treatments-xml/data/03/C4/87/03C487D0C459FFBD174FF9FC2635A52C.xml
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<document id="48FF815AD5B9CA69D2A0055497C9CE59" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625105362570" checkinUser="felipe" docAuthor="Arce-Pérez, Roberto &amp; Morón, Miguel-Angel" docDate="2005" docId="03C487D0C459FFBD174FF9FC2635A52C" docLanguage="en" docName="zt01012p037.pdf" docOrigin="Zootaxa 1012 (1012)" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Macrodactylus suavis Bates 1887" docType="treatment" docVersion="3" lastPageNumber="28" masterDocId="FFFDFFA8C45DFFB81647FFEA2418A64A" masterDocTitle="New species and new records of species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactyini) from Honduras and Nicaragua" masterLastPageNumber="37" masterPageNumber="23" pageNumber="27" updateTime="1698980141271" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3EA4BA3E5DCD166144E03D8430A45EE2">New species and new records of species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactyini) from Honduras and Nicaragua</mods:title>
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<mods:name id="928F15E7803D08EA1707C7D0D7204A2B" type="personal">
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<mods:namePart id="3A1EF7CE7403E9C4C2BBB6DCD86F381C">Arce-Pérez, Roberto</mods:namePart>
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<mods:namePart id="2735C82B583CCED09F09FBEE7E372273">Morón, Miguel-Angel</mods:namePart>
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<mods:date id="A98B1DEB80894286C773D5D978F5629F">2005</mods:date>
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<mods:number id="2A16C481CFB297043EC2651064F5B0AF">2005-06-28</mods:number>
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<mods:number id="8662CD4BADB02E884840D6D6D8299AEB">1012</mods:number>
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<treatment id="03C487D0C459FFBD174FF9FC2635A52C" ID-DOI="http://doi.org/10.5281/zenodo.5049184" ID-Zenodo-Dep="5049184" LSID="urn:lsid:plazi:treatment:03C487D0C459FFBD174FF9FC2635A52C" httpUri="http://treatment.plazi.org/id/03C487D0C459FFBD174FF9FC2635A52C" lastPageId="5" lastPageNumber="28" pageId="4" pageNumber="27">
<subSubSection id="C377654DC459FFBC174FF9FC26BBA07A" box="[264,675,1558,1584]" pageId="4" pageNumber="27" type="nomenclature">
<paragraph id="8BD236C6C459FFBC174FF9FC26BBA07A" blockId="4.[264,675,1558,1584]" box="[264,675,1558,1584]" pageId="4" pageNumber="27">
<heading id="D09A81AAC459FFBC174FF9FC26BBA07A" bold="true" box="[264,675,1558,1584]" fontSize="11" level="1" pageId="4" pageNumber="27" reason="1">
<taxonomicName id="4C6D4D45C459FFBC174FF9FC26BBA07A" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[264,675,1558,1584]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="suavis">
<emphasis id="B919EAD4C459FFBC174FF9FC26BBA07A" bold="true" box="[264,675,1558,1584]" pageId="4" pageNumber="27">
<emphasis id="B919EAD4C459FFBC174FF9FC2615A07A" bold="true" box="[264,525,1558,1584]" italics="true" pageId="4" pageNumber="27">Macrodactylus suavis</emphasis>
<bibRefCitation id="EFFC4B37C459FFBC1452F9FC26BBA07A" author="Bates, H. W." box="[533,675,1558,1584]" pageId="4" pageNumber="27" pagination="2" refId="ref6913" refString="Bates, H. W. (1887) Pectinicornia and Lamellicornia. In: Salvin, O. &amp; Godma, F. (Eds.), Biologia Centrali- Americana, Insecta, Coleoptera, Vol. II, parte 2. Taylor and Francis, London." type="book chapter" year="1887">Bates, 1887</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C377654DC459FFBD174FF98B200DA714" lastPageId="5" lastPageNumber="28" pageId="4" pageNumber="27" type="diagnosis">
<paragraph id="8BD236C6C459FFBD174FF98B200DA714" blockId="4.[264,1323,1633,1899]" lastBlockId="5.[264,1325,284,870]" lastPageId="5" lastPageNumber="28" pageId="4" pageNumber="27">
<emphasis id="B919EAD4C459FFBC174FF98B2598A031" bold="true" box="[264,384,1633,1659]" pageId="4" pageNumber="27">Diagnosis</emphasis>
. This species is distinguished by the following combination of characters: antennal club reddish­brown with the apex darkened; pronotum black with scattered scalelike setae mainly toward the borders of disc; scutellum with abundant, yellow, scale­like setae; elytra with numerous scale­like setae along the inner half and short setae distributed in two irregular stripes along the outer half, basal half reddish­brown and apical half black (
<figureCitation id="13562A43C459FFBC1756F8C32544A109" box="[273,348,1833,1859]" captionStart="FIGURES 16" captionStartId="6.[264,383,1501,1525]" captionTargetBox="[275,1312,313,1474]" captionTargetId="figure-83@6.[275,1312,309,1474]" captionTargetPageId="6" captionText="FIGURES 16. 1­­­ Macrodactylus costulatus Bates, male; 2­­­ M. sericeicollis Bates, male; 3­­­ M. sericinus Bates, male; 4­­­ M. suavis Bates, male; 5­­­ M. dimidiatus Guérin­Méneville, male; 6­ ­­ M. sylphis Bates, male." pageId="4" pageNumber="27">Fig. 4</figureCitation>
); pygidium black, dark brown or reddish­yellow with scattered scale­like setae near the center of disc; abdomen black; femora and tibiae reddish­brown, apex of tibiae darkened; tarsi black without white setae; parameres with lanceolate apex, without setae on the external borders; female vestiture and color similar to male.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC458FFBD177FFE8127C7A79C" pageId="5" pageNumber="28" type="materials_examined">
<paragraph id="8BD236C6C458FFBD177FFE8127C7A79C" blockId="5.[264,1325,284,870]" pageId="5" pageNumber="28">
<emphasis id="B919EAD4C458FFBD177FFE812630A7CF" bold="true" box="[312,552,363,389]" pageId="5" pageNumber="28">Material examined</emphasis>
(
<specimenCount id="9D6BFD4FC458FFBD147CFE862684A7CC" box="[571,668,364,390]" pageId="5" pageNumber="28" type="male">2 males</specimenCount>
,
<specimenCount id="9D6BFD4FC458FFBD14EBFE86270FA7CC" box="[684,791,364,390]" pageId="5" pageNumber="28" type="female">1 female</specimenCount>
)
<materialsCitation id="3B053C9BC458FFBD156EFE86258EA7E4" collectionCode="CNC" country="Nicaragua" location="Belt" pageId="5" pageNumber="28" specimenCount="1" stateProvince="Chontales" typeStatus="syntype">
<collectingCountry id="F37A7656C458FFBD156EFE8627C6A7CC" box="[809,990,364,390]" name="Nicaragua" pageId="5" pageNumber="28">NICARAGUA</collectingCountry>
:
<collectingRegion id="49A9F824C458FFBD15ABFE86207CA7CC" box="[1004,1124,364,390]" country="Nicaragua" name="Chontales" pageId="5" pageNumber="28">Chontales</collectingRegion>
,
<location id="8EB2601DC458FFBD1234FE8620BFA7CC" LSID="urn:lsid:plazi:treatment:03C487D0C459FFBD174FF9FC2635A52C:8EB2601DC458FFBD1234FE8620BFA7CC" box="[1139,1191,364,390]" country="Nicaragua" name="Belt" pageId="5" pageNumber="28" stateProvince="Chontales">Belt</location>
, male
<typeStatus id="54D68864C458FFBD12BDFE862522A7E4" pageId="5" pageNumber="28" type="syntype">syntype</typeStatus>
(
<collectionCode id="ED7CAE03C458FFBD170CFE7E2595A7E4" box="[331,397,404,430]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="5" pageNumber="28">CNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B053C9BC458FFBD17E6FE7E27C3A79C" collectingDate="1997-05-17" collectionCode="MXAL" collectorName="Col. F. A. Quezada" country="Costa Rica" elevation="1700" location="Quebrada Londres" municipality="San Antonio" pageId="5" pageNumber="28" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="San Jose" typeStatus="syntype">
<collectingCountry id="F37A7656C458FFBD17E6FE7E264AA7E4" box="[417,594,404,430]" name="Costa Rica" pageId="5" pageNumber="28">COSTA RICA</collectingCountry>
:
<collectingRegion id="49A9F824C458FFBD141AFE7E272CA7E4" box="[605,820,404,430]" country="Costa Rica" name="San Jose" pageId="5" pageNumber="28">San José province</collectingRegion>
,
<collectingMunicipality id="6BB6ACBCC458FFBD1507FE7E27CFA7E4" box="[832,983,404,430]" pageId="5" pageNumber="28">San Antonio</collectingMunicipality>
,
<location id="8EB2601DC458FFBD15A4FE7E20D8A7E4" LSID="urn:lsid:plazi:treatment:03C487D0C459FFBD174FF9FC2635A52C:8EB2601DC458FFBD15A4FE7E20D8A7E4" box="[995,1216,404,430]" country="Costa Rica" municipality="San Antonio" name="Quebrada Londres" pageId="5" pageNumber="28" stateProvince="San Jose">Quebrada Londres</location>
,
<quantity id="4C959B23C458FFBD128BFE7E2130A7E4" box="[1228,1320,404,430]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="28" unit="m" value="1700.0">
<elevation id="0040D1F5C458FFBD128BFE7E2130A7E4" box="[1228,1320,404,430]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="28" unit="m" value="1700.0">1700 m</elevation>
</quantity>
,
<date id="FFD31006C458FFBD174FFE5625BFA79C" box="[264,423,444,470]" pageId="5" pageNumber="28" value="1997-05-17">
<collectingDate id="EF97E9EEC458FFBD174FFE5625BFA79C" box="[264,423,444,470]" pageId="5" pageNumber="28" value="1997-05-17">17 May 1997</collectingDate>
</date>
,
<collectorName id="26985310C458FFBD17F4FE56268EA79C" box="[435,662,444,470]" pageId="5" pageNumber="28">Col. F. A. Quezada</collectorName>
,
<specimenCount id="9D6BFD4FC458FFBD14E4FE5626EDA79C" box="[675,757,444,470]" pageId="5" pageNumber="28" type="male">1 male</specimenCount>
,
<specimenCount id="9D6BFD4FC458FFBD1547FE56277EA79C" box="[768,870,444,470]" pageId="5" pageNumber="28" type="female">1 female</specimenCount>
(
<collectionCode id="ED7CAE03C458FFBD1531FE5627CBA79C" box="[886,979,444,470]" pageId="5" pageNumber="28">MXAL</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC458FFBD177FFE0926D7A46C" pageId="5" pageNumber="28" type="biology_ecology">
<paragraph id="8BD236C6C458FFBD177FFE0926D7A46C" blockId="5.[264,1325,284,870]" pageId="5" pageNumber="28">
<emphasis id="B919EAD4C458FFBD177FFE092584A7B7" bold="true" box="[312,412,483,509]" pageId="5" pageNumber="28">Biology.</emphasis>
This species inhabits wet montane forests located at altitudes from
<quantity id="4C959B23C458FFBD129AFE0E257CA46C" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.7" metricValueMin="0.8" pageId="5" pageNumber="28" unit="m" value="1250.0" valueMax="1700.0" valueMin="800.0">800 to 1700 m</quantity>
. Month of collection: May (2).
</paragraph>
</subSubSection>
<subSubSection id="C377654DC458FFBD177FFDD92547A43C" pageId="5" pageNumber="28" type="distribution">
<paragraph id="8BD236C6C458FFBD177FFDD92547A43C" blockId="5.[264,1325,284,870]" pageId="5" pageNumber="28">
<emphasis id="B919EAD4C458FFBD177FFDD925C9A407" bold="true" box="[312,465,563,589]" pageId="5" pageNumber="28">Distribution</emphasis>
. Southern
<collectingCountry id="F37A7656C458FFBD1411FDDE26D6A404" box="[598,718,564,590]" name="Nicaragua" pageId="5" pageNumber="28">Nicaragua</collectingCountry>
to central
<collectingCountry id="F37A7656C458FFBD1514FDDE27A8A404" box="[851,944,564,590]" name="Panama" pageId="5" pageNumber="28">Panama</collectingCountry>
(8° 50'
<geoCoordinate id="EE595001C458FFBD125CFDDE2088A404" box="[1051,1168,564,590]" degrees="12" direction="north" minutes="10" orientation="latitude" pageId="5" pageNumber="28" precision="925" value="12.166667">12° 10' N</geoCoordinate>
, 79° 30'
<geoCoordinate id="EE595001C458FFBD1342FDDE254AA43C" degrees="85" direction="west" minutes="20" orientation="longitude" pageId="5" pageNumber="28" precision="925" value="-85.333336">85° 20' W</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="C377654DC458FFBD177FFD6E2635A52C" pageId="5" pageNumber="28" type="discussion">
<paragraph id="8BD236C6C458FFBD177FFD6E2635A52C" blockId="5.[264,1325,284,870]" pageId="5" pageNumber="28">
<emphasis id="B919EAD4C458FFBD177FFD6E25A9A4D4" bold="true" box="[312,433,644,670]" pageId="5" pageNumber="28">Remarks.</emphasis>
This species was originally described with specimens from “
<collectingCountry id="F37A7656C458FFBD12C8FD6E2101A4D4" box="[1167,1305,644,670]" name="Costa Rica" pageId="5" pageNumber="28">Costa Rica</collectingCountry>
;” Volcán de Irazú, Cache,
<collectingCountry id="F37A7656C458FFBD1468FD4626ADA48C" box="[559,693,684,710]" name="Costa Rica" pageId="5" pageNumber="28">Costa Rica</collectingCountry>
; Bugaba,
<collectingCountry id="F37A7656C458FFBD1569FD462796A48C" box="[814,910,684,710]" name="Panama" pageId="5" pageNumber="28">Panama</collectingCountry>
; and
<collectingRegion id="49A9F824C458FFBD1597FD462050A48C" box="[976,1096,684,710]" country="Nicaragua" name="Chontales" pageId="5" pageNumber="28">Chontales</collectingRegion>
,
<collectingCountry id="F37A7656C458FFBD1211FD4620CAA48C" box="[1110,1234,684,710]" name="Nicaragua" pageId="5" pageNumber="28">Nicaragua</collectingCountry>
. Bates noted the variation of large body size (
<quantity id="4C959B23C458FFBD148FFD3E2703A4A4" box="[712,795,724,750]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="28" unit="mm" value="11.0">11 mm</quantity>
) and dark color of the middle and hind tibiae in the specimens from
<collectingRegion id="49A9F824C458FFBD1467FD162681A55C" box="[544,665,764,790]" country="Nicaragua" name="Chontales" pageId="5" pageNumber="28">Chontales</collectingRegion>
. Worn specimens are difficult to identify without the examination of male genitalia, because the distribution of setae on the elytra is an important diagnostic character.
</paragraph>
</subSubSection>
</treatment>
</document>