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<mods:title id="2C5C15479CADE2C2A43A0E6335B26488">Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)</mods:title>
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<mods:namePart id="A246FBF10BC71E0731E684059590DC67">Kim, Il-Hoi</mods:namePart>
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<mods:affiliation id="1A3F2194DE9E984EC2137F6F86FAF296">Korea Institute of Coastal Ecology, Inc., 802 - ho, 302 - dong, 397 Seokcheon-ro, Ojeong-gu, Bucheon, Gyeonggi-do 14449, Republic of Korea ® ihkim @ gwnu. ac. kr; https: // orcid. org / 0000 - 0002 - 7332 - 0043</mods:affiliation>
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<mods:namePart id="700D6F3AB53B5DCB1003A35FEAE4AF86">Boxshall, Geoff A.</mods:namePart>
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<mods:affiliation id="1233A7FF02B4E207B0B3FD64C0521F44">Korea Institute of Coastal Ecology, Inc., 802 - ho, 302 - dong, 397 Seokcheon-ro, Ojeong-gu, Bucheon, Gyeonggi-do 14449, Republic of Korea ® ihkim @ gwnu. ac. kr; https: // orcid. org / 0000 - 0002 - 7332 - 0043</mods:affiliation>
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<mods:title id="9D4AAD34153511E61AD1034CC9C040A9">Megataxa</mods:title>
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<treatment id="03C487CBEFF33A97FF4DFCFDFC36FAAC" ID-DOI="http://doi.org/10.5281/zenodo.5676116" ID-GBIF-Taxon="194475249" ID-Zenodo-Dep="5676116" LSID="urn:lsid:plazi:treatment:03C487CBEFF33A97FF4DFCFDFC36FAAC" httpUri="http://treatment.plazi.org/id/03C487CBEFF33A97FF4DFCFDFC36FAAC" lastPageNumber="163" pageId="162" pageNumber="163">
<subSubSection id="C3776556EFF33A97FF4DFCFDFD89FC8C" box="[136,588,819,846]" pageId="162" pageNumber="163" type="nomenclature">
<paragraph id="8BD236DDEFF33A97FF4DFCFDFD89FC8C" blockId="162.[136,588,819,846]" box="[136,588,819,846]" pageId="162" pageNumber="163">
<heading id="D09A81B1EFF33A97FF4DFCFDFD89FC8C" bold="true" box="[136,588,819,846]" fontSize="11" level="1" pageId="162" pageNumber="163" reason="1">
<emphasis id="B919EACFEFF33A97FF4DFCFDFD89FC8C" bold="true" box="[136,588,819,846]" pageId="162" pageNumber="163">
Genus
<taxonomicName id="4C6D4D5EEFF33A97FF1AFCFDFD89FC8C" authority="Buchholz, 1869" authorityName="Buchholz" authorityYear="1869" box="[223,588,819,846]" class="Maxillopoda" family="Notodelphyidae" genus="Goniodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="162" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEFF33A97FF1AFCFDFE40FC8F" bold="true" box="[223,389,819,846]" italics="true" pageId="162" pageNumber="163">Goniodelphys</emphasis>
<bibRefCitation id="EFFC4B2CEFF33A97FE48FCFDFD89FC8C" author="Buchholz, R." box="[397,588,819,845]" journalOrPublisher="Zeitschrift f ̡ r wissenschaftliche Zoologie" pageId="162" pageNumber="163" pagination="99 - 155" part="19" refId="ref327368" refString="Buchholz, R. (1869) Beitrage zur Kenntniss der innerhalb der Ascidien lebenden parasitischen Crustaceen des Mittelmeeres. Zeitschrift f ̡ r wissenschaftliche Zoologie, 19, 99 - 155, pls. 5 - 11. https: // doi. org / 10.5962 / bhl. title. 53715" title="Beitrage zur Kenntniss der innerhalb der Ascidien lebenden parasitischen Crustaceen des Mittelmeeres" type="journal article" year="1869">Buchholz, 1869</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3776556EFF33A97FF4DFCB9FBF2FF28" pageId="162" pageNumber="163" type="diagnosis">
<paragraph id="8BD236DDEFF33A97FF4DFCB9FDB2F8E6" blockId="162.[136,778,887,1832]" pageId="162" pageNumber="163">
<emphasis id="B919EACFEFF33A97FF4DFCB9FF3EFC50" bold="true" box="[136,251,887,913]" pageId="162" pageNumber="163">Diagnosis</emphasis>
: Body consisting of cephalosome, first to fifth pedigerous somites and free urosome. Fourth pedigerous somite fused to fifth and forming brood pouch as long as anterior part of prosome. Free urosome 5-segmented, comprising genital somite and 4 free abdominal somites. Anal somite with paired tapering posteroventral protuberances densely ornamented with minute spinules. Caudal rami armed with 2 claws and 4 setae. Antennule 8-segmented. Antenna 4-segmented; basis with large pinnate seta and small seta (representing exopod); compound distal endopodal segment armed with 10 setae plus terminal claw. Mandible with 5 teeth and 1 or 2 proximal setae on coxal gnathobase, basis with 1 seta; exopod 2-segmented and armedwith 3 and 2 setae on first and second segments, respectively, or unsegmented and armed with only 4 setae; endopod 2-segmented, armed with 4 or 5 and 9 setae on first and second segments, respectively. Maxillulewith 9 setaeon arthrite, 1 seta on coxal endite, 2 setae on epipodite, 3 setae on medial margin of basis; exopod with 3 distal setae; endopod with 4 distalsetae. Maxilla 5-segmented; syncoxawith 3, 1, 2, and 2/3 setae on first to fourth endites, respectively; basis with 3 setae; first to third endopodal segments with 1, 1, and 3/4 setae, respectively. Maxilliped lobate with 10 setae (4 + 4 medial plus 2 apical). Legs 1–4 with 3- segmented exopods and 2- or 3-segmented endopods. Inner coxal seta lacking in legs 1–4. Inner distal spine present on basis of leg 1. Armature formula:
</paragraph>
<paragraph id="8BD236DDEFF33A97FC9AFF63FBF2FF28" blockId="162.[810,1452,173,1390]" pageId="162" pageNumber="163">Leg 5 consistingof protopod with 1 seta and exopodal segment bearing 2 setae.</paragraph>
</subSubSection>
<subSubSection id="C3776556EFF33A97FC9AFF3FFBADFEEC" pageId="162" pageNumber="163" type="type_taxon">
<paragraph id="8BD236DDEFF33A97FC9AFF3FFBADFEEC" blockId="162.[810,1452,173,1390]" pageId="162" pageNumber="163">
<emphasis id="B919EACFEFF33A97FC9AFF3FFBC0FECB" bold="true" box="[863,1029,240,267]" pageId="162" pageNumber="163">
<typeStatus id="54D6887FEFF33A97FC9AFF3FFC5FFECA" box="[863,922,241,267]" pageId="162" pageNumber="163">Type</typeStatus>
Species
</emphasis>
:
<taxonomicName id="4C6D4D5EEFF33A97FBE7FF3FFCA0FEEC" authority="Buchholz, 1869" authorityName="Buchholz" authorityYear="1869" class="Maxillopoda" family="Notodelphyidae" genus="Goniodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="B919EACFEFF33A97FBE7FF3FFAE1FECA" box="[1058,1316,241,267]" italics="true" pageId="162" pageNumber="163">Goniodelphys trigona</emphasis>
<bibRefCitation id="EFFC4B2CEFF33A97FAFCFF3FFCA0FEEC" author="Buchholz, R." journalOrPublisher="Zeitschrift f ̡ r wissenschaftliche Zoologie" pageId="162" pageNumber="163" pagination="99 - 155" part="19" refId="ref327368" refString="Buchholz, R. (1869) Beitrage zur Kenntniss der innerhalb der Ascidien lebenden parasitischen Crustaceen des Mittelmeeres. Zeitschrift f ̡ r wissenschaftliche Zoologie, 19, 99 - 155, pls. 5 - 11. https: // doi. org / 10.5962 / bhl. title. 53715" title="Beitrage zur Kenntniss der innerhalb der Ascidien lebenden parasitischen Crustaceen des Mittelmeeres" type="journal article" year="1869">Buchholz, 1869</bibRefCitation>
</taxonomicName>
, by original monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C3776556EFF33A97FC9AFEFBFC36FAAC" pageId="162" pageNumber="163" type="discussion">
<paragraph id="8BD236DDEFF33A97FC9AFEFBFC06FC53" blockId="162.[810,1452,173,1390]" pageId="162" pageNumber="163">
<emphasis id="B919EACFEFF33A97FC9AFEFBFC09FE8E" bold="true" box="[863,972,309,335]" pageId="162" pageNumber="163">Remarks</emphasis>
. The original description of the
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species
<taxonomicName id="4C6D4D5EEFF33A97FC4FFE99FBC3FEB0" authorityName="Buchholz" authorityYear="1869" box="[906,1030,343,369]" class="Maxillopoda" family="Notodelphyidae" genus="Goniodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="trigona">
<emphasis id="B919EACFEFF33A97FC4FFE99FBC3FEB0" box="[906,1030,343,369]" italics="true" pageId="162" pageNumber="163">G. trigona</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFFC4B2CEFF33A97FB84FE99FAC3FEB3" author="Buchholz, R." box="[1089,1286,343,370]" journalOrPublisher="Zeitschrift f ̡ r wissenschaftliche Zoologie" pageId="162" pageNumber="163" pagination="99 - 155" part="19" refId="ref327368" refString="Buchholz, R. (1869) Beitrage zur Kenntniss der innerhalb der Ascidien lebenden parasitischen Crustaceen des Mittelmeeres. Zeitschrift f ̡ r wissenschaftliche Zoologie, 19, 99 - 155, pls. 5 - 11. https: // doi. org / 10.5962 / bhl. title. 53715" title="Beitrage zur Kenntniss der innerhalb der Ascidien lebenden parasitischen Crustaceen des Mittelmeeres" type="journal article" year="1869">Buchholz (1869)</bibRefCitation>
lacked detail by modern standards but it was redescribed by
<bibRefCitation id="EFFC4B2CEFF33A97FAA7FEB7FC17FE77" author="Illg, P. L. &amp; Dudley, P. L." journalOrPublisher="Pubblicazioni della Stazione Zoologica di Napoli" pageId="162" pageNumber="163" pagination="373 - 451" part="34" refId="ref329106" refString="Illg, P. L. &amp; Dudley, P. L. (1965) Notodelphyid copepods from the vicinity of Naples. Pubblicazioni della Stazione Zoologica di Napoli, 34, 373 - 451." title="Notodelphyid copepods from the vicinity of Naples" type="journal article" year="1965">
Illg &amp;
<collectingRegion id="49A9F83FEFF33A97FCEFFE55FCB9FE74" box="[810,892,411,437]" country="United Kingdom" name="Dudley" pageId="162" pageNumber="163">Dudley</collectingRegion>
(1965)
</bibRefCitation>
. However, their material was an immature female (Copepodid V) so, although their specimen came from Italian waters (Pozzuoli) close to the
<typeStatus id="54D6887FEFF33A97FAD8FE11FA88FE38" box="[1309,1357,479,505]" pageId="162" pageNumber="163">type</typeStatus>
locality (Naples Bay), their redescription provided insufficient morphological information for a robust species comparison. Subsequently Ooishi &amp; Illg (1973) obtained adults of both sexes from
<taxonomicName id="4C6D4D5EEFF33A97FB90FDA9FAE0FD40" authorityName=", MEDITS" authorityYear="2016" box="[1109,1317,615,641]" class="Ascidiacea" family="Ascidiidae" genus="Ascidiella" kingdom="Animalia" order="Enterogona" pageId="162" pageNumber="163" phylum="Chordata" rank="species" species="aspersa">
<emphasis id="B919EACFEFF33A97FB90FDA9FAE0FD40" box="[1109,1317,615,641]" italics="true" pageId="162" pageNumber="163">Ascidiella aspersa</emphasis>
</taxonomicName>
in the
<collectingRegion id="49A9F83FEFF33A97FABDFDA9FA6EFD40" box="[1400,1451,615,641]" country="Papua New Guinea" name="Gulf" pageId="162" pageNumber="163">Gulf</collectingRegion>
of Trieste. They commented that in the adult female leg 1 was functionally 2-segmented due to the suppression of the articulation between ancestral segments 2 and 3. Legs 2–4 exhibit the same segmentation as in the Copepodid V stage but the segments were more elongate and the setae were better developed. In addition, the 2 claws on the caudal ramus were much more robust than in the Copepodid V.
</paragraph>
<paragraph id="8BD236DDEFF33A97FC9AFC57FC36FAAC" blockId="162.[810,1452,173,1390]" pageId="162" pageNumber="163">
The genus
<taxonomicName id="4C6D4D5EEFF33A97FC22FC57FB46FC72" authorityName="Buchholz" authorityYear="1869" box="[999,1155,921,947]" class="Maxillopoda" family="Notodelphyidae" genus="Goniodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="162" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEFF33A97FC22FC57FB46FC72" box="[999,1155,921,947]" italics="true" pageId="162" pageNumber="163">Goniodelphys</emphasis>
</taxonomicName>
remained monotypic for more than a century until Ooishi &amp; Illg (1973) described a second species,
<taxonomicName id="4C6D4D5EEFF33A97FC19FC10FAF9FC36" authority="Ooishi &amp; Illg, 1973" authorityName="Ooishi &amp; Illg" authorityYear="1973" box="[988,1340,989,1016]" class="Maxillopoda" family="Notodelphyidae" genus="Ooishillgia" kingdom="Animalia" order="Cyclopoida" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="tokiokai">
<emphasis id="B919EACFEFF33A97FC19FC10FB9DFC36" box="[988,1112,989,1016]" italics="true" pageId="162" pageNumber="163">G. tokiokai</emphasis>
Ooishi &amp; Illg, 1973
</taxonomicName>
, collected from
<taxonomicName id="4C6D4D5EEFF33A97FCB4FC31FA94FBD8" authority="(Van Name, 1918)" baseAuthorityName="Van Name" baseAuthorityYear="1918" box="[881,1361,1023,1050]" class="Ascidiacea" family="Agneziidae" genus="Pterygascidia" kingdom="Animalia" order="Enterogona" pageId="162" pageNumber="163" phylum="Chordata" rank="species" species="longa">
<emphasis id="B919EACFEFF33A97FCB4FC31FB9BFBD8" box="[881,1118,1023,1049]" italics="true" pageId="162" pageNumber="163">Pterygascidia longa</emphasis>
(Van Name, 1918)
</taxonomicName>
in the
<collectingCountry id="F37A764DEFF33A97FCEFFBEFFC6DFBFA" box="[810,936,1057,1083]" name="Philippines" pageId="162" pageNumber="163">Philippines</collectingCountry>
at a depth of
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72 to
<specimenCount id="9D6BFD54EFF33A97FB49FBEFFB0FFBFD" box="[1164,1226,1057,1084]" pageId="162" pageNumber="163" type="generic">80 m</specimenCount>
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. Here we recognize
<taxonomicName id="4C6D4D5EEFF33A97FCEFFB8AFC60FB9C" authorityName="Ooishi &amp; Illg" authorityYear="1973" box="[810,933,1091,1118]" class="Maxillopoda" family="Notodelphyidae" genus="Ooishillgia" kingdom="Animalia" order="Cyclopoida" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="tokiokai">
<emphasis id="B919EACFEFF33A97FCEFFB8AFC60FB9C" box="[810,933,1091,1118]" italics="true" pageId="162" pageNumber="163">G. tokiokai</emphasis>
</taxonomicName>
as the
<typeStatus id="54D6887FEFF33A97FC2AFB8AFBDAFB9F" box="[1007,1055,1092,1118]" pageId="162" pageNumber="163">type</typeStatus>
species of a new genus,
<taxonomicName id="4C6D4D5EEFF33A97FAE8FB8DFA6EFB9C" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[1325,1451,1091,1117]" class="Maxillopoda" family="Notodelphyidae" genus="Ooishillgia" kingdom="Animalia" order="Cyclopoida" pageId="162" pageNumber="163" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B919EACFEFF33A97FAE8FB8DFA6EFB9C" box="[1325,1451,1091,1117]" italics="true" pageId="162" pageNumber="163">Ooishillgia</emphasis>
</taxonomicName>
<emphasis id="B919EACFEFF33A97FCEFFBABFC50FBBE" bold="true" box="[810,917,1125,1151]" pageId="162" pageNumber="163">
<taxonomicNameLabel id="A22A57B4EFF33A97FCEFFBABFC50FBBE" box="[810,917,1125,1151]" pageId="162" pageNumber="163" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(see below). In their wider discussion, Ooishi &amp; Illg (1973) highlighted the similarities between
<taxonomicName id="4C6D4D5EEFF33A97FCEFFB67FC03FB02" authorityName="Buchholz" authorityYear="1869" box="[810,966,1193,1219]" class="Maxillopoda" family="Notodelphyidae" genus="Goniodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="162" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEFF33A97FCEFFB67FC03FB02" box="[810,966,1193,1219]" italics="true" pageId="162" pageNumber="163">Goniodelphys</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6D4D5EEFF33A97FC19FB67FA6EFB02" authority="Schellenberg, 1922" authorityName="Schellenberg" authorityYear="1922" box="[988,1451,1193,1219]" class="Maxillopoda" family="Notodelphyidae" genus="Notopterophoroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="162" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEFF33A97FC19FB67FB7EFB02" box="[988,1211,1193,1219]" italics="true" pageId="162" pageNumber="163">Notopterophoroides</emphasis>
<bibRefCitation id="EFFC4B2CEFF33A97FB0EFB67FA6EFB02" author="Schellenberg, A." box="[1227,1451,1193,1219]" journalOrPublisher="Mitteilungen aus dem Zoologischen Museum in Berlin" pageId="162" pageNumber="163" pagination="217 - 274" part="10" refId="ref330676" refString="Schellenberg, A. (1922) Neue Notodelphyiden des Berliner und Hamburger Museums mit einer Uberscht der ascidienwohnenden Gattungen und Arten. Mitteilungen aus dem Zoologischen Museum in Berlin, 10 (2), 217 - 274. https: // doi. org / 10.1002 / mmnz. 4830100201" title="Neue Notodelphyiden des Berliner und Hamburger Museums mit einer Uberscht der ascidienwohnenden Gattungen und Arten" type="journal article" year="1922">Schellenberg, 1922</bibRefCitation>
</taxonomicName>
and the monotypic genus
<taxonomicName id="4C6D4D5EEFF33A97FBA0FB02FAF1FB27" authority="Illg, 1951" authorityName="Illg" authorityYear="1951" box="[1125,1332,1227,1254]" class="Maxillopoda" family="Notodelphyidae" genus="Ustina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="162" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEFF33A97FBA0FB02FB75FB24" box="[1125,1200,1228,1253]" italics="true" pageId="162" pageNumber="163">Ustina</emphasis>
<bibRefCitation id="EFFC4B2CEFF33A97FB78FB05FAF1FB27" author="Illg" box="[1213,1332,1227,1254]" journalOrPublisher="Journal of the Washington Academy of Sciences" pageId="162" pageNumber="163" pagination="30 - 34" part="41" publicationUrl="https://www.jstor.org/stable/24531659]" refId="ref328869" refString="Illg, P. L. (1951) A new genus and species of notodelphyoid copepod from Japan. Journal of the Washington Academy of Sciences, 41, 30 - 34. [https: // www. jstor. org / stable / 24531659]" title="A new genus and species of notodelphyoid copepod from Japan" type="journal article" year="1951">Illg, 1951</bibRefCitation>
</taxonomicName>
. They reexamined the
<typeStatus id="54D6887FEFF33A97FC00FB20FC30FAC9" box="[965,1013,1262,1288]" pageId="162" pageNumber="163">type</typeStatus>
material of
<taxonomicName id="4C6D4D5EEFF33A97FBBCFB20FAB8FAC9" authority="Illg, 1951" authorityName="Illg" authorityYear="1951" box="[1145,1405,1261,1288]" class="Maxillopoda" family="Notodelphyidae" genus="Ustina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="clarki">
<emphasis id="B919EACFEFF33A97FBBCFB20FACEFAC6" box="[1145,1291,1261,1287]" italics="true" pageId="162" pageNumber="163">Ustina clarki</emphasis>
<bibRefCitation id="EFFC4B2CEFF33A97FAD4FB23FAB8FAC9" author="Illg" box="[1297,1405,1261,1288]" journalOrPublisher="Journal of the Washington Academy of Sciences" pageId="162" pageNumber="163" pagination="30 - 34" part="41" publicationUrl="https://www.jstor.org/stable/24531659]" refId="ref328869" refString="Illg, P. L. (1951) A new genus and species of notodelphyoid copepod from Japan. Journal of the Washington Academy of Sciences, 41, 30 - 34. [https: // www. jstor. org / stable / 24531659]" title="A new genus and species of notodelphyoid copepod from Japan" type="journal article" year="1951">Illg, 1951</bibRefCitation>
</taxonomicName>
and concluded that it should be placed in
<taxonomicName id="4C6D4D5EEFF33A97FB0CFAC1FAA0FAE8" authorityName="Buchholz" authorityYear="1869" box="[1225,1381,1295,1321]" class="Maxillopoda" family="Notodelphyidae" genus="Goniodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="162" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEFF33A97FB0CFAC1FAA0FAE8" box="[1225,1381,1295,1321]" italics="true" pageId="162" pageNumber="163">Goniodelphys</emphasis>
</taxonomicName>
, as
<taxonomicName id="4C6D4D5EEFF33A97FA55FADEFC3DFA8D" authority="(Illg, 1951)" baseAuthorityName="Illg" baseAuthorityYear="1951" class="Maxillopoda" family="Notodelphyidae" genus="Ustina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="clarki">
<emphasis id="B919EACFEFF33A97FA55FADEFCAEFA8A" italics="true" pageId="162" pageNumber="163">G. clarki</emphasis>
(
<bibRefCitation id="EFFC4B2CEFF33A97FCBBFAFFFC35FA8D" author="Illg" box="[894,1008,1329,1356]" journalOrPublisher="Journal of the Washington Academy of Sciences" pageId="162" pageNumber="163" pagination="30 - 34" part="41" publicationUrl="https://www.jstor.org/stable/24531659]" refId="ref328869" refString="Illg, P. L. (1951) A new genus and species of notodelphyoid copepod from Japan. Journal of the Washington Academy of Sciences, 41, 30 - 34. [https: // www. jstor. org / stable / 24531659]" title="A new genus and species of notodelphyoid copepod from Japan" type="journal article" year="1951">Illg, 1951</bibRefCitation>
)
</taxonomicName>
which they formally recognized as a new combination.
</paragraph>
</subSubSection>
</treatment>
</document>
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