treatments-xml/data/4A/35/15/4A3515175B4E533AB88D40962DA2773C.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1029.63940" ID-GBIF-Dataset="27e8615f-ba3f-4d59-99a1-ecb8493d10f2" ID-PMC="PMC8050014" ID-Pensoft-Pub="1313-2970-1029-155" ID-Pensoft-UUID="50846BA2C2A85039BEFA6C1E2E3526FB" ID-PubMed="33935554" ID-ZooBank="9AE12DC8CDA44BA481C2B655C649AEC5" ModsDocID="1313-2970-1029-155" checkinTime="1617901766197" checkinUser="pensoft" docAuthor="Hava, Jiri, Abdel-Dayem, Mahmoud S. &amp; Aldhafer, Hathal M." docDate="2021" docId="4A3515175B4E533AB88D40962DA2773C" docLanguage="en" docName="ZooKeys 1029: 155-174" docOrigin="ZooKeys 1029" docPubDate="2021-04-08" docSource="http://dx.doi.org/10.3897/zookeys.1029.63940" docTitle="Thorictus shadensis Hava &amp; Abdel-Dayem 2021, sp. nov." docType="treatment" docUuid="B2CC6ACC-DA67-49F7-A14D-4BA09F608F0B" docUuidSource="ZooBank" docVersion="4" id="50846BA2C2A85039BEFA6C1E2E3526FB" lastPageNumber="155" masterDocId="50846BA2C2A85039BEFA6C1E2E3526FB" masterDocTitle="The subfamily Thorictinae (Coleoptera, Dermestidae) from Saudi Arabia" masterLastPageNumber="174" masterPageNumber="155" pageNumber="155" updateTime="1668152078424" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The subfamily Thorictinae (Coleoptera, Dermestidae) from Saudi Arabia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hava, Jiri</mods:namePart>
<mods:affiliation>Forestry and Game Management Research Institute, Strnady 136, CZ- 252 02 Praha 5 - Zbraslav, Czech Republic</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Abdel-Dayem, Mahmoud S.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6276-1740</mods:nameIdentifier>
<mods:affiliation>King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agricultral Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia</mods:affiliation>
<mods:nameIdentifier type="email">mseleem@ksu.edu.sa</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Aldhafer, Hathal M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4911-2332</mods:nameIdentifier>
<mods:affiliation>King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agricultral Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia</mods:affiliation>
</mods:name>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-08</mods:number>
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<mods:detail type="volume">
<mods:number>1029</mods:number>
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<mods:start>155</mods:start>
<mods:end>174</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1029.63940</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1029.63940</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1029-155</mods:identifier>
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<treatment ID-GBIF-Taxon="180881646" LSID="urn:lsid:zoobank.org:act:B2CC6ACC-DA67-49F7-A14D-4BA09F608F0B" httpUri="http://treatment.plazi.org/id/4A3515175B4E533AB88D40962DA2773C" lastPageNumber="155" pageId="0" pageNumber="155">
<subSubSection pageId="0" pageNumber="155" type="nomenclature">
<paragraph pageId="0" pageNumber="155">
<taxonomicName LSID="http://zoobank.org/B2CC6ACC-DA67-49F7-A14D-4BA09F608F0B" authority="Háva &amp; Abdel-Dayem" authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus shadensis" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="shadensis" status="sp. nov.">
Thorictus shadensis 
<normalizedToken originalValue="Háva">Hava</normalizedToken>
&amp; Abdel-Dayem
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="155">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus dorsal aspect of Thorictus shadensis Hava &amp; Abdel-Dayem, sp. nov." figureDoi="10.3897/zookeys.1029.63940.figure11" httpUri="https://binary.pensoft.net/fig/529728" pageId="0" pageNumber="155">Figs 11</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Thorictus shadensis Hava &amp; Abdel-Dayem, sp. nov. a head lateral eye b ventral setation on pronotum and metepisternum, T abyssinicus John, 1963 c head lateral eye d ventral setation on pronotum and metepisternum, T dilatipennis Reitter, 1881 e head lateral eye (according to John 1963) f ventral setation on pronotum and metepisternum (according to John 1963)." figureDoi="10.3897/zookeys.1029.63940.figure12" httpUri="https://binary.pensoft.net/fig/529729" pageId="0" pageNumber="155">, 12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="materials_examined">
<paragraph pageId="0" pageNumber="155">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="155">
<materialsCitation ID-GBIF-Occurrence="3069288306" collectingDate="2014-07-05" collectingMethod="pitfall trap" collectorName="Al Dhafer H, Fadl H, Abadel-Dayem M, El Torkey A, El Gharbawy A" country="Saudi Arabia" elevation="1474" latitude="19.845167" location="Shada Al Ala Nature Reserve" longLatPrecision="1" longitude="41.30445" specimenCount="1" stateProvince="Al Bahah" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="155">
<typeStatus>Holotype</typeStatus>
</emphasis>
. 
<collectingCountry name="Saudi Arabia">Saudi Arabia</collectingCountry>
• 
<specimenCount type="generic">1 ex</specimenCount>
; 
<collectingRegion country="Saudi Arabia" name="Al Bahah">Al Bahah</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:4A3515175B4E533AB88D40962DA2773C:174968B91192EA30A5533D01D6810732" country="Saudi Arabia" latitude="19.845167" longLatPrecision="1" longitude="41.30445" name="Shada Al Ala Nature Reserve" stateProvince="Al Bahah">Shada Al Ala Nature Reserve</location>
; 
<geoCoordinate degrees="19" direction="north" minutes="50.710" orientation="latitude" precision="1" value="19.845167">19°50.710'N</geoCoordinate>
, 
<geoCoordinate degrees="41" direction="east" minutes="18.267" orientation="longitude" precision="1" value="41.30445">41°18.267'E</geoCoordinate>
; alt. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.474" unit="m" value="1474.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.474" unit="m" value="1474.0">1474 m</elevation>
</quantity>
; 
<collectingDate value="2014-07-05">5 Jul. 2014</collectingDate>
; 
<collectorName>Al Dhafer H</collectorName>
, 
<collectorName>Fadl H</collectorName>
, 
<collectorName>Abadel-Dayem M</collectorName>
, 
<collectorName>El Torkey A</collectorName>
, 
<collectorName>El Gharbawy A</collectorName>
leg.; 
<collectingMethod>pitfall trap</collectingMethod>
; KSMA
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3069288305" collectingDate="2014-04-23" collectingMethod="pitfall trap" collectorName="Al Dhafer H, Fadl H, Abad Eidayem M, El Torkey A, El Gharbawy A" country="Saudi Arabia" elevation="1563" latitude="19.8627" location="Shada Al Ala Nature Reserve" longLatPrecision="1" longitude="41.301483" specimenCount="2" stateProvince="Al Bahah" typeStatus="Paratypes">
<emphasis bold="true" italics="true" pageId="0" pageNumber="155">
<typeStatus>Paratypes</typeStatus>
</emphasis>
. 
<collectingCountry name="Saudi Arabia">Saudi Arabia</collectingCountry>
• 
<specimenCount type="generic">1 ex</specimenCount>
; 
<collectingRegion country="Saudi Arabia" name="Al Bahah">Al Bahah</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:4A3515175B4E533AB88D40962DA2773C:CC5D18D7A5699BCDA6FB2F2C988AB0C7" country="Saudi Arabia" latitude="19.8627" longLatPrecision="1" longitude="41.301483" name="Shada Al Ala Nature Reserve" stateProvince="Al Bahah">Shada Al Ala Nature Reserve</location>
; 
<geoCoordinate degrees="19" direction="north" minutes="50.329" orientation="latitude" precision="1" value="19.838818">19°50.329'N</geoCoordinate>
, 
<geoCoordinate degrees="41" direction="east" minutes="18.604" orientation="longitude" precision="1" value="41.310066">41°18.604'E</geoCoordinate>
; alt. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.563" unit="m" value="1563.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.563" unit="m" value="1563.0">1563 m</elevation>
</quantity>
; 
<collectingDate value="2014-04-23">23 Apr. 2014</collectingDate>
; 
<collectorName>Al Dhafer H</collectorName>
, 
<collectorName>Fadl H</collectorName>
, 
<collectorName>Abad Eidayem M</collectorName>
, 
<collectorName>El Torkey A</collectorName>
, 
<collectorName>El Gharbawy A</collectorName>
leg.; 
<collectingMethod>pitfall trap</collectingMethod>
; KSMA • 
<specimenCount type="generic">1 ex</specimenCount>
; same collection data as preceding; 
<geoCoordinate degrees="19" direction="north" minutes="51.762" orientation="latitude" precision="1" value="19.8627">19°51.762'N</geoCoordinate>
, 
<geoCoordinate degrees="41" direction="east" minutes="18.089" orientation="longitude" precision="1" value="41.301483">41°18.089'E</geoCoordinate>
; alt. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.225" unit="m" value="1225.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.225" unit="m" value="1225.0">1225 m</elevation>
</quantity>
; 
<collectingDate value="2014-04-23">23 Apr. 2014</collectingDate>
; 
<collectorName>Al Dhafer H</collectorName>
, 
<collectorName>Fadl H</collectorName>
, 
<collectorName>Abad Eidayem M</collectorName>
, 
<collectorName>El Torkey A</collectorName>
, 
<collectorName>El Gharbawy A</collectorName>
leg.; 
<collectingMethod>pitfall trap</collectingMethod>
; JHAC
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1029.63940.figure11" httpUri="https://binary.pensoft.net/fig/529728" pageId="0" pageNumber="155" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="155">
<emphasis bold="true" pageId="0" pageNumber="155">Figure 11.</emphasis>
Habitus dorsal aspect of 
<taxonomicName authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus shadensis" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="shadensis">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus shadensis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Háva">Hava</normalizedToken>
&amp; Abdel-Dayem, sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="description">
<paragraph pageId="0" pageNumber="155">Description of holotype.</paragraph>
<paragraph pageId="0" pageNumber="155">Body small, brown covered by short yellow setae on dorsal surfaces and short setae on ventral surfaces. Measurements (mm): TL 2.1, PW 1.1, EW 1.2. Head finely punctate with short yellow setae. Labial palpi entirely brown. Antennae brown, with 11 antennomeres, antennal club compact with three antennomeres. Pronotum as finely punctate as head, covered by short yellow setation. Lateral margin of pronotum not dentate. Pronotum in posterior part near scutellum without bumps. Ventral posterior pronotal angles without long yellow setation. Scutellum not visible from above. Elytra very finely punctate covered by short yellow setation. Each elytron in anterior part near humeri with one very small bump. Epipleuron finely punctate, anterior angles with short yellow setation and small bump. Prosternum finely punctate. Mesosternum finely punctate, mesosternal bulge. Metasternum finely punctate. Visible abdominal ventrites very finely punctate, covered by short yellow setae. First abdominal ventrite with anterior, longitudinal striation. Legs brown, covered by yellow setae.</paragraph>
<caption doi="10.3897/zookeys.1029.63940.figure12" httpUri="https://binary.pensoft.net/fig/529729" pageId="0" pageNumber="155" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="155">
<emphasis bold="true" pageId="0" pageNumber="155">Figure 12.</emphasis>
<taxonomicName authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus shadensis" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="shadensis">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus shadensis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Háva">Hava</normalizedToken>
&amp; Abdel-Dayem, sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="155">a</emphasis>
head lateral eye 
<emphasis bold="true" pageId="0" pageNumber="155">b</emphasis>
ventral setation on pronotum and metepisternum, 
<taxonomicName class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T abyssinicus" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="abyssinicus">
<emphasis italics="true" pageId="0" pageNumber="155">T abyssinicus</emphasis>
</taxonomicName>
John, 1963 
<emphasis bold="true" pageId="0" pageNumber="155">c</emphasis>
head lateral eye 
<emphasis bold="true" pageId="0" pageNumber="155">d</emphasis>
ventral setation on pronotum and metepisternum, 
<taxonomicName class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T dilatipennis" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="dilatipennis">
<emphasis italics="true" pageId="0" pageNumber="155">T dilatipennis</emphasis>
</taxonomicName>
Reitter, 1881 
<emphasis bold="true" pageId="0" pageNumber="155">e</emphasis>
head lateral eye (according to 
<bibRefCitation author="John, H" journalOrPublisher="Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden" pageId="0" pageNumber="155" pagination="177 - 255" refId="B16" refString="John, H, 1963. Revision der Gattung Thorictus Germar (Thorictidae, Clavicornia Col.). Eine Bestandaufnahme, Kritik und Neubeschreibungen. Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden 28: 177 - 255" title="Revision der Gattung Thorictus Germar (Thorictidae, Clavicornia Col.). Eine Bestandaufnahme, Kritik und Neubeschreibungen." volume="28" year="1963">John 1963</bibRefCitation>
) 
<emphasis bold="true" pageId="0" pageNumber="155">f</emphasis>
ventral setation on pronotum and metepisternum (according to 
<bibRefCitation author="John, H" journalOrPublisher="Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden" pageId="0" pageNumber="155" pagination="177 - 255" refId="B16" refString="John, H, 1963. Revision der Gattung Thorictus Germar (Thorictidae, Clavicornia Col.). Eine Bestandaufnahme, Kritik und Neubeschreibungen. Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden 28: 177 - 255" title="Revision der Gattung Thorictus Germar (Thorictidae, Clavicornia Col.). Eine Bestandaufnahme, Kritik und Neubeschreibungen." volume="28" year="1963">John 1963</bibRefCitation>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="variability">
<paragraph pageId="0" pageNumber="155">Variability.</paragraph>
<paragraph pageId="0" pageNumber="155">Body measurements (mm): TL 2.1-2.2, PW 1.0-1.1, EW 1.1-1.2.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="differential diagnosis">
<paragraph pageId="0" pageNumber="155">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="155">
The new species belongs to the 
<taxonomicName class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus orientalis" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus orientalis</emphasis>
</taxonomicName>
species group and is very similar to two other species: 
<taxonomicName class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T abyssinicus" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="abyssinicus">
<emphasis italics="true" pageId="0" pageNumber="155">T abyssinicus</emphasis>
</taxonomicName>
John, 1963 (Ethiopia) and 
<taxonomicName class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T dilatipennis" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="dilatipennis">
<emphasis italics="true" pageId="0" pageNumber="155">T dilatipennis</emphasis>
</taxonomicName>
Reitter, 1881 (Egypt, Syria), but differs from them by the abovementioned characteristics.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="etymology">
<paragraph pageId="0" pageNumber="155">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="155">Named according to type locality: Shada Al Ala Nature Reserve.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="ecological notes">
<paragraph pageId="0" pageNumber="155">Ecological notes.</paragraph>
<paragraph pageId="0" pageNumber="155">
The adult beetles were found during April and July at elevations of 1225-1563 m in the Shada Al Ala Nature Reserve. The specimens were collected by pitfall traps in steep slopes covered with vegetation dominated by 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia" order="Fabales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="155">Acacia</emphasis>
</taxonomicName>
thorn woodlands and shrubs of Barbary fig or cactus pear, 
<taxonomicName class="Dicotyledoneae" family="Cactaceae" genus="Opuntia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Opuntia ficus-indica" order="Cactales" pageId="0" pageNumber="155" phylum="Angiospermae" rank="species" species="ficus-indica">
<emphasis italics="true" pageId="0" pageNumber="155">Opuntia ficus-indica</emphasis>
</taxonomicName>
(L.) Mill. (
<taxonomicName class="Dicotyledoneae" family="Cactaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Cactales" pageId="0" pageNumber="155" phylum="Angiospermae" rank="family">Cactaceae</taxonomicName>
) (Figs 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Acacia thorn woodlands, the type locality of Thorictus shadensis Hava &amp; Abdel-Dayem, sp. nov. holotype at Shada Al Ala Nature Reserve on the Shada Mountain, Baha Province, southwestern Saudi Arabia, at an elevation of 695 m." figureDoi="10.3897/zookeys.1029.63940.figure13" httpUri="https://binary.pensoft.net/fig/529730" pageId="0" pageNumber="155">13</figureCitation>
, 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Barbary fig or cactus pear shrubs, Opuntia ficus-indica (L.) Mill., the type locality of paratype of Thorictus shadensis Hava &amp; Abdel-Dayem, sp. nov. at Shada Al Ala Nature Reserve on the Shada Mountain, Baha Province, southwestern Saudi Arabia, at 1563 m elevation." figureDoi="10.3897/zookeys.1029.63940.figure14" httpUri="https://binary.pensoft.net/fig/529731" pageId="0" pageNumber="155">14</figureCitation>
). This new species was caught with ant species of 
<taxonomicName authorityName="Emery" authorityYear="1915" class="Insecta" family="Staphylinidae" genus="Camponotus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Camponotus aegyptiacus" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="aegyptiacus">
<emphasis italics="true" pageId="0" pageNumber="155">Camponotus aegyptiacus</emphasis>
</taxonomicName>
(Emery, 1915), 
<taxonomicName authorityName="Lund" authorityYear="1831" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster" order="Hymenoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="155">Crematogaster</emphasis>
</taxonomicName>
sp., 
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1901" class="Magnoliopsida" family="Asteraceae" genus="Lepisiota" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Lepisiota obtusa" order="Asterales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="species" species="obtusa">
<emphasis italics="true" pageId="0" pageNumber="155">Lepisiota obtusa</emphasis>
</taxonomicName>
(Emery, 1901), 
<taxonomicName authorityName="Collingwood &amp; Agosti" authorityYear="1996" class="Insecta" family="Formicidae" genus="Monomorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monomorium jizane" order="Hymenoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="jizane">
<emphasis italics="true" pageId="0" pageNumber="155">Monomorium jizane</emphasis>
</taxonomicName>
Collingwood &amp; Agosti, 1996, 
<taxonomicName authorityName="Bolton" authorityYear="1987" class="Insecta" family="Formicidae" genus="Monomorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monomorium rabirium" order="Hymenoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="rabirium">
<emphasis italics="true" pageId="0" pageNumber="155">Monomorium rabirium</emphasis>
</taxonomicName>
Bolton, 1987, 
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole" order="Hymenoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="155">Pheidole</emphasis>
</taxonomicName>
sp., and 
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium simillium" order="Hymenoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="simillium">
<emphasis italics="true" pageId="0" pageNumber="155">Tetramorium simillium</emphasis>
</taxonomicName>
(Smith, 1851) in the same pitfall traps.
</paragraph>
<caption doi="10.3897/zookeys.1029.63940.figure13" httpUri="https://binary.pensoft.net/fig/529730" pageId="0" pageNumber="155" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="155">
<emphasis bold="true" pageId="0" pageNumber="155">Figure 13.</emphasis>
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia" order="Fabales" pageId="0" pageNumber="155" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="155">Acacia</emphasis>
</taxonomicName>
thorn woodlands, the type locality of 
<taxonomicName authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus shadensis" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="shadensis">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus shadensis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Háva">Hava</normalizedToken>
&amp; Abdel-Dayem, sp. nov. holotype at Shada Al Ala Nature Reserve on the Shada Mountain, Baha Province, southwestern Saudi Arabia, at an elevation of 695 m.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1029.63940.figure14" httpUri="https://binary.pensoft.net/fig/529731" pageId="0" pageNumber="155" start="Figure 14" startId="F14">
<paragraph pageId="0" pageNumber="155">
<emphasis bold="true" pageId="0" pageNumber="155">Figure 14.</emphasis>
Barbary fig or cactus pear shrubs, 
<taxonomicName class="Dicotyledoneae" family="Cactaceae" genus="Opuntia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Opuntia ficus-indica" order="Cactales" pageId="0" pageNumber="155" phylum="Angiospermae" rank="species" species="ficus-indica">
<emphasis italics="true" pageId="0" pageNumber="155">Opuntia ficus-indica</emphasis>
</taxonomicName>
(L.) Mill., the type locality of paratype of 
<taxonomicName authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus shadensis" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="shadensis">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus shadensis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Háva">Hava</normalizedToken>
&amp; Abdel-Dayem, sp. nov. at Shada Al Ala Nature Reserve on the Shada Mountain, Baha Province, southwestern Saudi Arabia, at 1563 m elevation.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="distribution">
<paragraph pageId="0" pageNumber="155">Geographical distribution.</paragraph>
<paragraph pageId="0" pageNumber="155">
This species is only known from the type locality in the Shada Al Ala Nature Reserve, on the Shada Mountain, in the west of the Sarawat Mountains at Al Mekhwah, Baha Province (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of Saudi Arabian provinces and distribution of Thorictinae species." figureDoi="10.3897/zookeys.1029.63940.figure1" httpUri="https://binary.pensoft.net/fig/529719" pageId="0" pageNumber="155">1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>