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<document id="047742CE284EB2C464FF7856575C8614" ID-DOI="10.11646/zootaxa.4759.4.1" ID-GBIF-Dataset="4569c2ca-cbbb-49cb-8460-494916d5f3d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3740949" ID-ZooBank="131E86B0-BB4B-4D91-8F48-6F2A2207B424" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586154517882" checkinUser="plazi" docAuthor="Park, Kyu-Tek, Mey, Wolfram, Koo, Jun-Mo, Prins, Jurate De, Akite, Perpetra &amp; Cho, Soowon" docDate="2020" docId="03D46157FFA6FFB1FF58FF06FD70FB1A" docLanguage="en" docName="zootaxa.4759.4.1.pdf" docOrigin="Zootaxa 4759 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thubdora villosiphalla Park 2020, sp. nov." docType="treatment" docUuid="157ACA08-AAB4-4551-A8BF-844FFCE7B5BE" docUuidSource="ZooBank" docVersion="5" lastPageNumber="479" masterDocId="FFED192FFFBAFFADFFCFFF9EFF97FFF0" masterDocTitle="Fourteen new species of the genus Thubdora Park, 2018 (Lepidoptera Gelechioidea: Lecithoceridae) from Uganda, and three new combinations in Ptilothyris Walsingham, 1897 from DR Congo" masterLastPageNumber="487" masterPageNumber="451" pageNumber="479" updateTime="1698735467765" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1D4F26D2B5AB40C2E9975DE61ABE4683">Fourteen new species of the genus Thubdora Park, 2018 (Lepidoptera Gelechioidea: Lecithoceridae) from Uganda, and three new combinations in Ptilothyris Walsingham, 1897 from DR Congo</mods:title>
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<mods:namePart id="3A38C3488329DA4D2E53B75FFDE1ED9D">Park, Kyu-Tek</mods:namePart>
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<mods:namePart id="B7D3CF2B9FCCA14B823C5109ACC0C9D1">Mey, Wolfram</mods:namePart>
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<mods:namePart id="62C2EFE8846E63125C57B93670792BA7">Koo, Jun-Mo</mods:namePart>
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<mods:namePart id="6C3BC1AA6391FEC47C9B11975C1B3420">Prins, Jurate De</mods:namePart>
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<mods:namePart id="487BC67B86DECE3B06F88E85C7C29AC3">Akite, Perpetra</mods:namePart>
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<mods:namePart id="0A298E66F1986FFAEB60BF3E60B4E4B7">Cho, Soowon</mods:namePart>
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<subSubSection id="C36783CAFFA6FFB1FF58FF06FDEEFF43" box="[151,633,152,179]" pageId="28" pageNumber="479" type="nomenclature">
<paragraph id="8BC2D041FFA6FFB1FF58FF06FDEEFF43" blockId="28.[151,633,152,215]" box="[151,633,152,179]" pageId="28" pageNumber="479">
<heading id="D08A672DFFA6FFB1FF58FF06FDEEFF43" bold="true" box="[151,633,152,179]" fontSize="11" level="1" pageId="28" pageNumber="479" reason="1">
<emphasis id="B9090C53FFA6FFB1FF58FF06FDEEFF43" bold="true" box="[151,633,152,179]" pageId="28" pageNumber="479">
6.
<taxonomicName id="4C7DABC2FFA6FFB1FF08FF06FD84FF43" ID-CoL="9FSG8" authority="Park" authorityName="Park" authorityYear="2020" box="[199,531,152,179]" class="Insecta" family="Lecithoceridae" genus="Thubdora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="villosiphalla" status="sp. nov.">
<emphasis id="B9090C53FFA6FFB1FF08FF06FE58FF43" bold="true" box="[199,463,152,179]" italics="true" pageId="28" pageNumber="479">Thubdora villosiphalla</emphasis>
Park
</taxonomicName>
,
<taxonomicNameLabel id="A23AB128FFA6FFB1FDD0FF07FDEEFF43" box="[543,633,153,179]" pageId="28" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C36783CAFFA6FFB1FF58FF22FEA1FF27" box="[151,310,188,215]" pageId="28" pageNumber="479" type="description">
<paragraph id="8BC2D041FFA6FFB1FF58FF22FEA1FF27" blockId="28.[151,633,152,215]" box="[151,310,188,215]" pageId="28" pageNumber="479">
(
<figureCitation id="1346CCC4FFA6FFB1FF50FF22FF6EFF27" box="[159,249,188,215]" captionStart="FIGURE 19" captionStartId="27.[151,250,1933,1958]" captionTargetBox="[231,1350,687,1889]" captionTargetId="figure@27.[225,1363,669,1908]" captionTargetPageId="27" captionText="FIGURE 19. Thubdora villosiphalla Park, sp. nov.: A, adult, holotype; B, holotype label; C, antenna; D, male genitalia, holotype, gen. slide no. CIS-7353; E, aedeagus, holotype; F, abdominal segments VIVIII, holotype. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3740987" httpUri="https://zenodo.org/record/3740987/files/figure.png" pageId="28" pageNumber="479">Figs. 19</figureCitation>
AF)
</paragraph>
</subSubSection>
<subSubSection id="C36783CAFFA6FFB1FF58FE9BFBFBFEB3" pageId="28" pageNumber="479" type="materials_examined">
<paragraph id="8BC2D041FFA6FFB1FF58FE9BFBFBFEB3" blockId="28.[151,1437,260,1258]" pageId="28" pageNumber="479">
<emphasis id="B9090C53FFA6FFB1FF58FE9BFED8FEEF" bold="true" box="[151,335,260,287]" pageId="28" pageNumber="479">Type specimen.</emphasis>
<materialsCitation id="3B15DA1CFFA6FFB1FE98FE9AFBFBFEB3" ID-GBIF-Occurrence="2595809595" collectingDate="2019-04-27" collectingDateMax="2019-04-30" collectingDateMin="2019-04-27" collectionCode="NIBR" collectorName="KT Park &amp; JM Koo &amp; JD Kim" country="Uganda" location="Mpigi" pageId="28" pageNumber="479" specimenCount="1" stateProvince="Mpigi" typeStatus="holotype">
<typeStatus id="54C66EE3FFA6FFB1FE98FE9AFE53FEEE" box="[343,452,260,286]" pageId="28" pageNumber="479" type="holotype">Holotype</typeStatus>
: male,
<emphasis id="B9090C53FFA6FFB1FDDAFE9BFDE5FEEF" bold="true" box="[533,626,261,287]" pageId="28" pageNumber="479">
<collectingCountry id="F36A90D1FFA6FFB1FDDAFE9BFDE5FEEF" box="[533,626,261,287]" name="Uganda" pageId="28" pageNumber="479">Uganda</collectingCountry>
</emphasis>
,
<collectingRegion id="49B91EA3FFA6FFB1FD4EFE9AFD5FFEEE" box="[641,712,260,286]" country="Uganda" name="Mpigi" pageId="28" pageNumber="479">Mpigi</collectingRegion>
, Mpanga For., 0˚12ʹ24ʹʹN 32˚18ʹ05ʹʹE,
<date id="FFC3F681FFA6FFB1FB4DFE9BFABEFEEF" box="[1154,1321,260,287]" pageId="28" pageNumber="479" value="2019-04-27" valueMax="2019-04-30" valueMin="2019-04-27">
<collectingDate id="EF870F69FFA6FFB1FB4DFE9BFABEFEEF" box="[1154,1321,260,287]" pageId="28" pageNumber="479" value="2019-04-27" valueMax="2019-04-30" valueMin="2019-04-27">2730 iv 2019</collectingDate>
</date>
,
<collectorName id="2688B597FFA6FFB1FAFAFE9BFA0FFEEE" box="[1333,1432,260,287]" pageId="28" pageNumber="479">KT Park</collectorName>
,
<collectorName id="2688B597FFA6FFB1FF58FEB7FF62FEB3" box="[151,245,297,323]" pageId="28" pageNumber="479">JM Koo</collectorName>
,
<collectorName id="2688B597FFA6FFB1FECFFEB7FECCFEB2" box="[256,347,296,323]" pageId="28" pageNumber="479">JD Kim</collectorName>
; gen. slide no.CIS-7353, COI barcode CBNU119, deposited in
<collectionCode id="ED6C4884FFA6FFB1FBEDFEB7FBFEFEB3" box="[1058,1129,297,323]" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="28" pageNumber="479">NIBR</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36783CAFFA6FFB1FF08FED2FC1DFDEA" pageId="28" pageNumber="479" type="diagnosis">
<paragraph id="8BC2D041FFA6FFB1FF08FED2FC1DFDEA" blockId="28.[151,1437,260,1258]" pageId="28" pageNumber="479">
<emphasis id="B9090C53FFA6FFB1FF08FED2FEADFE97" bold="true" box="[199,314,332,359]" pageId="28" pageNumber="479">Diagnosis</emphasis>
. The new species is superficially similar to
<taxonomicName id="4C7DABC2FFA6FFB1FCD4FED3FC20FE96" authorityName="Park &amp; De Prins" authorityYear="2019" box="[795,951,332,358]" class="Insecta" family="Lecithoceridae" genus="Thubdora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="ghesquierei">
<emphasis id="B9090C53FFA6FFB1FCD4FED3FC20FE96" box="[795,951,332,358]" italics="true" pageId="28" pageNumber="479">T. ghesquierei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7DABC2FFA6FFB1FC24FED3FB12FE96" authorityName="Park" authorityYear="2020" box="[1003,1157,332,358]" class="Insecta" family="Lecithoceridae" genus="Thubdora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="trigonoides">
<emphasis id="B9090C53FFA6FFB1FC24FED3FB12FE96" box="[1003,1157,332,358]" italics="true" pageId="28" pageNumber="479">T. trigonoides</emphasis>
</taxonomicName>
, but can be distinguished from the latter by the more elongated or distinct costal patch on forewing. The male genitalia are similar to those of
<taxonomicName id="4C7DABC2FFA6FFB1FF7AFE0BFEC6FE5E" authorityName="Park" authorityYear="2020" box="[181,337,404,430]" class="Insecta" family="Lecithoceridae" genus="Thubdora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="trigonoides">
<emphasis id="B9090C53FFA6FFB1FF7AFE0BFEC6FE5E" box="[181,337,404,430]" italics="true" pageId="28" pageNumber="479">T. trigonoides</emphasis>
</taxonomicName>
, but they can be distinguished by the following: uncus more concaved on caudal margin medially, latero-caudal processes of the uncus more or less sharply acute apically, and aedeagus with a bundle of long hairs, longer than half of the aedeagus. It is notable that
<taxonomicName id="4C7DABC2FFA6FFB1FD0EFE43FCCBFE06" authorityName="Park" authorityYear="2020" box="[705,860,476,502]" class="Insecta" family="Lecithoceridae" genus="Thubdora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="trigonoides">
<emphasis id="B9090C53FFA6FFB1FD0EFE43FCCBFE06" box="[705,860,476,502]" italics="true" pageId="28" pageNumber="479">T. trigonoides</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7DABC2FFA6FFB1FC5EFE43FBABFE06" box="[913,1084,476,502]" class="Insecta" family="Lecithoceridae" genus="Thubdora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="villosephalla">
<emphasis id="B9090C53FFA6FFB1FC5EFE43FBABFE06" box="[913,1084,476,502]" italics="true" pageId="28" pageNumber="479">T. villosephalla</emphasis>
</taxonomicName>
are grouped as a sister group in our phylogenetic analyses while the two species are well separated.
</paragraph>
</subSubSection>
<subSubSection id="C36783CAFFA6FFB1FF08FDBAFE92FB8E" pageId="28" pageNumber="479" type="description">
<paragraph id="8BC2D041FFA6FFB1FF08FDBAFE1EFD92" blockId="28.[151,1437,260,1258]" pageId="28" pageNumber="479">
<emphasis id="B9090C53FFA6FFB1FF08FDBAFECEFDCF" bold="true" box="[199,345,548,575]" pageId="28" pageNumber="479">Description.</emphasis>
Male (
<figureCitation id="1346CCC4FFA6FFB1FE64FDBAFDD6FDCF" box="[427,577,548,575]" captionStart="FIGURE 19" captionStartId="27.[151,250,1933,1958]" captionTargetBox="[231,1350,687,1889]" captionTargetId="figure@27.[225,1363,669,1908]" captionTargetPageId="27" captionText="FIGURE 19. Thubdora villosiphalla Park, sp. nov.: A, adult, holotype; B, holotype label; C, antenna; D, male genitalia, holotype, gen. slide no. CIS-7353; E, aedeagus, holotype; F, abdominal segments VIVIII, holotype. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3740987" httpUri="https://zenodo.org/record/3740987/files/figure.png" pageId="28" pageNumber="479">Figs. 19A, C</figureCitation>
). Forewing length
<quantity id="4C857DA4FFA6FFB1FCD6FDBAFCE4FDCF" box="[793,883,548,575]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="28" pageNumber="479" unit="mm" value="6.5">6.5 mm</quantity>
.
<emphasis id="B9090C53FFA6FFB1FCB0FDBAFC2BFDCE" box="[895,956,548,574]" italics="true" pageId="28" pageNumber="479">Head</emphasis>
: no distinct separable characteristics from that of
<emphasis id="B9090C53FFA6FFB1FF29FDD7FE1EFD92" box="[230,393,584,610]" italics="true" pageId="28" pageNumber="479">
<taxonomicName id="4C7DABC2FFA6FFB1FF29FDD7FE12FD92" authorityName="Park" authorityYear="2020" box="[230,389,584,610]" class="Insecta" family="Lecithoceridae" genus="Thubdora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="trigonoides">T. trigonoides</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BC2D041FFA6FFB1FF08FDF2FCAAFCE6" blockId="28.[151,1437,260,1258]" pageId="28" pageNumber="479">
<emphasis id="B9090C53FFA6FFB1FF08FDF2FE8FFD76" box="[199,280,620,646]" italics="true" pageId="28" pageNumber="479">Thorax</emphasis>
: Notum and tegula dark brown; ventral surface of legs orange white. Forewing costa arched in basal 1/3, with more or less elongated costal patch near basal 3/4; then slightly oblique; dark brown scales irregularly scattered; antemedian fascia black, well-developed; apex obtuse; termen slightly oblique; fringe black, with narrow, orange-white basal line. Hind wing broader, paler than forewing; apex produced, slightly obtuse; termen concave medially; fringe black, with narrow, orange-white basal line.
</paragraph>
<paragraph id="8BC2D041FFA6FFB1FF08FCBEFE92FB8E" blockId="28.[151,1437,260,1258]" pageId="28" pageNumber="479">
<emphasis id="B9090C53FFA6FFB1FF08FCBEFEA6FCCA" box="[199,305,800,826]" italics="true" pageId="28" pageNumber="479">Abdomen</emphasis>
(
<figureCitation id="1346CCC4FFA6FFB1FE8CFCBEFE3CFCCB" box="[323,427,800,827]" captionStart="FIGURE 19" captionStartId="27.[151,250,1933,1958]" captionTargetBox="[231,1350,687,1889]" captionTargetId="figure@27.[225,1363,669,1908]" captionTargetPageId="27" captionText="FIGURE 19. Thubdora villosiphalla Park, sp. nov.: A, adult, holotype; B, holotype label; C, antenna; D, male genitalia, holotype, gen. slide no. CIS-7353; E, aedeagus, holotype; F, abdominal segments VIVIII, holotype. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3740987" httpUri="https://zenodo.org/record/3740987/files/figure.png" pageId="28" pageNumber="479">Fig. 19F</figureCitation>
): No hair-pencils developed in abdomen; tergite VIII with heavily sclerotized, crescent anterior margin, without spines; sternite VIII not specifically modified. Male genitalia (
<figureCitation id="1346CCC4FFA6FFB1FBBDFCDAFA91FCAF" box="[1138,1286,836,863]" captionStart="FIGURE 19" captionStartId="27.[151,250,1933,1958]" captionTargetBox="[231,1350,687,1889]" captionTargetId="figure@27.[225,1363,669,1908]" captionTargetPageId="27" captionText="FIGURE 19. Thubdora villosiphalla Park, sp. nov.: A, adult, holotype; B, holotype label; C, antenna; D, male genitalia, holotype, gen. slide no. CIS-7353; E, aedeagus, holotype; F, abdominal segments VIVIII, holotype. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3740987" httpUri="https://zenodo.org/record/3740987/files/figure.png" pageId="28" pageNumber="479">Figs. 19D, E</figureCitation>
): Uncus with large, latero-caudal processes, directed outwardly, slightly acute apically; anterior margin concave medially. Basal plate of gnathos with round caudal margin; median process curved pre-apically. Tegumen weakly sclerotized, incised in inverted V-shape on anterior margin. Valva expanded near base on costa, slightly concave beyond; ventral margin straight near base, expanded medially; cucullus narrowed toward apex, strongly upturned, densely setose. Juxta subparallelogramic with concave caudal and lateral sides and acute latero-caudal processes, with a narrow, long stalk antero-ventrally. Vinculum band-like, triangularly produced anteriorly. Aedeagus broad at base, gradually narrowed towards apex; cornuti with two adjacent bundles of long hair-like scales, about half the length of aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C36783CAFFA6FFB1FF08FB16FDFEFB53" box="[199,617,1160,1187]" pageId="28" pageNumber="479" type="distribution">
<paragraph id="8BC2D041FFA6FFB1FF08FB16FDFEFB53" blockId="28.[151,1437,260,1258]" box="[199,617,1160,1187]" pageId="28" pageNumber="479">
<emphasis id="B9090C53FFA6FFB1FF08FB16FECEFB53" bold="true" box="[199,345,1160,1187]" pageId="28" pageNumber="479">Distribution</emphasis>
.
<collectingCountry id="F36A90D1FFA6FFB1FEA8FB16FE29FB52" box="[359,446,1160,1186]" name="Uganda" pageId="28" pageNumber="479">Uganda</collectingCountry>
(
<collectingRegion id="49B91EA3FFA6FFB1FE02FB16FDCAFB53" box="[461,605,1160,1187]" country="Uganda" name="Mpigi" pageId="28" pageNumber="479">Mpigi Distr.</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C36783CAFFA6FFB1FF08FB33FD70FB1A" pageId="28" pageNumber="479" type="etymology">
<paragraph id="8BC2D041FFA6FFB1FF08FB33FD70FB1A" blockId="28.[151,1437,260,1258]" pageId="28" pageNumber="479">
<emphasis id="B9090C53FFA6FFB1FF08FB33FED9FB37" bold="true" box="[199,334,1197,1223]" pageId="28" pageNumber="479">Etymology.</emphasis>
This species name is derived from the Latin
<emphasis id="B9090C53FFA6FFB1FC86FB32FC11FB36" box="[841,902,1196,1222]" italics="true" pageId="28" pageNumber="479">villus</emphasis>
(= shaggy hair), referring to the bundle of long hair-like scales in the aedeagus of the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>