDF170D14095556D1DAF8B9FE00B117" box="[189,503,1792,1818]" italics="true" pageId="12" pageNumber="13">Orphnus parastrangulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65ECAA7614095556D366F8B9FDA9B117" box="[513,606,1792,1818]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to
<taxonomicName id="8BABB09C14095556D225F8B9FC0CB117" authorityName="Paulian" authorityYear="1948" box="[834,1019,1792,1818]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="strangulatus">
<emphasis id="7EDF170D14095556D225F8B9FC0CB117" box="[834,1019,1792,1818]" italics="true" pageId="12" pageNumber="13">O. strangulatus</emphasis>
</taxonomicName>
but can be separated from it in having a long transverse keel on frontoclypeus, relatively shorter pronotum and more rounded elytra, shorter and somewhat convex ventrally abdomen (in lateral view contour of abdominal sternites is somewhat curved).
</paragraph>
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<paragraph id="4C14CB1F14085557D1DAF8FFFC9DB1E0" blockId="13.[189,1399,1862,2030]" pageId="13" pageNumber="14">
<emphasis id="7EDF170D14085557D1DAF8FFFE77B16D" bold="true" box="[189,384,1862,1889]" pageId="13" pageNumber="14">
Fig. 5.
<emphasis id="7EDF170D14085557D079F8FEFECFB16D" bold="true" box="[286,312,1863,1888]" italics="true" pageId="13" pageNumber="14"></emphasis>
AH
</emphasis>
.
<taxonomicName id="8BABB09C14085557D0F3F8FEFD7FB16D" authority="Frolov &amp; Akhmetova, 2016" authorityName="Frolov &amp; Akhmetova" authorityYear="2016" box="[404,648,1862,1889]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="parastrangulatus" status="sp. nov.">
<emphasis id="7EDF170D14085557D0F3F8FEFD7FB16D" box="[404,648,1862,1889]" italics="true" pageId="13" pageNumber="14">O. parastrangulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65ECAA7614085557D3F1F8FEFD01B16C" box="[662,758,1863,1889]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7EDF170D14085557D264F8FFFCEFB16D" bold="true" box="[771,792,1862,1888]" pageId="13" pageNumber="14">A</emphasis>
. Habitus.
<emphasis id="7EDF170D14085557D2FAF8FFFC23B16D" bold="true" box="[925,980,1862,1888]" pageId="13" pageNumber="14">BC</emphasis>
. Head in dorsal and apical view.
<emphasis id="7EDF170D14085557D1DAF8D3FF25B189" bold="true" box="[189,210,1898,1924]" pageId="13" pageNumber="14">D</emphasis>
. Aedeagus in lateral view.
<emphasis id="7EDF170D14085557D375F8D3FDD1B189" bold="true" box="[530,550,1898,1924]" pageId="13" pageNumber="14">E</emphasis>
. Parameres in dorsal view.
<emphasis id="7EDF170D14085557D20AF8D3FC88B189" bold="true" box="[877,895,1898,1924]" pageId="13" pageNumber="14">F</emphasis>
. Label of holotype.
<emphasis id="7EDF170D14085557D517F8D0FB70B18E" bold="true" box="[1136,1159,1897,1923]" pageId="13" pageNumber="14">G</emphasis>
. Abdomen in dorsal view.
<emphasis id="7EDF170D14085557D064F834FEEDB1AA" bold="true" box="[259,282,1933,1959]" pageId="13" pageNumber="14">H</emphasis>
. Stridulatory field. —
<emphasis id="7EDF170D14085557D342F834FDA6B1AA" bold="true" box="[549,593,1933,1959]" pageId="13" pageNumber="14">IP</emphasis>
.
<taxonomicName id="8BABB09C14085557D306F834FC36B1AA" authority="Paulian, 1948" authorityName="Paulian" authorityYear="1948" box="[609,961,1933,1959]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="strangulatus">
<emphasis id="7EDF170D14085557D306F834FCEFB1AA" box="[609,792,1933,1959]" italics="true" pageId="13" pageNumber="14">O. strangulatus</emphasis>
<bibRefCitation id="283AB6EE14085557D278F834FC36B1AA" author="Paulian R." box="[799,961,1933,1959]" pageId="13" pageNumber="14" pagination="1 - 75" refId="ref8779" refString="Paulian R. 1948. Revision des Orphnus Africains (Coleoptera, Scarabaeidae). Annales de la Societe entomologique de France 117: 1 - 75." type="journal article" year="1948">Paulian, 1948</bibRefCitation>
</taxonomicName>
.
<emphasis id="7EDF170D14085557D2B7F834FC2CB1AA" bold="true" box="[976,987,1933,1959]" pageId="13" pageNumber="14">I</emphasis>
. Habitus.
<emphasis id="7EDF170D14085557D532F834FB7EB1AA" bold="true" box="[1109,1161,1933,1959]" pageId="13" pageNumber="14">JK</emphasis>
. Head in dorsal and apical view.
<emphasis id="7EDF170D14085557D02BF809FE97B1C7" bold="true" box="[332,352,1968,1994]" pageId="13" pageNumber="14">L</emphasis>
. Aedeagus in lateral view.
<emphasis id="7EDF170D14085557D3F5F809FD59B1C7" bold="true" box="[658,686,1968,1994]" pageId="13" pageNumber="14">M</emphasis>
. Parameres in dorsal view.
<emphasis id="7EDF170D14085557D281F809FC0CB1C7" bold="true" box="[998,1019,1968,1994]" pageId="13" pageNumber="14">N</emphasis>
. Label of holotype.
<emphasis id="7EDF170D14085557D586F809FB0FB1C7" bold="true" box="[1249,1272,1968,1994]" pageId="13" pageNumber="14">O</emphasis>
. Abdomen in dorsal view.
<emphasis id="7EDF170D14085557D017F86AFE75B1E0" bold="true" box="[368,386,2003,2029]" pageId="13" pageNumber="14">P</emphasis>
. Stridulatory field. AH, I, N. Holotypes.
</paragraph>
</caption>
<subSubSection id="04B19894140B5554D1DAFEB2FB93B759" box="[189,1124,267,340]" pageId="14" pageNumber="15" type="etymology">
<paragraph id="4C14CB1F140B5554D1DAFEB2FEB3B728" blockId="14.[189,324,267,293]" box="[189,324,267,293]" pageId="14" pageNumber="15">
<heading id="175C7C73140B5554D1DAFEB2FEB3B728" bold="true" box="[189,324,267,293]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="7EDF170D140B5554D1DAFEB2FEB3B728" bold="true" box="[189,324,267,293]" pageId="14" pageNumber="15">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="4C14CB1F140B5554D1DAFE83FB93B759" blockId="14.[189,1124,312,340]" box="[189,1124,312,340]" pageId="14" pageNumber="15">
From Greek “παρά” and “strangulatus” for close relations with
<taxonomicName id="8BABB09C140B5554D2C1FE83FBAAB75E" authorityName="Paulian" authorityYear="1948" box="[934,1117,313,340]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="strangulatus">
<emphasis id="7EDF170D140B5554D2C1FE83FBAAB75E" box="[934,1117,313,340]" italics="true" pageId="14" pageNumber="15">O. strangulatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="04B19894140B5554D1DAFEC2FCA3B7EA" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="4C14CB1F140B5554D1DAFEC2FE5DB798" blockId="14.[189,426,379,405]" box="[189,426,379,405]" pageId="14" pageNumber="15">
<heading id="175C7C73140B5554D1DAFEC2FE5DB798" bold="true" box="[189,426,379,405]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="7EDF170D140B5554D1DAFEC2FE5DB798" bold="true" box="[189,426,379,405]" pageId="14" pageNumber="15">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="FCC3C142140B5554D18DFE13FCA3B7EA" ID-GBIF-Occurrence="3761958325" collectionCode="ZMHUB" collectorName="D. Ost-Afrika" country="Tanzania" location="D. Ost-Afrika" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
<paragraph id="4C14CB1F140B5554D18DFE13FEABB7C9" blockId="14.[189,852,426,487]" box="[234,348,426,452]" pageId="14" pageNumber="15">
<emphasis id="7EDF170D140B5554D18DFE13FEABB7C9" bold="true" box="[234,348,426,452]" pageId="14" pageNumber="15">
<typeStatus id="931075BD140B5554D18DFE13FEABB7C9" box="[234,348,426,452]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="4C14CB1F140B5554D1DAFE74FCA3B7EA" blockId="14.[189,852,426,487]" box="[189,852,460,487]" pageId="14" pageNumber="15">
<collectingCountry id="34BC8B8F140B5554D1DAFE74FEA0B7EA" box="[189,343,461,487]" name="Tanzania" pageId="14" pageNumber="15">TANZANIA</collectingCountry>
:
<specimenCount id="5AAD0096140B5554D005FE74FE8EB7EA" box="[354,377,461,487]" pageId="14" pageNumber="15" type="male"></specimenCount>
, “
<location id="49749DC4140B5554D0F5FE74FDC1B7EA" LSID="urn:lsid:plazi:treatment:C4027A0914095554D361F9F0FB32B1C3:49749DC4140B5554D0F5FE74FDC1B7EA" box="[402,566,461,487]" country="Tanzania" name="D. Ost-Afrika" pageId="14" pageNumber="15">D. Ost-Afrika</location>
[Tanzania]” (
<collectionCode id="2ABA53DA140B5554D3B0FE74FCBFB7EA" box="[727,840,461,487]" pageId="14" pageNumber="15">ZMHUB</collectionCode>
).
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="04B19894140B5554D1DAFDB7FECEB136" pageId="14" pageNumber="15" type="description">
<paragraph id="4C14CB1F140B5554D1DAFDB7FEB9B425" blockId="14.[189,334,526,552]" box="[189,334,526,552]" pageId="14" pageNumber="15">
<heading id="175C7C73140B5554D1DAFDB7FEB9B425" bold="true" box="[189,334,526,552]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="7EDF170D140B5554D1DAFDB7FEB9B425" bold="true" box="[189,334,526,552]" pageId="14" pageNumber="15">Description</emphasis>
</heading>
</paragraph>
<paragraph id="4C14CB1F140B5554D18DFD84FE26B45A" blockId="14.[189,815,572,635]" box="[234,465,572,599]" pageId="14" pageNumber="15">
<emphasis id="7EDF170D140B5554D18DFD84FEABB45A" bold="true" box="[234,348,573,599]" pageId="14" pageNumber="15">
<typeStatus id="931075BD140B5554D18DFD84FEABB45A" box="[234,348,573,599]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="D490D79A140B5554D00BFD85FE3FB45A" box="[364,456,572,599]" captionStart="Fig" captionStartId="13.[189,232,1862,1888]" captionTargetBox="[190,1383,262,1815]" captionTargetId="figure@13.[189,1394,261,1821]" captionTargetPageId="13" captionText="Fig. 5. — AH. O. parastrangulatus sp. nov. A. Habitus. BC. Head in dorsal and apical view. D. Aedeagus in lateral view. E. Parameres in dorsal view. F. Label of holotype. G. Abdomen in dorsal view. H. Stridulatory field. — IP. O. strangulatus Paulian, 1948. I. Habitus. JK. Head in dorsal and apical view. L. Aedeagus in lateral view. M. Parameres in dorsal view. N. Label of holotype. O. Abdomen in dorsal view. P. Stridulatory field. AH, I, N. Holotypes." figureDoi="http://doi.org/10.5281/zenodo.3850319" httpUri="https://zenodo.org/record/3850319/files/figure.png" pageId="14" pageNumber="15">Fig. 5A</figureCitation>
)
</paragraph>
<paragraph id="4C14CB1F140B5554D1DAFDD9FCD8B477" blockId="14.[189,815,572,635]" box="[189,815,608,635]" pageId="14" pageNumber="15">
Body length
<quantity id="8B5366FA140B5554D032FDD9FE37B477" box="[341,448,608,635]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="14" pageNumber="15" unit="mm" value="12.5">12.5 mm</quantity>
. Colour uniformly dark brown.
</paragraph>
<paragraph id="4C14CB1F140B5554D1DAFD18FBC4B50F" blockId="14.[189,1398,673,770]" pageId="14" pageNumber="15">
Anterior margin of frontoclypeus convex, rounded laterally, with a keel-shaped border (
<figureCitation id="D490D79A140B5554D580FD18FADCB4B6" box="[1255,1323,673,699]" captionStart="Fig" captionStartId="4.[189,232,1860,1886]" captionTargetBox="[228,1350,266,1814]" captionTargetId="figure@4.[227,1357,265,1825]" captionTargetPageId="4" captionText="Fig. 1. — AM. Orphnus giganteus Paulian, 1948. AB. Habitus. CD. Head in dorsal and apical view. EG. Antennae. H. Aedeagus in lateral view. I. Parameres in dorsal view. J. Abdomen in dorsal view. KL. Stridulatory field. M. Habitus. — N. O. renaudi sp. nov., holotype, habitus. — O. O. ferrierei sp. nov., holotype, habitus. — P. O. valeriae sp. nov., holotype, habitus. — Q. O. parastrangulatus sp. nov., holotype, habitus. — R. O. strangulatus Paulian, 1948, habitus. MR = not to scale." figureDoi="http://doi.org/10.5281/zenodo.3850303" httpUri="https://zenodo.org/record/3850303/files/figure.png" pageId="14" pageNumber="15">Fig. 1</figureCitation>
BC). Frontoclypeus with low transverse carina approximately on the line connecting anterior margins of canthi. Frontoclypeus densely punctate anteriad and posteriad of the carina.
</paragraph>
<paragraph id="4C14CB1F140B5554D1DAFC93FDECB56A" blockId="14.[189,1398,809,871]" pageId="14" pageNumber="15">Eyes relatively large: width about 1/5.5 the distance between eyes in dorsal view.Antennae 10-segmented, without malformed segments.</paragraph>
<paragraph id="4C14CB1F140B5554D1DAFC37FB56B239" blockId="14.[189,1398,910,1077]" pageId="14" pageNumber="15">
Pronotum widely rounded laterally, narrower than elytra, 1.8 times wider than length, 0.5 length of elytra (
<figureCitation id="D490D79A140B5554D073FC08FE81B5C6" box="[276,374,945,971]" captionStart="Fig" captionStartId="4.[189,232,1860,1886]" captionTargetBox="[228,1350,266,1814]" captionTargetId="figure@4.[227,1357,265,1825]" captionTargetPageId="4" captionText="Fig. 1. — AM. Orphnus giganteus Paulian, 1948. AB. Habitus. CD. Head in dorsal and apical view. EG. Antennae. H. Aedeagus in lateral view. I. Parameres in dorsal view. J. Abdomen in dorsal view. KL. Stridulatory field. M. Habitus. — N. O. renaudi sp. nov., holotype, habitus. — O. O. ferrierei sp. nov., holotype, habitus. — P. O. valeriae sp. nov., holotype, habitus. — Q. O. parastrangulatus sp. nov., holotype, habitus. — R. O. strangulatus Paulian, 1948, habitus. MR = not to scale." figureDoi="http://doi.org/10.5281/zenodo.3850303" httpUri="https://zenodo.org/record/3850303/files/figure.png" pageId="14" pageNumber="15">Fig. 1Q</figureCitation>
). Anterior border wide, with almost smooth posterior margin. Basal border narrow, keel-shaped, separated from pronotal disc by a smooth groove. Pronotal disc anteromedially slightly flattened. Surface of pronotum covered with double puncturation composed of large rounded and minute punctures; large punctures cover sides of the pronotum and area near anterior border.
</paragraph>
<paragraph id="4C14CB1F140B5554D1DAFBE5FC21B27B" blockId="14.[189,982,1116,1142]" box="[189,982,1116,1142]" pageId="14" pageNumber="15">Scutellum narrowly rounded apically, about 1/16 the length of elytra.</paragraph>
<paragraph id="4C14CB1F140B5554D1DAFB24FEAAB32C" blockId="14.[189,1399,1181,1313]" pageId="14" pageNumber="15">Elytra about as wide as long, with feeble marked humeral humps. Elytra widest approximately in the middle, with widely rounded lateral margins. First six striae faintly visible as very shallow grooves, without distinct rows of punctures. Elytra covered with sparse punctures becoming slightly coarser towards base.</paragraph>
<paragraph id="4C14CB1F140B5554D1DAFAF1FC10B36F" blockId="14.[189,999,1351,1379]" box="[189,999,1351,1379]" pageId="14" pageNumber="15">Brachypterous. Wings vestigial, narrow, about ¾ the length of elytra.</paragraph>
<paragraph id="4C14CB1F140B5554D1DAFA33FB6AB3A8" blockId="14.[189,1181,1416,1445]" box="[189,1181,1416,1445]" pageId="14" pageNumber="15">
Stridulatory field: carinae separated by 1/30 length of the field in central ¼
<superScript id="BBDE6657140B5554D571FA31FBD3B39B" attach="left" box="[1046,1060,1416,1430]" fontSize="6" pageId="14" pageNumber="15">rd</superScript>
(
<figureCitation id="D490D79A140B5554D556FA33FB67B3A8" box="[1073,1168,1418,1445]" captionStart="Fig" captionStartId="13.[189,232,1862,1888]" captionTargetBox="[190,1383,262,1815]" captionTargetId="figure@13.[189,1394,261,1821]" captionTargetPageId="13" captionText="Fig. 5. — AH. O. parastrangulatus sp. nov. A. Habitus. BC. Head in dorsal and apical view. D. Aedeagus in lateral view. E. Parameres in dorsal view. F. Label of holotype. G. Abdomen in dorsal view. H. Stridulatory field. — IP. O. strangulatus Paulian, 1948. I. Habitus. JK. Head in dorsal and apical view. L. Aedeagus in lateral view. M. Parameres in dorsal view. N. Label of holotype. O. Abdomen in dorsal view. P. Stridulatory field. AH, I, N. Holotypes." figureDoi="http://doi.org/10.5281/zenodo.3850319" httpUri="https://zenodo.org/record/3850319/files/figure.png" pageId="14" pageNumber="15">Fig. 5H</figureCitation>
).
</paragraph>
<paragraph id="4C14CB1F140B5554D1DAFA72FDE1B07E" blockId="14.[189,1399,1483,1651]" pageId="14" pageNumber="15">
Abdomen 1.4 times wider than long (measured in ventral view from anterolateral margins of sternite 2 to apex of pygidium). Abdominal sternite 8 medially about as long as sternites 57 combined; sternite 7 very narrow medially. Pygidium visible from above, with rounded apex. Plectrum trapezoidal, elongated, with widely rounded apex (
<figureCitation id="D490D79A140B5554D361F98CFD91B05D" box="[518,614,1589,1616]" captionStart="Fig" captionStartId="13.[189,232,1862,1888]" captionTargetBox="[190,1383,262,1815]" captionTargetId="figure@13.[189,1394,261,1821]" captionTargetPageId="13" captionText="Fig. 5. — AH. O. parastrangulatus sp. nov. A. Habitus. BC. Head in dorsal and apical view. D. Aedeagus in lateral view. E. Parameres in dorsal view. F. Label of holotype. G. Abdomen in dorsal view. H. Stridulatory field. — IP. O. strangulatus Paulian, 1948. I. Habitus. JK. Head in dorsal and apical view. L. Aedeagus in lateral view. M. Parameres in dorsal view. N. Label of holotype. O. Abdomen in dorsal view. P. Stridulatory field. AH, I, N. Holotypes." figureDoi="http://doi.org/10.5281/zenodo.3850319" httpUri="https://zenodo.org/record/3850319/files/figure.png" pageId="14" pageNumber="15">Fig. 5G</figureCitation>
). Lateral plate of abdominal sternite relatively small, shorter than plectrum, with rounded apex.
</paragraph>
<paragraph id="4C14CB1F140B5554D1A5F920FC95B0DA" blockId="14.[189,1399,1689,1751]" pageId="14" pageNumber="15">
Aedeagus with relatively long parameres (0.5 length of phallobase), without excavations. Apices of the parameres rounded in dorsal and lateral view (
<figureCitation id="D490D79A140B5554D3BBF904FCEEB0DA" box="[732,793,1724,1751]" captionStart="Fig" captionStartId="13.[189,232,1862,1888]" captionTargetBox="[190,1383,262,1815]" captionTargetId="figure@13.[189,1394,261,1821]" captionTargetPageId="13" captionText="Fig. 5. — AH. O. parastrangulatus sp. nov. A. Habitus. BC. Head in dorsal and apical view. D. Aedeagus in lateral view. E. Parameres in dorsal view. F. Label of holotype. G. Abdomen in dorsal view. H. Stridulatory field. — IP. O. strangulatus Paulian, 1948. I. Habitus. JK. Head in dorsal and apical view. L. Aedeagus in lateral view. M. Parameres in dorsal view. N. Label of holotype. O. Abdomen in dorsal view. P. Stridulatory field. AH, I, N. Holotypes." figureDoi="http://doi.org/10.5281/zenodo.3850319" httpUri="https://zenodo.org/record/3850319/files/figure.png" pageId="14" pageNumber="15">Fig 5</figureCitation>
DE).
</paragraph>
<paragraph id="4C14CB1F140B5554D18DF947FEB2B115" blockId="14.[189,325,1790,1851]" box="[234,325,1790,1816]" pageId="14" pageNumber="15">
<emphasis id="7EDF170D140B5554D18DF947FEB2B115" bold="true" box="[234,325,1790,1816]" pageId="14" pageNumber="15">Female</emphasis>
</paragraph>
<paragraph id="4C14CB1F140B5554D1DAF898FECEB136" blockId="14.[189,325,1790,1851]" box="[189,313,1825,1851]" pageId="14" pageNumber="15">Unknown.</paragraph>
</subSubSection>
<subSubSection id="04B19894140B5554D1DAF8DAFB32B1C3" pageId="14" pageNumber="15" type="distribution">
<paragraph id="4C14CB1F140B5554D1DAF8DAFEA1B170" blockId="14.[189,342,1891,1917]" box="[189,342,1891,1917]" pageId="14" pageNumber="15">
<heading id="175C7C73140B5554D1DAF8DAFEA1B170" bold="true" box="[189,342,1891,1917]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="7EDF170D140B5554D1DAF8DAFEA1B170" bold="true" box="[189,342,1891,1917]" pageId="14" pageNumber="15">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="4C14CB1F140B5554D1DAF828FB32B1C3" blockId="14.[189,1399,1937,1999]" pageId="14" pageNumber="15">
The only
<typeStatus id="931075BD140B5554D054F82BFE91B1A1" box="[307,358,1938,1964]" pageId="14" pageNumber="15">type</typeStatus>
specimen lacks precise information of the collecting locality except for a reference to “Deutsch Ostafrika”, the area corresponding to modern
<collectingCountry id="34BC8B8F140B5554D22CF80DFC4FB1C3" box="[843,952,1972,1998]" name="Tanzania" pageId="14" pageNumber="15">Tanzania</collectingCountry>
,
<collectingCountry id="34BC8B8F140B5554D2A4F80DFBD4B1C3" box="[963,1059,1972,1998]" name="Burundi" pageId="14" pageNumber="15">Burundi</collectingCountry>
and
<collectingCountry id="34BC8B8F140B5554D53AF80DFB36B1C3" box="[1117,1217,1972,1998]" name="Rwanda" pageId="14" pageNumber="15">Rwanda</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>