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<document id="D4BC0DF51E6063F70F5D34BC9D6FA8EB" ID-DOI="10.5281/zenodo.4508304" ID-ISSN="2345-7600" ID-Zenodo-Dep="4508304" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612543142085" checkinUser="carolina" docAuthor="Barnes, R. S. K." docDate="2010" docId="03C48786FFBBFFA051DEF93EFAAEFC75" docLanguage="en" docName="RafflesBZool.58.1.31-49.pdf" docOrigin="Raffles Bulletin of Zoology 58 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Macrophthalmus (Paramareotis) Komai, Goshima &amp; Murai 1995" docType="treatment" docVersion="5" lastPageNumber="42" masterDocId="FFFDFFFEFFB0FFAB5126FF80FF89FFE0" masterDocTitle="A Review Of The Sentinel And Allied Crabs (Crustacea: Brachyura: Macrophthalmidae), With Particular Reference To The Genus Macrophthalmus" masterLastPageNumber="49" masterPageNumber="31" pageNumber="42" updateTime="1698867619473" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F5075895E8AAAC517ADD66BCF7020606">A Review Of The Sentinel And Allied Crabs (Crustacea: Brachyura: Macrophthalmidae), With Particular Reference To The Genus Macrophthalmus</mods:title>
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<mods:namePart id="76ED2680782CEBE01127E13C0B40962A">Barnes, R. S. K.</mods:namePart>
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<mods:title id="8809A46C81404BE3A76C6D1147284D4A">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="511B3D8217214F2AA063A8D0758ED193">2010</mods:date>
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<mods:number id="AC8B40A0732353F825A60B64E505EFE2">2010-02-28</mods:number>
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<mods:number id="80C264A42D13CB36C78FB3E3E0055639">58</mods:number>
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<mods:identifier id="E2EE1F3E82AD756C2D76118DF98F32AB" type="DOI">10.5281/zenodo.4508304</mods:identifier>
<mods:identifier id="EDBDF017AE148E8BEC30DE1D93284CE6" type="ISSN">2345-7600</mods:identifier>
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<treatment id="03C48786FFBBFFA051DEF93EFAAEFC75" ID-DOI="http://doi.org/10.5281/zenodo.4508917" ID-Zenodo-Dep="4508917" LSID="urn:lsid:plazi:treatment:03C48786FFBBFFA051DEF93EFAAEFC75" httpUri="http://treatment.plazi.org/id/03C48786FFBBFFA051DEF93EFAAEFC75" lastPageNumber="42" pageId="11" pageNumber="42">
<subSubSection id="C377651BFFBBFFA051DEF93EFDE1F916" pageId="11" pageNumber="42" type="nomenclature">
<paragraph id="8BD23690FFBBFFA051DEF93EFDE1F916" blockId="11.[244,616,1726,1813]" pageId="11" pageNumber="42">
<heading id="D09A81FCFFBBFFA051DEF93EFDE1F916" bold="true" centered="true" fontSize="10" level="2" pageId="11" pageNumber="42" reason="2">
<taxonomicName id="4C6D4D13FFBBFFA051DEF93EFDE1F916" ID-CoL="849FQ" authority="Komai, Goshima &amp; Murai, 1995" authorityName="Komai, Goshima &amp; Murai" authorityYear="1995" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="42" phylum="Arthropoda" rank="subGenus" subGenus="Paramareotis">
<emphasis id="B919EA82FFBBFFA051DEF93EFDE1F916" bold="true" pageId="11" pageNumber="42">
<emphasis id="B919EA82FFBBFFA051DEF93EFE3FF936" bold="true" box="[248,438,1726,1750]" italics="true" pageId="11" pageNumber="42">Macrophthalmus</emphasis>
(
<emphasis id="B919EA82FFBBFFA050E1F93EFDD2F936" bold="true" box="[455,603,1726,1750]" italics="true" pageId="11" pageNumber="42">Paramareotis</emphasis>
) Komai, Goshima &amp; Murai, 1995
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C377651BFFBBFFA0505AF97EFAAEFC75" pageId="11" pageNumber="42" type="description">
<paragraph id="8BD23690FFBBFFA0505AF97EFE69F8F5" blockId="11.[244,616,1726,1813]" box="[380,480,1790,1813]" pageId="11" pageNumber="42">
(
<figureCitation id="13562A15FFBBFFA050A2F97EFE51F8F5" box="[388,472,1790,1813]" captionStart="Fig" captionStartId="12.[134,169,2060,2081]" captionTargetBox="[147,1465,526,2019]" captionTargetId="figure-14@12.[128,1484,511,2037]" captionTargetPageId="12" captionText="Fig. 4. Form of the carapace and adult male chela of subgroups of the genus Macrophthalmus IV: (a) the M. japonicus group of the subgenus Mareotis (both the relatively parallel-sided and relatively rounded carapace types), and (b) subgenus Paramareotis. Scale bars = 5mm." figureDoi="http://doi.org/10.5281/zenodo.4508316" httpUri="https://zenodo.org/record/4508316/files/figure.png" pageId="11" pageNumber="42">Fig. 4B</figureCitation>
)
</paragraph>
<paragraph id="8BD23690FFBBFFA05149F8BEFA00FDB5" blockId="11.[111,750,1854,2101]" lastBlockId="11.[810,1449,190,597]" pageId="11" pageNumber="42">
Small, carapace breadth &lt;
<quantity id="4C959B75FFBBFFA050BEF8BEFE60F8B5" box="[408,489,1854,1877]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="42" unit="mm" value="15.0">15 mm</quantity>
; ocular peduncles short and stout, not projecting beyond lateral carapace margins, subequal in length to breadth of front or shorter; front broad, not constricted between bases of ocular peduncles, where its breadth is 20–30% distance between external orbital angles; ischium of external maxilliped ca. 1.25 times length of merus; carapace with breadth 1.3–1.6 times length, with lateral margins subparallel, with broad-based subrectangular or pointed anterolateral teeth, without conspicuous aggregations of granules into rows or clumps on branchial regions (indistinct rows present in some); central region of posterior border of epistome usually straight (with convexity in one species); males with stridulatory apparatus in all species (i.e. with short horny ridge on ventoroflexor margin, or on inner surface near ventoroflexor margin, of cheliped merus, and lower orbital border with 1–2 large triangular protuberances occupying at least one fifth of that margin); fingers of male chela short with index straight or slightly downflexed and with differentiated teeth on both fingers. Intertidal, with one species often occurring in holes in limestone rocks and the others often associated with mangrove pneumatophores.
</paragraph>
<paragraph id="8BD23690FFBBFFA0520CFDFEFAAEFC75" blockId="11.[810,1450,638,918]" pageId="11" pageNumber="42">
Four species are included:
<taxonomicName id="4C6D4D13FFBBFFA05572FDFFFA2CFD75" authority="(Sakai, 1969)" baseAuthorityName="Sakai" baseAuthorityYear="1969" box="[1108,1445,638,662]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="boteltobagoe">
<emphasis id="B919EA82FFBBFFA05572FDFFFA8CFD76" box="[1108,1285,639,662]" italics="true" pageId="11" pageNumber="42">M. boteltobagoe</emphasis>
(
<bibRefCitation id="EFFC4B61FFBBFFA05431FDFEFA14FD75" author="Sakai, T" box="[1303,1437,638,661]" pageId="11" pageNumber="42" pagination="243 - 280" refId="ref15171" refString="Sakai, T., 1969. Two genera and twenty two new species of crabs from Japan. Proceedings of the Biological Society of Washington, 82: 243 - 280." type="journal article" year="1969">Sakai, 1969</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C6D4D13FFBBFFA0520CFD1FFBA0FD55" authority="de Man, 1888" authorityName="de Man" authorityYear="1888" box="[810,1065,670,694]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="erato">
<emphasis id="B919EA82FFBBFFA0520CFD1FFC0EFD56" box="[810,903,671,694]" italics="true" pageId="11" pageNumber="42">M. erato</emphasis>
de
<bibRefCitation id="EFFC4B61FFBBFFA05294FD1EFBA0FD55" author="Man, J" box="[946,1065,670,693]" pageId="11" pageNumber="42" pagination="1 - 64" refId="ref14025" refString="Man, J. G. de, 1888. Report on the podophthalmus Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum. Journal of the Linnean Society of London, Zoology, 22 (136): 1 - 64." type="journal article" year="1888">Man, 1888</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6D4D13FFBBFFA05510FD1FFAD8FD55" authority="Serene, 1973" authorityName="Serene" authorityYear="1973" box="[1078,1361,670,694]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B919EA82FFBBFFA05510FD1FFB31FD56" box="[1078,1208,671,694]" italics="true" pageId="11" pageNumber="42">M. holthuisi</emphasis>
<bibRefCitation id="EFFC4B61FFBBFFA055E7FD1EFAD8FD55" author="Serene, R" box="[1217,1361,670,693]" pageId="11" pageNumber="42" pagination="99 - 116" refId="ref15606" refString="Serene, R., 1973. Notes on Indo-West Pacific species of Macrophthalmus. Zoologische Mededelingen, Leiden, 46 (8): 99 - 116." type="journal article" year="1973">Serène, 1973</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C6D4D13FFBBFFA054AAFD1FFB3BFD35" authority="A. Milne Edwards, 1873" authorityName="A. Milne Edwards" authorityYear="1873" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="quadratus">
<emphasis id="B919EA82FFBBFFA054AAFD1FFC1EFD36" italics="true" pageId="11" pageNumber="42">M. quadratus</emphasis>
A. Milne Edwards, 1873
</taxonomicName>
(the
<typeStatus id="54D68832FFBBFFA055D6FD3EFAABFD35" box="[1264,1314,702,725]" pageId="11" pageNumber="42">type</typeStatus>
), Kosuge &amp; Davie (2001) having demonstrated that
<taxonomicName id="4C6D4D13FFBBFFA055E8FD5FFAF5FD16" baseAuthorityName="Sakai" baseAuthorityYear="1969" box="[1230,1404,735,758]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="boteltobagoe">
<emphasis id="B919EA82FFBBFFA055E8FD5FFAF5FD16" box="[1230,1404,735,758]" italics="true" pageId="11" pageNumber="42">M. boteltobagoe</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6D4D13FFBBFFA0520CFD7FFC20FCF6" authorityName="Serene" authorityYear="1973" box="[810,937,767,790]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B919EA82FFBBFFA0520CFD7FFC20FCF6" box="[810,937,767,790]" italics="true" pageId="11" pageNumber="42">M. holthuisi</emphasis>
</taxonomicName>
are distinct species
<emphasis id="B919EA82FFBBFFA0555DFD7FFB48FCF6" box="[1147,1217,767,790]" italics="true" pageId="11" pageNumber="42">contra</emphasis>
the opinion of
<bibRefCitation id="EFFC4B61FFBBFFA05479FD7EFCFAFCD5" author="Barnes R" pageId="11" pageNumber="42" pagination="133 - 151" refId="ref11865" refString="Barnes R. S. K., 1976. Contributions towards a revision of Macrophthalmus (Crustacea: Brachyura), VIII. A re-examination of the M. telescopicus (Owen) complex; the status of M. laevis A. Milne Edwards; and the affinities of M. holthuisi Serene. Zoologische Mededelingen, 50: 133 - 151." type="journal article" year="1976">Barnes (1976)</bibRefCitation>
. Molecular sequence data (
<bibRefCitation id="EFFC4B61FFBBFFA055A1FC9EFAC7FCD5" author="Kitaura, J" box="[1159,1358,798,822]" pageId="11" pageNumber="42" pagination="45 - 59" refId="ref13284" refString="Kitaura, J., M. Nishida &amp; K. Wada, 2006. The evolution of social behaviour in sentinel crabs (Macrophthalmus): implications from molecular phylogeny. Biological Journal of the Linnean Society, 88: 45 - 59." type="journal article" year="2006">Kitaura et al., 2006</bibRefCitation>
) suggest that
<taxonomicName id="4C6D4D13FFBBFFA0527EFCBFFC3AFCB6" authorityName="de Man" authorityYear="1888" box="[856,947,831,854]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="erato">
<emphasis id="B919EA82FFBBFFA0527EFCBFFC3AFCB6" box="[856,947,831,854]" italics="true" pageId="11" pageNumber="42">M. erato</emphasis>
</taxonomicName>
+
<taxonomicName id="4C6D4D13FFBBFFA052E9FCBFFBD7FCB6" authorityName="A. Milne Edwards" authorityYear="1873" box="[975,1118,831,854]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="quadratus">
<emphasis id="B919EA82FFBBFFA052E9FCBFFBD7FCB6" box="[975,1118,831,854]" italics="true" pageId="11" pageNumber="42">M. quadratus</emphasis>
</taxonomicName>
are only distantly related to
<taxonomicName id="4C6D4D13FFBBFFA054AAFCBFFC0EFC96" authorityName="Serene" authorityYear="1973" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B919EA82FFBBFFA054AAFCBFFC0EFC96" italics="true" pageId="11" pageNumber="42">M. holthuisi</emphasis>
</taxonomicName>
+
<taxonomicName id="4C6D4D13FFBBFFA05284FCDFFBD9FC96" baseAuthorityName="Sakai" baseAuthorityYear="1969" box="[930,1104,863,886]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="boteltobagoe">
<emphasis id="B919EA82FFBBFFA05284FCDFFBD9FC96" box="[930,1104,863,886]" italics="true" pageId="11" pageNumber="42">M. boteltobagoe</emphasis>
</taxonomicName>
, but instead are related to the
<taxonomicName id="4C6D4D13FFBBFFA054AAFCDFFCE3FC76" baseAuthorityName="Herbst" baseAuthorityYear="1804" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="B919EA82FFBBFFA054AAFCDFFCE3FC76" italics="true" pageId="11" pageNumber="42">M. brevis</emphasis>
</taxonomicName>
group of the subgenus
<taxonomicName id="4C6D4D13FFBBFFA0554CFCFFFA96FC76" box="[1130,1311,895,918]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B919EA82FFBBFFA0554CFCFFFA96FC76" box="[1130,1311,895,918]" italics="true" pageId="11" pageNumber="42">Macrophthalmus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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