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<document id="B1FDB0CF6648D971D91A2F4C4585E873" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625105362570" checkinUser="felipe" docAuthor="Arce-Pérez, Roberto &amp; Morón, Miguel-Angel" docDate="2005" docId="03C487D0C45EFFBC174FFBD6259AA404" docLanguage="en" docName="zt01012p037.pdf" docOrigin="Zootaxa 1012 (1012)" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Macrodactylus sericeicollis Bates 1887" docType="treatment" docVersion="2" lastPageNumber="27" masterDocId="FFFDFFA8C45DFFB81647FFEA2418A64A" masterDocTitle="New species and new records of species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactyini) from Honduras and Nicaragua" masterLastPageNumber="37" masterPageNumber="23" pageNumber="26" updateTime="1698980141271" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9370AE7A05B90BB608A72E388F7BAB64">New species and new records of species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactyini) from Honduras and Nicaragua</mods:title>
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<mods:name id="5F0F6F42CDB5AEB3E64F5E0420452D07" type="personal">
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<mods:namePart id="9987E06532549D64C18A0A7D65C3D6C6">Arce-Pérez, Roberto</mods:namePart>
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<mods:namePart id="6869C431534DFB438A30614D25F29D4B">Morón, Miguel-Angel</mods:namePart>
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<mods:title id="6AE9A951C466F5BF7A5665E50E998A7E">Zootaxa</mods:title>
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<mods:date id="7B52AE73975603CBFC487B0A849561AB">2005</mods:date>
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<mods:number id="51E4C5D06A2E7FB233CB466D402E854E">2005-06-28</mods:number>
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<treatment id="03C487D0C45EFFBC174FFBD6259AA404" LSID="urn:lsid:plazi:treatment:03C487D0C45EFFBC174FFBD6259AA404" httpUri="http://treatment.plazi.org/id/03C487D0C45EFFBC174FFBD6259AA404" lastPageId="4" lastPageNumber="27" pageId="3" pageNumber="26">
<subSubSection id="C377654DC45EFFBB174FFBD626FDA21C" box="[264,741,1084,1110]" pageId="3" pageNumber="26" type="nomenclature">
<paragraph id="8BD236C6C45EFFBB174FFBD626FDA21C" blockId="3.[264,741,1084,1110]" box="[264,741,1084,1110]" pageId="3" pageNumber="26">
<heading id="D09A81AAC45EFFBB174FFBD626FDA21C" bold="true" box="[264,741,1084,1110]" fontSize="11" level="1" pageId="3" pageNumber="26" reason="1">
<taxonomicName id="4C6D4D45C45EFFBB174FFBD626FDA21C" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[264,741,1084,1110]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="sericeicollis">
<emphasis id="B919EAD4C45EFFBB174FFBD626FDA21C" bold="true" box="[264,741,1084,1110]" pageId="3" pageNumber="26">
<emphasis id="B919EAD4C45EFFBB174FFBD62648A21C" bold="true" box="[264,592,1084,1110]" italics="true" pageId="3" pageNumber="26">Macrodactylus sericeicollis</emphasis>
<bibRefCitation id="EFFC4B37C45EFFBB1410FBD626FDA21C" author="Bates, H. W." box="[599,741,1084,1110]" pageId="3" pageNumber="26" pagination="2" refId="ref6913" refString="Bates, H. W. (1887) Pectinicornia and Lamellicornia. In: Salvin, O. &amp; Godma, F. (Eds.), Biologia Centrali- Americana, Insecta, Coleoptera, Vol. II, parte 2. Taylor and Francis, London." type="book chapter" year="1887">Bates, 1887</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C377654DC45EFFBB174FFB6C27DDA3F3" pageId="3" pageNumber="26" type="diagnosis">
<paragraph id="8BD236C6C45EFFBB174FFB6C27DDA3F3" blockId="3.[264,1323,1158,1905]" pageId="3" pageNumber="26">
<emphasis id="B919EAD4C45EFFBB174FFB6C2598A2EA" bold="true" box="[264,384,1158,1184]" pageId="3" pageNumber="26">Diagnosis</emphasis>
. This species is distinguished by the following combination of characters: antennal club reddish­brown with distal half darkened; pronotum black with numerous scale­like setae uniformly distributed on the disc; elytra reddish­brown with numerous scale­like setae along the inner half and scattered, short setae irregularly distributed along the outer half (
<figureCitation id="13562A43C45EFFBB17F4FACD25E3A30B" box="[435,507,1319,1345]" captionStart="FIGURES 16" captionStartId="6.[264,383,1501,1525]" captionTargetBox="[275,1312,313,1474]" captionTargetId="figure-83@6.[275,1312,309,1474]" captionTargetPageId="6" captionText="FIGURES 16. 1­­­ Macrodactylus costulatus Bates, male; 2­­­ M. sericeicollis Bates, male; 3­­­ M. sericinus Bates, male; 4­­­ M. suavis Bates, male; 5­­­ M. dimidiatus Guérin­Méneville, male; 6­ ­­ M. sylphis Bates, male." pageId="3" pageNumber="26">Fig. 2</figureCitation>
); pygidium black or dark brown with scattered scale­like setae; abdomen black; femora and tibiae reddish­brown, apex of tibiae noticeably darkened; tarsi black with scattered white setae; parameres with lanceolate apex, without setae on the external borders; female vestiture and color similar to male.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC45EFFBB177FFA2C2074A09B" pageId="3" pageNumber="26" type="materials_examined">
<paragraph id="8BD236C6C45EFFBB177FFA2C2074A09B" blockId="3.[264,1323,1158,1905]" pageId="3" pageNumber="26">
<emphasis id="B919EAD4C45EFFBB177FFA2C263EA3AA" bold="true" box="[312,550,1478,1504]" pageId="3" pageNumber="26">Material examined</emphasis>
(
<specimenCount id="9D6BFD4FC45EFFBB1404FA2D26A8A3AB" box="[579,688,1479,1505]" pageId="3" pageNumber="26" type="male">32 males</specimenCount>
,
<specimenCount id="9D6BFD4FC45EFFBB14F9FA2D275AA3AB" box="[702,834,1479,1505]" pageId="3" pageNumber="26" type="female">30 females</specimenCount>
)
<emphasis id="B919EAD4C45EFFBB1514FA2C20C0A3AA" bold="true" box="[851,1240,1478,1504]" pageId="3" pageNumber="26">
<collectionCode id="ED7CAE03C45EFFBB1514FA2C2781A3AA" box="[851,921,1478,1504]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="3" pageNumber="26">NEW</collectionCode>
<collectionCode id="ED7CAE03C45EFFBB15E4FA2C20CBA3AA" box="[931,1235,1478,1504]" pageId="3" pageNumber="26">COUNTRY RECORDS</collectionCode>
:
</emphasis>
<collectionCode id="ED7CAE03C45EFFBB12A6FA2D2568A043" pageId="3" pageNumber="26">HON­ DURAS</collectionCode>
:
<materialsCitation id="3B053C9BC45EFFBB17C5FA052083A0CB" collectingDate="1957-09-01" collectingDateMax="1994-12-28" collectingDateMin="1957-09-01" collectionCode="AMNH, HAHC, IEXA, MXAL" collectorName="Col. Baires &amp; Francisco Morazan &amp; El Zamorano &amp; Cols. P. H. Freytag &amp; H. J. Harlan &amp; Cols. J. Maes &amp; J Hernandez" country="Nicaragua" elevation="2000" location="Fuerte Pura" municipality="Mt. Uyuca" pageId="3" pageNumber="26" specimenCount="45" specimenCount-female="21" specimenCount-male="24" stateProvince="Matagalpa">
Siguatepec,
<date id="FFD31006C45EFFBB145FFA0526E6A043" box="[536,766,1519,1545]" pageId="3" pageNumber="26" value="1957-09-01">
<collectingDate id="EF97E9EEC45EFFBB145FFA0526E6A043" box="[536,766,1519,1545]" pageId="3" pageNumber="26" value="1957-09-01">1 September 1957</collectingDate>
</date>
,
<collectorName id="26985310C45EFFBB1555FA0527B8A043" box="[786,928,1519,1545]" pageId="3" pageNumber="26">Col. Baires</collectorName>
,
<specimenCount id="9D6BFD4FC45EFFBB1586FA052000A043" box="[961,1048,1519,1545]" pageId="3" pageNumber="26" type="male">1 male</specimenCount>
(
<collectionCode id="ED7CAE03C45EFFBB127AFA052085A043" box="[1085,1181,1519,1545]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="26">AMNH</collectionCode>
);
<collectorName id="26985310C45EFFBB12FEFA05256BA07B" pageId="3" pageNumber="26">Francisco Morazán</collectorName>
,
<collectorName id="26985310C45EFFBB1738F9FD2605A07B" box="[383,541,1559,1585]" pageId="3" pageNumber="26">El Zamorano</collectorName>
,
<collectingMunicipality id="6BB6ACBCC45EFFBB1475F9FD26AAA07B" box="[562,690,1559,1585]" pageId="3" pageNumber="26">Mt. Uyuca</collectingMunicipality>
, cloud forest
<quantity id="4C959B23C45EFFBB1514F9FD27B7A07B" box="[851,943,1559,1585]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="26" unit="m" value="2000.0">
<elevation id="0040D1F5C45EFFBB1514F9FD27B7A07B" box="[851,943,1559,1585]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="26" unit="m" value="2000.0">2000 m</elevation>
</quantity>
,
<date id="FFD31006C45EFFBB15FBF9FD2087A07B" box="[956,1183,1559,1585]" pageId="3" pageNumber="26" value="1965-12-24">
<collectingDate id="EF97E9EEC45EFFBB15FBF9FD2087A07B" box="[956,1183,1559,1585]" pageId="3" pageNumber="26" value="1965-12-24">24 December 1965</collectingDate>
</date>
,
<collectorName id="26985310C45EFFBB12EBF9FD2579A013" pageId="3" pageNumber="26">Cols. P. H. Freytag</collectorName>
&amp;
<collectorName id="26985310C45EFFBB17CBF9D5263BA013" box="[396,547,1599,1625]" pageId="3" pageNumber="26">H. J. Harlan</collectorName>
,
<specimenCount id="9D6BFD4FC45EFFBB1476F9D526B8A013" box="[561,672,1599,1625]" pageId="3" pageNumber="26" type="male">23 males</specimenCount>
,
<specimenCount id="9D6BFD4FC45EFFBB14E9F9D52729A013" box="[686,817,1599,1625]" pageId="3" pageNumber="26" type="female">21 females</specimenCount>
(
<collectionCode id="ED7CAE03C45EFFBB1504F9D52782A013" box="[835,922,1599,1625]" pageId="3" pageNumber="26">HAHC</collectionCode>
,
<collectionCode id="ED7CAE03C45EFFBB15EEF9D527EAA013" box="[937,1010,1599,1625]" pageId="3" pageNumber="26">IEXA</collectionCode>
,
<collectionCode id="ED7CAE03C45EFFBB1247F9D52044A013" box="[1024,1116,1599,1625]" pageId="3" pageNumber="26">MXAL</collectionCode>
).
<collectingCountry id="F37A7656C45EFFBB1235F9D5213EA013" box="[1138,1318,1599,1625]" name="Nicaragua" pageId="3" pageNumber="26">NICARAGUA</collectingCountry>
:
<collectingRegion id="49A9F824C45EFFBB174FF98D259FA0CB" box="[264,391,1639,1665]" country="Nicaragua" name="Matagalpa" pageId="3" pageNumber="26">Matagalpa</collectingRegion>
,
<location id="8EB2601DC45EFFBB17D4F98D2604A0CB" LSID="urn:lsid:plazi:treatment:03C487D0C45EFFBC174FFBD6259AA404:8EB2601DC45EFFBB17D4F98D2604A0CB" box="[403,540,1639,1665]" country="Nicaragua" municipality="Mt. Uyuca" name="Fuerte Pura" pageId="3" pageNumber="26" stateProvince="Matagalpa">Fuerte Pura</location>
<date id="FFD31006C45EFFBB1463F98D271EA0CB" box="[548,774,1639,1665]" pageId="3" pageNumber="26" value="1994-12-28">
<collectingDate id="EF97E9EEC45EFFBB1463F98D271EA0CB" box="[548,774,1639,1665]" pageId="3" pageNumber="26" value="1994-12-28">28 December 1994</collectingDate>
</date>
,
<collectorName id="26985310C45EFFBB1554F98D27B5A0CB" box="[787,941,1639,1665]" pageId="3" pageNumber="26">Cols. J.Maes</collectorName>
,
<collectorName id="26985310C45EFFBB15FDF98D2054A0CB" box="[954,1100,1639,1665]" pageId="3" pageNumber="26">J Hernández</collectorName>
and L
</materialsCitation>
.
<collectionCode id="ED7CAE03C45EFFBB12EFF98D20A7A0CB" box="[1192,1215,1639,1665]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="26">R</collectionCode>
.
<materialsCitation id="3B053C9BC45EFFBB128CF98D207FA09B" collectingDate="1994-11-20" collectionCode="MELN, MXAL" collectorName="Hernandez &amp; Tellez &amp; Johnson" country="Nicaragua" location="Cols. Maes" pageId="3" pageNumber="26" specimenCount="17" specimenCount-female="9" specimenCount-male="8" stateProvince="Jinotega">
<collectorName id="26985310C45EFFBB128CF98D2528A0E3" pageId="3" pageNumber="26">Hernández</collectorName>
<specimenCount id="9D6BFD4FC45EFFBB1770F965258BA0E3" box="[311,403,1679,1705]" pageId="3" pageNumber="26" type="male">5 males</specimenCount>
,
<specimenCount id="9D6BFD4FC45EFFBB17D8F9652617A0E3" box="[415,527,1679,1705]" pageId="3" pageNumber="26" type="female">4 females</specimenCount>
(
<collectionCode id="ED7CAE03C45EFFBB1459F965266DA0E3" box="[542,629,1679,1705]" pageId="3" pageNumber="26">MELN</collectionCode>
,
<collectionCode id="ED7CAE03C45EFFBB14C6F96526C5A0E3" box="[641,733,1679,1705]" pageId="3" pageNumber="26">MXAL</collectionCode>
); km 147 carr. Matagalpa­
<collectingRegion id="49A9F824C45EFFBB125CF9652067A0E3" box="[1051,1151,1679,1705]" country="Nicaragua" name="Jinotega" pageId="3" pageNumber="26">Jinotega</collectingRegion>
,
<date id="FFD31006C45EFFBB12CCF965255DA09B" pageId="3" pageNumber="26" value="1994-11-20">
<collectingDate id="EF97E9EEC45EFFBB12CCF965255DA09B" pageId="3" pageNumber="26" value="1994-11-20">20 November 1994</collectingDate>
</date>
,
<location id="8EB2601DC45EFFBB1716F95D25CFA09B" LSID="urn:lsid:plazi:treatment:03C487D0C45EFFBC174FFBD6259AA404:8EB2601DC45EFFBB1716F95D25CFA09B" box="[337,471,1719,1745]" country="Nicaragua" name="Cols. Maes" pageId="3" pageNumber="26" stateProvince="Jinotega">Cols. Maes</location>
,
<collectorName id="26985310C45EFFBB17A3F95D2633A09B" box="[484,555,1719,1745]" pageId="3" pageNumber="26">Tellez</collectorName>
y
<collectorName id="26985310C45EFFBB140FF95D26B3A09B" box="[584,683,1719,1745]" pageId="3" pageNumber="26">Johnson</collectorName>
,
<specimenCount id="9D6BFD4FC45EFFBB14F0F95D270CA09B" box="[695,788,1719,1745]" pageId="3" pageNumber="26" type="male">3 males</specimenCount>
,
<specimenCount id="9D6BFD4FC45EFFBB1567F95D2789A09B" box="[800,913,1719,1745]" pageId="3" pageNumber="26" type="female">5 females</specimenCount>
(
<collectionCode id="ED7CAE03C45EFFBB15E7F95D27E0A09B" box="[928,1016,1719,1745]" pageId="3" pageNumber="26">MELN</collectionCode>
,
<collectionCode id="ED7CAE03C45EFFBB1243F95D2078A09B" box="[1028,1120,1719,1745]" pageId="3" pageNumber="26">MXAL</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC45EFFBB177FF9342004A13B" pageId="3" pageNumber="26" type="biology_ecology">
<paragraph id="8BD236C6C45EFFBB177FF9342004A13B" blockId="3.[264,1323,1158,1905]" pageId="3" pageNumber="26">
<emphasis id="B919EAD4C45EFFBB177FF9342584A0B2" bold="true" box="[312,412,1758,1784]" pageId="3" pageNumber="26">Biology.</emphasis>
This species inhabits cloud forests and coffee plantations located at altitudes from
<quantity id="4C959B23C45EFFBB170AF8ED260EA16B" box="[333,534,1799,1825]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="3" pageNumber="26" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">1800 to 2000 m</quantity>
. Months of collection: September (1), November (8), December (53). The adults of this species fly during late autumn, which is notable because other Central American species are usually active during spring and summer.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC459FFBC177FFEF125D8A714" pageId="4" pageNumber="27" type="distribution">
<paragraph id="8BD236C6C459FFBC177FFEF125D8A714" blockId="4.[264,1324,283,590]" pageId="4" pageNumber="27">
<emphasis id="B919EAD4C459FFBC177FFEF125C9A77F" bold="true" box="[312,465,283,309]" pageId="4" pageNumber="27">Distribution</emphasis>
. Central
<collectingCountry id="F37A7656C459FFBC1405FEF626D9A77C" box="[578,705,284,310]" name="Guatemala" pageId="4" pageNumber="27">Guatemala</collectingCountry>
to northwestern
<collectingCountry id="F37A7656C459FFBC15C9FEF6201FA77C" box="[910,1031,284,310]" name="Nicaragua" pageId="4" pageNumber="27">Nicaragua</collectingCountry>
(13° 00'
<geoCoordinate id="EE595001C459FFBC12C6FEF620EDA77C" box="[1153,1269,284,310]" degrees="15" direction="north" minutes="24" orientation="latitude" pageId="4" pageNumber="27" precision="925" value="15.4">15° 24' N</geoCoordinate>
, 85° 50'
<geoCoordinate id="EE595001C459FFBC177CFEAE25ABA714" box="[315,435,324,350]" degrees="91" direction="west" minutes="10" orientation="longitude" pageId="4" pageNumber="27" precision="925" value="-91.166664">91° 10' W</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="C377654DC459FFBC177FFE81259AA404" pageId="4" pageNumber="27" type="discussion">
<paragraph id="8BD236C6C459FFBC177FFE81259AA404" blockId="4.[264,1324,283,590]" pageId="4" pageNumber="27">
<emphasis id="B919EAD4C459FFBC177FFE8125A9A7CF" bold="true" box="[312,433,363,389]" pageId="4" pageNumber="27">Remarks.</emphasis>
This species was originally described using a large series from San Gerónimo, Purula, Coban, and Cubilguitz from Vera Paz,
<collectingCountry id="F37A7656C459FFBC1539FE7E2019A7E4" box="[894,1025,404,430]" name="Guatemala" pageId="4" pageNumber="27">Guatemala</collectingCountry>
. The specimens studied from
<collectingCountry id="F37A7656C459FFBC170FFE5625A2A79C" box="[328,442,444,470]" name="Honduras" pageId="4" pageNumber="27">Honduras</collectingCountry>
and
<collectingCountry id="F37A7656C459FFBC17B3FE562674A79C" box="[500,620,444,470]" name="Nicaragua" pageId="4" pageNumber="27">Nicaragua</collectingCountry>
do not exhibit any major differences when compared to the Guatemalan specimens. The new records in
<collectingCountry id="F37A7656C459FFBC155EFE0E2789A7B4" box="[793,913,484,510]" name="Nicaragua" pageId="4" pageNumber="27">Nicaragua</collectingCountry>
extend the distribution
<quantity id="4C959B23C459FFBC12F7FE0E2112A7B4" box="[1200,1290,484,510]" metricMagnitude="5" metricUnit="m" metricValue="2.6" pageId="4" pageNumber="27" unit="km" value="260.0">260 km</quantity>
to the southeast of Coban,
<collectingCountry id="F37A7656C459FFBC146EFDE626B0A46C" box="[553,680,524,550]" name="Guatemala" pageId="4" pageNumber="27">Guatemala</collectingCountry>
across several chains of mountains in
<collectingCountry id="F37A7656C459FFBC123EFDE620E0A46C" box="[1145,1272,524,550]" name="Guatemala" pageId="4" pageNumber="27">Guatemala</collectingCountry>
and
<collectingCountry id="F37A7656C459FFBC174FFDDE2566A404" box="[264,382,564,590]" name="Honduras" pageId="4" pageNumber="27">Honduras</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>