1299 lines
157 KiB
XML
1299 lines
157 KiB
XML
<document id="5ACFBAB16CF7145B146A8545F1CEDE1F" ID-CLB-Dataset="9394" ID-DOI="10.11646/zootaxa.4890.4.5" ID-GBIF-Dataset="ea102704-86a3-47e5-8320-3424157f7232" ID-ISSN="1175-5326" ID-Zenodo-Dep="4306554" ID-ZooBank="919D8D2B-1FE0-45EB-A6CC-9B48FF6F22E7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1607103483912" checkinUser="plazi" docAuthor="Martínez-Revelo, Diego Esteban, Lopera-Toro, Alejandro & Medina, Claudia A." docDate="2020" docId="03D46376FFA7EA2F2FF0F1A5FD8AFA62" docLanguage="en" docName="zootaxa.4890.4.5.pdf" docOrigin="Zootaxa 4890 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scatimus strenua Martinez-Revelo, Lopera-Toro, & Medina 2020, new species" docType="treatment" docVersion="10" lastPageNumber="529" masterDocId="FFED1B0EFFA5EA272F67F252FFE6FFFF" masterDocTitle="Review of Scatimus Erichson (Coleoptera: Scarabaeidae) in Colombia with the description of a new species" masterLastPageNumber="534" masterPageNumber="521" pageNumber="523" updateTime="1698847185713" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="73E05E2AC5D1A5991B5559541DE3184A">Review of Scatimus Erichson (Coleoptera: Scarabaeidae) in Colombia with the description of a new species</mods:title>
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<mods:namePart id="2F8BEC255E0913246466A057EDDB9B7B">Martínez-Revelo, Diego Esteban</mods:namePart>
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<mods:affiliation id="77347D898657A4C7DA3A0111CCFFC6B0">Asociación Grupo de Amigos para la Investigación y Conservación de las Aves, Calle 11 A # No. 32 - 21, Pasto, Nariño.</mods:affiliation>
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<mods:namePart id="2248C7F925867A848761FA6B25EA1251">Lopera-Toro, Alejandro</mods:namePart>
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<mods:affiliation id="0CA294DDF0828B67C89D688FF1196BEF">Asociación Grupo de Amigos para la Investigación y Conservación de las Aves, Calle 11 A # No. 32 - 21, Pasto, Nariño. & alejandro. lopera @ gmail. com; https: // orcid. org / 0000 - 0001 - 5097 - 1655</mods:affiliation>
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<mods:namePart id="5B5CCD354BAECA7773B3DD8BB0BCC4B0">Medina, Claudia A.</mods:namePart>
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<mods:affiliation id="1F0BC0248E30C5CF2377E10EFEC0B607">Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, Claustro de San Agustín, Carrera 8 No. 15 - 08, Villa de Leyva, Boyacá, Colombia. camedina @ humboldt. org. co; https: // orcid. org / 0000 - 0002 - 7714 - 9220</mods:affiliation>
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<mods:title id="B61C2C0CADBB187A2DAD23D9972E7E74">Zootaxa</mods:title>
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<mods:part id="18DB61E7BA0F8358E184B009D7B23D5B">
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<mods:date id="CB46A6B9C4C34C93ED959CDFB3E7A985">2020</mods:date>
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<taxonomicName id="4C7DA9E3FFA7EA252FF0F1A5FC67FBEF" ID-CoL="7YYB3" authority="Martinez-Revelo, Lopera-Toro, & Medina" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[151,897,1014,1041]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="strenua" status="sp. nov.">
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<emphasis id="B9090E72FFA7EA252FF0F1A5FE8EFBEE" bold="true" box="[151,360,1015,1041]" italics="true" pageId="2" pageNumber="523">Scatimus strenua</emphasis>
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Martínez-Revelo, Lopera-Toro, & Medina
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,
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(
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A–B, 2A–B, 3B, D–F, H, 4D, H, L, P, 5–6)
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</paragraph>
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<emphasis id="B9090E72FFA7EA252FF0F631FED8FB83" bold="true" box="[151,318,1122,1149]" pageId="2" pageNumber="523">Type material</emphasis>
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.
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. “
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:
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:
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<collectingCounty id="62A3AAECFFA7EA252DA8F631FCAEFB82" box="[719,840,1123,1149]" pageId="2" pageNumber="523">El Carmen</collectingCounty>
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/ de Viboral,
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<collectingMunicipality id="6BA6481AFFA7EA252C86F631FB70FB82" box="[993,1174,1123,1149]" pageId="2" pageNumber="523">Finca la Meseta</collectingMunicipality>
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montano bajo, / 5° 52’/
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,
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.Exc.H. #
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_24, 4.xii. / 2016,
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<collectorName id="2688B7B6FFA7EA252B16F6D5FB00FB5E" box="[1137,1254,1159,1185]" pageId="2" pageNumber="523">A. Lopera</collectorName>
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,
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<collectorName id="2688B7B6FFA7EA252B95F6D5FA90FB5E" box="[1266,1398,1159,1185]" pageId="2" pageNumber="523">J. Cárdenas</collectorName>
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”— male (IAvH-E-196594). Printed on red label: “
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<taxonomicName id="4C7DA9E3FFA7EA252DF9F6F9FAD5FB3A" authority="Martinez-Revelo, Lopera-Toro & Medina" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[670,1331,1195,1221]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="strenua" status="sp. nov.">
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<emphasis id="B9090E72FFA7EA252DF9F6F9FCBAFB3B" box="[670,860,1195,1221]" italics="true" pageId="2" pageNumber="523">Scatimus strenua</emphasis>
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Martínez-Revelo, Lopera-Toro & Medina
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”
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<typeStatus id="54C66CC2FFA7EA252FF0F69DFEE9FB16" box="[151,271,1231,1257]" pageId="2" pageNumber="523" type="paratype">Paratypes</typeStatus>
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</emphasis>
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[
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<specimenCount id="9D7B19E9FFA7EA252E7BF69DFE93FB16" box="[284,373,1231,1257]" pageId="2" pageNumber="523" type="male">7 males</specimenCount>
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,
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<specimenCount id="9D7B19E9FFA7EA252E18F69CFE17FB16" box="[383,497,1230,1257]" pageId="2" pageNumber="523" type="female">5 females</specimenCount>
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].
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:
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<collectingRegion id="49B91C82FFA7EA252D1DF69DFD09FB16" box="[634,751,1231,1257]" country="Colombia" name="Antioquia" pageId="2" pageNumber="523">Antioquia</collectingRegion>
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:
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,
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<collectorName id="2688B7B6FFA7EA252C93F69DFB7CFB16" box="[1012,1178,1231,1257]" pageId="2" pageNumber="523">Finca San José</collectorName>
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,
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<collectorName id="2688B7B6FFA7EA252BC3F69DFB1EFB16" box="[1188,1272,1231,1257]" pageId="2" pageNumber="523">Bosque</collectorName>
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montano bajo,
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<location id="8EA284BBFFA7EA252FF0F6A1FDE0FAF2" LSID="urn:lsid:plazi:treatment:03D46376FFA7EA2F2FF0F1A5FD8AFA62:8EA284BBFFA7EA252FF0F6A1FDE0FAF2" box="[151,518,1267,1293]" country="Colombia" municipality="El Carmen de Viboral" name="Trampa de Excremento Humano" pageId="2" pageNumber="523" stateProvince="Antioquia">Trampa de Excremento Humano</location>
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</materialsCitation>
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, #
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_48,
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,
|
||
<materialsCitation id="3B15D83DFFA7EA252B58F6A1FD7CFACE" ID-GBIF-Occurrence="2990035330" collectingDate="2016-12-02" collectionCode="WGS" country="Colombia" pageId="2" pageNumber="523" specimenCode="WGS84" specimenCount="2" specimenCount-female="2" stateProvince="Antioquia" typeStatus="paratype">
|
||
<specimenCode id="DBDB7A1BFFA7EA252B58F6A1FB7AFAF2" box="[1087,1180,1267,1293]" collectionCode="WGS" pageId="2" pageNumber="523">WGS84</specimenCode>
|
||
,
|
||
<date id="FFC3F4A0FFA7EA252BC0F6A1FAFCFAF2" box="[1191,1306,1267,1293]" pageId="2" pageNumber="523" value="2016-12-02">
|
||
<collectingDate id="EF870D48FFA7EA252BC0F6A1FAFCFAF2" box="[1191,1306,1267,1293]" pageId="2" pageNumber="523" value="2016-12-02">2.xii.2016</collectingDate>
|
||
</date>
|
||
, A. Lopera, J. Cárdenas—
|
||
<specimenCount id="9D7B19E9FFA7EA252E51F745FE47FACE" box="[310,417,1303,1329]" pageId="2" pageNumber="523" type="female">2 females</specimenCount>
|
||
(IAvH-E-196589-90)
|
||
</materialsCitation>
|
||
; #
|
||
<materialsCitation id="3B15D83DFFA7EA252DD3F745FB28FACE" ID-GBIF-Occurrence="2990035320" box="[692,1230,1302,1329]" collectionCode="T" country="Colombia" elevation="1214" latitude="5.8879137" longLatPrecision="1" longitude="-75.19461" pageId="2" pageNumber="523" specimenCode="T6" specimenCount="1" stateProvince="Antioquia" typeStatus="paratype">
|
||
<specimenCode id="DBDB7A1BFFA7EA252DD3F745FD35FACE" box="[692,723,1303,1329]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="523">T6</specimenCode>
|
||
_24,
|
||
<geoCoordinate id="EE49B4A7FFA7EA252C6CF744FC4BFACE" box="[779,941,1302,1329]" degrees="5" direction="north" minutes="53" orientation="latitude" pageId="2" pageNumber="523" precision="1" seconds="16.49" value="5.8879137">5°53’16.49’’N</geoCoordinate>
|
||
<geoCoordinate id="EE49B4A7FFA7EA252CD2F744FB8CFACE" box="[949,1130,1302,1329]" degrees="75" direction="west" minutes="11" orientation="longitude" pageId="2" pageNumber="523" precision="1" seconds="40.59" value="-75.19461">75°11’40.59’’W</geoCoordinate>
|
||
,
|
||
<quantity id="4C857F85FFA7EA252B11F745FB28FACE" box="[1142,1230,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.214" pageId="2" pageNumber="523" unit="m" value="1214.0">
|
||
<elevation id="00503553FFA7EA252B11F745FB28FACE" box="[1142,1230,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.214" pageId="2" pageNumber="523" unit="m" value="1214.0">1214 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
,
|
||
<date id="FFC3F4A0FFA7EA252BBDF745FAAAFACE" box="[1242,1356,1303,1329]" pageId="2" pageNumber="523" value="2016-12-01">1.xii.2016</date>
|
||
—
|
||
<specimenCount id="9D7B19E9FFA7EA252A0BF745FF36FAAA" pageId="2" pageNumber="523" type="female">1 female</specimenCount>
|
||
, (IAvH-E-196652); #
|
||
<materialsCitation id="3B15D83DFFA7EA252EA5F769FC3FFAAA" ID-GBIF-Occurrence="2990035305" box="[450,985,1338,1365]" collectionCode="T" country="Colombia" elevation="1237" latitude="5.887372" longLatPrecision="1" longitude="-75.194824" pageId="2" pageNumber="523" specimenCode="T7" specimenCount="1" stateProvince="Antioquia" typeStatus="paratype">
|
||
<specimenCode id="DBDB7A1BFFA7EA252EA5F769FE06FAAA" box="[450,480,1339,1365]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="523">T7</specimenCode>
|
||
_24,
|
||
<geoCoordinate id="EE49B4A7FFA7EA252D7FF768FD5CFAAA" box="[536,698,1338,1365]" degrees="5" direction="north" minutes="53" orientation="latitude" pageId="2" pageNumber="523" precision="1" seconds="14.54" value="5.887372">5°53’14.54’’N</geoCoordinate>
|
||
<geoCoordinate id="EE49B4A7FFA7EA252DA6F768FC91FAAA" box="[705,887,1338,1365]" degrees="75" direction="west" minutes="11" orientation="longitude" pageId="2" pageNumber="523" precision="1" seconds="41.36" value="-75.194824">75°11’41.36’’W</geoCoordinate>
|
||
,
|
||
<quantity id="4C857F85FFA7EA252CE5F769FC3FFAAA" box="[898,985,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="1.237" pageId="2" pageNumber="523" unit="m" value="1237.0">
|
||
<elevation id="00503553FFA7EA252CE5F769FC3FFAAA" box="[898,985,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="1.237" pageId="2" pageNumber="523" unit="m" value="1237.0">1237 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
,
|
||
<date id="FFC3F4A0FFA7EA252C82F769FBB1FAAA" box="[997,1111,1339,1365]" pageId="2" pageNumber="523" value="2016-12-01">1.xii.2016</date>
|
||
—
|
||
<specimenCount id="9D7B19E9FFA7EA252B10F769FB27FAAA" box="[1143,1217,1339,1365]" pageId="2" pageNumber="523" type="male">1 male</specimenCount>
|
||
, (IAvH-E-196588); #
|
||
<materialsCitation id="3B15D83DFFA7EA252FC1F70DFD57FA86" ID-GBIF-Occurrence="2990035331" box="[166,689,1374,1401]" collectingDate="2016-12-02" collectionCode="T" country="Colombia" pageId="2" pageNumber="523" specimenCode="T7" specimenCount="1" specimenCount-male="1" stateProvince="Antioquia" typeStatus="paratype">
|
||
<specimenCode id="DBDB7A1BFFA7EA252FC1F70DFF23FA86" box="[166,197,1375,1401]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="523">T7</specimenCode>
|
||
_48,
|
||
<date id="FFC3F4A0FFA7EA252F98F70DFE94FA86" box="[255,370,1375,1401]" pageId="2" pageNumber="523" value="2016-12-02">
|
||
<collectingDate id="EF870D48FFA7EA252F98F70DFE94FA86" box="[255,370,1375,1401]" pageId="2" pageNumber="523" value="2016-12-02">2.xii.2016</collectingDate>
|
||
</date>
|
||
—
|
||
<specimenCount id="9D7B19E9FFA7EA252EF5F70DFE3DFA86" box="[402,475,1375,1401]" pageId="2" pageNumber="523" type="male">1 male</specimenCount>
|
||
(IAvH-E-196591)
|
||
</materialsCitation>
|
||
; #
|
||
<materialsCitation id="3B15D83DFFA7EA252DA8F70DFB14FA86" ID-GBIF-Occurrence="2990035309" box="[719,1266,1374,1401]" collectionCode="T" country="Colombia" elevation="1250" latitude="5.886947" longLatPrecision="1" longitude="-75.1949" pageId="2" pageNumber="523" specimenCode="T8" specimenCount="1" stateProvince="Antioquia" typeStatus="paratype">
|
||
<specimenCode id="DBDB7A1BFFA7EA252DA8F70DFD0AFA86" box="[719,748,1375,1401]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="523">T8</specimenCode>
|
||
_48,
|
||
<geoCoordinate id="EE49B4A7FFA7EA252C40F70CFC2CFA86" box="[807,970,1374,1401]" degrees="5" direction="north" minutes="53" orientation="latitude" pageId="2" pageNumber="523" precision="1" seconds="13.01" value="5.886947">5°53’13.01’’N</geoCoordinate>
|
||
<geoCoordinate id="EE49B4A7FFA7EA252CB3F70CFB6FFA86" box="[980,1161,1374,1401]" degrees="75" direction="west" minutes="11" orientation="longitude" pageId="2" pageNumber="523" precision="1" seconds="41.63" value="-75.1949">75°11’41.63’’W</geoCoordinate>
|
||
,
|
||
<quantity id="4C857F85FFA7EA252BFFF70CFB14FA86" box="[1176,1266,1374,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="523" unit="m" value="1250.0">
|
||
<elevation id="00503553FFA7EA252BFFF70CFB14FA86" box="[1176,1266,1374,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="523" unit="m" value="1250.0">1250 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
,
|
||
<date id="FFC3F4A0FFA7EA252A66F70DFA92FA86" box="[1281,1396,1375,1401]" pageId="2" pageNumber="523" value="2016-12-02">2.xii.2016</date>
|
||
—
|
||
<specimenCount id="9D7B19E9FFA7EA252AF4F70DFF2BFA62" pageId="2" pageNumber="523" type="male">1 male</specimenCount>
|
||
(IAvH-E-196653); #
|
||
<materialsCitation id="3B15D83DFFA7EA252ED9F7D1FC3DFA62" ID-GBIF-Occurrence="2990035323" box="[446,987,1410,1437]" collectionCode="T" country="Colombia" elevation="1125" latitude="5.8869667" longLatPrecision="1" longitude="-75.19438" pageId="2" pageNumber="523" specimenCode="T9" specimenCount="1" stateProvince="Antioquia" typeStatus="paratype">
|
||
<specimenCode id="DBDB7A1BFFA7EA252ED9F7D1FE3BFA62" box="[446,477,1411,1437]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="523">T9</specimenCode>
|
||
_24,
|
||
<geoCoordinate id="EE49B4A7FFA7EA252D71F7D0FD5EFA62" box="[534,696,1410,1437]" degrees="5" direction="north" minutes="53" orientation="latitude" pageId="2" pageNumber="523" precision="1" seconds="13.08" value="5.8869667">5°53’13.08’’N</geoCoordinate>
|
||
<geoCoordinate id="EE49B4A7FFA7EA252DA6F7D0FC90FA62" box="[705,886,1410,1437]" degrees="75" direction="west" minutes="11" orientation="longitude" pageId="2" pageNumber="523" precision="1" seconds="39.77" value="-75.19438">75°11’39.77’’W</geoCoordinate>
|
||
,
|
||
<quantity id="4C857F85FFA7EA252CE3F7D0FC3DFA62" box="[900,987,1410,1437]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="2" pageNumber="523" unit="m" value="1125.0">
|
||
<elevation id="00503553FFA7EA252CE3F7D0FC3DFA62" box="[900,987,1410,1437]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="2" pageNumber="523" unit="m" value="1125.0">1125 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
,
|
||
<date id="FFC3F4A0FFA7EA252C8EF7D1FBBCFA62" box="[1001,1114,1411,1437]" pageId="2" pageNumber="523" value="2016-12-01">1.xii.2016</date>
|
||
—
|
||
<specimenCount id="9D7B19E9FFA7EA252B1CF7D1FB25FA62" box="[1147,1219,1411,1437]" pageId="2" pageNumber="523" type="male">1 male</specimenCount>
|
||
(IAvH-E-196586); #
|
||
<materialsCitation id="3B15D83DFFA7EA252FC1F7F5FB68FA3E" ID-GBIF-Occurrence="2990035303" box="[166,1166,1446,1473]" collectingDate="2016-12-02" collectionCode="T" collectorName="Bosque" country="Colombia" location="Finca la Meseta" pageId="2" pageNumber="523" specimenCode="T9" specimenCount="2" specimenCount-male="2" stateProvince="Antioquia" typeStatus="paratype">
|
||
<specimenCode id="DBDB7A1BFFA7EA252FC1F7F5FF23FA3E" box="[166,197,1447,1473]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="523">T9</specimenCode>
|
||
_48,
|
||
<date id="FFC3F4A0FFA7EA252F9DF7F5FE8DFA3E" box="[250,363,1447,1473]" pageId="2" pageNumber="523" value="2016-12-02">
|
||
<collectingDate id="EF870D48FFA7EA252F9DF7F5FE8DFA3E" box="[250,363,1447,1473]" pageId="2" pageNumber="523" value="2016-12-02">2.xii.2016</collectingDate>
|
||
</date>
|
||
,—
|
||
<specimenCount id="9D7B19E9FFA7EA252EE8F7F5FE05FA3E" box="[399,483,1447,1473]" pageId="2" pageNumber="523" type="male">2 males</specimenCount>
|
||
(IAvH-E-196592-93);
|
||
<location id="8EA284BBFFA7EA252D85F7F5FC74FA3E" LSID="urn:lsid:plazi:treatment:03D46376FFA7EA2F2FF0F1A5FD8AFA62:8EA284BBFFA7EA252D85F7F5FC74FA3E" box="[738,914,1447,1473]" country="Colombia" name="Finca la Meseta" pageId="2" pageNumber="523" stateProvince="Antioquia">Finca la Meseta</location>
|
||
,
|
||
<collectorName id="2688B7B6FFA7EA252CFCF7F5FC09FA3E" box="[923,1007,1447,1473]" pageId="2" pageNumber="523">Bosque</collectorName>
|
||
montano bajo
|
||
</materialsCitation>
|
||
, #
|
||
<materialsCitation id="3B15D83DFFA7EA252BC0F7F5FEAAFA1A" ID-GBIF-Occurrence="2990035304" collectionCode="T" country="Colombia" latitude="5.8827972" longLatPrecision="1" longitude="-75.1987" pageId="2" pageNumber="523" specimenCode="T1" specimenCount="1" stateProvince="Antioquia" typeStatus="paratype">
|
||
<specimenCode id="DBDB7A1BFFA7EA252BC0F7F5FB24FA3E" box="[1191,1218,1447,1473]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="523">T1</specimenCode>
|
||
_48,
|
||
<geoCoordinate id="EE49B4A7FFA7EA252B9DF7F4FA7BFA3E" box="[1274,1437,1446,1473]" degrees="5" direction="north" minutes="52" orientation="latitude" pageId="2" pageNumber="523" precision="1" seconds="58.07" value="5.8827972">5°52’58.07’’N</geoCoordinate>
|
||
<geoCoordinate id="EE49B4A7FFA7EA252FF0F798FEAAFA1A" box="[151,332,1482,1509]" degrees="75" direction="west" minutes="11" orientation="longitude" pageId="2" pageNumber="523" precision="1" seconds="55.32" value="-75.1987">75°11’55.32’’W</geoCoordinate>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B15D83DFFA7EA252E30F799FC2DFA1A" ID-GBIF-Occurrence="2990035306" box="[343,971,1482,1509]" collectingDate="2016-12-05" collectionCode="WGS" country="Colombia" pageId="2" pageNumber="523" specimenCode="WGS84, 1437" specimenCount="1" specimenCount-male="1" stateProvince="Antioquia" typeStatus="paratype">
|
||
<specimenCode id="DBDB7A1BFFA7EA252E30F799FE52FA1A" box="[343,436,1483,1509]" collectionCode="WGS" pageId="2" pageNumber="523">WGS84</specimenCode>
|
||
,
|
||
<specimenCode id="DBDB7A1BFFA7EA252ED8F799FE11FA1A" box="[447,503,1483,1509]" collectionCode="WGS" pageId="2" pageNumber="523">1437</specimenCode>
|
||
m,
|
||
<date id="FFC3F4A0FFA7EA252D47F798FD74FA1A" box="[544,658,1482,1509]" pageId="2" pageNumber="523" value="2016-12-05">
|
||
<collectingDate id="EF870D48FFA7EA252D47F798FD74FA1A" box="[544,658,1482,1509]" pageId="2" pageNumber="523" value="2016-12-05">5.xii.2016</collectingDate>
|
||
</date>
|
||
—
|
||
<specimenCount id="9D7B19E9FFA7EA252DD4F798FD1EFA1A" box="[691,760,1482,1509]" pageId="2" pageNumber="523" type="male">1 male</specimenCount>
|
||
(IAvH-E-196595)
|
||
</materialsCitation>
|
||
; #
|
||
<materialsCitation id="3B15D83DFFA7EA252C83F799FF09F9F6" ID-GBIF-Occurrence="2990035326" collectionCode="T" country="Colombia" elevation="1494" latitude="5.882336" longLatPrecision="1" longitude="-75.19853" pageId="2" pageNumber="523" specimenCode="T2" specimenCount="1" stateProvince="Antioquia" typeStatus="paratype">
|
||
<specimenCode id="DBDB7A1BFFA7EA252C83F799FBE2FA1A" box="[996,1028,1483,1509]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="523">T2</specimenCode>
|
||
_48,
|
||
<geoCoordinate id="EE49B4A7FFA7EA252B5DF798FB3AFA1A" box="[1082,1244,1482,1509]" degrees="5" direction="north" minutes="52" orientation="latitude" pageId="2" pageNumber="523" precision="1" seconds="56.41" value="5.882336">5°52’56.41’’N</geoCoordinate>
|
||
<geoCoordinate id="EE49B4A7FFA7EA252B85F798FA7EFA1A" box="[1250,1432,1482,1509]" degrees="75" direction="west" minutes="11" orientation="longitude" pageId="2" pageNumber="523" precision="1" seconds="54.70" value="-75.19853">75°11’54.70’’W</geoCoordinate>
|
||
,
|
||
<quantity id="4C857F85FFA7EA252FF0F7BDFF09F9F6" box="[151,239,1519,1545]" metricMagnitude="3" metricUnit="m" metricValue="1.494" pageId="2" pageNumber="523" unit="m" value="1494.0">
|
||
<elevation id="00503553FFA7EA252FF0F7BDFF09F9F6" box="[151,239,1519,1545]" metricMagnitude="3" metricUnit="m" metricValue="1.494" pageId="2" pageNumber="523" unit="m" value="1494.0">1494 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B15D83DFFA7EA252F9AF7BCFB73F9F6" ID-GBIF-Occurrence="2990035327" box="[253,1173,1518,1545]" collectingDate="2016-12-05" collectorName="Bosque" country="Colombia" location="Finca la Samaria" pageId="2" pageNumber="523" specimenCount="1" specimenCount-female="1" stateProvince="Antioquia" typeStatus="paratype">
|
||
<date id="FFC3F4A0FFA7EA252F9AF7BCFE89F9F6" box="[253,367,1518,1545]" pageId="2" pageNumber="523" value="2016-12-05">
|
||
<collectingDate id="EF870D48FFA7EA252F9AF7BCFE89F9F6" box="[253,367,1518,1545]" pageId="2" pageNumber="523" value="2016-12-05">5.xii.2016</collectingDate>
|
||
</date>
|
||
—
|
||
<specimenCount id="9D7B19E9FFA7EA252EE8F7BCFE0AF9F6" box="[399,492,1518,1545]" pageId="2" pageNumber="523" type="female">1 female</specimenCount>
|
||
(IAvH-E-196596);
|
||
<location id="8EA284BBFFA7EA252DAAF7BDFC69F9F6" LSID="urn:lsid:plazi:treatment:03D46376FFA7EA2F2FF0F1A5FD8AFA62:8EA284BBFFA7EA252DAAF7BDFC69F9F6" box="[717,911,1519,1545]" country="Colombia" name="Finca la Samaria" pageId="2" pageNumber="523" stateProvince="Antioquia">Finca la Samaria</location>
|
||
,
|
||
<collectorName id="2688B7B6FFA7EA252CFCF7BDFC09F9F6" box="[923,1007,1519,1545]" pageId="2" pageNumber="523">Bosque</collectorName>
|
||
montano bajo
|
||
</materialsCitation>
|
||
, #
|
||
<materialsCitation id="3B15D83DFFA7EA252BD7F7BDFE79F9D2" ID-GBIF-Occurrence="2990035315" collectionCode="T" country="Colombia" elevation="1166" latitude="5.888889" longLatPrecision="1" longitude="-75.187004" pageId="2" pageNumber="523" specimenCode="T2" specimenCount="1" stateProvince="Antioquia" typeStatus="paratype">
|
||
<specimenCode id="DBDB7A1BFFA7EA252BD7F7BDFB29F9F6" box="[1200,1231,1519,1545]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="523">T2</specimenCode>
|
||
_72,
|
||
<geoCoordinate id="EE49B4A7FFA7EA252A6FF7BCFA7AF9F6" box="[1288,1436,1518,1545]" degrees="5" direction="north" minutes="53" orientation="latitude" pageId="2" pageNumber="523" precision="1" seconds="20.0" value="5.888889">5°53’20.0’’N</geoCoordinate>
|
||
<geoCoordinate id="EE49B4A7FFA7EA252FF0F440FED8F9D2" box="[151,318,1554,1581]" degrees="75" direction="west" minutes="11" orientation="longitude" pageId="2" pageNumber="523" precision="1" seconds="13.2" value="-75.187004">75°11’13.2’’W</geoCoordinate>
|
||
,
|
||
<quantity id="4C857F85FFA7EA252E2DF441FE79F9D2" box="[330,415,1555,1581]" metricMagnitude="3" metricUnit="m" metricValue="1.166" pageId="2" pageNumber="523" unit="m" value="1166.0">
|
||
<elevation id="00503553FFA7EA252E2DF441FE79F9D2" box="[330,415,1555,1581]" metricMagnitude="3" metricUnit="m" metricValue="1.166" pageId="2" pageNumber="523" unit="m" value="1166.0">1166 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
,
|
||
<date id="FFC3F4A0FFA7EA252ECDF441FDFBF9D2" box="[426,541,1555,1581]" pageId="2" pageNumber="523" value="2016-12-02">2.xii.2016</date>
|
||
—
|
||
<specimenCount id="9D7B19E9FFA7EA252D5AF441FD7EF9D2" box="[573,664,1555,1581]" pageId="2" pageNumber="523" type="female">1 female</specimenCount>
|
||
(IAvH-E-196585). Printed on yellow labels: “
|
||
<taxonomicName id="4C7DA9E3FFA7EA252BFBF441FABDF9D3" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[1180,1371,1555,1581]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA7EA252BFBF441FABDF9D3" box="[1180,1371,1555,1581]" italics="true" pageId="2" pageNumber="523">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="54C66CC2FFA7EA252A06F441FF21F9AE" pageId="2" pageNumber="523" type="paratype">Paratype</typeStatus>
|
||
Martínez-Revelo, Lopera-Toro & Medina
|
||
<taxonomicNameLabel id="A23AB309FFA7EA252DC0F465FCE6F9AE" box="[679,768,1591,1617]" pageId="2" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
|
||
”
|
||
</paragraph>
|
||
<paragraph id="8BC2D260FFA7EA252FA0F409FC1EF966" blockId="2.[151,1437,1122,2013]" pageId="2" pageNumber="523">
|
||
<emphasis id="B9090E72FFA7EA252FA0F409FDCCF98B" bold="true" box="[199,554,1626,1653]" pageId="2" pageNumber="523">Non-type material examined.</emphasis>
|
||
<materialsCitation id="3B15D83DFFA7EA252D53F409FC12F966" ID-GBIF-Occurrence="2990035324" collectingDate="2009-05-03" collectionCode="MEFLG" collectorName="Grisales, S." country="Colombia" elevation="1310" latitude="6.975528" location="Vereda Salazar" longLatPrecision="1" longitude="-75.084145" municipality="Amalfi" pageId="2" pageNumber="523" specimenCount="1" stateProvince="Antioquia">
|
||
<collectingCountry id="F36A92F0FFA7EA252D53F409FD41F98A" box="[564,679,1627,1653]" name="Colombia" pageId="2" pageNumber="523">Colombia</collectingCountry>
|
||
:
|
||
<collectingRegion id="49B91C82FFA7EA252DD4F409FCCEF98A" box="[691,808,1627,1653]" country="Colombia" name="Antioquia" pageId="2" pageNumber="523">Antioquia</collectingRegion>
|
||
:
|
||
<collectingMunicipality id="6BA6481AFFA7EA252C52F409FC61F98A" box="[821,903,1627,1653]" pageId="2" pageNumber="523">Amalfi</collectingMunicipality>
|
||
,
|
||
<location id="8EA284BBFFA7EA252CF2F409FBD9F98A" LSID="urn:lsid:plazi:treatment:03D46376FFA7EA2F2FF0F1A5FD8AFA62:8EA284BBFFA7EA252CF2F409FBD9F98A" box="[917,1087,1627,1653]" country="Colombia" latitude="6.975528" longLatPrecision="1" longitude="-75.084145" municipality="Amalfi" name="Vereda Salazar" pageId="2" pageNumber="523" stateProvince="Antioquia">Vereda Salazar</location>
|
||
(Bodega Vieja),
|
||
<geoCoordinate id="EE49B4A7FFA7EA252A6FF408FA7AF98A" box="[1288,1436,1626,1653]" degrees="6" direction="north" minutes="58" orientation="latitude" pageId="2" pageNumber="523" precision="1" seconds="31.9" value="6.975528">6°58’31.9’’N</geoCoordinate>
|
||
<geoCoordinate id="EE49B4A7FFA7EA252FF0F42CFEC5F966" box="[151,291,1662,1689]" degrees="75" direction="west" minutes="5" orientation="longitude" pageId="2" pageNumber="523" precision="1" seconds="2.9" value="-75.084145">75°5’2.9’’W</geoCoordinate>
|
||
,
|
||
<quantity id="4C857F85FFA7EA252E48F42DFE60F966" box="[303,390,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="2" pageNumber="523" unit="m" value="1310.0">
|
||
<elevation id="00503553FFA7EA252E48F42DFE60F966" box="[303,390,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="2" pageNumber="523" unit="m" value="1310.0">1310 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFC3F4A0FFA7EA252EF5F42DFE12F966" box="[402,500,1663,1689]" pageId="2" pageNumber="523" value="2009-05-03">
|
||
<collectingDate id="EF870D48FFA7EA252EF5F42DFE12F966" box="[402,500,1663,1689]" pageId="2" pageNumber="523" value="2009-05-03">3.v.2009</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="2688B7B6FFA7EA252D67F42DFD98F966" box="[512,638,1663,1689]" pageId="2" pageNumber="523">Grisales, S.</collectorName>
|
||
—1 sex undetermined (
|
||
<collectionCode id="ED6C4AA5FFA7EA252CE1F42DFC0AF966" box="[902,1004,1663,1689]" country="Colombia" httpUri="http://grbio.org/cool/ct7t-ipa9" name="Museo Entomologico Francisco Luis Gallego" pageId="2" pageNumber="523">MEFLG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36781EBFFA7EA242FA0F4F0FD6AFF06" lastPageId="3" lastPageNumber="524" pageId="2" pageNumber="523" type="diagnosis">
|
||
<paragraph id="8BC2D260FFA7EA242FA0F4F0FD6AFF06" blockId="2.[151,1437,1122,2013]" lastBlockId="3.[151,1437,150,2013]" lastPageId="3" lastPageNumber="524" pageId="2" pageNumber="523">
|
||
<emphasis id="B9090E72FFA7EA252FA0F4F0FEA4F943" bold="true" box="[199,322,1698,1724]" pageId="2" pageNumber="523">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C7DA9E3FFA7EA252E2FF4F1FDE1F943" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[328,519,1699,1725]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA7EA252E2FF4F1FDE1F943" box="[328,519,1699,1725]" italics="true" pageId="2" pageNumber="523">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
is close to
|
||
<taxonomicName id="4C7DA9E3FFA7EA252DE3F4F1FD09F942" authorityName="Balthasar" authorityYear="1939" box="[644,751,1699,1725]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B9090E72FFA7EA252DE3F4F1FD74F942" box="[644,658,1699,1725]" italics="true" pageId="2" pageNumber="523">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA7EA252DC7F4F1FD09F942" box="[672,751,1699,1725]" italics="true" pageId="2" pageNumber="523">strandi</emphasis>
|
||
</taxonomicName>
|
||
but can be distinguished from this and other species of the
|
||
<taxonomicName id="4C7DA9E3FFA7EA252AE0F4F1FF00F91E" authorityName="Balthasar" authorityYear="1939" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B9090E72FFA7EA252AE0F4F1FA73F942" box="[1415,1429,1699,1725]" italics="true" pageId="2" pageNumber="523">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA7EA252FF0F495FF00F91E" box="[151,230,1735,1761]" italics="true" pageId="2" pageNumber="523">strandi</emphasis>
|
||
</taxonomicName>
|
||
clade by the following characters: anterior margin of metafemur enlarged and very sharp in
|
||
<taxonomicName id="4C7DA9E3FFA7EA252B8DF495FABFF91F" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[1258,1369,1735,1761]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA7EA252B8DF495FB1EF91E" box="[1258,1272,1735,1761]" italics="true" pageId="2" pageNumber="523">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA7EA252A61F495FABFF91F" box="[1286,1369,1735,1760]" italics="true" pageId="2" pageNumber="523">strenua</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<superScript id="7C087F28FFA7EA252A0FF495FA6AF92B" attach="none" box="[1384,1420,1734,1761]" fontSize="6" pageId="2" pageNumber="523">411</superScript>
|
||
), simply rounded in
|
||
<taxonomicName id="4C7DA9E3FFA7EA252E0EF4B9FE05F8FA" box="[361,483,1771,1797]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="cribosus">
|
||
<emphasis id="B9090E72FFA7EA252E0EF4B9FE05F8FA" box="[361,483,1771,1797]" italics="true" pageId="2" pageNumber="523">S. cribosus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<superScript id="7C087F28FFA7EA252E95F4B9FDF1F907" attach="none" box="[498,535,1770,1797]" fontSize="6" pageId="2" pageNumber="523">410</superScript>
|
||
). Median lobe of metaventrites anterior edge arcuate medially in
|
||
<taxonomicName id="4C7DA9E3FFA7EA252B94F4B9FA87F8FB" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[1267,1377,1771,1797]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA7EA252B94F4B9FAE7F8FA" box="[1267,1281,1771,1797]" italics="true" pageId="2" pageNumber="523">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA7EA252A69F4B9FA87F8FB" box="[1294,1377,1771,1796]" italics="true" pageId="2" pageNumber="523">strenua</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<superScript id="7C087F28FFA7EA252A17F4B8FA73F907" attach="none" box="[1392,1429,1770,1797]" fontSize="6" pageId="2" pageNumber="523">251</superScript>
|
||
,
|
||
<figureCitation id="1346CEE5FFA7EA252FF0F55DFF0AF8D6" box="[151,236,1807,1833]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[354,1233,187,1858]" captionTargetId="figure-22@6.[354,1233,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Scatimus pronotum, mesepimeron, metasternum, abdomen, and metafemur. A, Scatimus strandi pronotum (lateral view); B, Scatimus strenua pronotum (lateral view); C, Scatimus strandi mesepimeron (ventral view); D, Scatimus strenua mesepimeron (ventral view); E, Scatimus strenua metasternum (ventral view); F, Scatimus strenua abdomen; G, Scatimus erinnyos metafemur (ventral view); H, Scatimus strenua metafemur (ventral view). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4306560" httpUri="https://zenodo.org/record/4306560/files/figure.png" pageId="2" pageNumber="523">Fig. 3E</figureCitation>
|
||
), distinctly angular medially in
|
||
<taxonomicName id="4C7DA9E3FFA7EA252D34F55DFD32F8D6" authorityName="Genier & Kohlmann" authorityYear="2003" box="[595,724,1807,1833]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="pacificus">
|
||
<emphasis id="B9090E72FFA7EA252D34F55DFD32F8D6" box="[595,724,1807,1833]" italics="true" pageId="2" pageNumber="523">S. pacificus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA7EA252D85F55DFCBCF8D6" authorityName="Balthasar" authorityYear="1939" box="[738,858,1807,1833]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="furcatus">
|
||
<emphasis id="B9090E72FFA7EA252D85F55DFCBCF8D6" box="[738,858,1807,1833]" italics="true" pageId="2" pageNumber="523">S. furcatus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C7DA9E3FFA7EA252CF0F55DFBA3F8D6" authorityName="Genier & Kohlmann" authorityYear="2003" box="[919,1093,1807,1833]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="quadricuspis">
|
||
<emphasis id="B9090E72FFA7EA252CF0F55DFC43F8D6" box="[919,933,1807,1833]" italics="true" pageId="2" pageNumber="523">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA7EA252CD3F55DFBA3F8D6" box="[948,1093,1807,1833]" italics="true" pageId="2" pageNumber="523">quadricuspis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<superScript id="7C087F28FFA7EA252B33F55CFB9FF8E3" attach="none" box="[1108,1145,1806,1833]" fontSize="6" pageId="2" pageNumber="523">250</superScript>
|
||
). Body elongate in dorsal view in
|
||
<taxonomicName id="4C7DA9E3FFA7EA252F96F561FE86F8B3" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[241,352,1843,1869]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA7EA252F96F561FF19F8B2" box="[241,255,1843,1869]" italics="true" pageId="2" pageNumber="523">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA7EA252E6AF561FE86F8B3" box="[269,352,1843,1868]" italics="true" pageId="2" pageNumber="523">strenua</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<superScript id="7C087F28FFA7EA252E15F561FE63F8BF" attach="none" box="[370,389,1842,1869]" fontSize="6" pageId="2" pageNumber="523">11</superScript>
|
||
,
|
||
<figureCitation id="1346CEE5FFA7EA252EF3F561FE36F8B2" box="[404,464,1843,1869]" captionStart="FIGURE 1" captionStartId="4.[151,250,1877,1901]" captionTargetBox="[166,1426,636,1837]" captionTargetId="figure-253@4.[151,1436,630,1850]" captionTargetPageId="4" captionText="FIGURE 1. Species of Scatimus from Colombia.A, Scatimus strenua holotype łmale (Antioquia, 5°52’45.05’’N 75°12’2.33’’W); B, Scatimus strenua, paratype female (Antioquia, 5°53’16.68’’N 75°11’37.11’’W); C, Scatimus fernandezi male (Casanare, 5°16’20.9”N 72°24’2.6”W); D, Scatimus ovatus male (Tolima, 5°8’58.3’’N 74°47’6.1’’W); E, Scatimus strandi male (Nariño, 0°30’N 77°13’W). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4306556" httpUri="https://zenodo.org/record/4306556/files/figure.png" pageId="2" pageNumber="523">Fig. 1</figureCitation>
|
||
A–B, E), body oval in
|
||
<taxonomicName id="4C7DA9E3FFA7EA252DB4F561FCD1F8B3" authorityName="Genier & Kohlmann" authorityYear="2003" box="[723,823,1843,1869]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="onorei">
|
||
<emphasis id="B9090E72FFA7EA252DB4F561FD07F8B2" box="[723,737,1843,1869]" italics="true" pageId="2" pageNumber="523">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA7EA252D88F561FCD1F8B3" box="[751,823,1843,1868]" italics="true" pageId="2" pageNumber="523">onorei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7DA9E3FFA7EA252C0AF561FC0FF8B3" authorityName="Kohlmann & Solis" authorityYear="1996" box="[877,1001,1843,1869]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="erinnyos">
|
||
<emphasis id="B9090E72FFA7EA252C0AF561FC9DF8B2" box="[877,891,1843,1869]" italics="true" pageId="2" pageNumber="523">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA7EA252CEEF561FC0FF8B3" box="[905,1001,1843,1868]" italics="true" pageId="2" pageNumber="523">erinnyos</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<superScript id="7C087F28FFA7EA252C9BF561FBF6F8BF" attach="none" box="[1020,1040,1842,1869]" fontSize="6" pageId="2" pageNumber="523">10</superScript>
|
||
,
|
||
<figureCitation id="1346CEE5FFA7EA252B79F561FBBDF8B2" box="[1054,1115,1843,1869]" captionStart="FIGURE 1" captionStartId="4.[151,250,1877,1901]" captionTargetBox="[166,1426,636,1837]" captionTargetId="figure-253@4.[151,1436,630,1850]" captionTargetPageId="4" captionText="FIGURE 1. Species of Scatimus from Colombia.A, Scatimus strenua holotype łmale (Antioquia, 5°52’45.05’’N 75°12’2.33’’W); B, Scatimus strenua, paratype female (Antioquia, 5°53’16.68’’N 75°11’37.11’’W); C, Scatimus fernandezi male (Casanare, 5°16’20.9”N 72°24’2.6”W); D, Scatimus ovatus male (Tolima, 5°8’58.3’’N 74°47’6.1’’W); E, Scatimus strandi male (Nariño, 0°30’N 77°13’W). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4306556" httpUri="https://zenodo.org/record/4306556/files/figure.png" pageId="2" pageNumber="523">Fig. 1</figureCitation>
|
||
C–D). Surface of abdominal ventrites with weakly defined microsculpture on segment 3–4 laterally in
|
||
<taxonomicName id="4C7DA9E3FFA7EA252CAAF505FBDDF88F" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[973,1083,1879,1905]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA7EA252CAAF505FC3DF88E" box="[973,987,1879,1905]" italics="true" pageId="2" pageNumber="523">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA7EA252C8FF505FBDDF88F" box="[1000,1083,1879,1904]" italics="true" pageId="2" pageNumber="523">strenua</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<superScript id="7C087F28FFA7EA252B2DF505FB89F89B" attach="none" box="[1098,1135,1878,1905]" fontSize="6" pageId="2" pageNumber="523">473</superScript>
|
||
,
|
||
<figureCitation id="1346CEE5FFA7EA252B1AF505FB36F88E" box="[1149,1232,1879,1905]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[354,1233,187,1858]" captionTargetId="figure-22@6.[354,1233,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Scatimus pronotum, mesepimeron, metasternum, abdomen, and metafemur. A, Scatimus strandi pronotum (lateral view); B, Scatimus strenua pronotum (lateral view); C, Scatimus strandi mesepimeron (ventral view); D, Scatimus strenua mesepimeron (ventral view); E, Scatimus strenua metasternum (ventral view); F, Scatimus strenua abdomen; G, Scatimus erinnyos metafemur (ventral view); H, Scatimus strenua metafemur (ventral view). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4306560" httpUri="https://zenodo.org/record/4306560/files/figure.png" pageId="2" pageNumber="523">Fig. 3F</figureCitation>
|
||
), only on segment
|
||
<quantity id="4C857F85FFA7EA252FF0F529FF24F86A" box="[151,194,1915,1941]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="2" pageNumber="523" unit="in" value="3.0">3 in</quantity>
|
||
<taxonomicName id="4C7DA9E3FFA7EA252FAEF529FECBF86B" authorityName="Genier & Kohlmann" authorityYear="2003" box="[201,301,1915,1941]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="onorei">
|
||
<emphasis id="B9090E72FFA7EA252FAEF529FF31F86A" box="[201,215,1915,1941]" italics="true" pageId="2" pageNumber="523">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA7EA252F82F529FECBF86B" box="[229,301,1915,1940]" italics="true" pageId="2" pageNumber="523">onorei</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<superScript id="7C087F28FFA7EA252E5BF529FE87F877" attach="none" box="[316,353,1914,1941]" fontSize="6" pageId="2" pageNumber="523">474</superScript>
|
||
). Lateral surface of mesepimeron lacking transverse carina medially in
|
||
<taxonomicName id="4C7DA9E3FFA7EA252BE2F529FB12F86B" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[1157,1268,1915,1941]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA7EA252BE2F529FB75F86A" box="[1157,1171,1915,1941]" italics="true" pageId="2" pageNumber="523">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA7EA252BC6F529FB12F86B" box="[1185,1268,1915,1940]" italics="true" pageId="2" pageNumber="523">strenua</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<superScript id="7C087F28FFA7EA252A64F529FACEF877" attach="none" box="[1283,1320,1914,1941]" fontSize="6" pageId="2" pageNumber="523">272</superScript>
|
||
,
|
||
<figureCitation id="1346CEE5FFA7EA252A51F529FA68F86A" box="[1334,1422,1915,1941]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[354,1233,187,1858]" captionTargetId="figure-22@6.[354,1233,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Scatimus pronotum, mesepimeron, metasternum, abdomen, and metafemur. A, Scatimus strandi pronotum (lateral view); B, Scatimus strenua pronotum (lateral view); C, Scatimus strandi mesepimeron (ventral view); D, Scatimus strenua mesepimeron (ventral view); E, Scatimus strenua metasternum (ventral view); F, Scatimus strenua abdomen; G, Scatimus erinnyos metafemur (ventral view); H, Scatimus strenua metafemur (ventral view). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4306560" httpUri="https://zenodo.org/record/4306560/files/figure.png" pageId="2" pageNumber="523">Fig. 3D</figureCitation>
|
||
), present in
|
||
<taxonomicName id="4C7DA9E3FFA7EA252E6CF5CDFE61F847" authorityName="Kohlmann & Solis" authorityYear="1996" box="[267,391,1951,1977]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="erinnyos">
|
||
<emphasis id="B9090E72FFA7EA252E6CF5CDFEFFF846" box="[267,281,1951,1977]" italics="true" pageId="2" pageNumber="523">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA7EA252E40F5CDFE61F847" box="[295,391,1951,1976]" italics="true" pageId="2" pageNumber="523">erinnyos</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7DA9E3FFA7EA252EDAF5CDFDCEF846" authorityName="Balthasar" authorityYear="1939" box="[445,552,1951,1977]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B9090E72FFA7EA252EDAF5CDFE2DF846" box="[445,459,1951,1977]" italics="true" pageId="2" pageNumber="523">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA7EA252EBEF5CDFDCEF846" box="[473,552,1951,1977]" italics="true" pageId="2" pageNumber="523">strandi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<superScript id="7C087F28FFA7EA252D50F5CDFDBAF853" attach="none" box="[567,604,1950,1977]" fontSize="6" pageId="2" pageNumber="523">271</superScript>
|
||
,
|
||
<figureCitation id="1346CEE5FFA7EA252D0DF5CDFD27F846" box="[618,705,1951,1977]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[354,1233,187,1858]" captionTargetId="figure-22@6.[354,1233,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Scatimus pronotum, mesepimeron, metasternum, abdomen, and metafemur. A, Scatimus strandi pronotum (lateral view); B, Scatimus strenua pronotum (lateral view); C, Scatimus strandi mesepimeron (ventral view); D, Scatimus strenua mesepimeron (ventral view); E, Scatimus strenua metasternum (ventral view); F, Scatimus strenua abdomen; G, Scatimus erinnyos metafemur (ventral view); H, Scatimus strenua metafemur (ventral view). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4306560" httpUri="https://zenodo.org/record/4306560/files/figure.png" pageId="2" pageNumber="523">Fig. 3C</figureCitation>
|
||
). Surface around dorsal portions of eye lacking microsculpture in
|
||
<taxonomicName id="4C7DA9E3FFA7EA252FF0F591FEE3F823" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[151,261,1987,2013]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA7EA252FF0F591FF43F822" box="[151,165,1987,2013]" italics="true" pageId="2" pageNumber="523">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA7EA252FD5F591FEE3F823" box="[178,261,1987,2012]" italics="true" pageId="2" pageNumber="523">strenua</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<superScript id="7C087F28FFA7EA252E73F591FECDF82F" attach="none" box="[276,299,1986,2013]" fontSize="6" pageId="2" pageNumber="523">90</superScript>
|
||
), present in
|
||
<taxonomicName id="4C7DA9E3FFA7EA252ED2F591FDD6F823" authorityName="Kohlmann & Solis" authorityYear="1996" box="[437,560,1987,2013]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="erinnyos">
|
||
<emphasis id="B9090E72FFA7EA252ED2F591FE25F822" box="[437,451,1987,2013]" italics="true" pageId="2" pageNumber="523">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA7EA252EB7F591FDD6F823" box="[464,560,1987,2012]" italics="true" pageId="2" pageNumber="523">erinnyos</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<superScript id="7C087F28FFA7EA252D27F591FDB1F82F" attach="none" box="[576,599,1986,2013]" fontSize="6" pageId="2" pageNumber="523">91</superScript>
|
||
) and surface of mesofemur and metafemur with minute puncturesthrough-out in
|
||
<taxonomicName id="4C7DA9E3FFA6EA242FB8F2C5FEA8FF4F" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[223,334,151,177]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA6EA242FB8F2C5FF0BFF4E" box="[223,237,151,177]" italics="true" pageId="3" pageNumber="524">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA6EA242F9CF2C5FEA8FF4F" box="[251,334,151,176]" italics="true" pageId="3" pageNumber="524">strenua</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<superScript id="7C087F28FFA6EA242E3AF2C5FE64FF5B" attach="none" box="[349,386,150,177]" fontSize="6" pageId="3" pageNumber="524">380</superScript>
|
||
,
|
||
<figureCitation id="1346CEE5FFA6EA242EF6F2C5FE0FFF4E" box="[401,489,151,177]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[354,1233,187,1858]" captionTargetId="figure-22@6.[354,1233,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Scatimus pronotum, mesepimeron, metasternum, abdomen, and metafemur. A, Scatimus strandi pronotum (lateral view); B, Scatimus strenua pronotum (lateral view); C, Scatimus strandi mesepimeron (ventral view); D, Scatimus strenua mesepimeron (ventral view); E, Scatimus strenua metasternum (ventral view); F, Scatimus strenua abdomen; G, Scatimus erinnyos metafemur (ventral view); H, Scatimus strenua metafemur (ventral view). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4306560" httpUri="https://zenodo.org/record/4306560/files/figure.png" pageId="3" pageNumber="524">Fig. 3H</figureCitation>
|
||
), with several coarse and umbilicate punctures at apex anteriorly in
|
||
<taxonomicName id="4C7DA9E3FFA6EA242B83F2C5FA86FF4F" authorityName="Kohlmann & Solis" authorityYear="1996" box="[1252,1376,151,177]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="erinnyos">
|
||
<emphasis id="B9090E72FFA6EA242B83F2C5FB14FF4E" box="[1252,1266,151,177]" italics="true" pageId="3" pageNumber="524">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA6EA242A67F2C5FA86FF4F" box="[1280,1376,151,176]" italics="true" pageId="3" pageNumber="524">erinnyos</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<superScript id="7C087F28FFA6EA242A17F2C5FA73FF5B" attach="none" box="[1392,1429,150,177]" fontSize="6" pageId="3" pageNumber="524">381</superScript>
|
||
,
|
||
<figureCitation id="1346CEE5FFA6EA242FF0F2E9FF14FF2A" box="[151,242,187,213]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[354,1233,187,1858]" captionTargetId="figure-22@6.[354,1233,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Scatimus pronotum, mesepimeron, metasternum, abdomen, and metafemur. A, Scatimus strandi pronotum (lateral view); B, Scatimus strenua pronotum (lateral view); C, Scatimus strandi mesepimeron (ventral view); D, Scatimus strenua mesepimeron (ventral view); E, Scatimus strenua metasternum (ventral view); F, Scatimus strenua abdomen; G, Scatimus erinnyos metafemur (ventral view); H, Scatimus strenua metafemur (ventral view). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4306560" httpUri="https://zenodo.org/record/4306560/files/figure.png" pageId="3" pageNumber="524">Fig. 3G</figureCitation>
|
||
). Lateral marginal bead of pronotum distinctly reduced medially in
|
||
<taxonomicName id="4C7DA9E3FFA6EA242C9DF2E9FB8DFF2B" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[1018,1131,187,213]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA6EA242C9DF2E9FBEEFF2A" box="[1018,1032,187,213]" italics="true" pageId="3" pageNumber="524">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA6EA242B7FF2E9FB8DFF2B" box="[1048,1131,187,212]" italics="true" pageId="3" pageNumber="524">strenua</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<superScript id="7C087F28FFA6EA242B1BF2E9FB47FF37" attach="none" box="[1148,1185,186,213]" fontSize="6" pageId="3" pageNumber="524">160</superScript>
|
||
,
|
||
<figureCitation id="1346CEE5FFA6EA242BD6F2E9FAE1FF2A" box="[1201,1287,187,213]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[354,1233,187,1858]" captionTargetId="figure-22@6.[354,1233,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Scatimus pronotum, mesepimeron, metasternum, abdomen, and metafemur. A, Scatimus strandi pronotum (lateral view); B, Scatimus strenua pronotum (lateral view); C, Scatimus strandi mesepimeron (ventral view); D, Scatimus strenua mesepimeron (ventral view); E, Scatimus strenua metasternum (ventral view); F, Scatimus strenua abdomen; G, Scatimus erinnyos metafemur (ventral view); H, Scatimus strenua metafemur (ventral view). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4306560" httpUri="https://zenodo.org/record/4306560/files/figure.png" pageId="3" pageNumber="524">Fig. 3B</figureCitation>
|
||
) and sharply defined medially in
|
||
<taxonomicName id="4C7DA9E3FFA6EA242E1FF28DFE05FF06" authorityName="Balthasar" authorityYear="1939" box="[376,483,223,249]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B9090E72FFA6EA242E1FF28DFE60FF06" box="[376,390,223,249]" italics="true" pageId="3" pageNumber="524">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFA6EA242EF3F28DFE05FF06" box="[404,483,223,249]" italics="true" pageId="3" pageNumber="524">strandi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<superScript id="7C087F28FFA6EA242E94F28DFDFEFF13" attach="none" box="[499,536,222,249]" fontSize="6" pageId="3" pageNumber="524">161</superScript>
|
||
,
|
||
<figureCitation id="1346CEE5FFA6EA242D41F28DFD99FF06" box="[550,639,223,249]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[354,1233,187,1858]" captionTargetId="figure-22@6.[354,1233,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Scatimus pronotum, mesepimeron, metasternum, abdomen, and metafemur. A, Scatimus strandi pronotum (lateral view); B, Scatimus strenua pronotum (lateral view); C, Scatimus strandi mesepimeron (ventral view); D, Scatimus strenua mesepimeron (ventral view); E, Scatimus strenua metasternum (ventral view); F, Scatimus strenua abdomen; G, Scatimus erinnyos metafemur (ventral view); H, Scatimus strenua metafemur (ventral view). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4306560" httpUri="https://zenodo.org/record/4306560/files/figure.png" pageId="3" pageNumber="524">Fig. 3A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36781EBFFA6EA222FA0F350FDA8FF06" lastPageId="5" lastPageNumber="526" pageId="3" pageNumber="524" type="description">
|
||
<paragraph id="8BC2D260FFA6EA232FA0F350FBCCFE0A" blockId="3.[151,1437,150,2013]" lastBlockId="4.[151,1437,150,609]" lastPageId="4" lastPageNumber="525" pageId="3" pageNumber="524">
|
||
<emphasis id="B9090E72FFA6EA242FA0F350FEBFFEE3" bold="true" box="[199,345,258,284]" pageId="3" pageNumber="524">Description.</emphasis>
|
||
<typeStatus id="54C66CC2FFA6EA242E38F351FE21FEE2" box="[351,455,259,285]" pageId="3" pageNumber="524" type="holotype">Holotype</typeStatus>
|
||
male, (
|
||
<figureCitation id="1346CEE5FFA6EA242D7DF351FD94FEE2" box="[538,626,259,285]" captionStart="FIGURE 1" captionStartId="4.[151,250,1877,1901]" captionTargetBox="[166,1426,636,1837]" captionTargetId="figure-253@4.[151,1436,630,1850]" captionTargetPageId="4" captionText="FIGURE 1. Species of Scatimus from Colombia.A, Scatimus strenua holotype łmale (Antioquia, 5°52’45.05’’N 75°12’2.33’’W); B, Scatimus strenua, paratype female (Antioquia, 5°53’16.68’’N 75°11’37.11’’W); C, Scatimus fernandezi male (Casanare, 5°16’20.9”N 72°24’2.6”W); D, Scatimus ovatus male (Tolima, 5°8’58.3’’N 74°47’6.1’’W); E, Scatimus strandi male (Nariño, 0°30’N 77°13’W). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4306556" httpUri="https://zenodo.org/record/4306556/files/figure.png" pageId="3" pageNumber="524">Fig. 1A</figureCitation>
|
||
). Pinned.
|
||
<emphasis id="B9090E72FFA6EA242D85F351FC77FEE2" bold="true" box="[738,913,259,285]" pageId="3" pageNumber="524">Measurements</emphasis>
|
||
. Body length
|
||
<quantity id="4C857F85FFA6EA242B49F350FB75FEE2" box="[1070,1171,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="5.84" pageId="3" pageNumber="524" unit="mm" value="5.84">5.84 mm</quantity>
|
||
, elytral width
|
||
<quantity id="4C857F85FFA6EA242A55F351FA7FFEE2" box="[1330,1433,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="3.73" pageId="3" pageNumber="524" unit="mm" value="3.73">3.73 mm</quantity>
|
||
. Body elongate, lateral edges parallel on median third in dorsal view (
|
||
<superScript id="7C087F28FFA6EA242CD7F375FC22FECB" attach="none" box="[944,964,294,321]" fontSize="6" pageId="3" pageNumber="524">11</superScript>
|
||
,
|
||
<figureCitation id="1346CEE5FFA6EA242CB4F375FBF7FEBE" box="[979,1041,295,321]" captionStart="FIGURE 1" captionStartId="4.[151,250,1877,1901]" captionTargetBox="[166,1426,636,1837]" captionTargetId="figure-253@4.[151,1436,630,1850]" captionTargetPageId="4" captionText="FIGURE 1. Species of Scatimus from Colombia.A, Scatimus strenua holotype łmale (Antioquia, 5°52’45.05’’N 75°12’2.33’’W); B, Scatimus strenua, paratype female (Antioquia, 5°53’16.68’’N 75°11’37.11’’W); C, Scatimus fernandezi male (Casanare, 5°16’20.9”N 72°24’2.6”W); D, Scatimus ovatus male (Tolima, 5°8’58.3’’N 74°47’6.1’’W); E, Scatimus strandi male (Nariño, 0°30’N 77°13’W). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4306556" httpUri="https://zenodo.org/record/4306556/files/figure.png" pageId="3" pageNumber="524">Fig. 1</figureCitation>
|
||
A–B, E).
|
||
<emphasis id="B9090E72FFA6EA242BE2F374FB28FEBF" bold="true" box="[1157,1230,294,320]" pageId="3" pageNumber="524">Color.</emphasis>
|
||
Black, with some brown reflections.
|
||
<emphasis id="B9090E72FFA6EA242E00F319FE41FE9A" bold="true" box="[359,423,331,357]" pageId="3" pageNumber="524">Head</emphasis>
|
||
(
|
||
<figureCitation id="1346CEE5FFA6EA242ED2F319FDE7FE9A" box="[437,513,331,357]" captionStart="FIGURE 2" captionStartId="5.[151,250,1655,1679]" captionTargetBox="[159,1434,387,1632]" captionTargetId="figure-121@5.[151,1436,381,1632]" captionTargetPageId="5" captionText="FIGURE 2. Scatimus heads in dorsal view. A, Scatimus strenua male (Antioquia, 5°52’45.05’’N 75°12’2.33’’W); B, Scatimus strenua female (Antioquia, 5°53’16.68’’N 75°11’37.11’’W); C, Scatimus erinnyos male (Costa Rica, Parque Nacional Guanacaste); D, Scatimus strandi male (Nariño, 0°30’N 77°13’W); E, Scatimus fernandezi (Casanare, 5°16’20.9”N 72°24’2.6”W); F, Scatimus ovatus (Tolima, 05°08’58.3’’N 74°47’06.1’’W). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4306558" httpUri="https://zenodo.org/record/4306558/files/figure.png" pageId="3" pageNumber="524">Figs. 2</figureCitation>
|
||
A–B). Dorsal surface of head glabrous, lacking setiferous punctures (
|
||
<superScript id="7C087F28FFA6EA242B9AF318FAF2FEA7" attach="right" box="[1277,1300,330,357]" fontSize="6" pageId="3" pageNumber="524">51</superScript>
|
||
). Surface of clypeus and gena smooth, lacking wrinkles or transverse rugulae (
|
||
<superScript id="7C087F28FFA6EA242C00F33DFC9BFE83" attach="none" box="[871,893,366,393]" fontSize="6" pageId="3" pageNumber="524">30</superScript>
|
||
), with scattered, fine punctures more pronounced towards the margins. Head with a single frontal carina between the eyes, arcuate medially, with lateral extremities anteriorly directed (
|
||
<superScript id="7C087F28FFA6EA242E14F3E5FE6CFE3B" attach="none" box="[371,394,438,465]" fontSize="6" pageId="3" pageNumber="524">63</superScript>
|
||
). Anterior edge of clypeus with two moderately developed and reflexed teeth, clypeal median emargination angulate (
|
||
<superScript id="7C087F28FFA6EA242EFAF389FE52FE17" attach="none" box="[413,436,474,501]" fontSize="6" pageId="3" pageNumber="524">40</superScript>
|
||
), lateral margins arcuate between teeth and gena, disc of clypeus concave, vertical surface of clypeus V-shaped with a small central depression in the lower edge. Clypeogenal suture pronounced, surface of gena adjacent to the eye lacking coarse punctures (
|
||
<superScript id="7C087F28FFA6EA242DB4F071FD0FFDCF" attach="none" box="[723,745,546,573]" fontSize="6" pageId="3" pageNumber="524">70</superScript>
|
||
). Smooth eyes completely divided by canthus, dorsal portion of the eye approximately twice as long as wide (
|
||
<superScript id="7C087F28FFA6EA242DD9F015FD32FDAB" attach="none" box="[702,724,582,609]" fontSize="6" pageId="3" pageNumber="524">80</superScript>
|
||
), surface around dorsal portions of eye lacking microsculpture (
|
||
<superScript id="7C087F28FFA6EA242FC7F039FF51FD87" attach="none" box="[160,183,618,645]" fontSize="6" pageId="3" pageNumber="524">90</superScript>
|
||
). Antennomeres 7–8 with pit on distal surface (
|
||
<superScript id="7C087F28FFA6EA242DB3F039FD1FFD87" attach="none" box="[724,761,618,645]" fontSize="6" pageId="3" pageNumber="524">101</superScript>
|
||
).
|
||
<emphasis id="B9090E72FFA6EA242C75F039FC6DFD7A" bold="true" box="[786,907,619,645]" pageId="3" pageNumber="524">Pronotum</emphasis>
|
||
(
|
||
<figureCitation id="1346CEE5FFA6EA242CFCF039FC12FD7A" box="[923,1012,619,645]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[354,1233,187,1858]" captionTargetId="figure-22@6.[354,1233,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Scatimus pronotum, mesepimeron, metasternum, abdomen, and metafemur. A, Scatimus strandi pronotum (lateral view); B, Scatimus strenua pronotum (lateral view); C, Scatimus strandi mesepimeron (ventral view); D, Scatimus strenua mesepimeron (ventral view); E, Scatimus strenua metasternum (ventral view); F, Scatimus strenua abdomen; G, Scatimus erinnyos metafemur (ventral view); H, Scatimus strenua metafemur (ventral view). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4306560" httpUri="https://zenodo.org/record/4306560/files/figure.png" pageId="3" pageNumber="524">Fig. 3B</figureCitation>
|
||
). Surface smooth. Anterior edge with membranous projection at head insertion (
|
||
<superScript id="7C087F28FFA6EA242D0AF0DDFD69FD63" attach="none" box="[621,655,654,681]" fontSize="6" pageId="3" pageNumber="524">111</superScript>
|
||
). Lateral edge of pronotum simple, lacking setiferous punctures (
|
||
<superScript id="7C087F28FFA6EA242A00F0DDFA6AFD63" attach="none" box="[1383,1420,654,681]" fontSize="6" pageId="3" pageNumber="524">121</superScript>
|
||
), with eight coarse setiferous punctures present in lateral fossae and three anteriorly to fossae (
|
||
<superScript id="7C087F28FFA6EA242BD8F0E1FB05FD3F" attach="none" box="[1215,1251,690,717]" fontSize="6" pageId="3" pageNumber="524">131</superScript>
|
||
), non-setiferous punctures each with a minute granule medially (
|
||
<superScript id="7C087F28FFA6EA242DA3F085FD0EFD1B" attach="none" box="[708,744,726,753]" fontSize="6" pageId="3" pageNumber="524">141</superScript>
|
||
). Lateral declivity of pronotum lacking setiferous punctures (
|
||
<superScript id="7C087F28FFA6EA242FC7F0A8FF23FCF7" attach="none" box="[160,197,762,789]" fontSize="6" pageId="3" pageNumber="524">151</superScript>
|
||
), lateral marginal bead of pronotum distinctly reduced medially (
|
||
<superScript id="7C087F28FFA6EA242CD5F0A9FC31FCF7" attach="none" box="[946,983,762,789]" fontSize="6" pageId="3" pageNumber="524">161</superScript>
|
||
), posterior margin of pronotum lacking coarse punctures (
|
||
<superScript id="7C087F28FFA6EA242E03F14DFE6FFCD3" attach="none" box="[356,393,798,825]" fontSize="6" pageId="3" pageNumber="524">171</superScript>
|
||
).
|
||
<emphasis id="B9090E72FFA6EA242EC6F14DFE0BFCC6" bold="true" box="[417,493,799,825]" pageId="3" pageNumber="524">Elytra</emphasis>
|
||
. Elytral disc smooth and shiny, with scattered minute punctures (3.5 X). Anterolateral angle of elytron slightly produced (
|
||
<superScript id="7C087F28FFA6EA242D3DF111FD98FCAF" attach="none" box="[602,638,834,861]" fontSize="6" pageId="3" pageNumber="524">180</superScript>
|
||
). Striae distinctly impressed with oval to rounded punctures separated by 3–4 times diameters on disc and lacking setiferous punctures on interstriae. Striae 1–3 with punctures larger, deeper and more pronounced and fused on apical declivity, striae 7 incomplete on posterior third, striae 8 deeply impressed with interstria lacking setiferous punctures (
|
||
<superScript id="7C087F28FFA6EA242D9CF1FDFCF9FC43" attach="none" box="[763,799,942,969]" fontSize="6" pageId="3" pageNumber="524">192</superScript>
|
||
). Apical declivity of elytron lacking setiferous punctures on interstriae (
|
||
<superScript id="7C087F28FFA6EA242E5FF181FEBBFC1F" attach="none" box="[312,349,978,1005]" fontSize="6" pageId="3" pageNumber="524">201</superScript>
|
||
). Proepipleural sulcus shallower on anterior half with a distinct depression (fovea) adjacent to the anterior angle (
|
||
<superScript id="7C087F28FFA6EA242E58F1A5FE85FBFB" attach="none" box="[319,355,1014,1041]" fontSize="6" pageId="3" pageNumber="524">211</superScript>
|
||
).
|
||
<emphasis id="B9090E72FFA6EA242E1EF1A4FDB4FBEF" bold="true" box="[377,594,1014,1040]" pageId="3" pageNumber="524">Thoracic ventrites</emphasis>
|
||
(
|
||
<figureCitation id="1346CEE5FFA6EA242D07F1A5FD33FBEE" box="[608,725,1015,1041]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[354,1233,187,1858]" captionTargetId="figure-22@6.[354,1233,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Scatimus pronotum, mesepimeron, metasternum, abdomen, and metafemur. A, Scatimus strandi pronotum (lateral view); B, Scatimus strenua pronotum (lateral view); C, Scatimus strandi mesepimeron (ventral view); D, Scatimus strenua mesepimeron (ventral view); E, Scatimus strenua metasternum (ventral view); F, Scatimus strenua abdomen; G, Scatimus erinnyos metafemur (ventral view); H, Scatimus strenua metafemur (ventral view). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4306560" httpUri="https://zenodo.org/record/4306560/files/figure.png" pageId="3" pageNumber="524">Fig. 3D, F</figureCitation>
|
||
). Prosternum behind procoxa lacking setiferous punctures (
|
||
<superScript id="7C087F28FFA6EA242A01F1A5FA6AFBFB" attach="right" box="[1382,1420,1014,1041]" fontSize="6" pageId="3" pageNumber="524">223</superScript>
|
||
), with few rounded punctures aligned with the posterior edge, margin of the punctures incomplete. Anterior region of propleuron strongly excavated and delimited posteriorly by a complete propleural carina, surface of excavated portion with rivose microsculpture and fine setae. Area posterior of propleuron with rounded punctures located on the posterior edge and reaching half the margin around procoxa. Mesoventrite with a moderately deep anteriomedian fossa and with setae anteriorly (
|
||
<superScript id="7C087F28FFA6EA242D52F6F9FDBCFB47" attach="none" box="[565,602,1194,1221]" fontSize="6" pageId="3" pageNumber="524">231</superScript>
|
||
), lacking a median longitudinal carina (
|
||
<superScript id="7C087F28FFA6EA242B4EF6F9FBA9FB47" attach="right" box="[1065,1103,1194,1221]" fontSize="6" pageId="3" pageNumber="524">240</superScript>
|
||
). Mesoepisternal suture well defined, thicker near the procoxa, and thinner towards the lateral edge. Disc of mesoventrite with dense ocellate punctures. Median lobe of metaventrite anterior edge arcuate medially (
|
||
<superScript id="7C087F28FFA6EA242CA0F6A0FC0AFAFF" attach="none" box="[967,1004,1266,1293]" fontSize="6" pageId="3" pageNumber="524">251</superScript>
|
||
), with straight lateral borders. Medial edge of mesocoxal cavity at least almost straight on a short distance (
|
||
<superScript id="7C087F28FFA6EA242CC6F745FC21FADB" attach="right" box="[929,967,1302,1329]" fontSize="6" pageId="3" pageNumber="524">260</superScript>
|
||
). Lateral surface of mesepimeron lacking transverse carina medially (
|
||
<superScript id="7C087F28FFA6EA242EB6F769FE10FAB7" attach="none" box="[465,502,1338,1365]" fontSize="6" pageId="3" pageNumber="524">272</superScript>
|
||
), anteromedial surface punctate (
|
||
<superScript id="7C087F28FFA6EA242C13F769FC7FFAB7" attach="none" box="[884,921,1338,1365]" fontSize="6" pageId="3" pageNumber="524">280</superScript>
|
||
). Posterior portion of metepisternon rounded (
|
||
<superScript id="7C087F28FFA6EA242FF8F70DFF23FA93" attach="right" box="[159,197,1374,1401]" fontSize="6" pageId="3" pageNumber="524">291</superScript>
|
||
), posteromedial area of metepisternon flat (
|
||
<superScript id="7C087F28FFA6EA242DCCF70DFD36FA93" attach="none" box="[683,720,1374,1401]" fontSize="6" pageId="3" pageNumber="524">301</superScript>
|
||
), lateral lobes of metaventrite lacking a posterior transverse row of punctures (
|
||
<superScript id="7C087F28FFA6EA242E53F7D1FEBEFA6F" attach="none" box="[308,344,1410,1437]" fontSize="6" pageId="3" pageNumber="524">310</superScript>
|
||
).
|
||
<emphasis id="B9090E72FFA6EA242E17F7D1FE4EFA62" bold="true" box="[368,424,1411,1437]" pageId="3" pageNumber="524">Legs</emphasis>
|
||
(
|
||
<figureCitation id="1346CEE5FFA6EA242EDEF7D1FDF5FA62" box="[441,531,1411,1437]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[354,1233,187,1858]" captionTargetId="figure-22@6.[354,1233,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Scatimus pronotum, mesepimeron, metasternum, abdomen, and metafemur. A, Scatimus strandi pronotum (lateral view); B, Scatimus strenua pronotum (lateral view); C, Scatimus strandi mesepimeron (ventral view); D, Scatimus strenua mesepimeron (ventral view); E, Scatimus strenua metasternum (ventral view); F, Scatimus strenua abdomen; G, Scatimus erinnyos metafemur (ventral view); H, Scatimus strenua metafemur (ventral view). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4306560" httpUri="https://zenodo.org/record/4306560/files/figure.png" pageId="3" pageNumber="524">Fig. 3H</figureCitation>
|
||
). Protibiae tridentate on the outer margin, basal tooth smallest. External teeth of protibiae with additional small indentations at basal angles (
|
||
<superScript id="7C087F28FFA6EA242C49F7F4FCB5FA4B" attach="none" box="[814,851,1446,1473]" fontSize="6" pageId="3" pageNumber="524">351</superScript>
|
||
), dorsoapical transverse carina of protibiae emarginate (
|
||
<superScript id="7C087F28FFA6EA242FAFF799FF0BFA27" attach="none" box="[200,237,1482,1509]" fontSize="6" pageId="3" pageNumber="524">361</superScript>
|
||
). Ventral surface of protibiae with two rows of setae, the outer row with large curved apex setae, and the inner row with smaller straight setae. Ventral surface of profemora lacking setiferous punctures (
|
||
<superScript id="7C087F28FFA6EA242B84F7BDFAEEFA03" attach="none" box="[1251,1288,1518,1545]" fontSize="6" pageId="3" pageNumber="524">321</superScript>
|
||
), with a row of long setae on the anterior margin and scattered coarse punctures in the posterior margin. Trochanto-femoral pit of prothoracic leg rounded (
|
||
<superScript id="7C087F28FFA6EA242EA8F465FE12F9BB" attach="none" box="[463,500,1590,1617]" fontSize="6" pageId="3" pageNumber="524">340</superScript>
|
||
). Procoxa with anteromarginal sulcus (
|
||
<superScript id="7C087F28FFA6EA242CCBF465FC37F9BB" attach="none" box="[940,977,1590,1617]" fontSize="6" pageId="3" pageNumber="524">331</superScript>
|
||
). Mesotibiae with two transverse carinas on the external surface (
|
||
<superScript id="7C087F28FFA6EA242ED7F409FE30F997" attach="right" box="[432,470,1626,1653]" fontSize="6" pageId="3" pageNumber="524">421</superScript>
|
||
). Surface of mesofemur with minute punctures throughout (
|
||
<superScript id="7C087F28FFA6EA242BF7F409FB53F997" attach="none" box="[1168,1205,1626,1653]" fontSize="6" pageId="3" pageNumber="524">380</superScript>
|
||
). Ventral surface of mesocoxa with coarse setiferous punctures, setae longer than punctures diameter (
|
||
<superScript id="7C087F28FFA6EA242B52F42DFBBCF973" attach="none" box="[1077,1114,1662,1689]" fontSize="6" pageId="3" pageNumber="524">372</superScript>
|
||
). Metatibiae with two transverse carinae on the external surface. Dorsal surface of metatibiae with a longitudinal medial row of straight setae and strong microsculpture on external half only (
|
||
<superScript id="7C087F28FFA6EA242DA5F495FD01F92B" attach="none" box="[706,743,1734,1761]" fontSize="6" pageId="3" pageNumber="524">431</superScript>
|
||
), lacking dorsal accessory setiferous punctures (
|
||
<superScript id="7C087F28FFA6EA242A69F495FAD5F92B" attach="none" box="[1294,1331,1734,1761]" fontSize="6" pageId="3" pageNumber="524">441</superScript>
|
||
). Ventral portion of metatibiae broadly arcuate in cross section near apex (
|
||
<superScript id="7C087F28FFA6EA242C00F4B8FC6AF907" attach="none" box="[871,908,1770,1797]" fontSize="6" pageId="3" pageNumber="524">451</superScript>
|
||
). Anterior edge of metafemur lacking marginal bead (
|
||
<superScript id="7C087F28FFA6EA242FBCF55DFEE6F8E3" attach="none" box="[219,256,1806,1833]" fontSize="6" pageId="3" pageNumber="524">400</superScript>
|
||
), with anterior margin enlarged and very sharp (
|
||
<superScript id="7C087F28FFA6EA242C78F55DFCA5F8E3" attach="none" box="[799,835,1806,1833]" fontSize="6" pageId="3" pageNumber="524">411</superScript>
|
||
). Dorsal surface of metafemur with minute punctures throughout. Longitudinal median sulcus of metacoxa sharply defined posteriorly (
|
||
<superScript id="7C087F28FFA6EA242B54F561FBBEF8BF" attach="none" box="[1075,1112,1842,1869]" fontSize="6" pageId="3" pageNumber="524">391</superScript>
|
||
).
|
||
<emphasis id="B9090E72FFA6EA242B17F560FA80F8B3" bold="true" box="[1136,1382,1842,1868]" pageId="3" pageNumber="524">Abdominal ventrites</emphasis>
|
||
(
|
||
<figureCitation id="1346CEE5FFA6EA242A12F561FF5EF88E" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[354,1233,187,1858]" captionTargetId="figure-22@6.[354,1233,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Scatimus pronotum, mesepimeron, metasternum, abdomen, and metafemur. A, Scatimus strandi pronotum (lateral view); B, Scatimus strenua pronotum (lateral view); C, Scatimus strandi mesepimeron (ventral view); D, Scatimus strenua mesepimeron (ventral view); E, Scatimus strenua metasternum (ventral view); F, Scatimus strenua abdomen; G, Scatimus erinnyos metafemur (ventral view); H, Scatimus strenua metafemur (ventral view). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4306560" httpUri="https://zenodo.org/record/4306560/files/figure.png" pageId="3" pageNumber="524">Fig 3F</figureCitation>
|
||
). Median portion of abdominal ventrites unmodified, each suture clearly visible throughout (
|
||
<superScript id="7C087F28FFA6EA242B81F505FAEAF89B" attach="right" box="[1254,1292,1878,1905]" fontSize="6" pageId="3" pageNumber="524">461</superScript>
|
||
). Surface of abdominal ventrites with weakly defined microsculpture laterally on ventrites 3 and 4 (
|
||
<superScript id="7C087F28FFA6EA242B19F529FB45F877" attach="none" box="[1150,1187,1914,1941]" fontSize="6" pageId="3" pageNumber="524">473</superScript>
|
||
), abdominal ventrites 7 and 8 feebly grooved along suture (49:1). Pre-pygidium is clearly visible, with a central, longitudinal, deep sulcus.
|
||
<emphasis id="B9090E72FFA6EA242FACF590FEA0F823" bold="true" box="[203,326,1986,2012]" pageId="3" pageNumber="524">Pygidium.</emphasis>
|
||
Basal sulcus of pygidium narrowed and shallowly impressed medially (
|
||
<superScript id="7C087F28FFA6EA242B11F590FB7AF82F" attach="right" box="[1142,1180,1986,2013]" fontSize="6" pageId="3" pageNumber="524">500</superScript>
|
||
). Surface of pygidium lacking setiferous punctures (
|
||
<superScript id="7C087F28FFA1EA232E83F2C4FDEEFF5B" attach="none" box="[484,520,150,177]" fontSize="6" pageId="4" pageNumber="525">512</superScript>
|
||
). Transversal sulcus well defined with the superior border thickened towards the middle.
|
||
<emphasis id="B9090E72FFA1EA232F95F2E8FE83FF2B" bold="true" box="[242,357,186,212]" pageId="4" pageNumber="525">Aedeagus</emphasis>
|
||
(
|
||
<figureCitation id="1346CEE5FFA1EA232E14F2E9FE0CFF2A" box="[371,490,187,213]" captionStart="FIGURE 4" captionStartId="7.[151,250,1653,1677]" captionTargetBox="[194,1410,204,1617]" captionTargetId="figure-22@7.[177,1410,181,1629]" captionTargetPageId="7" captionText="FIGURE 4. Male genitalia of Scatimus from Colombia. A, Scatimus fernandezi aedeagus (lateral view, scale bar = 0.5 mm); B, Scatimus ovatus aedeagus (lateral view, scale bar = 0.5 mm); C, Scatimus strandi aedeagus (lateral view, scale bar = 0.5 mm); D, Scatimus strenua aedeagus (lateral view, scale bar = 0.5 mm); E, Scatimus fernandezi parameres (ventral view, scale bar = 0.25 mm); F, Scatimus ovatus parameres (ventral view, scale bar = 0.25 mm); G, Scatimus strandi parameres (ventral view, scale bar = 0.25 mm); H, Scatimus strenua parameres (ventral view, scale bar = 0.25 mm); I, Scatimus fernandezi superior right peripheral sclerite (SRP, scale = 0.25 mm); J, Scatimus ovatus superior right peripheral sclerite (SRP, scale = 0.25 mm); K, Scatimus strandi superior right peripheral sclerite (SRP, scale = 0.25 mm); L, Scatimus strenua superior right peripheral sclerite (SRP, scale = 0.25 mm); M, Scatimus fernandezi additional (AS), subaxial (SA), and axial (A) sclerites superposed, scale bar = 0.25 mm; N, Scatimus ovatus additional (AS), subaxial (SA), and axial (A) sclerites superposed, scale bar = 0.25 mm; O, Scatimus strandi additional (AS), subaxial (SA), and axial (A) sclerites superposed, scale bar = 0.25 mm; P, Scatimus strenua additional (AS), subaxial (SA), and axial (A) sclerites superposed, scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4306562" httpUri="https://zenodo.org/record/4306562/files/figure.png" pageId="4" pageNumber="525">Fig. 4D, H</figureCitation>
|
||
). Parameres approximately conical (
|
||
<superScript id="7C087F28FFA1EA232C18F2E8FC43FF37" attach="right" box="[895,933,186,213]" fontSize="6" pageId="4" pageNumber="525">521</superScript>
|
||
). Ventral surface of paramere with strong microsculpture (
|
||
<superScript id="7C087F28FFA1EA232E57F28CFEB3FF13" attach="none" box="[304,341,222,249]" fontSize="6" pageId="4" pageNumber="525">531</superScript>
|
||
), parameres apex not projecting ventrally (
|
||
<superScript id="7C087F28FFA1EA232C54F28CFCBEFF13" attach="none" box="[819,856,222,249]" fontSize="6" pageId="4" pageNumber="525">540</superScript>
|
||
). Dorsal portion of paramere sclerotized throughout (
|
||
<superScript id="7C087F28FFA1EA232FF8F350FF23FEEF" attach="right" box="[159,197,258,285]" fontSize="6" pageId="4" pageNumber="525">550</superScript>
|
||
). Paramere hook feebly developed (
|
||
<superScript id="7C087F28FFA1EA232D32F350FD9CFEEF" attach="none" box="[597,634,258,285]" fontSize="6" pageId="4" pageNumber="525">561</superScript>
|
||
).
|
||
<emphasis id="B9090E72FFA1EA232DF7F351FCC1FEE2" bold="true" box="[656,807,259,285]" pageId="4" pageNumber="525">Endophallus</emphasis>
|
||
(
|
||
<figureCitation id="1346CEE5FFA1EA232C52F351FC54FEE2" box="[821,946,259,285]" captionStart="FIGURE 4" captionStartId="7.[151,250,1653,1677]" captionTargetBox="[194,1410,204,1617]" captionTargetId="figure-22@7.[177,1410,181,1629]" captionTargetPageId="7" captionText="FIGURE 4. Male genitalia of Scatimus from Colombia. A, Scatimus fernandezi aedeagus (lateral view, scale bar = 0.5 mm); B, Scatimus ovatus aedeagus (lateral view, scale bar = 0.5 mm); C, Scatimus strandi aedeagus (lateral view, scale bar = 0.5 mm); D, Scatimus strenua aedeagus (lateral view, scale bar = 0.5 mm); E, Scatimus fernandezi parameres (ventral view, scale bar = 0.25 mm); F, Scatimus ovatus parameres (ventral view, scale bar = 0.25 mm); G, Scatimus strandi parameres (ventral view, scale bar = 0.25 mm); H, Scatimus strenua parameres (ventral view, scale bar = 0.25 mm); I, Scatimus fernandezi superior right peripheral sclerite (SRP, scale = 0.25 mm); J, Scatimus ovatus superior right peripheral sclerite (SRP, scale = 0.25 mm); K, Scatimus strandi superior right peripheral sclerite (SRP, scale = 0.25 mm); L, Scatimus strenua superior right peripheral sclerite (SRP, scale = 0.25 mm); M, Scatimus fernandezi additional (AS), subaxial (SA), and axial (A) sclerites superposed, scale bar = 0.25 mm; N, Scatimus ovatus additional (AS), subaxial (SA), and axial (A) sclerites superposed, scale bar = 0.25 mm; O, Scatimus strandi additional (AS), subaxial (SA), and axial (A) sclerites superposed, scale bar = 0.25 mm; P, Scatimus strenua additional (AS), subaxial (SA), and axial (A) sclerites superposed, scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4306562" httpUri="https://zenodo.org/record/4306562/files/figure.png" pageId="4" pageNumber="525">Figs. 4L, P</figureCitation>
|
||
). Internal sac with four apical endophallites, the basal semicircular (Bsc), frontolateral peripheral (FLP) endophallites and lamella copulatrix (LC) are absent. The superior right peripheral endophallite (SRP) is semicircular, with a broader and rounded extremity, narrow and curved in its middle part, and ending in a handle-shaped projection with bilobed apex. The additional (
|
||
<collectingCountry id="F36A92F0FFA1EA232A71F33DFADAFE76" box="[1302,1340,367,393]" name="American Samoa" pageId="4" pageNumber="525">AS</collectingCountry>
|
||
) sclerite is short, with a forked apex. The subaxial (SA) and axial (A) endophallites are superposed. The SA endophallite is elongate, one extremity with a truncated apex and the other one with a forked apex. The A endophallite is elongate the same size as SA, one extremity with an acute apex and the other one rounded.
|
||
</paragraph>
|
||
<paragraph id="8BC2D260FFA1EA232FA0F3ADFCC2FD9E" blockId="4.[151,1437,150,609]" pageId="4" pageNumber="525">
|
||
<emphasis id="B9090E72FFA1EA232FA0F3ADFEC3FDE6" bold="true" box="[199,293,511,537]" pageId="4" pageNumber="525">Female.</emphasis>
|
||
(
|
||
<figureCitation id="1346CEE5FFA1EA232E51F3ADFE6BFDE6" box="[310,397,511,537]" captionStart="FIGURE 1" captionStartId="4.[151,250,1877,1901]" captionTargetBox="[166,1426,636,1837]" captionTargetId="figure-253@4.[151,1436,630,1850]" captionTargetPageId="4" captionText="FIGURE 1. Species of Scatimus from Colombia.A, Scatimus strenua holotype łmale (Antioquia, 5°52’45.05’’N 75°12’2.33’’W); B, Scatimus strenua, paratype female (Antioquia, 5°53’16.68’’N 75°11’37.11’’W); C, Scatimus fernandezi male (Casanare, 5°16’20.9”N 72°24’2.6”W); D, Scatimus ovatus male (Tolima, 5°8’58.3’’N 74°47’6.1’’W); E, Scatimus strandi male (Nariño, 0°30’N 77°13’W). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4306556" httpUri="https://zenodo.org/record/4306556/files/figure.png" pageId="4" pageNumber="525">Fig. 1B</figureCitation>
|
||
). Body length
|
||
<quantity id="4C857F85FFA1EA232D50F3ADFD78FDE6" box="[567,670,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="6.63" pageId="4" pageNumber="525" unit="mm" value="6.63">6.63 mm</quantity>
|
||
, elytral width
|
||
<quantity id="4C857F85FFA1EA232C25F3ADFC4CFDE6" box="[834,938,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="4.18" pageId="4" pageNumber="525" unit="mm" value="4.18">4.18 mm</quantity>
|
||
. Similar to the male except in: clypeus with four teeth (
|
||
<superScript id="7C087F28FFA1EA232E74F071FEC1FDCF" attach="none" box="[275,295,546,573]" fontSize="6" pageId="4" pageNumber="525">11</superScript>
|
||
), middle ones larger, pronotum with one to seven large rounded punctures anterior to and five to ten in the lateral fossae abdominal ventrite 7 parallel sided (
|
||
<superScript id="7C087F28FFA1EA232D89F015FCF5FDAB" attach="none" box="[750,787,582,609]" fontSize="6" pageId="4" pageNumber="525">481</superScript>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF0282E8FFA1EA232FF0F507FDF7F826" ID-DOI="http://doi.org/10.5281/zenodo.4306556" ID-Zenodo-Dep="4306556" httpUri="https://zenodo.org/record/4306556/files/figure.png" pageId="4" pageNumber="525" startId="4.[151,250,1877,1901]" targetBox="[166,1426,636,1837]" targetPageId="4">
|
||
<paragraph id="8BC2D260FFA1EA232FF0F507FDF7F826" blockId="4.[151,1436,1877,2009]" pageId="4" pageNumber="525">
|
||
<emphasis id="B9090E72FFA1EA232FF0F507FEF7F892" bold="true" box="[151,273,1877,1901]" pageId="4" pageNumber="525">FIGURE 1.</emphasis>
|
||
Species of
|
||
<taxonomicName id="4C7DA9E3FFA1EA232EE6F504FE3AF892" authorityName="Erichson" authorityYear="1847" box="[385,476,1878,1901]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="525" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9090E72FFA1EA232EE6F504FE3AF892" box="[385,476,1878,1901]" italics="true" pageId="4" pageNumber="525">Scatimus</emphasis>
|
||
</taxonomicName>
|
||
from Colombia.
|
||
<emphasis id="B9090E72FFA1EA232DE2F507FD71F892" bold="true" box="[645,663,1877,1901]" pageId="4" pageNumber="525">A</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA1EA232DC5F504FCABF892" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[674,845,1878,1901]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="525" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA1EA232DC5F504FCABF892" box="[674,845,1878,1901]" italics="true" pageId="4" pageNumber="525">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
holotype łmale (Antioquia, 5°52’45.05’’N 75°12’2.33’’W);
|
||
<emphasis id="B9090E72FFA1EA232FF0F52BFF4EF86E" bold="true" box="[151,168,1913,1937]" pageId="4" pageNumber="525">B</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA1EA232FDFF528FE81F86E" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[184,359,1914,1937]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="525" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA1EA232FDFF528FE81F86E" box="[184,359,1914,1937]" italics="true" pageId="4" pageNumber="525">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
, paratype female (Antioquia, 5°53’16.68’’N 75°11’37.11’’W);
|
||
<emphasis id="B9090E72FFA1EA232C9CF52BFBEBF86E" bold="true" box="[1019,1037,1913,1937]" pageId="4" pageNumber="525">C</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA1EA232B7AF528FB0BF86E" authorityName="Martinez" authorityYear="1988" box="[1053,1261,1913,1937]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="525" phylum="Arthropoda" rank="species" species="fernandezi">
|
||
<emphasis id="B9090E72FFA1EA232B7AF528FB0BF86E" box="[1053,1261,1913,1937]" italics="true" pageId="4" pageNumber="525">Scatimus fernandezi</emphasis>
|
||
</taxonomicName>
|
||
male (Casanare, 5°16’20.9”N 72°24’2.6”W);
|
||
<emphasis id="B9090E72FFA1EA232ED8F5CFFE37F84A" bold="true" box="[447,465,1949,1973]" pageId="4" pageNumber="525">D</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA1EA232EB9F5CCFD64F84A" authorityName="Harold" authorityYear="1862" box="[478,642,1950,1973]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="525" phylum="Arthropoda" rank="species" species="ovatus">
|
||
<emphasis id="B9090E72FFA1EA232EB9F5CCFD64F84A" box="[478,642,1950,1973]" italics="true" pageId="4" pageNumber="525">Scatimus ovatus</emphasis>
|
||
</taxonomicName>
|
||
male (Tolima, 5°8’58.3’’N 74°47’6.1’’W);
|
||
<emphasis id="B9090E72FFA1EA232B27F5CFFBB7F84A" bold="true" box="[1088,1105,1949,1973]" pageId="4" pageNumber="525">E</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA1EA232B39F5CCFAEFF84A" authorityName="Balthasar" authorityYear="1939" box="[1118,1289,1949,1973]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="525" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B9090E72FFA1EA232B39F5CCFAEFF84A" box="[1118,1289,1949,1973]" italics="true" pageId="4" pageNumber="525">Scatimus strandi</emphasis>
|
||
</taxonomicName>
|
||
male (Nariño, 0°30’N 77°13’W). Scale bars: 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC2D260FFA0EA222FA0F2C4FDA8FF06" blockId="5.[151,1437,150,357]" pageId="5" pageNumber="526">
|
||
<emphasis id="B9090E72FFA0EA222FA0F2C4FED9FF4F" bold="true" box="[199,319,150,176]" pageId="5" pageNumber="526">Variation.</emphasis>
|
||
Male: body length
|
||
<quantity id="4C857F85FFA0EA222D43F2C4FD2DFF4E" box="[548,715,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="5.9399999999999995" metricValueMax="6.27" metricValueMin="5.61" pageId="5" pageNumber="526" unit="mm" value="5.9399999999999995" valueMax="6.27" valueMin="5.61">5.61–6.27 mm</quantity>
|
||
, elytral width
|
||
<quantity id="4C857F85FFA0EA222C13F2C4FBFAFF4E" box="[884,1052,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="4.06" metricValueMax="4.35" metricValueMin="3.77" pageId="5" pageNumber="526" unit="mm" value="4.06" valueMax="4.35" valueMin="3.77">3.77–4.35 mm</quantity>
|
||
. Pronotum with minimum two to maximum eight large rounded punctures anterior to and five to ten in the lateral fossae. Female: body length
|
||
<quantity id="4C857F85FFA0EA222A39F2E8FF18FF06" metricMagnitude="-3" metricUnit="m" metricValue="5.87" metricValueMax="6.63" metricValueMin="5.11" pageId="5" pageNumber="526" unit="mm" value="5.87" valueMax="6.63" valueMin="5.11">5.11– 6.63 mm</quantity>
|
||
; Elytral width
|
||
<quantity id="4C857F85FFA0EA222EC3F28DFDACFF06" box="[420,586,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="3.71" metricValueMax="4.18" metricValueMin="3.24" pageId="5" pageNumber="526" unit="mm" value="3.71" valueMax="4.18" valueMin="3.24">3.24–4.18 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36781EBFFA0EA222FA0F351FAEDFE9A" pageId="5" pageNumber="526" type="etymology">
|
||
<paragraph id="8BC2D260FFA0EA222FA0F351FAEDFE9A" blockId="5.[151,1437,150,357]" pageId="5" pageNumber="526">
|
||
<emphasis id="B9090E72FFA0EA222FA0F351FEA8FEE2" bold="true" box="[199,334,259,285]" pageId="5" pageNumber="526">Etymology.</emphasis>
|
||
The species epithet “
|
||
<taxonomicName id="4C7DA9E3FFA0EA222D5CF351FD68FEE3" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[571,654,259,284]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA0EA222D5CF351FD68FEE3" box="[571,654,259,284]" italics="true" pageId="5" pageNumber="526">strenua</emphasis>
|
||
</taxonomicName>
|
||
” refers to a Roman goddess of the new year, purification, and well-being, and should be treated as a noun in apposition. The name is dedicated to the courageous women from this area of
|
||
<collectingCountry id="F36A92F0FFA0EA222FD1F319FEC1FE9A" box="[182,295,331,357]" name="Colombia" pageId="5" pageNumber="526">Colombia</collectingCountry>
|
||
, that have overcome the armed conflict that affected their region in the 1980s and 1990s.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF0282E8FFA0EA222FF0F425FC5AF904" ID-DOI="http://doi.org/10.5281/zenodo.4306558" ID-Zenodo-Dep="4306558" httpUri="https://zenodo.org/record/4306558/files/figure.png" pageId="5" pageNumber="526" startId="5.[151,250,1655,1679]" targetBox="[159,1434,387,1632]" targetPageId="5">
|
||
<paragraph id="8BC2D260FFA0EA222FF0F425FC5AF904" blockId="5.[151,1437,1655,1787]" pageId="5" pageNumber="526">
|
||
<emphasis id="B9090E72FFA0EA222FF0F425FEF5F970" bold="true" box="[151,275,1655,1679]" pageId="5" pageNumber="526">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C7DA9E3FFA0EA222E7DF42AFE93F970" authorityName="Erichson" authorityYear="1847" box="[282,373,1656,1679]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="526" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9090E72FFA0EA222E7DF42AFE93F970" box="[282,373,1656,1679]" italics="true" pageId="5" pageNumber="526">Scatimus</emphasis>
|
||
</taxonomicName>
|
||
heads in dorsal view.
|
||
<emphasis id="B9090E72FFA0EA222D30F425FD8FF970" bold="true" box="[599,617,1655,1679]" pageId="5" pageNumber="526">A</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA0EA222D11F42AFCC5F970" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[630,803,1656,1679]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA0EA222D11F42AFCC5F970" box="[630,803,1656,1679]" italics="true" pageId="5" pageNumber="526">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
male (Antioquia, 5°52’45.05’’N 75°12’2.33’’W);
|
||
<emphasis id="B9090E72FFA0EA222A44F425FAD2F970" bold="true" box="[1315,1332,1655,1679]" pageId="5" pageNumber="526">B</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA0EA222A26F42AFF04F94C" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA0EA222A26F42AFF04F94C" italics="true" pageId="5" pageNumber="526">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
female (Antioquia, 5°53’16.68’’N 75°11’37.11’’W);
|
||
<emphasis id="B9090E72FFA0EA222C63F4C9FCF0F94C" bold="true" box="[772,790,1691,1715]" pageId="5" pageNumber="526">C</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA0EA222C43F4CEFC3BF94C" authorityName="Kohlmann & Solis" authorityYear="1996" box="[804,989,1692,1715]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="erinnyos">
|
||
<emphasis id="B9090E72FFA0EA222C43F4CEFC3BF94C" box="[804,989,1692,1715]" italics="true" pageId="5" pageNumber="526">Scatimus erinnyos</emphasis>
|
||
</taxonomicName>
|
||
male (Costa Rica, Parque Nacional Guanacaste);
|
||
<emphasis id="B9090E72FFA0EA222FB8F4EDFF17F928" bold="true" box="[223,241,1727,1751]" pageId="5" pageNumber="526">D</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA0EA222F9AF492FE41F928" authorityName="Balthasar" authorityYear="1939" box="[253,423,1727,1751]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B9090E72FFA0EA222F9AF492FE41F928" box="[253,423,1727,1751]" italics="true" pageId="5" pageNumber="526">Scatimus strandi</emphasis>
|
||
</taxonomicName>
|
||
male (Nariño, 0°30’N 77°13’W);
|
||
<emphasis id="B9090E72FFA0EA222C66F4EDFCF4F928" bold="true" box="[769,786,1727,1751]" pageId="5" pageNumber="526">E</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA0EA222C79F492FC0CF928" authorityName="Martinez" authorityYear="1988" box="[798,1002,1727,1751]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="fernandezi">
|
||
<emphasis id="B9090E72FFA0EA222C79F492FC0CF928" box="[798,1002,1727,1751]" italics="true" pageId="5" pageNumber="526">Scatimus fernandezi</emphasis>
|
||
</taxonomicName>
|
||
(Casanare, 5°16’20.9”N 72°24’2.6”W);
|
||
<emphasis id="B9090E72FFA0EA222AE1F4EDFA73F928" bold="true" box="[1414,1429,1727,1751]" pageId="5" pageNumber="526">F</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA0EA222FF0F4B6FEDDF904" authorityName="Harold" authorityYear="1862" box="[151,315,1764,1787]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="ovatus">
|
||
<emphasis id="B9090E72FFA0EA222FF0F4B6FEDDF904" box="[151,315,1764,1787]" italics="true" pageId="5" pageNumber="526">Scatimus ovatus</emphasis>
|
||
</taxonomicName>
|
||
(Tolima, 05°08’58.3’’N 74°47’06.1’’W). Scale bars = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C36781EBFFA0EA2F2FA0F578FD8AFA62" lastPageId="8" lastPageNumber="529" pageId="5" pageNumber="526" type="biology_ecology">
|
||
<paragraph id="8BC2D260FFA0EA2F2FA0F578FE48FE9A" blockId="5.[151,1437,1833,2040]" lastBlockId="8.[151,1437,150,1437]" lastPageId="8" lastPageNumber="529" pageId="5" pageNumber="526">
|
||
<emphasis id="B9090E72FFA0EA222FA0F578FE1DF8BC" bold="true" box="[199,507,1833,1860]" pageId="5" pageNumber="526">Ecology and distribution.</emphasis>
|
||
The landscape inhabited by
|
||
<taxonomicName id="4C7DA9E3FFA0EA222C2FF578FC5FF8BC" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[840,953,1834,1860]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA0EA222C2FF578FC5FF8BC" box="[840,953,1834,1860]" italics="true" pageId="5" pageNumber="526">S. strenua</emphasis>
|
||
</taxonomicName>
|
||
is part of the “Rio Melcocho” watershed. Vegetation cover is mostly forest classified by Espinel (2011) as Very Humid pre-Montane forest with influence of sub-Andean and Andean forests (van der Hammen & Rangel 1997). Scattered pastures and plantations are located on the few flat areas of these mountains. The tree canopy average height is
|
||
<quantity id="4C857F85FFA0EA222CB9F5C7FBF1F84F" box="[990,1047,1941,1968]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="526" unit="m" value="25.0">25 m</quantity>
|
||
with emergent trees of up to
|
||
<specimenCount id="9D7B19E9FFA0EA222A3AF5C7FA7BF84F" box="[1373,1437,1941,1968]" pageId="5" pageNumber="526" type="generic">
|
||
<quantity id="4C857F85FFA0EA222A3AF5C7FA7EF84F" box="[1373,1432,1941,1968]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="526" unit="m" value="35.0">35 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
Although the canopy is mostly continuous with abundant epiphytes, most timber producing trees have been already harvested, and the remaining forest can be considered as a mature secondary forest. The soils are well drained and covered by a leaf litter layer
|
||
<quantity id="4C857F85FFADEA2F2D6CF2C4FD9DFF4E" box="[523,635,150,177]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="8" pageNumber="529" unit="cm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 cm</quantity>
|
||
thick. Biogeographically, these forests belong to the
|
||
<collectingRegion id="49B91C82FFADEA2F2BB6F2C5FAABFF4E" box="[1233,1357,151,177]" country="Colombia" name="Magdalena" pageId="8" pageNumber="529">Magdalena</collectingRegion>
|
||
Valley mountain forests in the Tumbes-Chocó-Magdalena province, a key biological hotspot. The new species is part of a dung beetle (
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2E41F28DFE5BFF05" authorityName="Latreille" authorityYear="1802" box="[294,445,223,250]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="subFamily" subFamily="Scarabaeinae">Scarabaeinae</taxonomicName>
|
||
) ensemble of approximately 45 species including other endemic beetles such as
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2A5EF28DFDE6FEE2" authority="Sarmiento & Amat" authorityName="Sarmiento & Amat" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="andresi">
|
||
<emphasis id="B9090E72FFADEA2F2A5EF28DFEC0FEE2" italics="true" pageId="8" pageNumber="529">Dichotomius andresi</emphasis>
|
||
Sarmiento & Amat
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2D51F351FC7FFEE2" authority="Howden" authorityName="Howden" box="[566,921,259,285]" class="Insecta" family="Scarabaeidae" genus="Cryptocanthon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="parvus">
|
||
<emphasis id="B9090E72FFADEA2F2D51F351FCD6FEE3" box="[566,816,259,285]" italics="true" pageId="8" pageNumber="529">Cryptocanthon parvus</emphasis>
|
||
Howden
|
||
</taxonomicName>
|
||
, and moist forest beetles such as
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2A6BF351FE79FEBD" authority="(Harold)" baseAuthorityName="Harold" baseAuthorityYear="1868" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="aequinoctialis">
|
||
<emphasis id="B9090E72FFADEA2F2A6BF351FED0FEBE" italics="true" pageId="8" pageNumber="529">Sylvicanthon aequinoctialis</emphasis>
|
||
(Harold)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2ECEF375FCD6FEBE" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[425,816,295,321]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="moniliatus">
|
||
<emphasis id="B9090E72FFADEA2F2ECEF375FD31FEBE" box="[425,727,295,321]" italics="true" pageId="8" pageNumber="529">Scybalocanthon moniliatus</emphasis>
|
||
(Bates)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2C0EF375FB52FEBE" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[873,1204,295,321]" class="Insecta" family="Scarabaeidae" genus="Sulcophanaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="noctis">
|
||
<emphasis id="B9090E72FFADEA2F2C0EF375FBBDFEBF" box="[873,1115,295,321]" italics="true" pageId="8" pageNumber="529">Sulcophanaeus noctis</emphasis>
|
||
(Bates)
|
||
</taxonomicName>
|
||
, frequently collected in the
|
||
<collectingRegion id="49B91C82FFADEA2F2FBAF319FEBFFE9A" box="[221,345,331,357]" country="Colombia" name="Magdalena" pageId="8" pageNumber="529">Magdalena</collectingRegion>
|
||
Valley.
|
||
</paragraph>
|
||
<caption id="DF0282E8FFA3EA212FF0F531FB6AF819" ID-DOI="http://doi.org/10.5281/zenodo.4306560" ID-Zenodo-Dep="4306560" httpUri="https://zenodo.org/record/4306560/files/figure.png" pageId="6" pageNumber="527" startId="6.[151,250,1891,1915]" targetBox="[354,1233,187,1858]" targetPageId="6">
|
||
<paragraph id="8BC2D260FFA3EA212FF0F531FB6AF819" blockId="6.[151,1436,1891,2023]" pageId="6" pageNumber="527">
|
||
<emphasis id="B9090E72FFA3EA212FF0F531FEF5F884" bold="true" box="[151,275,1891,1915]" pageId="6" pageNumber="527">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C7DA9E3FFA3EA212E7EF531FE92F885" authorityName="Erichson" authorityYear="1847" box="[281,372,1891,1914]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9090E72FFA3EA212E7EF531FE92F885" box="[281,372,1891,1914]" italics="true" pageId="6" pageNumber="527">Scatimus</emphasis>
|
||
</taxonomicName>
|
||
pronotum, mesepimeron, metasternum, abdomen, and metafemur.
|
||
<emphasis id="B9090E72FFA3EA212B7AF531FBC9F884" bold="true" box="[1053,1071,1891,1915]" pageId="6" pageNumber="527">A</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA3EA212B5BF531FB03F884" authorityName="Balthasar" authorityYear="1939" box="[1084,1253,1891,1915]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B9090E72FFA3EA212B5BF531FB03F884" box="[1084,1253,1891,1915]" italics="true" pageId="6" pageNumber="527">Scatimus strandi</emphasis>
|
||
</taxonomicName>
|
||
pronotum (lateral view);
|
||
<emphasis id="B9090E72FFA3EA212F86F5D5FF14F860" bold="true" box="[225,242,1927,1951]" pageId="6" pageNumber="527">B</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA3EA212E66F5D5FE57F860" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[257,433,1927,1951]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA3EA212E66F5D5FE57F860" box="[257,433,1927,1951]" italics="true" pageId="6" pageNumber="527">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
pronotum (lateral view);
|
||
<emphasis id="B9090E72FFA3EA212DA7F5D5FD34F860" bold="true" box="[704,722,1927,1951]" pageId="6" pageNumber="527">C</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA3EA212D85F5D5FC69F860" authorityName="Balthasar" authorityYear="1939" box="[738,911,1927,1951]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B9090E72FFA3EA212D85F5D5FC69F860" box="[738,911,1927,1951]" italics="true" pageId="6" pageNumber="527">Scatimus strandi</emphasis>
|
||
</taxonomicName>
|
||
mesepimeron (ventral view);
|
||
<emphasis id="B9090E72FFA3EA212BACF5D5FB3BF860" bold="true" box="[1227,1245,1927,1951]" pageId="6" pageNumber="527">D</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA3EA212B8AF5D5FA7AF860" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[1261,1436,1927,1951]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA3EA212B8AF5D5FA7AF860" box="[1261,1436,1927,1951]" italics="true" pageId="6" pageNumber="527">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
mesepimeron (ventral view);
|
||
<emphasis id="B9090E72FFA3EA212EA7F5F9FE37F83C" bold="true" box="[448,465,1963,1987]" pageId="6" pageNumber="527">E</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA3EA212EBAF5F9FD6CF83C" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[477,650,1963,1987]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA3EA212EBAF5F9FD6CF83C" box="[477,650,1963,1987]" italics="true" pageId="6" pageNumber="527">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
metasternum (ventral view);
|
||
<emphasis id="B9090E72FFA3EA212CD4F5F9FC24F83C" bold="true" box="[947,962,1963,1987]" pageId="6" pageNumber="527">F</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA3EA212CA9F5F9FB9DF83C" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[974,1147,1963,1987]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA3EA212CA9F5F9FB9DF83C" box="[974,1147,1963,1987]" italics="true" pageId="6" pageNumber="527">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
abdomen;
|
||
<emphasis id="B9090E72FFA3EA212B8DF5F9FB18F83C" bold="true" box="[1258,1278,1963,1987]" pageId="6" pageNumber="527">G</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA3EA212A6DF5F9FF23F818" authorityName="Kohlmann & Solis" authorityYear="1996" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="erinnyos">
|
||
<emphasis id="B9090E72FFA3EA212A6DF5F9FF23F818" italics="true" pageId="6" pageNumber="527">Scatimus erinnyos</emphasis>
|
||
</taxonomicName>
|
||
metafemur (ventral view);
|
||
<emphasis id="B9090E72FFA3EA212EBBF59DFE16F818" bold="true" box="[476,496,1999,2023]" pageId="6" pageNumber="527">H</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA3EA212E9AF59DFD4FF818" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[509,681,1999,2023]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA3EA212E9AF59DFD4FF818" box="[509,681,1999,2023]" italics="true" pageId="6" pageNumber="527">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
metafemur (ventral view). Scale bar = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0282E8FFA2EA202FF0F427FC41F80A" ID-DOI="http://doi.org/10.5281/zenodo.4306562" ID-Zenodo-Dep="4306562" httpUri="https://zenodo.org/record/4306562/files/figure.png" pageId="7" pageNumber="528" startId="7.[151,250,1653,1677]" targetBox="[194,1410,204,1617]" targetPageId="7">
|
||
<paragraph id="8BC2D260FFA2EA202FF0F427FC41F80A" blockId="7.[151,1436,1653,2037]" pageId="7" pageNumber="528">
|
||
<emphasis id="B9090E72FFA2EA202FF0F427FEF4F972" bold="true" box="[151,274,1653,1677]" pageId="7" pageNumber="528">FIGURE 4.</emphasis>
|
||
Male genitalia of
|
||
<taxonomicName id="4C7DA9E3FFA2EA202EADF424FDC3F972" authorityName="Erichson" authorityYear="1847" box="[458,549,1654,1677]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9090E72FFA2EA202EADF424FDC3F972" box="[458,549,1654,1677]" italics="true" pageId="7" pageNumber="528">Scatimus</emphasis>
|
||
</taxonomicName>
|
||
from Colombia.
|
||
<emphasis id="B9090E72FFA2EA202DB4F427FD03F972" bold="true" box="[723,741,1653,1677]" pageId="7" pageNumber="528">A</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA2EA202D95F424FC58F972" authorityName="Martinez" authorityYear="1988" box="[754,958,1653,1677]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="fernandezi">
|
||
<emphasis id="B9090E72FFA2EA202D95F424FC58F972" box="[754,958,1653,1677]" italics="true" pageId="7" pageNumber="528">Scatimus fernandezi</emphasis>
|
||
</taxonomicName>
|
||
aedeagus (lateral view, scale bar = 0.5 mm);
|
||
<emphasis id="B9090E72FFA2EA202AE2F427FA70F972" bold="true" box="[1413,1430,1653,1677]" pageId="7" pageNumber="528">B</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA2EA202FF0F4C8FEDCF94E" authorityName="Harold" authorityYear="1862" box="[151,314,1690,1713]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="ovatus">
|
||
<emphasis id="B9090E72FFA2EA202FF0F4C8FEDCF94E" box="[151,314,1690,1713]" italics="true" pageId="7" pageNumber="528">Scatimus ovatus</emphasis>
|
||
</taxonomicName>
|
||
aedeagus (lateral view, scale bar = 0.5 mm);
|
||
<emphasis id="B9090E72FFA2EA202D9BF4CBFCE8F94E" bold="true" box="[764,782,1689,1713]" pageId="7" pageNumber="528">C</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA2EA202C7DF4C8FC24F94E" authorityName="Balthasar" authorityYear="1939" box="[794,962,1689,1713]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B9090E72FFA2EA202C7DF4C8FC24F94E" box="[794,962,1689,1713]" italics="true" pageId="7" pageNumber="528">Scatimus strandi</emphasis>
|
||
</taxonomicName>
|
||
aedeagus (lateral view, scale bar = 0.5 mm);
|
||
<emphasis id="B9090E72FFA2EA202AE3F4CBFA70F94E" bold="true" box="[1412,1430,1689,1713]" pageId="7" pageNumber="528">D</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA2EA202FF0F4ECFEA2F92A" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[151,324,1726,1749]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA2EA202FF0F4ECFEA2F92A" box="[151,324,1726,1749]" italics="true" pageId="7" pageNumber="528">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
aedeagus (lateral view, scale bar = 0.5 mm);
|
||
<emphasis id="B9090E72FFA2EA202C6AF4EFFCF8F92A" bold="true" box="[781,798,1725,1749]" pageId="7" pageNumber="528">E</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA2EA202C4CF4ECFC11F92A" authorityName="Martinez" authorityYear="1988" box="[811,1015,1725,1749]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="fernandezi">
|
||
<emphasis id="B9090E72FFA2EA202C4CF4ECFC11F92A" box="[811,1015,1725,1749]" italics="true" pageId="7" pageNumber="528">Scatimus fernandezi</emphasis>
|
||
</taxonomicName>
|
||
parameres (ventral view, scale bar = 0.25 mm);
|
||
<emphasis id="B9090E72FFA2EA202FB3F4B3FF05F906" bold="true" box="[212,227,1761,1785]" pageId="7" pageNumber="528">F</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA2EA202F88F4B0FE74F906" authorityName="Harold" authorityYear="1862" box="[239,402,1762,1785]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="ovatus">
|
||
<emphasis id="B9090E72FFA2EA202F88F4B0FE74F906" box="[239,402,1762,1785]" italics="true" pageId="7" pageNumber="528">Scatimus ovatus</emphasis>
|
||
</taxonomicName>
|
||
parameres (ventral view, scale bar = 0.25 mm);
|
||
<emphasis id="B9090E72FFA2EA202C10F4B3FC6DF906" bold="true" box="[887,907,1761,1785]" pageId="7" pageNumber="528">G</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA2EA202CF0F4B0FBA6F906" authorityName="Balthasar" authorityYear="1939" box="[919,1088,1761,1785]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B9090E72FFA2EA202CF0F4B0FBA6F906" box="[919,1088,1761,1785]" italics="true" pageId="7" pageNumber="528">Scatimus strandi</emphasis>
|
||
</taxonomicName>
|
||
parameres (ventral view, scale bar = 0.25 mm);
|
||
<emphasis id="B9090E72FFA2EA202E7AF557FED7F8E2" bold="true" box="[285,305,1797,1821]" pageId="7" pageNumber="528">H</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA2EA202E5AF554FE0DF8E2" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[317,491,1798,1821]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA2EA202E5AF554FE0DF8E2" box="[317,491,1798,1821]" italics="true" pageId="7" pageNumber="528">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
parameres (ventral view, scale bar = 0.25 mm);
|
||
<emphasis id="B9090E72FFA2EA202CBFF557FC04F8E2" bold="true" box="[984,994,1797,1821]" pageId="7" pageNumber="528">I</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA2EA202C88F554FB5AF8E2" authorityName="Martinez" authorityYear="1988" box="[1007,1212,1797,1821]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="fernandezi">
|
||
<emphasis id="B9090E72FFA2EA202C88F554FB5AF8E2" box="[1007,1212,1797,1821]" italics="true" pageId="7" pageNumber="528">Scatimus fernandezi</emphasis>
|
||
</taxonomicName>
|
||
superior right peripheral sclerite (SRP, scale = 0.25 mm);
|
||
<emphasis id="B9090E72FFA2EA202D76F57BFDF8F8BE" bold="true" box="[529,542,1833,1857]" pageId="7" pageNumber="528">J</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA2EA202D4CF578FD29F8BE" authorityName="Harold" authorityYear="1862" box="[555,719,1834,1857]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="ovatus">
|
||
<emphasis id="B9090E72FFA2EA202D4CF578FD29F8BE" box="[555,719,1834,1857]" italics="true" pageId="7" pageNumber="528">Scatimus ovatus</emphasis>
|
||
</taxonomicName>
|
||
superior right peripheral sclerite (SRP, scale = 0.25 mm);
|
||
<emphasis id="B9090E72FFA2EA202A47F57BFAD2F8BE" bold="true" box="[1312,1332,1833,1857]" pageId="7" pageNumber="528">K</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA2EA202A26F578FF39F89A" authorityName="Balthasar" authorityYear="1939" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B9090E72FFA2EA202A26F578FF39F89A" italics="true" pageId="7" pageNumber="528">Scatimus strandi</emphasis>
|
||
</taxonomicName>
|
||
superior right peripheral sclerite (SRP, scale = 0.25 mm);
|
||
<emphasis id="B9090E72FFA2EA202C5BF51FFCABF89A" bold="true" box="[828,845,1869,1893]" pageId="7" pageNumber="528">L</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA2EA202C3CF51CFBECF89A" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[859,1034,1870,1893]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA2EA202C3CF51CFBECF89A" box="[859,1034,1870,1893]" italics="true" pageId="7" pageNumber="528">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
superior right peripheral sclerite (SRP, scale = 0.25 mm);
|
||
<emphasis id="B9090E72FFA2EA202E30F523FE89F876" bold="true" box="[343,367,1905,1929]" pageId="7" pageNumber="528">M</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA2EA202E1BF520FDAFF876" authorityName="Martinez" authorityYear="1988" box="[380,585,1905,1929]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="fernandezi">
|
||
<emphasis id="B9090E72FFA2EA202E1BF520FDAFF876" box="[380,585,1905,1929]" italics="true" pageId="7" pageNumber="528">Scatimus fernandezi</emphasis>
|
||
</taxonomicName>
|
||
additional (AS), subaxial (SA), and axial (A) sclerites superposed, scale bar = 0.25 mm;
|
||
<emphasis id="B9090E72FFA2EA202FABF5C7FF38F852" bold="true" box="[204,222,1941,1965]" pageId="7" pageNumber="528">N</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA2EA202F8BF5C4FE76F852" authorityName="Harold" authorityYear="1862" box="[236,400,1942,1965]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="ovatus">
|
||
<emphasis id="B9090E72FFA2EA202F8BF5C4FE76F852" box="[236,400,1942,1965]" italics="true" pageId="7" pageNumber="528">Scatimus ovatus</emphasis>
|
||
</taxonomicName>
|
||
additional (AS), subaxial (SA), and axial (A) sclerites superposed, scale bar = 0.25 mm;
|
||
<emphasis id="B9090E72FFA2EA202A47F5C7FAD2F852" bold="true" box="[1312,1332,1941,1965]" pageId="7" pageNumber="528">O</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA2EA202A26F5C4FF39F82E" authorityName="Balthasar" authorityYear="1939" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B9090E72FFA2EA202A26F5C4FF39F82E" italics="true" pageId="7" pageNumber="528">Scatimus strandi</emphasis>
|
||
</taxonomicName>
|
||
additional (AS), subaxial (SA), and axial (A) sclerites superposed, scale bar = 0.25 mm;
|
||
<emphasis id="B9090E72FFA2EA202B0FF5EBFB91F82E" bold="true" box="[1128,1143,1977,2001]" pageId="7" pageNumber="528">P</emphasis>
|
||
,
|
||
<taxonomicName id="4C7DA9E3FFA2EA202BE3F5E8FAD6F82E" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[1156,1328,1978,2001]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFA2EA202BE3F5E8FAD6F82E" box="[1156,1328,1978,2001]" italics="true" pageId="7" pageNumber="528">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
additional (AS), subaxial (SA), and axial (A) sclerites superposed, scale bar = 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC2D260FFADEA2F2FA0F33CFAE2FDC2" blockId="8.[151,1437,150,1437]" pageId="8" pageNumber="529">
|
||
<emphasis id="B9090E72FFADEA2F2FA0F33CFB9EFE76" bold="true" box="[199,1144,366,393]" pageId="8" pageNumber="529">
|
||
On the phylogenetic placement of
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2D01F33DFCC9FE76" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[614,815,367,393]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFADEA2F2D01F33DFCC9FE76" bold="true" box="[614,815,367,393]" italics="true" pageId="8" pageNumber="529">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
within the genus
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2B6FF33DFB94FE76" authorityName="Erichson" authorityYear="1847" box="[1032,1138,367,393]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9090E72FFADEA2F2B6FF33DFB94FE76" bold="true" box="[1032,1138,367,393]" italics="true" pageId="8" pageNumber="529">Scatimus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The cladistic analysis resulted in a single parsimonious cladogram with length: 122 steps, CI = 0.680, RI = 0.811. The topology of the strict consensus cladogram (
|
||
<figureCitation id="1346CEE5FFADEA2F2EFDF3E5FE39FE2E" box="[410,479,438,465]" captionStart="FIGURE 5" captionStartId="9.[151,250,1361,1385]" captionTargetBox="[160,1436,526,1338]" captionTargetId="figure-141@9.[151,1436,518,1338]" captionTargetPageId="9" captionText="FIGURE 5. Results of the phylogenetic analyses. A, The single most parsimonious tree from TNT, with character evolution of Scatimus; all characters states are treated as unordered and equally weighted. Characters were mapped on branches using unambiguous character changes in Winclada (black circles, non homoplasious changes; white circles, homoplasious changes); numbers above the circles indicate characters, and number below circles indicate their states. B), Strict consensus of the five most parsimonious trees (length = 122 steps; consistency index = 0.680; retention index = 0.811) obtained with heuristic search. Clade support is indicated on branches (Bremer support values/bootstrap proportions)." figureDoi="http://doi.org/10.5281/zenodo.4306564" httpUri="https://zenodo.org/record/4306564/files/figure.png" pageId="8" pageNumber="529">Fig. 5</figureCitation>
|
||
) is similar to the one obtained by
|
||
<bibRefCitation id="EFECAF91FFADEA2F2C15F3E5FB2EFE2E" author="Genier, F. & Kohlmann, B." box="[882,1224,439,465]" pageId="8" pageNumber="529" pagination="57 - 111" refId="ref8928" refString="Genier, F. & Kohlmann, B. (2003) Revision of the Neotropical dung beetle genera Scatimus Erichson and Scatrichus gen. nov. (Coleoptera: Scarabaeidae: Scarabaeinae). Fabreries, 28, 57 - 111." type="journal article" year="2003">Génier and Kohlmann (2003)</bibRefCitation>
|
||
. The three species groups were recovered, the
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2EBAF389FDA3FE0B" authorityName="Harold" authorityYear="1862" box="[477,581,475,501]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="ovatus">
|
||
<emphasis id="B9090E72FFADEA2F2EBAF389FE0DFE0A" box="[477,491,475,501]" italics="true" pageId="8" pageNumber="529">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFADEA2F2E9BF389FDA3FE0B" box="[508,581,475,500]" italics="true" pageId="8" pageNumber="529">ovatus</emphasis>
|
||
</taxonomicName>
|
||
clade was recovered as monophyletic with the following non-ambiguous synapomorphies: mesoventrites with a deep anteromedian fossa and with setae medially (
|
||
<superScript id="7C087F28FFADEA2F2BFAF3ADFB24FDF3" attach="none" box="[1181,1218,510,537]" fontSize="6" pageId="8" pageNumber="529">232</superScript>
|
||
), dorsal portion of paramere widely membranous at juncture medially (
|
||
<superScript id="7C087F28FFADEA2F2D87F070FCE3FDCF" attach="none" box="[736,773,546,573]" fontSize="6" pageId="8" pageNumber="529">551</superScript>
|
||
), hook of paramere strongly developed (
|
||
<superScript id="7C087F28FFADEA2F2BA9F070FB12FDCF" attach="right" box="[1230,1268,546,573]" fontSize="6" pageId="8" pageNumber="529">562</superScript>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC2D260FFADEA2F2FA0F015FD49FD56" blockId="8.[151,1437,150,1437]" pageId="8" pageNumber="529">
|
||
The
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2F9AF015FE68FD9E" authorityName="Erichson" authorityYear="1847" box="[253,398,583,609]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="cucullatus">
|
||
<emphasis id="B9090E72FFADEA2F2F9AF015FEEDFD9E" box="[253,267,583,609]" italics="true" pageId="8" pageNumber="529">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFADEA2F2E7CF015FE68FD9E" box="[283,398,583,609]" italics="true" pageId="8" pageNumber="529">cucullatus</emphasis>
|
||
</taxonomicName>
|
||
clade also was recovered as monophyletic, but with ambiguous synapomorphies: body in dorsal view elongate, lateral edges parallel on median third (
|
||
<superScript id="7C087F28FFADEA2F2C24F039FCB1FD87" attach="none" box="[835,855,618,645]" fontSize="6" pageId="8" pageNumber="529">11</superScript>
|
||
), head with a single straight frontal carina (
|
||
<superScript id="7C087F28FFADEA2F2A22F039FABAFD87" attach="none" box="[1349,1372,618,645]" fontSize="6" pageId="8" pageNumber="529">60</superScript>
|
||
), and anterolateral angle of elytron subquadrate (
|
||
<superScript id="7C087F28FFADEA2F2D1EF0DDFD78FD63" attach="right" box="[633,670,654,681]" fontSize="6" pageId="8" pageNumber="529">181</superScript>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC2D260FFADEA2F2FA0F0E1FD29FCA2" blockId="8.[151,1437,150,1437]" pageId="8" pageNumber="529">
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2FA0F0E1FE6EFD33" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[199,392,691,717]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFADEA2F2FA0F0E1FE6EFD33" box="[199,392,691,717]" italics="true" pageId="8" pageNumber="529">Scatimus strenua</emphasis>
|
||
</taxonomicName>
|
||
clustered within the
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2D1DF0E1FD00FD32" authorityName="Balthasar" authorityYear="1939" box="[634,742,691,717]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B9090E72FFADEA2F2D1DF0E1FD6EFD32" box="[634,648,691,717]" italics="true" pageId="8" pageNumber="529">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFADEA2F2DF0F0E1FD00FD32" box="[663,742,691,717]" italics="true" pageId="8" pageNumber="529">strandi</emphasis>
|
||
</taxonomicName>
|
||
clade, supported by a single non-ambiguous synapomorphy: head with a single frontal arcuate carina with lateral extremities anteriorly directed (
|
||
<superScript id="7C087F28FFADEA2F2B52F085FBAAFD1B" attach="none" box="[1077,1100,726,753]" fontSize="6" pageId="8" pageNumber="529">63</superScript>
|
||
), and forming a monophyletic unit with
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2E65F0A9FE83FCEB" authorityName="Genier & Kohlmann" authorityYear="2003" box="[258,357,763,789]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="onorei">
|
||
<emphasis id="B9090E72FFADEA2F2E65F0A9FEF6FCEA" box="[258,272,763,789]" italics="true" pageId="8" pageNumber="529">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFADEA2F2E7AF0A9FE83FCEB" box="[285,357,763,788]" italics="true" pageId="8" pageNumber="529">onorei</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2EE5F0A9FE18FCEB" authorityName="Kohlmann & Solis" authorityYear="1996" box="[386,510,763,789]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="erinnyos">
|
||
<emphasis id="B9090E72FFADEA2F2EE5F0A9FE76FCEA" box="[386,400,763,789]" italics="true" pageId="8" pageNumber="529">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFADEA2F2EF9F0A9FE18FCEB" box="[414,510,763,788]" italics="true" pageId="8" pageNumber="529">erinnyos</emphasis>
|
||
</taxonomicName>
|
||
+ (
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2D43F0A9FD69FCEA" authorityName="Balthasar" authorityYear="1939" box="[548,655,763,789]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B9090E72FFADEA2F2D43F0A9FDD4FCEA" box="[548,562,763,789]" italics="true" pageId="8" pageNumber="529">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFADEA2F2D27F0A9FD69FCEA" box="[576,655,763,789]" italics="true" pageId="8" pageNumber="529">strandi</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2DCBF0A9FCFDFCEB" authorityName="Martinez-Revelo, Lopera-Toro, & Medina" authorityYear="2020" box="[684,795,763,789]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="strenua">
|
||
<emphasis id="B9090E72FFADEA2F2DCBF0A9FD5CFCEA" box="[684,698,763,789]" italics="true" pageId="8" pageNumber="529">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFADEA2F2DAFF0A9FCFDFCEB" box="[712,795,763,788]" italics="true" pageId="8" pageNumber="529">strenua</emphasis>
|
||
</taxonomicName>
|
||
), supported by the following two ambiguous synapomorphies: non setiferous punctures of pronotal lateral declivity each with a minute granule medially (
|
||
<superScript id="7C087F28FFADEA2F2BB2F14DFB1CFCD3" attach="none" box="[1237,1274,798,825]" fontSize="6" pageId="8" pageNumber="529">141</superScript>
|
||
), and posteromedial area of metepisternon flat or concave (
|
||
<superScript id="7C087F28FFADEA2F2DFEF111FD59FCAF" attach="right" box="[665,703,834,861]" fontSize="6" pageId="8" pageNumber="529">301</superScript>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC2D260FFADEA2F2FA0F135FDCAFB16" blockId="8.[151,1437,150,1437]" pageId="8" pageNumber="529">
|
||
<emphasis id="B9090E72FFADEA2F2FA0F135FE19FC7F" bold="true" box="[199,511,870,897]" pageId="8" pageNumber="529">Updated species checklist.</emphasis>
|
||
Two species of the
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2D85F135FCAEFC7F" authorityName="Harold" authorityYear="1862" box="[738,840,871,897]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="ovatus">
|
||
<emphasis id="B9090E72FFADEA2F2D85F135FD16FC7E" box="[738,752,871,897]" italics="true" pageId="8" pageNumber="529">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFADEA2F2D98F135FCAEFC7F" box="[767,840,871,896]" italics="true" pageId="8" pageNumber="529">ovatus</emphasis>
|
||
</taxonomicName>
|
||
clade,
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2CFEF135FC19FC7F" authorityName="Harold" authorityYear="1862" box="[921,1023,871,897]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="ovatus">
|
||
<emphasis id="B9090E72FFADEA2F2CFEF135FC19FC7F" box="[921,1023,871,897]" italics="true" pageId="8" pageNumber="529">S. ovatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2B50F135FB2CFC7E" authorityName="Martinez" authorityYear="1988" box="[1079,1226,871,897]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="fernandezi">
|
||
<emphasis id="B9090E72FFADEA2F2B50F135FB2CFC7E" box="[1079,1226,871,897]" italics="true" pageId="8" pageNumber="529">S. fernandezi</emphasis>
|
||
</taxonomicName>
|
||
were already confirmed for
|
||
<collectingCountry id="F36A92F0FFADEA2F2E74F1D9FE67FC5A" box="[275,385,907,933]" name="Colombia" pageId="8" pageNumber="529">Colombia</collectingCountry>
|
||
by
|
||
<bibRefCitation id="EFECAF91FFADEA2F2EC9F1D9FD7CFC5A" author="Medina, C. A. & Lopera-Toro, A. & Vitolo, A. & Gill, B." box="[430,666,907,933]" pageId="8" pageNumber="529" pagination="131 - 144" refId="ref9454" refString="Medina, C. A., Lopera-Toro, A., Vitolo, A. & Gill, B. (2001) Escarabajos coprofagos (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. Biota Colombiana, 2, 131 - 144." type="journal article" year="2001">
|
||
Medina
|
||
<emphasis id="B9090E72FFADEA2F2D6CF1D9FDD8FC5A" box="[523,574,907,933]" italics="true" pageId="8" pageNumber="529">et al</emphasis>
|
||
. (2001)
|
||
</bibRefCitation>
|
||
.
|
||
<bibRefCitation id="EFECAF91FFADEA2F2DC0F1D9FCB2FC5A" author="Escobar, F." box="[679,852,907,933]" pageId="8" pageNumber="529" pagination="197 - 210" refId="ref8707" refString="Escobar, F. (2000) Diversidad y distribucion de los escarabajos del estiercol (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. In: Piera, F. M., Morrone, J. J. & Melic, A. (Eds.), Hacia un proyecto CYTED para el inventario y estimacion de la diversidad entomologica en Iberoamerica: PrIBES- 2000. Monografias Tercer Milenio I. SEA, Zaragoza, pp. 197 - 210." type="book chapter" year="2000">Escobar (2000)</bibRefCitation>
|
||
first reported
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2C90F1D9FB85FC5A" authorityName="Balthasar" authorityYear="1939" box="[1015,1123,907,933]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B9090E72FFADEA2F2C90F1D9FB85FC5A" box="[1015,1123,907,933]" italics="true" pageId="8" pageNumber="529">S. strandi</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F36A92F0FFADEA2F2BEEF1D9FB11FC5A" box="[1161,1271,907,933]" name="Colombia" pageId="8" pageNumber="529">Colombia</collectingCountry>
|
||
but neither he nor
|
||
<bibRefCitation id="EFECAF91FFADEA2F2FA3F1FDFE4BFC36" author="Medina, C. A. & Lopera-Toro, A. & Vitolo, A. & Gill, B." box="[196,429,943,969]" pageId="8" pageNumber="529" pagination="131 - 144" refId="ref9454" refString="Medina, C. A., Lopera-Toro, A., Vitolo, A. & Gill, B. (2001) Escarabajos coprofagos (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. Biota Colombiana, 2, 131 - 144." type="journal article" year="2001">
|
||
Medina
|
||
<emphasis id="B9090E72FFADEA2F2E45F1FDFEB2FC36" box="[290,340,943,969]" italics="true" pageId="8" pageNumber="529">et al</emphasis>
|
||
. (2001)
|
||
</bibRefCitation>
|
||
included a locality, possibly causing
|
||
<bibRefCitation id="EFECAF91FFADEA2F2C31F1FDFB76FC36" author="Genier, F. & Kohlmann, B." box="[854,1168,943,969]" pageId="8" pageNumber="529" pagination="57 - 111" refId="ref8928" refString="Genier, F. & Kohlmann, B. (2003) Revision of the Neotropical dung beetle genera Scatimus Erichson and Scatrichus gen. nov. (Coleoptera: Scarabaeidae: Scarabaeinae). Fabreries, 28, 57 - 111." type="journal article" year="2003">Génier & Kohlmann (2003)</bibRefCitation>
|
||
to miss the presence of the species in
|
||
<collectingCountry id="F36A92F0FFADEA2F2E5FF181FE4CFC12" box="[312,426,979,1005]" name="Colombia" pageId="8" pageNumber="529">Colombia</collectingCountry>
|
||
. Recently, the presence of
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2DBEF181FCA3FC12" authorityName="Balthasar" authorityYear="1939" box="[729,837,979,1005]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="strandi">
|
||
<emphasis id="B9090E72FFADEA2F2DBEF181FCA3FC12" box="[729,837,979,1005]" italics="true" pageId="8" pageNumber="529">S. strandi</emphasis>
|
||
</taxonomicName>
|
||
was confirmed in the department of
|
||
<collectingRegion id="49B91C82FFADEA2F2B8EF181FAC5FC12" box="[1257,1315,979,1005]" country="Colombia" name="Meta" pageId="8" pageNumber="529">Meta</collectingRegion>
|
||
(Cárdenas
|
||
<emphasis id="B9090E72FFADEA2F2FF0F1A5FF2EFBEE" box="[151,200,1015,1041]" italics="true" pageId="8" pageNumber="529">et al</emphasis>
|
||
. 2020) and we here report three additional localities for the species in the departments of
|
||
<collectingRegion id="49B91C82FFADEA2F2BD2F1A5FAF7FBEE" box="[1205,1297,1015,1041]" country="Colombia" name="Caqueta" pageId="8" pageNumber="529">Caquetá</collectingRegion>
|
||
and
|
||
<collectingRegion id="49B91C82FFADEA2F2A20F1A5FA7EFBEE" box="[1351,1432,1015,1041]" country="Colombia" name="Narino" pageId="8" pageNumber="529">Nariño</collectingRegion>
|
||
.
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2FF0F649FE9CFBCA" authorityName="Martinez" authorityYear="1988" box="[151,378,1051,1077]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="fernandezi">
|
||
<emphasis id="B9090E72FFADEA2F2FF0F649FE9CFBCA" box="[151,378,1051,1077]" italics="true" pageId="8" pageNumber="529">Scatimus fernandezi</emphasis>
|
||
</taxonomicName>
|
||
previously was recorded in
|
||
<collectingRegion id="49B91C82FFADEA2F2DDEF649FCC6FBCA" box="[697,800,1051,1077]" country="Colombia" name="Casanare" pageId="8" pageNumber="529">Casanare</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EFECAF91FFADEA2F2C49F649FBEEFBCA" author="Medina, C. A. & Lopera-Toro, A. & Vitolo, A. & Gill, B." box="[814,1032,1051,1077]" pageId="8" pageNumber="529" pagination="131 - 144" refId="ref9454" refString="Medina, C. A., Lopera-Toro, A., Vitolo, A. & Gill, B. (2001) Escarabajos coprofagos (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. Biota Colombiana, 2, 131 - 144." type="journal article" year="2001">
|
||
Medina
|
||
<emphasis id="B9090E72FFADEA2F2CEAF649FC59FBCA" box="[909,959,1051,1077]" italics="true" pageId="8" pageNumber="529">et al</emphasis>
|
||
. 2001
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFECAF91FFADEA2F2B73F649FADBFBCA" author="Genier, F. & Kohlmann, B." box="[1044,1341,1051,1077]" pageId="8" pageNumber="529" pagination="57 - 111" refId="ref8928" refString="Genier, F. & Kohlmann, B. (2003) Revision of the Neotropical dung beetle genera Scatimus Erichson and Scatrichus gen. nov. (Coleoptera: Scarabaeidae: Scarabaeinae). Fabreries, 28, 57 - 111." type="journal article" year="2003">Génier & Kohlmann 2003</bibRefCitation>
|
||
) and we here add new departmental records from
|
||
<collectingRegion id="49B91C82FFADEA2F2D38F66DFD7FFBA6" box="[607,665,1087,1113]" country="Colombia" name="Meta" pageId="8" pageNumber="529">Meta</collectingRegion>
|
||
and
|
||
<collectingRegion id="49B91C82FFADEA2F2DAAF66DFCA4FBA6" box="[717,834,1087,1113]" country="Colombia" name="Putumayo" pageId="8" pageNumber="529">Putumayo</collectingRegion>
|
||
.
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2C2BF66DFBE7FBA7" authorityName="Harold" authorityYear="1862" box="[844,1025,1087,1113]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="ovatus">
|
||
<emphasis id="B9090E72FFADEA2F2C2BF66DFBE7FBA7" box="[844,1025,1087,1113]" italics="true" pageId="8" pageNumber="529">Scatimus ovatus</emphasis>
|
||
</taxonomicName>
|
||
was reported in
|
||
<bibRefCitation id="EFECAF91FFADEA2F2BDFF66DFA7AFBA6" author="Medina, C. A. & Lopera-Toro, A. & Vitolo, A. & Gill, B." box="[1208,1436,1087,1113]" pageId="8" pageNumber="529" pagination="131 - 144" refId="ref9454" refString="Medina, C. A., Lopera-Toro, A., Vitolo, A. & Gill, B. (2001) Escarabajos coprofagos (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. Biota Colombiana, 2, 131 - 144." type="journal article" year="2001">
|
||
Medina
|
||
<emphasis id="B9090E72FFADEA2F2A73F66DFAA3FBA6" box="[1300,1349,1087,1113]" italics="true" pageId="8" pageNumber="529">et al</emphasis>
|
||
. (2001)
|
||
</bibRefCitation>
|
||
without a precise locality;
|
||
<bibRefCitation id="EFECAF91FFADEA2F2EA2F631FCF5FB82" author="Genier, F. & Kohlmann, B." box="[453,787,1123,1149]" pageId="8" pageNumber="529" pagination="57 - 111" refId="ref8928" refString="Genier, F. & Kohlmann, B. (2003) Revision of the Neotropical dung beetle genera Scatimus Erichson and Scatrichus gen. nov. (Coleoptera: Scarabaeidae: Scarabaeinae). Fabreries, 28, 57 - 111." type="journal article" year="2003">Génier and Kohlmann (2003)</bibRefCitation>
|
||
registered it in
|
||
<collectingRegion id="49B91C82FFADEA2F2CA0F631FBA1FB82" box="[967,1095,1123,1149]" country="Colombia" name="Magdalena" pageId="8" pageNumber="529">Magdalena</collectingRegion>
|
||
;
|
||
<bibRefCitation id="EFECAF91FFADEA2F2B34F631FAF8FB82" author="Solis, C. & Noriega, J. & Herrera, G." box="[1107,1310,1123,1149]" pageId="8" pageNumber="529" pagination="33 - 41" refId="ref9803" refString="Solis, C., Noriega, J. & Herrera, G. (2011) Escarabajos coprofagos (Coleoptera: Scarabaeinae) en tres bosques secos del departamento del Atlantico-Colombia. Boletin del Museo de Entomologia de la Universidad del Valle, 12, 33 - 41." type="journal article" year="2011">
|
||
Solís
|
||
<emphasis id="B9090E72FFADEA2F2BF4F631FB23FB82" box="[1171,1221,1123,1149]" italics="true" pageId="8" pageNumber="529">et al</emphasis>
|
||
. (2011)
|
||
</bibRefCitation>
|
||
mentioned “confer”
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2F98F6D5FE52FB5F" authorityName="Harold" authorityYear="1862" box="[255,436,1159,1185]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="ovatus">
|
||
<emphasis id="B9090E72FFADEA2F2F98F6D5FE52FB5F" box="[255,436,1159,1185]" italics="true" pageId="8" pageNumber="529">Scatimus ovatus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingRegion id="49B91C82FFADEA2F2EB0F6D5FDA5FB5E" box="[471,579,1159,1185]" country="Colombia" name="Atlantico" pageId="8" pageNumber="529">Atlántico</collectingRegion>
|
||
;
|
||
<bibRefCitation id="EFECAF91FFADEA2F2D29F6D5FC13FB5E" author="Gonzalez-Alvarado, A. & Medina, C. A." box="[590,1013,1158,1185]" pageId="8" pageNumber="529" pagination="36 - 44" refId="ref9016" refString="Gonzalez-Alvarado, A. & Medina, C. A. (2015) Listado de especies de escarabajos coprofagos (Coleoptera: Scarabaeidae: Scarabaeinae) de bosque seco de Colombia. Biota Colombiana, 16, 36 - 44." type="journal article" year="2015">González-Alvarado & Medina (2015)</bibRefCitation>
|
||
reported the species in Bolívar,
|
||
<collectingRegion id="49B91C82FFADEA2F2A07F6D5FF05FB3A" country="Colombia" name="Magdalena" pageId="8" pageNumber="529">Magdalena</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B91C82FFADEA2F2F89F6F9FED7FB3A" box="[238,305,1195,1221]" country="Colombia" name="Sucre" pageId="8" pageNumber="529">Sucre</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B91C82FFADEA2F2E5DF6F9FE6AFB3A" box="[314,396,1195,1221]" country="Colombia" name="Tolima" pageId="8" pageNumber="529">Tolima</collectingRegion>
|
||
, and
|
||
<collectingRegion id="49B91C82FFADEA2F2EA2F6F9FDDFFB3A" box="[453,569,1195,1221]" country="Colombia" name="Santander" pageId="8" pageNumber="529">Santander</collectingRegion>
|
||
; and
|
||
<bibRefCitation id="EFECAF91FFADEA2F2D15F6F9FCB2FB3A" author="Mendivil, J. A. & Giraldo-Echeverri, C. & Quevedo-Vega, C. J." box="[626,852,1195,1221]" pageId="8" pageNumber="529" refId="ref9567" refString="Mendivil, J. A., Giraldo-Echeverri, C. & Quevedo-Vega, C. J. (2020) Escarabajos estercoleros asociados a sistemas de ganaderia sostenible en diferentes regiones de Colombia. Version 2.3. Fundacion Centro para la Investigacion en Sistemas Sostenibles de Produccion Agropecuaria-Cipav. Dataset / Occurrence. https: // doi. org / 10.15472 / yje 84 v" type="book" year="2020">
|
||
Mendivil
|
||
<emphasis id="B9090E72FFADEA2F2DB9F6F9FCE9FB3A" box="[734,783,1195,1221]" italics="true" pageId="8" pageNumber="529">et al</emphasis>
|
||
. 2020
|
||
</bibRefCitation>
|
||
from
|
||
<collectingRegion id="49B91C82FFADEA2F2CF0F6F9FC01FB3A" box="[919,999,1195,1221]" country="Colombia" name="Caldas" pageId="8" pageNumber="529">Caldas</collectingRegion>
|
||
.
|
||
<collectingRegion id="49B91C82FFADEA2F2C88F6F9FB84FB3A" box="[1007,1122,1195,1221]" country="Colombia" name="Antioquia" pageId="8" pageNumber="529">Antioquia</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B91C82FFADEA2F2B0BF6F9FB25FB3A" box="[1132,1219,1195,1221]" country="Colombia" name="Boyaca" pageId="8" pageNumber="529">Boyacá</collectingRegion>
|
||
, and
|
||
<collectingRegion id="49B91C82FFADEA2F2B9BF6F9FA92FB3A" box="[1276,1396,1195,1221]" country="Colombia" name="La Guajira" pageId="8" pageNumber="529">La Guajira</collectingRegion>
|
||
are new departmental records (
|
||
<tableCitation id="C6FFE7DBFFADEA2F2EA0F69DFDFDFB16" box="[455,539,1231,1257]" captionStart="TABLE 1" captionStartId="10.[151,239,260,284]" captionTargetPageId="10" captionText="TABLE 1. Checklist of the genus Scatimus from Colombia, departmental distribution based on examined specimens." httpUri="http://table.plazi.org/id/DF0282E8FFAFEA2D2FF0F356FA9DFEE3" pageId="8" pageNumber="529" tableUuid="DF0282E8FFAFEA2D2FF0F356FA9DFEE3">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC2D260FFADEA2F2FA0F6A1FD8AFA62" blockId="8.[151,1437,150,1437]" pageId="8" pageNumber="529">
|
||
A fifth species could be added to the Colombian list when expeditions are performed on the border with
|
||
<collectingCountry id="F36A92F0FFADEA2F2A0EF6A1FF2FFACE" name="Panama" pageId="8" pageNumber="529">Panama</collectingCountry>
|
||
.
|
||
<materialsCitation id="3B15D83DFFADEA2F2FB1F745FF15FAAA" ID-GBIF-Occurrence="2990035319" collectorName="Cerro Pirre & Estacion Ambiental Cana" country="Panama" latitude="7.7553334" location="Panama" longLatPrecision="13" longitude="-77.6845" pageId="8" pageNumber="529" specimenCount="1">
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2FB1F745FE43FACF" authorityName="Kohlmann & Solis" authorityYear="1996" box="[214,421,1303,1329]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="erinnyos">
|
||
<emphasis id="B9090E72FFADEA2F2FB1F745FE43FACF" box="[214,421,1303,1329]" italics="true" pageId="8" pageNumber="529">Scatimus erinnyos</emphasis>
|
||
</taxonomicName>
|
||
is registered in
|
||
<collectorName id="2688B7B6FFADEA2F2D06F745FD07FACE" box="[609,737,1303,1329]" pageId="8" pageNumber="529">Cerro Pirre</collectorName>
|
||
and the
|
||
<collectorName id="2688B7B6FFADEA2F2C2FF745FB8CFACE" box="[840,1130,1303,1329]" pageId="8" pageNumber="529">Estación Ambiental Cana</collectorName>
|
||
(
|
||
<geoCoordinate id="EE49B4A7FFADEA2F2B1CF744FB12FACE" box="[1147,1268,1302,1329]" degrees="7" direction="north" minutes="45.32" orientation="latitude" pageId="8" pageNumber="529" precision="9" value="7.7553334">7°45.32’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE49B4A7FFADEA2F2A65F745FA69FACE" box="[1282,1423,1303,1329]" degrees="77" direction="west" minutes="41.07" orientation="longitude" pageId="8" pageNumber="529" precision="9" value="-77.6845">77°41.07’W</geoCoordinate>
|
||
),
|
||
<collectingCountry id="F36A92F0FFADEA2F2FF0F769FF15FAAA" box="[151,243,1339,1365]" name="Panama" pageId="8" pageNumber="529">Panama</collectingCountry>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B15D83DFFADEA2F2F9BF769FC24FAAA" ID-GBIF-Occurrence="2990035307" box="[252,962,1339,1365]" country="Colombia" location="Choco" pageId="8" pageNumber="529" specimenCount="1" stateProvince="Embera-Wounaan">
|
||
approximately
|
||
<quantity id="4C857F85FFADEA2F2EC3F769FE3AFAAA" box="[420,476,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="529" unit="km" value="8.0">8 km</quantity>
|
||
from the department of
|
||
<collectingRegion id="49B91C82FFADEA2F2D81F769FCC9FAAA" box="[742,815,1339,1365]" country="Panama" name="Embera-Wounaan" pageId="8" pageNumber="529">Chocó</collectingRegion>
|
||
in
|
||
<collectingCountry id="F36A92F0FFADEA2F2C37F769FC24FAAA" box="[848,962,1339,1365]" name="Colombia" pageId="8" pageNumber="529">Colombia</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B15D83DFFADEA2F2CACF769FD81FA62" ID-GBIF-Occurrence="2990035328" collectorName="It & Choco-Darien" country="Colombia" location="Choco" pageId="8" pageNumber="529" specimenCount="1" stateProvince="Antioquia">
|
||
<collectorName id="2688B7B6FFADEA2F2CACF769FC3AFAAA" box="[971,988,1339,1365]" pageId="8" pageNumber="529">It</collectorName>
|
||
is possible that
|
||
<taxonomicName id="4C7DA9E3FFADEA2F2BECF769FAE0FAAB" authorityName="Kohlmann & Solis" authorityYear="1996" box="[1163,1286,1339,1365]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="erinnyos">
|
||
<emphasis id="B9090E72FFADEA2F2BECF769FB7FFAAA" box="[1163,1177,1339,1365]" italics="true" pageId="8" pageNumber="529">S</emphasis>
|
||
.
|
||
<emphasis id="B9090E72FFADEA2F2BC1F769FAE0FAAB" box="[1190,1286,1339,1364]" italics="true" pageId="8" pageNumber="529">erinnyos</emphasis>
|
||
</taxonomicName>
|
||
is distributed throughout the continuous
|
||
<collectorName id="2688B7B6FFADEA2F2EAFF70DFD81FA86" box="[456,615,1375,1401]" pageId="8" pageNumber="529">Chocó-Darién</collectorName>
|
||
moist forest, that includes the departments of
|
||
<collectingRegion id="49B91C82FFADEA2F2B10F70DFB0CFA86" box="[1143,1258,1375,1401]" country="Colombia" name="Antioquia" pageId="8" pageNumber="529">Antioquia</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B91C82FFADEA2F2B90F70DFAA6FA86" box="[1271,1344,1375,1401]" country="Colombia" name="Cauca" pageId="8" pageNumber="529">Cauca</collectingRegion>
|
||
,
|
||
<location id="8EA284BBFFADEA2F2A2BF70DFA7EFA86" LSID="urn:lsid:plazi:treatment:03D46376FFA7EA2F2FF0F1A5FD8AFA62:8EA284BBFFADEA2F2A2BF70DFA7EFA86" box="[1356,1432,1375,1401]" country="Colombia" name="Choco" pageId="8" pageNumber="529" stateProvince="Antioquia">Chocó</location>
|
||
,
|
||
<collectingRegion id="49B91C82FFADEA2F2FF0F7D1FF0EFA62" box="[151,232,1411,1437]" country="Colombia" name="Narino" pageId="8" pageNumber="529">Nariño</collectingRegion>
|
||
, and
|
||
<collectingRegion id="49B91C82FFADEA2F2E45F7D1FE34FA62" box="[290,466,1411,1437]" country="Colombia" name="Valle del Cauca" pageId="8" pageNumber="529">Valle del Cauca</collectingRegion>
|
||
in
|
||
<collectingCountry id="F36A92F0FFADEA2F2E92F7D1FD81FA62" box="[501,615,1411,1437]" name="Colombia" pageId="8" pageNumber="529">Colombia</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |