treatments-xml/data/03/FE/87/03FE87CE6204FFB5A6AA56B52109F984.xml

<document id="DAE7ACDC18D8969F46DD45A42E369140" ID-CLB-Dataset="277864" ID-DOI="10.11646/zootaxa.5369.3.6" ID-GBIF-Dataset="ee47ab31-d6c0-4592-8c1f-04103c00a26c" ID-ISSN="1175-5326" ID-Zenodo-Dep="10151690" ID-ZooBank="26FA0386-41F5-4C5C-A9AC-A3014CE71799" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1700256274454" checkinUser="plazi" docAuthor="Anker, Arthur &amp; Assayie, Aymere Awoke" docDate="2023" docId="03FE87CE6204FFB5A6AA56B52109F984" docLanguage="en" docName="zootaxa.5369.3.6.pdf" docOrigin="Zootaxa 5369 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5369.3.6/52251" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Palaemonella yalla Anker &amp; Assayie 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="434" masterDocId="FFC7FFB66205FFB2A63D56222315FFA9" masterDocTitle="A further new species of Palaemonella Dana, 1852 from the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Palaemonidae)" masterLastPageNumber="436" masterPageNumber="427" pageNumber="428" updateTime="1700657320102" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0E789541F2E524870ED4445B3A30420E">A further new species of Palaemonella Dana, 1852 from the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Palaemonidae)</mods:title>
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<mods:namePart id="053B19B15051B305DBD0AACA2B476842">Anker, Arthur</mods:namePart>
<mods:affiliation id="4C506AF71C329A3AF869FC1FB63F9A92">Universidade Federal de Pelotas (UFPEL), Departamento de Ecologia, Zoologia e Genética, Instituto de Biologia, Campus Universitário Capão do Leão, RS, 96010 - 610, Brazil. &amp; Red Sea Research Center, Biological and Environmental Science &amp; Engineering Division, King Abdullah University of Science and Technology (KAUST), Thuwal, Saudi Arabia.</mods:affiliation>
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<mods:namePart id="C84BAEAF635B75AD8E87ED7BEFFCA631">Assayie, Aymere Awoke</mods:namePart>
<mods:affiliation id="6F2763FDD60B0154A07415B6194F66DC">Red Sea Research Center, Biological and Environmental Science &amp; Engineering Division, King Abdullah University of Science and Technology (KAUST), Thuwal, Saudi Arabia.</mods:affiliation>
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<treatment id="03FE87CE6204FFB5A6AA56B52109F984" ID-DOI="http://doi.org/10.5281/zenodo.10169119" ID-GBIF-Taxon="217779639" ID-Zenodo-Dep="10169119" LSID="urn:lsid:plazi:treatment:03FE87CE6204FFB5A6AA56B52109F984" httpUri="http://treatment.plazi.org/id/03FE87CE6204FFB5A6AA56B52109F984" lastPageId="7" lastPageNumber="434" pageId="1" pageNumber="428">
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<paragraph id="8BE836D86204FFB3A6AA56B522F1FF18" blockId="1.[151,484,151,213]" box="[151,484,151,178]" pageId="1" pageNumber="428">
<heading id="D0A081B46204FFB3A6AA56B522F1FF18" bold="true" box="[151,484,151,178]" fontSize="11" level="1" pageId="1" pageNumber="428" reason="1">
<emphasis id="B923EACA6204FFB3A6AA56B522F1FF18" bold="true" box="[151,484,151,178]" pageId="1" pageNumber="428">
<taxonomicName id="4C574D5B6204FFB3A6AA56B52295FF1B" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[151,384,151,178]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="428" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6204FFB3A6AA56B52295FF1B" bold="true" box="[151,384,151,178]" italics="true" pageId="1" pageNumber="428">Palaemonella yalla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21057B16204FFB3A7BA56B522F1FF18" box="[391,484,151,177]" pageId="1" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C34D65536204FFB3A6AA56992207FF7C" box="[151,274,187,213]" pageId="1" pageNumber="428" type="description">
<paragraph id="8BE836D86204FFB3A6AA56992207FF7C" blockId="1.[151,484,151,213]" box="[151,274,187,213]" pageId="1" pageNumber="428">
(
<figureCitation id="136C2A5D6204FFB3A6A25699221CFF7C" box="[159,265,187,213]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,150,174]" captionStartId-1="4.[151,250,1795,1819]" captionStartId-2="6.[151,250,150,174]" captionTargetBox-1="[167,1406,209,1747]" captionTargetId-0="figure-21@2.[151,1436,179,1873]" captionTargetId-1="figure-21@4.[151,1436,181,1771]" captionTargetId-2="figure-17@5.[157,1430,166,1990]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm." captionText-1="FIGURE 2. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446:A, right first pereiopod, lateral; B, same, distal portion of carpus and chela, mesial; C, right second pereiopod, lateral; D, same, distal portion of merus, latteral; E, same, distal portion of carpus, lateral; F, same, distal portion of carpus, mesial; G, same, chela, lateral; H, same, chela fingers slightly open, lateral; I, right third pereiopod, lateral; J, same, distal portion of propodus and dactylus, lateral; K, right fourth pereiopod, lateral; L, right fifth pereiopod, lateral; M, same, distal portion of propodus and dactylus, mesial. Scales are in mm." captionText-2="FIGURE 3. Palaemonella yalla sp. nov., living shrimps in dorsal view; A, holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446; B, non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445. Photographs by A. Anker." figureDoi-0="http://doi.org/10.5281/zenodo.10185395" figureDoi-1="http://doi.org/10.5281/zenodo.10151692" figureDoi-2="http://doi.org/10.5281/zenodo.10185402" httpUri-0="https://zenodo.org/record/10185395/files/figure.png" httpUri-1="https://zenodo.org/record/10151692/files/figure.png" httpUri-2="https://zenodo.org/record/10185402/files/figure.png" pageId="1" pageNumber="428">Figs. 1–3</figureCitation>
)
</paragraph>
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<emphasis id="B923EACA6204FFB3A6AA5721222AFEB4" bold="true" box="[151,319,259,285]" pageId="1" pageNumber="428">Type material</emphasis>
. 
<materialsCitation id="3B3F3C856204FFB3A7735721223CFE04" ID-GBIF-Occurrence="4441120302" collectingDate="2022-12-29" collectionCode="FLMNH, KAUST" collectorName="A. Anker &amp; A. Assayie" country="Saudi Arabia" county="Thuwal" elevation="7" latitude="22.340612" location="Technology" longLatPrecision="1" longitude="39.08753" municipality="King Abdullah University of Science" pageId="1" pageNumber="428" specimenCode="UF 71446" specimenCount="1" stateProvince="Makkah" typeStatus="holotype">
<typeStatus id="54EC887A6204FFB3A773572122AEFEB4" box="[334,443,259,285]" pageId="1" pageNumber="428" type="holotype">Holotype</typeStatus>
: ovigerous female, pocl 
<quantity id="4CAF9B3D6204FFB3A4F257212032FEB7" box="[719,807,259,286]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="1" pageNumber="428" unit="mm" value="3.9">3.9 mm</quantity>
, cl 
<quantity id="4CAF9B3D6204FFB3A56D572120BCFEB7" box="[848,937,259,286]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="1" pageNumber="428" unit="mm" value="5.2">5.2 mm</quantity>
, 
<collectionCode id="ED46AE1D6204FFB3A58857262702FEB7" box="[949,1047,260,286]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="1" pageNumber="428" type="Museum">FLMNH</collectionCode>
<specimenCode id="DBF19EA36204FFB3A22257262781FEB4" box="[1055,1172,259,286]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="1" pageNumber="428" type="Museum">UF 71446</specimenCode>
, 
<collectingCountry id="F34076486204FFB3A2A257212621FEB4" box="[1183,1332,259,285]" name="Saudi Arabia" pageId="1" pageNumber="428">Saudi Arabia</collectingCountry>
, 
<collectingRegion id="4993F83A6204FFB3A37D572123E8FEE8" country="Saudi Arabia" name="Makkah" pageId="1" pageNumber="428">Makkah Province</collectingRegion>
, 
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, 
<collectingMunicipality id="6B8CACA26204FFB3A75457052017FEE8" box="[361,770,295,321]" pageId="1" pageNumber="428">King Abdullah University of Science</collectingMunicipality>
and 
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(
<collectionCode id="ED46AE1D6204FFB3A5FB5705273CFEE8" box="[966,1065,295,321]" pageId="1" pageNumber="428">KAUST</collectionCode>
), near 
<location id="8E8860036204FFB3A24C5705268DFEEB" LSID="urn:lsid:plazi:treatment:03FE87CE6204FFB5A6AA56B52109F984:8E8860036204FFB3A24C5705268DFEEB" box="[1137,1432,295,322]" country="Saudi Arabia" county="Thuwal" latitude="22.340612" longLatPrecision="1" longitude="39.08753" municipality="King Abdullah University of Science" name="King Abdullah Monument" pageId="1" pageNumber="428" stateProvince="Makkah">King Abdullah Monument</location>
, 
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/ 
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, shallow sandflat between small mangrove and deeper channel, some coral rubble and seagrass present, in burrow, suction (yabby) pump, depth 
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, leg. 
<collectorName id="26A2530E6204FFB3A5E3574D2751FE20" box="[990,1092,367,393]" pageId="1" pageNumber="428">A. Anker</collectorName>
&amp; 
<collectorName id="26A2530E6204FFB3A258574D27F5FE20" box="[1125,1248,367,393]" pageId="1" pageNumber="428">A. Assayie</collectorName>
, 
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<collectingDate id="EFADE9F06204FFB3A2D6574D267CFE20" box="[1259,1385,367,393]" pageId="1" pageNumber="428" value="2022-12-29">29.12.2022</collectingDate>
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[fcn AA-22-437].
</materialsCitation>
</paragraph>
<paragraph id="8BE836D86204FFB3A6FA57952682FDB0" blockId="1.[151,1437,259,1906]" pageId="1" pageNumber="428">
Additional material: 
<materialsCitation id="3B3F3C856204FFB3A78957952686FDB0" ID-GBIF-Occurrence="4441120301" collectingDate="2022-10-11" collectionCode="FLMNH" collectorName="A. Anker &amp; J. Meiburg" country="Oman" elevation="2" latitude="20.251833" location="Masirah Island" longLatPrecision="1" longitude="58.624554" municipality="Ghab" pageId="1" pageNumber="428" specimenCode="UF 71445" specimenCount="1">
ovigerous female, pocl 
<quantity id="4CAF9B3D6204FFB3A4FD5795200FFE7B" box="[704,794,439,466]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="428" unit="mm" value="3.4">3.4 mm</quantity>
, cl 
<quantity id="4CAF9B3D6204FFB3A5795795208BFE7B" box="[836,926,439,466]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="1" pageNumber="428" unit="mm" value="5.6">5.6 mm</quantity>
, 
<collectionCode id="ED46AE1D6204FFB3A596579A2718FE7B" box="[939,1037,440,466]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="1" pageNumber="428" type="Museum">FLMNH</collectionCode>
<specimenCode id="DBF19EA36204FFB3A22B579A279EFE7B" box="[1046,1163,439,466]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="1" pageNumber="428" type="Museum">UF 71445</specimenCode>
, 
<collectingCountry id="F34076486204FFB3A2A5579527CAFE78" box="[1176,1247,439,465]" name="Oman" pageId="1" pageNumber="428">Oman</collectingCountry>
, 
<location id="8E8860036204FFB3A2D15795268DFE78" LSID="urn:lsid:plazi:treatment:03FE87CE6204FFB5A6AA56B52109F984:8E8860036204FFB3A2D15795268DFE78" box="[1260,1432,439,465]" country="Oman" latitude="20.251833" longLatPrecision="1" longitude="58.624554" municipality="Ghab" name="Masirah Island" pageId="1" pageNumber="428">Masirah Island</location>
, 
<collectingMunicipality id="6B8CACA26204FFB3A6AA57F923C3FE5C" box="[151,214,475,501]" pageId="1" pageNumber="428">Ghab</collectingMunicipality>
, 
<geoCoordinate id="EE63501F6204FFB3A6DA57F92290FE5F" box="[231,389,475,502]" degrees="20" direction="north" minutes="15" orientation="latitude" pageId="1" pageNumber="428" precision="1" seconds="06.6" value="20.251833">20°15’06.6”N</geoCoordinate>
/ 
<geoCoordinate id="EE63501F6204FFB3A79857F92157FE5F" box="[421,578,475,502]" degrees="58" direction="east" minutes="37" orientation="longitude" pageId="1" pageNumber="428" precision="1" seconds="28.4" value="58.624554">58°37’28.4”E</geoCoordinate>
, silty sandflat with numerous massive corals, sieving muddy water after flipping large coral rock, depth 
<quantity id="4CAF9B3D6204FFB3A7C457DD212CFDB3" box="[505,569,511,538]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="428" unit="m" value="0.2">
<elevation id="007AD1EB6204FFB3A7C457DD212CFDB3" box="[505,569,511,538]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="428" unit="m" value="0.2">0.2 m</elevation>
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(at low tide), leg. 
<collectorName id="26A2530E6204FFB3A53857DD2079FDB0" box="[773,876,511,537]" pageId="1" pageNumber="428">A. Anker</collectorName>
&amp; 
<collectorName id="26A2530E6204FFB3A5AD54222719FDB0" box="[912,1036,511,538]" pageId="1" pageNumber="428">J. Meiburg</collectorName>
, 
<date id="FFE910186204FFB3A22A57DD2786FDB0" box="[1047,1171,511,537]" pageId="1" pageNumber="428" value="2022-10-11">
<collectingDate id="EFADE9F06204FFB3A22A57DD2786FDB0" box="[1047,1171,511,537]" pageId="1" pageNumber="428" value="2022-10-11">10.11.2022</collectingDate>
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[fcn BOMAN-13448]
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.
</paragraph>
</subSubSection>
<subSubSection id="C34D65536204FFB1A6FA540121F6F8C6" lastPageId="3" lastPageNumber="430" pageId="1" pageNumber="428" type="description">
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<emphasis id="B923EACA6204FFB3A6FA54012244FD94" bold="true" box="[199,337,547,573]" pageId="1" pageNumber="428">Description</emphasis>
. Medium-sized palaemonid shrimp with moderately slender, subcylindrical, not particularly compressed or flattened body. Carapace (
<figureCitation id="136C2A5D6204FFB3A443546521EEFDC8" box="[638,763,583,609]" captionStart="FIGURE 1" captionStartId="3.[151,250,150,174]" captionTargetId="figure-21@2.[151,1436,179,1873]" captionTargetPageId="2" captionText="FIGURE 1. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10185395" httpUri="https://zenodo.org/record/10185395/files/figure.png" pageId="1" pageNumber="428">Fig. 1A–C</figureCitation>
) smooth; orbit moderately shallow; inferior orbital angle strongly produced anteriorly; postorbital area with semicircular depression abruptly delimited posteriorly by unarmed, non-elevated ridge; supra-orbital tooth or tubercle absent; antennal tooth marginal, distally sharp, long, almost reaching distodorsal margin of basicerite; hepatic tooth well-developed, slightly smaller than antennal tooth and located distinctly below it; pterygostomial angle broadly rounded. Rostrum (
<figureCitation id="136C2A5D6204FFB3A20654F527C2FD58" box="[1083,1239,727,753]" captionStart="FIGURE 1" captionStartId="3.[151,250,150,174]" captionTargetId="figure-21@2.[151,1436,179,1873]" captionTargetPageId="2" captionText="FIGURE 1. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10185395" httpUri="https://zenodo.org/record/10185395/files/figure.png" pageId="1" pageNumber="428">Fig. 1A–C, P</figureCitation>
) well developed, almost straight, horizontal, about 0.7 times length of carapace; tip sharp, overreaching mid-length of scaphocerite and distal margin of second article of antennular peduncle; dorsal margin and continuing postrostral carina with eight large, equidistant, anteriorly directed, sharp teeth, five of them anterior to posterior orbital margin, one at about same level and two posterior to it in 
<typeStatus id="54EC887A6204FFB3A41355452185FC28" box="[558,656,871,897]" pageId="1" pageNumber="428" type="holotype">holotype</typeStatus>
(six anterior to posterior orbital margin and two posterior in non-type specimen), posterior-most tooth somewhat more separated from remaining teeth; ventral margin with two widely separated teeth both located in its distal half; lateral ridge fading at about mid-length of rostrum.
</paragraph>
<paragraph id="8BE836D86204FFB3A6FA55F123CEFBF0" blockId="1.[151,1437,259,1906]" pageId="1" pageNumber="428">
Pleon (
<figureCitation id="136C2A5D6204FFB3A72855F12262FC44" box="[277,375,979,1005]" captionStart="FIGURE 1" captionStartId="3.[151,250,150,174]" captionTargetId="figure-21@2.[151,1436,179,1873]" captionTargetPageId="2" captionText="FIGURE 1. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10185395" httpUri="https://zenodo.org/record/10185395/files/figure.png" pageId="1" pageNumber="428">Figs. 1D</figureCitation>
, 
<figureCitation id="136C2A5D6204FFB3A7BE55F12284FC44" box="[387,401,979,1005]" captionStart="FIGURE 3" captionStartId="6.[151,250,150,174]" captionTargetId="figure-17@5.[157,1430,166,1990]" captionTargetPageId="5" captionText="FIGURE 3. Palaemonella yalla sp. nov., living shrimps in dorsal view; A, holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446; B, non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.10185402" httpUri="https://zenodo.org/record/10185402/files/figure.png" pageId="1" pageNumber="428">3</figureCitation>
) typical for genus; third pleonite not humped or posteriorly produced; pleura of first to fourth pleonites rounded; fifth pleonite posteroventrally subacutely produced; sixth pleonite 1.3 times as long as fifth pleonite (measured along mid-dorsal line), with sharply produced posterolateral angle and subacute posteroventral angle.
</paragraph>
<paragraph id="8BE836D86204FFB3A6FA524122CDFA9B" blockId="1.[151,1437,259,1906]" pageId="1" pageNumber="428">
Telson (
<figureCitation id="136C2A5D6204FFB3A71F52412282FBD4" box="[290,407,1123,1149]" captionStart="FIGURE 1" captionStartId="3.[151,250,150,174]" captionTargetId="figure-21@2.[151,1436,179,1873]" captionTargetPageId="2" captionText="FIGURE 1. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10185395" httpUri="https://zenodo.org/record/10185395/files/figure.png" pageId="1" pageNumber="428">Fig. 1E, F</figureCitation>
) about twice as long as maximal width, tapering smoothly posteriorly; dorsal surface with two pairs of long, stout, submarginal spiniform setae, at about 0.4 and 0.7 times of telson length, respectively; posterior margin 0.3 times of proximal margin width, subtriangular, with central minute tooth, and three pairs of stout spiniform setae, as follows: short but stout lateral spiniform setae, stouter and much longer intermediate spiniform setae, and slenderer submedian spiniform setae, latter about half as long as intermediate ones and three times as long as lateral ones.
</paragraph>
<paragraph id="8BE836D86204FFB3A6FA5319269EFAD0" blockId="1.[151,1437,259,1906]" pageId="1" pageNumber="428">
Eyestalks (
<figureCitation id="136C2A5D6204FFB3A778531922D7FAFC" box="[325,450,1339,1365]" captionStart="FIGURE 1" captionStartId="3.[151,250,150,174]" captionTargetId="figure-21@2.[151,1436,179,1873]" captionTargetPageId="2" captionText="FIGURE 1. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10185395" httpUri="https://zenodo.org/record/10185395/files/figure.png" pageId="1" pageNumber="428">Fig. 1A, B</figureCitation>
) with anterior margin convex, slightly wider than long (without cornea); cornea well developed and pigmented, as wide as eyestalk, hemispherical, with distinct accessory pigment spot (Nebenauge).
</paragraph>
<paragraph id="8BE836D86204FFB3A6FA53A122EBF9F8" blockId="1.[151,1437,259,1906]" pageId="1" pageNumber="428">
Antennule (
<figureCitation id="136C2A5D6204FFB3A77253A122DBFA34" box="[335,462,1411,1437]" captionStart="FIGURE 1" captionStartId="3.[151,250,150,174]" captionTargetId="figure-21@2.[151,1436,179,1873]" captionTargetPageId="2" captionText="FIGURE 1. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10185395" httpUri="https://zenodo.org/record/10185395/files/figure.png" pageId="1" pageNumber="428">Fig. 1A, B</figureCitation>
) not particularly modified; first article of antennular peduncle stoutest, longest and widest, with moderately developed, sharp distolateral tooth; statocyst visible dorsally, rounded; stylocerite slender, acute distally, its tip reaching mid-length of first article; ventromesial tooth small, acute; second and third articles subequal in length, together about 0.6 of dorsally visible portion of first article; flagella long, slender; dorsal (= lateral) flagellum biramous, fused portion composed of 11 subdivisions, free ramus well developed, slender, with five or so groups of aesthetascs.
</paragraph>
<paragraph id="8BE836D86204FFB3A6FA507923CEF8AC" blockId="1.[151,1437,259,1906]" pageId="1" pageNumber="428">
Antenna (
<figureCitation id="136C2A5D6204FFB3A70A507922A7F9DC" box="[311,434,1627,1653]" captionStart="FIGURE 1" captionStartId="3.[151,250,150,174]" captionTargetId="figure-21@2.[151,1436,179,1873]" captionTargetPageId="2" captionText="FIGURE 1. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10185395" httpUri="https://zenodo.org/record/10185395/files/figure.png" pageId="1" pageNumber="428">Fig. 1A, B</figureCitation>
) typical for genus; basicerite moderately stout, with acute distolateral tooth; carpocerite subcylindrical, short, not reaching 0.3 of scaphocerite length; flagellum not thickened; scaphocerite extending well beyond end of antennular peduncle, about 3.5 times as long as maximal width; blade relatively narrow, with broadly rounded distal margin; lateral margin almost straight; distolateral tooth strong, distinctly exceeding distal margin of blade.
</paragraph>
<paragraph id="8BE836D86204FFB3A6FA512D20AFF8D8" blockId="1.[151,1437,259,1906]" pageId="1" pageNumber="428">Epistome and labrum without specific features. Fourth thoracic sternite with strong, subacute, median process (not illustrated). Fifth thoracic sternite with two lateral lobes adjacent to second pereiopods, each with acute triangular submedian process. Sixth to eight thoracic sternites unarmed.</paragraph>
<subSection id="E2D82F336207FFB0A6AA51AD22AFF801" box="[151,442,1935,1960]" pageId="2" pageNumber="429" type="multiple">
<footnote id="E84C2AD66207FFB0A6AA51AD22AFF801" box="[151,442,1935,1960]" pageId="2" pageNumber="429">
<paragraph id="8BE836D86207FFB0A6AA51AD22AFF801" blockId="2.[151,442,1935,1960]" box="[151,442,1935,1960]" pageId="2" pageNumber="429">
<heading id="D0A081B46207FFB0A6AA51AD22AFF801" bold="true" box="[151,442,1935,1960]" fontSize="10" level="2" pageId="2" pageNumber="429" reason="2">
<emphasis id="B923EACA6207FFB0A6AA51AD22AFF801" bold="true" box="[151,442,1935,1960]" pageId="2" pageNumber="429">Figure legend on next page</emphasis>
</heading>
</paragraph>
</footnote>
</subSection>
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<paragraph id="8BE836D86206FFB1A6AA56B42199FE2F" blockId="3.[151,1437,150,391]" pageId="3" pageNumber="430">
<emphasis id="B923EACA6206FFB1A6AA56B42207FF07" bold="true" box="[151,274,150,174]" pageId="3" pageNumber="430">FIGURE 1.</emphasis>
<taxonomicName id="4C574D5B6206FFB1A72A56B522C9FF06" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[279,476,151,175]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6206FFB1A72A56B522C9FF06" box="[279,476,151,175]" italics="true" pageId="3" pageNumber="430">Palaemonella yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6206FFB1A7DC56B52125FF06" bold="true" box="[481,560,151,175]" pageId="3" pageNumber="430">
<taxonomicNameLabel id="A21057B16206FFB1A7DC56B52125FF06" box="[481,560,151,175]" pageId="3" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm.
</paragraph>
</caption>
<paragraph id="8BE836D86206FFB1A6FA579723EBFD62" blockId="3.[151,1437,437,1903]" pageId="3" pageNumber="430">
Mouthparts typical for genus, as illustrated (
<figureCitation id="136C2A5D6206FFB1A4FA57972053FE66" box="[711,838,437,463]" captionStart="FIGURE 1" captionStartId="3.[151,250,150,174]" captionTargetId="figure-21@2.[151,1436,179,1873]" captionTargetPageId="2" captionText="FIGURE 1. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10185395" httpUri="https://zenodo.org/record/10185395/files/figure.png" pageId="3" pageNumber="430">Fig. 1G–N</figureCitation>
). Mandible (
<figureCitation id="136C2A5D6206FFB1A5E757972726FE66" box="[986,1075,437,463]" captionStart="FIGURE 1" captionStartId="3.[151,250,150,174]" captionTargetId="figure-21@2.[151,1436,179,1873]" captionTargetPageId="2" captionText="FIGURE 1. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10185395" httpUri="https://zenodo.org/record/10185395/files/figure.png" pageId="3" pageNumber="430">Fig. 1G</figureCitation>
) with small, biarticulated palp; incisor process with three stout teeth. Second maxilliped (
<figureCitation id="136C2A5D6206FFB1A52057FB2086FE5A" box="[797,915,473,499]" captionStart="FIGURE 1" captionStartId="3.[151,250,150,174]" captionTargetId="figure-21@2.[151,1436,179,1873]" captionTargetPageId="2" captionText="FIGURE 1. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10185395" httpUri="https://zenodo.org/record/10185395/files/figure.png" pageId="3" pageNumber="430">Fig. 1K, L</figureCitation>
) with epipod bearing rudimentary podobranch consisting of single finger-like lamella. Third maxilliped (
<figureCitation id="136C2A5D6206FFB1A51257DF20A7FDBE" box="[815,946,509,535]" captionStart="FIGURE 1" captionStartId="3.[151,250,150,174]" captionTargetId="figure-21@2.[151,1436,179,1873]" captionTargetPageId="2" captionText="FIGURE 1. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10185395" httpUri="https://zenodo.org/record/10185395/files/figure.png" pageId="3" pageNumber="430">Fig. 1M, N</figureCitation>
) moderately stout, setose; coxa with large, rounded lateral plate (epipod); basis distinctly separated from endopod; antepenultimate article relatively stout, somewhat twisted and flattened laterally, widening distally, ventrolateral margin with row of small spiniform setae; penultimate article 0.7 length of antepenultimate article, about six times as long as wide; ultimate article 0.7 times as long as penultimate, strongly tapering distally, apex with stout spiniform seta; arthrobranch reduced to few lamellae.
</paragraph>
<paragraph id="8BE836D86206FFB1A6FA54F721B6FCD6" blockId="3.[151,1437,437,1903]" pageId="3" pageNumber="430">
First pereiopods (
<figureCitation id="136C2A5D6206FFB1A7AC54F7211BFD46" box="[401,526,725,751]" captionStart="FIGURE 2" captionStartId="4.[151,250,1795,1819]" captionTargetBox="[167,1406,209,1747]" captionTargetId="figure-21@4.[151,1436,181,1771]" captionTargetPageId="4" captionText="FIGURE 2. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446:A, right first pereiopod, lateral; B, same, distal portion of carpus and chela, mesial; C, right second pereiopod, lateral; D, same, distal portion of merus, latteral; E, same, distal portion of carpus, lateral; F, same, distal portion of carpus, mesial; G, same, chela, lateral; H, same, chela fingers slightly open, lateral; I, right third pereiopod, lateral; J, same, distal portion of propodus and dactylus, lateral; K, right fourth pereiopod, lateral; L, right fifth pereiopod, lateral; M, same, distal portion of propodus and dactylus, mesial. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10151692" httpUri="https://zenodo.org/record/10151692/files/figure.png" pageId="3" pageNumber="430">Fig. 2A, B</figureCitation>
) slender; coxa with ventral process; ischium about five times as long as wide; merus about 1.8 times as long as ischium, about eight times as long as wide; carpus 1.2 times as long as merus, widening distally; carpo-propodal brush well developed; chela slender, about 0.6 length of carpus; palm smooth, subcylindrical; fingers subequal in length, 1.2 times as long as palm, with spaced tufts of setae; fingertips curved, faintly crossing; finger cutting edges unarmed.
</paragraph>
<paragraph id="8BE836D86206FFB1A6FA55AB22A7FB36" blockId="3.[151,1437,437,1903]" pageId="3" pageNumber="430">
Second pereiopods (
<figureCitation id="136C2A5D6206FFB1A79655AB2125FC0A" box="[427,560,905,931]" captionStart="FIGURE 2" captionStartId="4.[151,250,1795,1819]" captionTargetBox="[167,1406,209,1747]" captionTargetId="figure-21@4.[151,1436,181,1771]" captionTargetPageId="4" captionText="FIGURE 2. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446:A, right first pereiopod, lateral; B, same, distal portion of carpus and chela, mesial; C, right second pereiopod, lateral; D, same, distal portion of merus, latteral; E, same, distal portion of carpus, lateral; F, same, distal portion of carpus, mesial; G, same, chela, lateral; H, same, chela fingers slightly open, lateral; I, right third pereiopod, lateral; J, same, distal portion of propodus and dactylus, lateral; K, right fourth pereiopod, lateral; L, right fifth pereiopod, lateral; M, same, distal portion of propodus and dactylus, mesial. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10151692" httpUri="https://zenodo.org/record/10151692/files/figure.png" pageId="3" pageNumber="430">Figs. 2C–H</figureCitation>
, 
<figureCitation id="136C2A5D6206FFB1A40155AB215FFC0A" box="[572,586,905,931]" captionStart="FIGURE 3" captionStartId="6.[151,250,150,174]" captionTargetId="figure-17@5.[157,1430,166,1990]" captionTargetPageId="5" captionText="FIGURE 3. Palaemonella yalla sp. nov., living shrimps in dorsal view; A, holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446; B, non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.10185402" httpUri="https://zenodo.org/record/10185402/files/figure.png" pageId="3" pageNumber="430">3</figureCitation>
) equal in size and similar in shape and armature of chela, slender, elongate; coxa with ventral process; ischium almost five times as long as wide; merus 1.2 times as long as ischium, about 6.5 times as long as wide, distolateral margin produced as stout sharp tooth; carpus 1.2 times as long as merus, distally noticeably widening, except for shallow subdistal constriction, distal margin with three bluntly angular (one subacute) lobes; chela 1.6 times as long as carpus, feebly swollen; palm smooth, subcylindrical, about three times as long as high; fingers subequal in length, 0.9 times as long as palm, fingertips curved and crossing distally; cutting margins of pollex and dactylus each with two low, subtriangular teeth proximally fitting into gape between similar teeth on opposed margin.
</paragraph>
<paragraph id="8BE836D86206FFB1A6FA528B2639F9AE" blockId="3.[151,1437,437,1903]" pageId="3" pageNumber="430">
Third pereiopod (
<figureCitation id="136C2A5D6206FFB1A7B3528B22E3FB6A" box="[398,502,1193,1219]" captionStart="FIGURE 2" captionStartId="4.[151,250,1795,1819]" captionTargetBox="[167,1406,209,1747]" captionTargetId="figure-21@4.[151,1436,181,1771]" captionTargetPageId="4" captionText="FIGURE 2. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446:A, right first pereiopod, lateral; B, same, distal portion of carpus and chela, mesial; C, right second pereiopod, lateral; D, same, distal portion of merus, latteral; E, same, distal portion of carpus, lateral; F, same, distal portion of carpus, mesial; G, same, chela, lateral; H, same, chela fingers slightly open, lateral; I, right third pereiopod, lateral; J, same, distal portion of propodus and dactylus, lateral; K, right fourth pereiopod, lateral; L, right fifth pereiopod, lateral; M, same, distal portion of propodus and dactylus, mesial. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10151692" httpUri="https://zenodo.org/record/10151692/files/figure.png" pageId="3" pageNumber="430">Fig. 2I, J</figureCitation>
) very slender, elongate; ischium slightly curved posteriorly, more than eight times as long as wide; merus about 1.8 times as long as ischium, 13 times as long as wide, unarmed; carpus almost 0.7 length of merus, unarmed; propodus 1.6 times as long as carpus, subequal to merus in length, ventral margin with four widely spaced, short spiniform setae, in addition to distal pair of longer spiniform setae flanking base of dactylus; dactylus about 0.3 length of propodus, slender, gently curved, simple, dorsal margin slightly crenulate and with short row of setae. Fourth pereiopod (
<figureCitation id="136C2A5D6206FFB1A448537F21D8FADE" box="[629,717,1373,1399]" captionStart="FIGURE 2" captionStartId="4.[151,250,1795,1819]" captionTargetBox="[167,1406,209,1747]" captionTargetId="figure-21@4.[151,1436,181,1771]" captionTargetPageId="4" captionText="FIGURE 2. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446:A, right first pereiopod, lateral; B, same, distal portion of carpus and chela, mesial; C, right second pereiopod, lateral; D, same, distal portion of merus, latteral; E, same, distal portion of carpus, lateral; F, same, distal portion of carpus, mesial; G, same, chela, lateral; H, same, chela fingers slightly open, lateral; I, right third pereiopod, lateral; J, same, distal portion of propodus and dactylus, lateral; K, right fourth pereiopod, lateral; L, right fifth pereiopod, lateral; M, same, distal portion of propodus and dactylus, mesial. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10151692" httpUri="https://zenodo.org/record/10151692/files/figure.png" pageId="3" pageNumber="430">Fig. 2K</figureCitation>
) generally similar to third pereiopod, with comparatively longer propodus; dactylus about 0.3 length of propodus. Fifth pereiopod (
<figureCitation id="136C2A5D6206FFB1A54653A320E0FA32" box="[891,1013,1409,1435]" captionStart="FIGURE 2" captionStartId="4.[151,250,1795,1819]" captionTargetBox="[167,1406,209,1747]" captionTargetId="figure-21@4.[151,1436,181,1771]" captionTargetPageId="4" captionText="FIGURE 2. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446:A, right first pereiopod, lateral; B, same, distal portion of carpus and chela, mesial; C, right second pereiopod, lateral; D, same, distal portion of merus, latteral; E, same, distal portion of carpus, lateral; F, same, distal portion of carpus, mesial; G, same, chela, lateral; H, same, chela fingers slightly open, lateral; I, right third pereiopod, lateral; J, same, distal portion of propodus and dactylus, lateral; K, right fourth pereiopod, lateral; L, right fifth pereiopod, lateral; M, same, distal portion of propodus and dactylus, mesial. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10151692" httpUri="https://zenodo.org/record/10151692/files/figure.png" pageId="3" pageNumber="430">Fig. 2L, M</figureCitation>
) longest and more slender than fourth pereiopod, with relatively longer ischium and propodus; propodus with five spiniform setae on ventral margin and longer spiniform seta at distomesial angle adjacent to base of dactylus; grooming brush short, composed of two rows of microserrulate setae distally on ventrolateral surface; dactylus barely reaching 0.3 length of propodus.
</paragraph>
<paragraph id="8BE836D86206FFB1A6FA50332759F912" blockId="3.[151,1437,437,1903]" pageId="3" pageNumber="430">
Female pleopods without noteworthy features; second pleopod (not figured) and remaining pleopods each with appendix interna; male characters currently unknown. Uropod (
<figureCitation id="136C2A5D6206FFB1A551501720D0F9E6" box="[876,965,1589,1615]" captionStart="FIGURE 1" captionStartId="3.[151,250,150,174]" captionTargetId="figure-21@2.[151,1436,179,1873]" captionTargetPageId="2" captionText="FIGURE 1. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10185395" httpUri="https://zenodo.org/record/10185395/files/figure.png" pageId="3" pageNumber="430">Fig. 1O</figureCitation>
) extending well beyond telson; protopod robust, lateral lobe distally produced as acute tooth; exopod with lateral margin faintly convex, terminating in small distolateral tooth; adjacent distolateral spiniform seta well developed, far exceeding distolateral tooth, not reaching distal margin of exopod; endopod subequal in length to exopod, more slender, ovate.
</paragraph>
<paragraph id="8BE836D86206FFB1A6FA50E721F6F8C6" blockId="3.[151,1437,437,1903]" pageId="3" pageNumber="430">
<emphasis id="B923EACA6206FFB1A6FA50E72295F976" bold="true" box="[199,384,1733,1759]" pageId="3" pageNumber="430">Colour pattern</emphasis>
. Body largely translucent with pale yellow tinge; carapace with scattered, small, reddish spots; pleon with similar reddish spots, some arranged in diffuse transverse bands along posterior margin of each pleonite; eyestalks with pale reddish bands dorsally, cornea golden; antennules with some pale reddish spots; second pereiopods semi-hyaline with pale orange tinge; remaining appendages largely translucent and colourless, sometimes with yellowish tinge; eggs green (
<figureCitation id="136C2A5D6206FFB1A4AC517721C3F8C6" box="[657,726,1877,1903]" captionStart="FIGURE 3" captionStartId="6.[151,250,150,174]" captionTargetId="figure-17@5.[157,1430,166,1990]" captionTargetPageId="5" captionText="FIGURE 3. Palaemonella yalla sp. nov., living shrimps in dorsal view; A, holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446; B, non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.10185402" httpUri="https://zenodo.org/record/10185402/files/figure.png" pageId="3" pageNumber="430">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF2866506201FFB6A6AA51212199F866" ID-DOI="http://doi.org/10.5281/zenodo.10151692" ID-Zenodo-Dep="10151692" httpUri="https://zenodo.org/record/10151692/files/figure.png" pageId="4" pageNumber="431" startId="4.[151,250,1795,1819]" targetBox="[167,1406,209,1747]" targetPageId="4" targetType="figure">
<paragraph id="8BE836D86201FFB6A6AA51212199F866" blockId="4.[151,1436,1795,1999]" pageId="4" pageNumber="431">
<emphasis id="B923EACA6201FFB6A6AA51212207F8B2" bold="true" box="[151,274,1795,1819]" pageId="4" pageNumber="431">FIGURE 2.</emphasis>
<taxonomicName id="4C574D5B6201FFB6A72A512122C9F8B2" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[279,476,1795,1819]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6201FFB6A72A512122C9F8B2" box="[279,476,1795,1819]" italics="true" pageId="4" pageNumber="431">Palaemonella yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6201FFB6A7DC51212125F8B2" bold="true" box="[481,560,1795,1819]" pageId="4" pageNumber="431">
<taxonomicNameLabel id="A21057B16201FFB6A7DC51212125F8B2" box="[481,560,1795,1819]" pageId="4" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446: A, right first pereiopod, lateral; B, same, distal portion of carpus and chela, mesial; C, right second pereiopod, lateral; D, same, distal portion of merus, latteral; E, same, distal portion of carpus, lateral; F, same, distal portion of carpus, mesial; G, same, chela, lateral; H, same, chela fingers slightly open, lateral; I, right third pereiopod, lateral; J, same, distal portion of propodus and dactylus, lateral; K, right fourth pereiopod, lateral; L, right fifth pereiopod, lateral; M, same, distal portion of propodus and dactylus, mesial. Scales are in mm.
</paragraph>
</caption>
<subSection id="E2D82F336200FFB7A6AA51C222AFF850" box="[151,442,2016,2041]" pageId="5" pageNumber="432" type="multiple">
<footnote id="E84C2AD66200FFB7A6AA51C222AFF850" box="[151,442,2016,2041]" pageId="5" pageNumber="432">
<paragraph id="8BE836D86200FFB7A6AA51C222AFF850" blockId="5.[151,442,2016,2041]" box="[151,442,2016,2041]" pageId="5" pageNumber="432">
<heading id="D0A081B46200FFB7A6AA51C222AFF850" bold="true" box="[151,442,2016,2041]" fontSize="10" level="2" pageId="5" pageNumber="432" reason="2">
<emphasis id="B923EACA6200FFB7A6AA51C222AFF850" bold="true" box="[151,442,2016,2041]" pageId="5" pageNumber="432">Figure legend on next page</emphasis>
</heading>
</paragraph>
</footnote>
</subSection>
<caption id="DF2866506203FFB4A6AA56B421AFFF5E" ID-DOI="http://doi.org/10.5281/zenodo.10185402" ID-Zenodo-Dep="10185402" httpUri="https://zenodo.org/record/10185402/files/figure.png" pageId="6" pageNumber="433" startId="6.[151,250,150,174]" targetBox="[152,1435,166,2011]" targetPageId="5" targetType="figure">
<paragraph id="8BE836D86203FFB4A6AA56B421AFFF5E" blockId="6.[151,1437,150,247]" pageId="6" pageNumber="433">
<emphasis id="B923EACA6203FFB4A6AA56B42200FF07" bold="true" box="[151,277,150,174]" pageId="6" pageNumber="433">FIGURE 3.</emphasis>
<taxonomicName id="4C574D5B6203FFB4A72056B522F1FF06" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[285,484,151,175]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6203FFB4A72056B522F1FF06" box="[285,484,151,175]" italics="true" pageId="6" pageNumber="433">Palaemonella yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6203FFB4A7D156B5212BFF06" bold="true" box="[492,574,151,175]" pageId="6" pageNumber="433">
<taxonomicNameLabel id="A21057B16203FFB4A7D156B5212BFF06" box="[492,574,151,175]" pageId="6" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, living shrimps in dorsal view; A, holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446; B, non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445. Photographs by A. Anker.
</paragraph>
</caption>
<subSubSection id="C34D65536203FFB4A6FA57072130FE02" pageId="6" pageNumber="433" type="etymology">
<paragraph id="8BE836D86203FFB4A6FA57072130FE02" blockId="6.[151,1437,293,787]" pageId="6" pageNumber="433">
<emphasis id="B923EACA6203FFB4A6FA5707225DFE96" bold="true" box="[199,328,293,319]" pageId="6" pageNumber="433">Etymology</emphasis>
. The name of the new species derives from the common and popular Arabic word 
<emphasis id="B923EACA6203FFB4A2D757072625FE96" box="[1258,1328,293,319]" italics="true" pageId="6" pageNumber="433">yallah</emphasis>
(with the subtly pronounced terminal “h” omitted for consonance), meaning “let’s go”, “hurry up” or “alright”, and used in many situations depending on the context (such as yallah! – “let’s go! (for fieldwork)” and yallah! – “alright, let’s go!”); used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C34D65536203FFB4A6FA579727B6FE5A" pageId="6" pageNumber="433" type="distribution">
<paragraph id="8BE836D86203FFB4A6FA579727B6FE5A" blockId="6.[151,1437,293,787]" pageId="6" pageNumber="433">
<emphasis id="B923EACA6203FFB4A6FA5797224CFE66" bold="true" box="[199,345,437,463]" pageId="6" pageNumber="433">Distribution</emphasis>
. Presently known only from two localities in the north-western Indian Ocean, viz. the Saudi Arabian coast of the Red Sea near Thuwal (
<typeStatus id="54EC887A6203FFB4A44257FB21A4FE5A" box="[639,689,473,499]" pageId="6" pageNumber="433">type</typeStatus>
locality) and Masirah Island, eastern 
<collectingCountry id="F34076486203FFB4A26A57FB278AFE5A" box="[1111,1183,473,499]" name="Oman" pageId="6" pageNumber="433">Oman</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D65536203FFB4A6FA57DF2252FCBA" pageId="6" pageNumber="433" type="biology_ecology">
<paragraph id="8BE836D86203FFB4A6FA57DF2252FCBA" blockId="6.[151,1437,293,787]" pageId="6" pageNumber="433">
<emphasis id="B923EACA6203FFB4A6FA57DF2233FDBE" bold="true" box="[199,294,509,535]" pageId="6" pageNumber="433">Ecology</emphasis>
. The 
<typeStatus id="54EC887A6203FFB4A75A57DF2282FDBE" box="[359,407,509,535]" pageId="6" pageNumber="433">type</typeStatus>
locality of 
<taxonomicName id="4C574D5B6203FFB4A42557DF21E6FDBE" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[536,755,509,535]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6203FFB4A42557DF21E6FDBE" box="[536,755,509,535]" italics="true" pageId="6" pageNumber="433">Palaemonella yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6203FFB4A4C757DF2041FDBE" bold="true" box="[762,852,509,535]" pageId="6" pageNumber="433">
<taxonomicNameLabel id="A21057B16203FFB4A4C757DF2041FDBE" box="[762,852,509,535]" pageId="6" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in 
<collectingCountry id="F34076486203FFB4A54557DF271FFDBE" box="[888,1034,509,535]" name="Saudi Arabia" pageId="6" pageNumber="433">Saudi Arabia</collectingCountry>
is the same as that of 
<taxonomicName id="4C574D5B6203FFB4A33957DC267DFDBE" authorityName="Anker &amp; Benzoni" authorityYear="2023" box="[1284,1384,509,535]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="jamila">
<emphasis id="B923EACA6203FFB4A33957DC267DFDBE" box="[1284,1384,509,535]" italics="true" pageId="6" pageNumber="433">P. jamila</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFC64B296203FFB4A6AA540322BCFD92" author="Anker, A. &amp; Benzoni, F." box="[151,425,545,571]" pageId="6" pageNumber="433" pagination="78 - 102" refId="ref4589" refString="Anker, A. &amp; Benzoni, F. (2023) Hidden in plain sight: two new species of decapod crustaceans (Palaemonidae and Porcellanidae) discovered in the Red Sea on a university campus in Saudi Arabia. Zootaxa, 5278 (1), 78 - 102. https: // doi. org / 10.11646 / zootaxa. 5278.1.3" type="journal article" year="2023">Anker &amp; Benzoni 2023</bibRefCitation>
). Both species were collected from burrows of unknown hosts on a shallow (less than 
<quantity id="4CAF9B3D6203FFB4A3B3540323A5FDF6" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="433" unit="m" value="1.0">1 m</quantity>
), near-shore sandflat with patchy seagrass and fragmented coral rubble, not far from mangrove stands (
<figureCitation id="136C2A5D6203FFB4A30854672685FDF6" box="[1333,1424,581,607]" captionStart="FIGURE 4" captionStartId="6.[151,250,1331,1355]" captionTargetBox="[151,1436,811,1308]" captionTargetId="figure-300@6.[151,1436,811,1308]" captionTargetPageId="6" captionText="FIGURE 4. Habitat of Palaemonella yalla sp. nov.: A, type locality, shallow near-shore sandflat with some rocks, coral rubble and patchy seagrass near Thuwal, Red Sea coast of Saudi Arabia, with one of the authors filling a collection vial with water; B, collection locality of the non-type specimen, shallow tidal sandflat with living massive corals and coral rubble on the eastern coast of Masirah Island, Oman. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.10151694" httpUri="https://zenodo.org/record/10151694/files/figure.png" pageId="6" pageNumber="433">Fig. 4A</figureCitation>
). The Omani specimen was collected on a silty sandflat dominated by living, mostly massive corals, near large rocky outcrops (
<figureCitation id="136C2A5D6203FFB4A73554AF2274FD0E" box="[264,353,653,679]" captionStart="FIGURE 4" captionStartId="6.[151,250,1331,1355]" captionTargetBox="[151,1436,811,1308]" captionTargetId="figure-300@6.[151,1436,811,1308]" captionTargetPageId="6" captionText="FIGURE 4. Habitat of Palaemonella yalla sp. nov.: A, type locality, shallow near-shore sandflat with some rocks, coral rubble and patchy seagrass near Thuwal, Red Sea coast of Saudi Arabia, with one of the authors filling a collection vial with water; B, collection locality of the non-type specimen, shallow tidal sandflat with living massive corals and coral rubble on the eastern coast of Masirah Island, Oman. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.10151694" httpUri="https://zenodo.org/record/10151694/files/figure.png" pageId="6" pageNumber="433">Fig. 4B</figureCitation>
). More precisely, this specimen was found by sieving murky water after turning over a massive coral standing at a depth of 
<quantity id="4CAF9B3D6203FFB4A7E85493210EFD62" box="[469,539,689,715]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="433" unit="cm" value="20.0">20 cm</quantity>
at incoming tide; however, it cannot be excluded that the shrimp was dwelling inside some kind of burrow excavated by a larger animal (such as a pair of snapping shrimps or a ghost-shrimp) under the coral.
</paragraph>
</subSubSection>
<caption id="DF2866506203FFB4A6AA531121F7FA1E" ID-DOI="http://doi.org/10.5281/zenodo.10151694" ID-Zenodo-Dep="10151694" httpUri="https://zenodo.org/record/10151694/files/figure.png" pageId="6" pageNumber="433" startId="6.[151,250,1331,1355]" targetBox="[151,1436,811,1308]" targetPageId="6" targetType="figure">
<paragraph id="8BE836D86203FFB4A6AA531121F7FA1E" blockId="6.[151,1437,1331,1464]" pageId="6" pageNumber="433">
<emphasis id="B923EACA6203FFB4A6AA53112206FAE2" bold="true" box="[151,275,1331,1355]" pageId="6" pageNumber="433">FIGURE 4.</emphasis>
Habitat of 
<taxonomicName id="4C574D5B6203FFB4A7B85311215EFAE2" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[389,587,1331,1355]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6203FFB4A7B85311215EFAE2" box="[389,587,1331,1355]" italics="true" pageId="6" pageNumber="433">Palaemonella yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6203FFB4A46C531121B4FAE2" bold="true" box="[593,673,1331,1355]" pageId="6" pageNumber="433">
<taxonomicNameLabel id="A21057B16203FFB4A46C531121B4FAE2" box="[593,673,1331,1355]" pageId="6" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A, type locality, shallow near-shore sandflat with some rocks, coral rubble and patchy seagrass near Thuwal, Red Sea coast of Saudi Arabia, with one of the authors filling a collection vial with water; B, collection locality of the non-type specimen, shallow tidal sandflat with living massive corals and coral rubble on the eastern coast of Masirah Island, Oman. Photographs by A. Anker.
</paragraph>
</caption>
<subSubSection id="C34D65536203FFB5A6FA53C42109F984" lastPageId="7" lastPageNumber="434" pageId="6" pageNumber="433" type="discussion">
<paragraph id="8BE836D86203FFB4A6FA53C42247F87D" blockId="6.[151,1437,1509,2004]" pageId="6" pageNumber="433">
<emphasis id="B923EACA6203FFB4A6FA53C42221F9A9" bold="true" box="[199,308,1510,1536]" pageId="6" pageNumber="433">Remarks</emphasis>
. 
<taxonomicName id="4C574D5B6203FFB4A77553C7213DFA56" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[328,552,1509,1535]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6203FFB4A77553C7213DFA56" box="[328,552,1509,1535]" italics="true" pageId="6" pageNumber="433">Palaemonella yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6203FFB4A40853C42181F9A9" bold="true" box="[565,660,1510,1536]" pageId="6" pageNumber="433">
<taxonomicNameLabel id="A21057B16203FFB4A40853C42181F9A9" box="[565,660,1510,1536]" pageId="6" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is morphologically and probably also phylogenetically closest to 
<taxonomicName id="4C574D5B6203FFB4A6AA5028213EF98D" authority="Komai &amp; Yamada, 2015" authorityName="Komai &amp; Yamada" authorityYear="2015" box="[151,555,1545,1572]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="okunoi">
<emphasis id="B923EACA6203FFB4A6AA50282217F98A" box="[151,258,1545,1571]" italics="true" pageId="6" pageNumber="433">P. okunoi</emphasis>
<bibRefCitation id="EFC64B296203FFB4A730502B213EF98D" author="Komai, T. &amp; Yamada, Y." box="[269,555,1545,1572]" pageId="6" pageNumber="433" pagination="352 - 362" refId="ref5907" refString="Komai, T. &amp; Yamada, Y. (2015) A new species of the palaemonid shrimp genus Palaemonella Dana, 1852 (Crustacea: Decapoda: Caridea) from Okinawa Island, Ryukyu Islands, Japan. Zootaxa, 3964 (3), 352 - 362. https: // doi. org / 10.11646 / zootaxa. 3964.3.4" type="journal article" year="2015">Komai &amp; Yamada, 2015</bibRefCitation>
</taxonomicName>
, which is presently known only from the Ryukyu Islands, 
<collectingCountry id="F34076486203FFB4A2E150282609F98D" box="[1244,1308,1546,1572]" name="Japan" pageId="6" pageNumber="433">Japan</collectingCountry>
(
<bibRefCitation id="EFC64B296203FFB4A313502B2220F9E1" author="Komai, T. &amp; Yamada, Y." pageId="6" pageNumber="433" pagination="352 - 362" refId="ref5907" refString="Komai, T. &amp; Yamada, Y. (2015) A new species of the palaemonid shrimp genus Palaemonella Dana, 1852 (Crustacea: Decapoda: Caridea) from Okinawa Island, Ryukyu Islands, Japan. Zootaxa, 3964 (3), 352 - 362. https: // doi. org / 10.11646 / zootaxa. 3964.3.4" type="journal article" year="2015">Komai &amp; Yamada 2015</bibRefCitation>
). The two species share many morphological features, including the presence of a moderate, unarmed postorbital ridge, and also have very similar colour patterns (cf. 
<figureCitation id="136C2A5D6203FFB4A54D507320A3F9C5" box="[880,950,1617,1644]" captionStart="FIGURE 4" captionStartId="6.[151,250,1331,1355]" captionTargetBox="[151,1436,811,1308]" captionTargetId="figure-300@6.[151,1436,811,1308]" captionTargetPageId="6" captionText="FIGURE 4. Habitat of Palaemonella yalla sp. nov.: A, type locality, shallow near-shore sandflat with some rocks, coral rubble and patchy seagrass near Thuwal, Red Sea coast of Saudi Arabia, with one of the authors filling a collection vial with water; B, collection locality of the non-type specimen, shallow tidal sandflat with living massive corals and coral rubble on the eastern coast of Masirah Island, Oman. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.10151694" httpUri="https://zenodo.org/record/10151694/files/figure.png" pageId="6" pageNumber="433">Fig. 4</figureCitation>
; 
<bibRefCitation id="EFC64B296203FFB4A5FE507327C7F9C5" author="Komai, T. &amp; Yamada, Y." box="[963,1234,1617,1644]" pageId="6" pageNumber="433" pagination="352 - 362" refId="ref5907" refString="Komai, T. &amp; Yamada, Y. (2015) A new species of the palaemonid shrimp genus Palaemonella Dana, 1852 (Crustacea: Decapoda: Caridea) from Okinawa Island, Ryukyu Islands, Japan. Zootaxa, 3964 (3), 352 - 362. https: // doi. org / 10.11646 / zootaxa. 3964.3.4" type="journal article" year="2015">Komai &amp; Yamada 2015</bibRefCitation>
: fig. 1). The most obvious and reliable morphological feature to separate them is the presence in 
<taxonomicName id="4C574D5B6203FFB4A22350542766F926" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[1054,1139,1653,1679]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6203FFB4A22350542766F926" box="[1054,1139,1653,1679]" italics="true" pageId="6" pageNumber="433">P. yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6203FFB4A241505427CDF939" bold="true" box="[1148,1240,1654,1680]" pageId="6" pageNumber="433">
<taxonomicNameLabel id="A21057B16203FFB4A241505427CDF939" box="[1148,1240,1654,1680]" pageId="6" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B923EACA6203FFB4A2DD505427EAF926" box="[1248,1279,1654,1679]" italics="true" pageId="6" pageNumber="433">vs.</emphasis>
absence in 
<taxonomicName id="4C574D5B6203FFB4A3B5505423F6F91A" authorityName="Komai &amp; Yamada" authorityYear="2015" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="okunoi">
<emphasis id="B923EACA6203FFB4A3B5505423F6F91A" italics="true" pageId="6" pageNumber="433">P. okunoi</emphasis>
</taxonomicName>
of a stout sharp tooth on the distoventral margin of the merus of the second pereiopod (cf. 
<figureCitation id="136C2A5D6203FFB4A32550BB2682F91D" box="[1304,1431,1689,1716]" captionStart="FIGURE 2" captionStartId="4.[151,250,1795,1819]" captionTargetBox="[167,1406,209,1747]" captionTargetId="figure-21@4.[151,1436,181,1771]" captionTargetPageId="4" captionText="FIGURE 2. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446:A, right first pereiopod, lateral; B, same, distal portion of carpus and chela, mesial; C, right second pereiopod, lateral; D, same, distal portion of merus, latteral; E, same, distal portion of carpus, lateral; F, same, distal portion of carpus, mesial; G, same, chela, lateral; H, same, chela fingers slightly open, lateral; I, right third pereiopod, lateral; J, same, distal portion of propodus and dactylus, lateral; K, right fourth pereiopod, lateral; L, right fifth pereiopod, lateral; M, same, distal portion of propodus and dactylus, mesial. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10151692" httpUri="https://zenodo.org/record/10151692/files/figure.png" pageId="6" pageNumber="433">Fig. 2C, D</figureCitation>
; 
<bibRefCitation id="EFC64B296203FFB4A6AA509F22B1F971" author="Komai, T. &amp; Yamada, Y." box="[151,420,1725,1752]" pageId="6" pageNumber="433" pagination="352 - 362" refId="ref5907" refString="Komai, T. &amp; Yamada, Y. (2015) A new species of the palaemonid shrimp genus Palaemonella Dana, 1852 (Crustacea: Decapoda: Caridea) from Okinawa Island, Ryukyu Islands, Japan. Zootaxa, 3964 (3), 352 - 362. https: // doi. org / 10.11646 / zootaxa. 3964.3.4" type="journal article" year="2015">Komai &amp; Yamada 2015</bibRefCitation>
: fig. 5). Further differences are the presence in 
<taxonomicName id="4C574D5B6203FFB4A58A509C271EF97E" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[951,1035,1725,1751]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6203FFB4A58A509C271EF97E" box="[951,1035,1725,1751]" italics="true" pageId="6" pageNumber="433">P. yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6203FFB4A22F509C2779F971" bold="true" box="[1042,1132,1726,1752]" pageId="6" pageNumber="433">
<taxonomicNameLabel id="A21057B16203FFB4A22F509C2779F971" box="[1042,1132,1726,1752]" pageId="6" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
of an apical spiniform seta on the ultimate article of the third maxilliped, which seems to be absent in 
<taxonomicName id="4C574D5B6203FFB4A5DB50C0275BF952" authorityName="Komai &amp; Yamada" authorityYear="2015" box="[998,1102,1761,1787]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="okunoi">
<emphasis id="B923EACA6203FFB4A5DB50C0275BF952" box="[998,1102,1761,1787]" italics="true" pageId="6" pageNumber="433">P. okunoi</emphasis>
</taxonomicName>
; the number of articles in the mandibular palp, i.e., two in 
<taxonomicName id="4C574D5B6203FFB4A7E251242126F8B6" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[479,563,1797,1823]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6203FFB4A7E251242126F8B6" box="[479,563,1797,1823]" italics="true" pageId="6" pageNumber="433">P. yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6203FFB4A40651242183F889" bold="true" box="[571,662,1798,1824]" pageId="6" pageNumber="433">
<taxonomicNameLabel id="A21057B16203FFB4A40651242183F889" box="[571,662,1798,1824]" pageId="6" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B923EACA6203FFB4A4A3512421A9F8B6" box="[670,700,1798,1823]" italics="true" pageId="6" pageNumber="433">vs.</emphasis>
one in 
<taxonomicName id="4C574D5B6203FFB4A52E5124206EF8B6" authorityName="Komai &amp; Yamada" authorityYear="2015" box="[787,891,1797,1823]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="okunoi">
<emphasis id="B923EACA6203FFB4A52E5124206EF8B6" box="[787,891,1797,1823]" italics="true" pageId="6" pageNumber="433">P. okunoi</emphasis>
</taxonomicName>
; and the proportions of the dactylus of the third pereiopod, which is relatively longer (0.4 length of propodus) and more slender (almost eight times as long as wide at base) in the new species (
<emphasis id="B923EACA6203FFB4A7E7516C22E4F8CE" box="[474,497,1870,1895]" italics="true" pageId="6" pageNumber="433">vs</emphasis>
. 0.25 length of propodus and less than six times as long as wide in 
<taxonomicName id="4C574D5B6203FFB4A2C6516C2671F8CE" authorityName="Komai &amp; Yamada" authorityYear="2015" box="[1275,1380,1869,1895]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="okunoi">
<emphasis id="B923EACA6203FFB4A2C6516C2671F8CE" box="[1275,1380,1869,1895]" italics="true" pageId="6" pageNumber="433">P. okunoi</emphasis>
</taxonomicName>
) (cf. 
<figureCitation id="136C2A5D6203FFB4A6AA51532235F825" box="[151,288,1905,1932]" captionStart="FIGURE 1" captionStartId="3.[151,250,150,174]" captionTargetId="figure-21@2.[151,1436,179,1873]" captionTargetPageId="2" captionText="FIGURE 1. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10185395" httpUri="https://zenodo.org/record/10185395/files/figure.png" pageId="6" pageNumber="433">Figs. 1G, M</figureCitation>
, 
<figureCitation id="136C2A5D6203FFB4A71651502248F825" box="[299,349,1906,1932]" captionStart="FIGURE 2" captionStartId="4.[151,250,1795,1819]" captionTargetBox="[167,1406,209,1747]" captionTargetId="figure-21@4.[151,1436,181,1771]" captionTargetPageId="4" captionText="FIGURE 2. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446:A, right first pereiopod, lateral; B, same, distal portion of carpus and chela, mesial; C, right second pereiopod, lateral; D, same, distal portion of merus, latteral; E, same, distal portion of carpus, lateral; F, same, distal portion of carpus, mesial; G, same, chela, lateral; H, same, chela fingers slightly open, lateral; I, right third pereiopod, lateral; J, same, distal portion of propodus and dactylus, lateral; K, right fourth pereiopod, lateral; L, right fifth pereiopod, lateral; M, same, distal portion of propodus and dactylus, mesial. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10151692" httpUri="https://zenodo.org/record/10151692/files/figure.png" pageId="6" pageNumber="433">2I, J</figureCitation>
; 
<bibRefCitation id="EFC64B296203FFB4A754515321BEF825" author="Komai, T. &amp; Yamada, Y." box="[361,683,1905,1932]" pageId="6" pageNumber="433" pagination="352 - 362" refId="ref5907" refString="Komai, T. &amp; Yamada, Y. (2015) A new species of the palaemonid shrimp genus Palaemonella Dana, 1852 (Crustacea: Decapoda: Caridea) from Okinawa Island, Ryukyu Islands, Japan. Zootaxa, 3964 (3), 352 - 362. https: // doi. org / 10.11646 / zootaxa. 3964.3.4" type="journal article" year="2015">Komai &amp; Yamada 2015: 354</bibRefCitation>
, figs. 3D, 4A, F, G, note: palp missing in fig. 3E, F). Noteworthy, 
<taxonomicName id="4C574D5B6203FFB4A3B5515023F6F806" authorityName="Komai &amp; Yamada" authorityYear="2015" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="okunoi">
<emphasis id="B923EACA6203FFB4A3B5515023F6F806" italics="true" pageId="6" pageNumber="433">P. okunoi</emphasis>
</taxonomicName>
was collected at depths of 
<quantity id="4CAF9B3D6203FFB4A42D51B72172F819" box="[528,615,1941,1968]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="6" pageNumber="433" unit="m" value="17.5" valueMax="30.0" valueMin="5.0">5–30 m</quantity>
, among coral rubble, whereas 
<taxonomicName id="4C574D5B6203FFB4A58151B4271AF806" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[956,1039,1941,1967]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6203FFB4A58151B4271AF806" box="[956,1039,1941,1967]" italics="true" pageId="6" pageNumber="433">P. yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6203FFB4A22851B4277BF819" bold="true" box="[1045,1134,1942,1968]" pageId="6" pageNumber="433">
<taxonomicNameLabel id="A21057B16203FFB4A22851B4277BF819" box="[1045,1134,1942,1968]" pageId="6" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was collected in shallower water (
<quantity id="4CAF9B3D6203FFB4A6D851982253F87D" box="[229,326,1978,2004]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" metricValueMax="10.0" metricValueMin="2.0" pageId="6" pageNumber="433" unit="m" value="0.6" valueMax="1.0" valueMin="0.2">0.2–1 m</quantity>
).
</paragraph>
<paragraph id="8BE836D86202FFB5A6FA56B521E0FB9C" blockId="7.[151,1437,151,1581]" pageId="7" pageNumber="434">
Other species that are morphologically similar to 
<taxonomicName id="4C574D5B6202FFB5A4C256B52041FF18" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[767,852,151,177]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6202FFB5A4C256B52041FF18" box="[767,852,151,177]" italics="true" pageId="7" pageNumber="434">P. yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6202FFB5A56056B520ADFF18" bold="true" box="[861,952,151,177]" pageId="7" pageNumber="434">
<taxonomicNameLabel id="A21057B16202FFB5A56056B520ADFF18" box="[861,952,151,177]" pageId="7" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are: 
<taxonomicName id="4C574D5B6202FFB5A5C956B52610FF18" authority="Okuno, 1999" authorityName="Okuno" authorityYear="1999" box="[1012,1285,151,177]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="hachijo">
<emphasis id="B923EACA6202FFB5A5C956B52770FF18" box="[1012,1125,151,177]" italics="true" pageId="7" pageNumber="434">P. hachijo</emphasis>
<bibRefCitation id="EFC64B296202FFB5A25356B52610FF18" author="Okuno, J." box="[1134,1285,151,177]" pageId="7" pageNumber="434" pagination="739 - 745" refId="ref6127" refString="Okuno, J. (1999) Palaemonella hachijo, a new species of shrimp (Crustacea: Decapoda: Palaemonidae) from a submarine cave in southern Japan. Proceedings of the Biological Society of Washington, 112, 739 - 745." type="journal article" year="1999">Okuno, 1999</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C574D5B6202FFB5A32E56B5227FFF7C" authority="(Borradaile, 1898)" baseAuthorityName="Borradaile" baseAuthorityYear="1898" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="rotumana">
<emphasis id="B923EACA6202FFB5A32E56B52689FF19" box="[1299,1436,151,176]" italics="true" pageId="7" pageNumber="434">P. rotumana</emphasis>
(
<bibRefCitation id="EFC64B296202FFB5A6A256992277FF7C" author="Borradaile, L. A." box="[159,354,187,213]" pageId="7" pageNumber="434" pagination="376 - 391" refId="ref4744" refString="Borradaile, L. A. (1898) A revision of the Pontoniidae. The Annals and Magazine of Natural History, Series 7, 2 (11), 376 - 391. https: // doi. org / 10.1080 / 00222939808678058" type="journal article" year="1898">Borradaile, 1898</bibRefCitation>
)
</taxonomicName>
; 
<taxonomicName id="4C574D5B6202FFB5A74A569921B9FF7C" authority="(Borradaile, 1915)" baseAuthorityName="Borradaile" baseAuthorityYear="1915" box="[375,684,186,213]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="pottsi">
<emphasis id="B923EACA6202FFB5A74A569922C4FF7D" box="[375,465,187,212]" italics="true" pageId="7" pageNumber="434">P. pottsi</emphasis>
(
<bibRefCitation id="EFC64B296202FFB5A7DC569921B1FF7C" author="Borradaile, L. A." box="[481,676,186,213]" pageId="7" pageNumber="434" pagination="205 - 213" refId="ref4789" refString="Borradaile, L. A. (1915) Notes on Carides. The Annals and Magazine of Natural History, Series 8, 15 (86), 205 - 213. https: // doi. org / 10.1080 / 00222931508693629" type="journal article" year="1915">Borradaile, 1915</bibRefCitation>
)
</taxonomicName>
; 
<taxonomicName id="4C574D5B6202FFB5A48456992082FF7C" authority="Kemp, 1922" authorityName="Kemp" authorityYear="1922" box="[697,919,187,213]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="B923EACA6202FFB5A48456992014FF7C" box="[697,769,187,213]" italics="true" pageId="7" pageNumber="434">P. lata</emphasis>
<bibRefCitation id="EFC64B296202FFB5A53456992082FF7C" author="Kemp, S." box="[777,919,187,213]" pageId="7" pageNumber="434" pagination="113 - 228" refId="ref5855" refString="Kemp, S. (1922) Notes on Crustacea Decapoda in the Indian Museum. XV. Pontoniinae. Records of the Indian Museum, 24, 113 - 228. https: // doi. org / 10.26515 / rzsi / v 24 / i 2 / 1922 / 163464" type="journal article" year="1922">Kemp, 1922</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C574D5B6202FFB5A599569927BFFF7C" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[932,1194,186,213]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B923EACA6202FFB5A59956992708FF7D" box="[932,1053,187,212]" italics="true" pageId="7" pageNumber="434">P. tenuipes</emphasis>
<bibRefCitation id="EFC64B296202FFB5A218569927BFFF7C" author="Dana, J. D." box="[1061,1194,186,213]" pageId="7" pageNumber="434" pagination="10 - 28" refId="ref5362" refString="Dana, J. D. (1852) Conspectus Crustaceorum quae in Orbis Terrarum circumnavigatione, Carolo Wilkes e Classe Republicae Foederatae e Duce, lexit et descripsit. Proceedings of the Academy of Natural Sciences of Philadelphia, 1852, 10 - 28." type="journal article" year="1852">Dana, 1852</bibRefCitation>
</taxonomicName>
; and 
<taxonomicName id="4C574D5B6202FFB5A2DA569923DAFF50" authority="Bruce, 2002" authorityName="Bruce" authorityYear="2002" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="B923EACA6202FFB5A2DA5699265EFF7C" box="[1255,1355,187,213]" italics="true" pageId="7" pageNumber="434">P. foresti</emphasis>
Bruce, 2002
</taxonomicName>
(
<bibRefCitation id="EFC64B296202FFB5A6E056FD2249FF50" author="Dana, J. D." box="[221,348,222,249]" pageId="7" pageNumber="434" pagination="10 - 28" refId="ref5362" refString="Dana, J. D. (1852) Conspectus Crustaceorum quae in Orbis Terrarum circumnavigatione, Carolo Wilkes e Classe Republicae Foederatae e Duce, lexit et descripsit. Proceedings of the Academy of Natural Sciences of Philadelphia, 1852, 10 - 28." type="journal article" year="1852">Dana 1852</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A75A56FD210AFF50" author="Borradaile, L. A." box="[359,543,222,249]" pageId="7" pageNumber="434" pagination="205 - 213" refId="ref4789" refString="Borradaile, L. A. (1915) Notes on Carides. The Annals and Magazine of Natural History, Series 8, 15 (86), 205 - 213. https: // doi. org / 10.1080 / 00222931508693629" type="journal article" year="1915">Borradaile 1915</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A41756FD21A5FF50" author="Kemp, S." box="[554,688,223,249]" pageId="7" pageNumber="434" pagination="113 - 228" refId="ref5855" refString="Kemp, S. (1922) Notes on Crustacea Decapoda in the Indian Museum. XV. Pontoniinae. Records of the Indian Museum, 24, 113 - 228. https: // doi. org / 10.26515 / rzsi / v 24 / i 2 / 1922 / 163464" type="journal article" year="1922">Kemp 1922</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A48156FD205FFF50" author="Okuno, J." box="[700,842,223,249]" pageId="7" pageNumber="434" pagination="739 - 745" refId="ref6127" refString="Okuno, J. (1999) Palaemonella hachijo, a new species of shrimp (Crustacea: Decapoda: Palaemonidae) from a submarine cave in southern Japan. Proceedings of the Biological Society of Washington, 112, 739 - 745." type="journal article" year="1999">Okuno 1999</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A56856FD20FDFF50" author="Bruce, A. J." box="[853,1000,223,249]" pageId="7" pageNumber="434" pagination="277 - 298" refId="ref5091" refString="Bruce, A. J. (2002 a) Notes on some Indo-Pacific Pontoniinae, XLVI. Palaemonella foresti sp. nov., a new pontoniine shrimp from western Australia (Decapoda, Palaemonidae), with a review of the Indo-West Pacific species of the genus Palaemonella Dana, 1852. Crustaceana, 75, 277 - 298. https: // doi. org / 10.1163 / 156854002760095390" type="journal article" year="2002">Bruce 2002a</bibRefCitation>
). The new species can be distinguished from 
<taxonomicName id="4C574D5B6202FFB5A6E857212251FEB4" authorityName="Okuno" authorityYear="1999" box="[213,324,259,285]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="hachijo">
<emphasis id="B923EACA6202FFB5A6E857212251FEB4" box="[213,324,259,285]" italics="true" pageId="7" pageNumber="434">P. hachijo</emphasis>
</taxonomicName>
by the biarticulate mandibular palp (
<emphasis id="B923EACA6202FFB5A4D957212017FEB5" box="[740,770,259,284]" italics="true" pageId="7" pageNumber="434">vs.</emphasis>
unarticulated in 
<taxonomicName id="4C574D5B6202FFB5A5825721273AFEB4" authorityName="Okuno" authorityYear="1999" box="[959,1071,259,285]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="hachijo">
<emphasis id="B923EACA6202FFB5A5825721273AFEB4" box="[959,1071,259,285]" italics="true" pageId="7" pageNumber="434">P. hachijo</emphasis>
</taxonomicName>
); the ultimate article of the third maxilliped apically armed with a stout spiniform seta (
<emphasis id="B923EACA6202FFB5A53857052031FEE9" box="[773,804,295,320]" italics="true" pageId="7" pageNumber="434">vs.</emphasis>
with slender setae in 
<taxonomicName id="4C574D5B6202FFB5A2205705279BFEE8" authorityName="Okuno" authorityYear="1999" box="[1053,1166,295,321]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="hachijo">
<emphasis id="B923EACA6202FFB5A2205705279BFEE8" box="[1053,1166,295,321]" italics="true" pageId="7" pageNumber="434">P. hachijo</emphasis>
</taxonomicName>
); the proximal tooth on the pollex of the second pereiopod being subtriangular (
<emphasis id="B923EACA6202FFB5A53557692032FECD" box="[776,807,331,356]" italics="true" pageId="7" pageNumber="434">vs.</emphasis>
distally serrated in 
<taxonomicName id="4C574D5B6202FFB5A23E57692767FECC" authorityName="Okuno" authorityYear="1999" box="[1027,1138,331,357]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="hachijo">
<emphasis id="B923EACA6202FFB5A23E57692767FECC" box="[1027,1138,331,357]" italics="true" pageId="7" pageNumber="434">P. hachijo</emphasis>
</taxonomicName>
); and the dorsal surface of the telson with much larger spiniform setae, which are also more distant from the lateral margin than in 
<taxonomicName id="4C574D5B6202FFB5A31B574D2680FE20" authorityName="Okuno" authorityYear="1999" box="[1318,1429,367,393]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="hachijo">
<emphasis id="B923EACA6202FFB5A31B574D2680FE20" box="[1318,1429,367,393]" italics="true" pageId="7" pageNumber="434">P. hachijo</emphasis>
</taxonomicName>
, where they are submarginal (cf. 
<bibRefCitation id="EFC64B296202FFB5A7C257B12198FE04" author="Okuno, J." box="[511,653,403,429]" pageId="7" pageNumber="434" pagination="739 - 745" refId="ref6127" refString="Okuno, J. (1999) Palaemonella hachijo, a new species of shrimp (Crustacea: Decapoda: Palaemonidae) from a submarine cave in southern Japan. Proceedings of the Biological Society of Washington, 112, 739 - 745." type="journal article" year="1999">Okuno 1999</bibRefCitation>
: figs. 1–3). The new species can be easily separated from the common and widespread 
<taxonomicName id="4C574D5B6202FFB5A772579522C2FE79" baseAuthorityName="Borradaile" baseAuthorityYear="1898" box="[335,471,439,464]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="rotumana">
<emphasis id="B923EACA6202FFB5A772579522C2FE79" box="[335,471,439,464]" italics="true" pageId="7" pageNumber="434">P. rotumana</emphasis>
</taxonomicName>
by the absence of a small tubercle at the dorsal end of the postrostral ridge (typically present in 
<taxonomicName id="4C574D5B6202FFB5A73157F92286FE5D" baseAuthorityName="Borradaile" baseAuthorityYear="1898" box="[268,403,475,500]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="rotumana">
<emphasis id="B923EACA6202FFB5A73157F92286FE5D" box="[268,403,475,500]" italics="true" pageId="7" pageNumber="434">P. rotumana</emphasis>
</taxonomicName>
); the much longer and slenderer second pereiopods (cf. 
<bibRefCitation id="EFC64B296202FFB5A23557F92798FE5C" author="Bruce, A. J." box="[1032,1165,475,501]" pageId="7" pageNumber="434" pagination="233 - 284" refId="ref4962" refString="Bruce, A. J. (1982) The pontoniine shrimp fauna or Hong Kong. In: Morton, B. &amp; Tseng, C. K. (Eds.), Proceedings of the First International Marine Biological Workshop: The marine Fauna of Hong Kong and southern China, Hong Kong 1980, I, pp. 233 - 284." type="journal article" year="1982">Bruce 1982</bibRefCitation>
: fig. 1); and the absence of reddish brown or pale orange transverse bands on each of the second pereiopod fingers, which are typical for 
<taxonomicName id="4C574D5B6202FFB5A3B557DD2216FD95" baseAuthorityName="Borradaile" baseAuthorityYear="1898" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="rotumana">
<emphasis id="B923EACA6202FFB5A3B557DD2216FD95" italics="true" pageId="7" pageNumber="434">P. rotumana</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="EFC64B296202FFB5A70554012140FD94" author="Anker, A. &amp; De Grave, S." box="[312,597,547,573]" pageId="7" pageNumber="434" pagination="343 - 454" refId="ref4653" refString="Anker, A. &amp; De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, 34, 343 - 454." type="journal article" year="2016">Anker &amp; De Grave 2016</bibRefCitation>
: fig. 96). Furthermore, 
<taxonomicName id="4C574D5B6202FFB5A563540120A6FD94" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[862,947,547,573]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6202FFB5A563540120A6FD94" box="[862,947,547,573]" italics="true" pageId="7" pageNumber="434">P. yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6202FFB5A58654012703FD94" bold="true" box="[955,1046,547,573]" pageId="7" pageNumber="434">
<taxonomicNameLabel id="A21057B16202FFB5A58654012703FD94" box="[955,1046,547,573]" pageId="7" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="4C574D5B6202FFB5A29054012612FD95" baseAuthorityName="Borradaile" baseAuthorityYear="1915" box="[1197,1287,547,572]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="pottsi">
<emphasis id="B923EACA6202FFB5A29054012612FD95" box="[1197,1287,547,572]" italics="true" pageId="7" pageNumber="434">P. pottsi</emphasis>
</taxonomicName>
by the much shorter rostrum and distinctly more slender second to fifth pereiopods (cf. 
<bibRefCitation id="EFC64B296202FFB5A5CF5465276EFDC8" author="Bruce, A. J." box="[1010,1147,583,609]" pageId="7" pageNumber="434" pagination="273 - 287" refId="ref4832" refString="Bruce, A. J. (1970) Observations on the Indo-West-Pacific species of the genus Palaemonella Dana, 1852 (Decapoda, Pontoniinae). Crustaceana, 19, 273 - 287. https: // doi. org / 10.1163 / 156854070 X 00365" type="journal article" year="1970">Bruce 1970</bibRefCitation>
), and from 
<taxonomicName id="4C574D5B6202FFB5A33F54652659FDC8" authorityName="Kemp" authorityYear="1922" box="[1282,1356,583,609]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="B923EACA6202FFB5A33F54652659FDC8" box="[1282,1356,583,609]" italics="true" pageId="7" pageNumber="434">P. lata</emphasis>
</taxonomicName>
by the much longer fused portion of the lateral antennular flagellum (11 subdivisions in the new species 
<emphasis id="B923EACA6202FFB5A2CF5449261CFD2D" box="[1266,1289,619,644]" italics="true" pageId="7" pageNumber="434">vs</emphasis>
. only five in 
<taxonomicName id="4C574D5B6202FFB5A6AA54AD23CAFD00" authorityName="Kemp" authorityYear="1922" box="[151,223,655,681]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="B923EACA6202FFB5A6AA54AD23CAFD00" box="[151,223,655,681]" italics="true" pageId="7" pageNumber="434">P. lata</emphasis>
</taxonomicName>
), the distally armed merus of the second pereiopod (
<emphasis id="B923EACA6202FFB5A50C54AD205DFD01" box="[817,840,655,680]" italics="true" pageId="7" pageNumber="434">vs</emphasis>
. unarmed in 
<taxonomicName id="4C574D5B6202FFB5A5E354AD2733FD00" authorityName="Kemp" authorityYear="1922" box="[990,1062,655,681]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="B923EACA6202FFB5A5E354AD2733FD00" box="[990,1062,655,681]" italics="true" pageId="7" pageNumber="434">P. lata</emphasis>
</taxonomicName>
), and the much slenderer third to fifth pereiopods (cf. 
<bibRefCitation id="EFC64B296202FFB5A74054912110FD64" author="Kemp, S." box="[381,517,691,717]" pageId="7" pageNumber="434" pagination="113 - 228" refId="ref5855" refString="Kemp, S. (1922) Notes on Crustacea Decapoda in the Indian Museum. XV. Pontoniinae. Records of the Indian Museum, 24, 113 - 228. https: // doi. org / 10.26515 / rzsi / v 24 / i 2 / 1922 / 163464" type="journal article" year="1922">Kemp 1922</bibRefCitation>
). In addition, 
<taxonomicName id="4C574D5B6202FFB5A49D549121ECFD65" baseAuthorityName="Borradaile" baseAuthorityYear="1915" box="[672,761,691,716]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="pottsi">
<emphasis id="B923EACA6202FFB5A49D549121ECFD65" box="[672,761,691,716]" italics="true" pageId="7" pageNumber="434">P. pottsi</emphasis>
</taxonomicName>
is an obligate associate of crinoids, and both 
<taxonomicName id="4C574D5B6202FFB5A2C554912644FD65" baseAuthorityName="Borradaile" baseAuthorityYear="1915" box="[1272,1361,691,716]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="pottsi">
<emphasis id="B923EACA6202FFB5A2C554912644FD65" box="[1272,1361,691,716]" italics="true" pageId="7" pageNumber="434">P. pottsi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C574D5B6202FFB5A3B5549123D6FD58" authorityName="Kemp" authorityYear="1922" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="B923EACA6202FFB5A3B5549123D6FD58" italics="true" pageId="7" pageNumber="434">P. lata</emphasis>
</taxonomicName>
have different colour patterns (cf. 
<bibRefCitation id="EFC64B296202FFB5A47854FA21D0FD58" author="Duris, Z." box="[581,709,727,755]" pageId="7" pageNumber="434" pagination="1147 - 1161" refId="ref5408" refString="Duris, Z. (2017) Palaemonid shrimps (Crustacea: Decapoda) of Saudi Arabia from the ' Red Sea Biodiversity Survey' 2011 - 2013, with 11 new records for the Red Sea. Marine Biodiversity, 47, 1147 - 1161. https: // doi. org / 10.1007 / s 12526 - 017 - 0681 - 8" type="journal article" year="2017">Ďuriš 2017</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A4F354F520F0FD58" author="Anker, A. &amp; De Grave, S." box="[718,997,727,753]" pageId="7" pageNumber="434" pagination="343 - 454" refId="ref4653" refString="Anker, A. &amp; De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, 34, 343 - 454." type="journal article" year="2016">Anker &amp; De Grave 2016</bibRefCitation>
: figs. 94, 95). The new species is easily distinguishable from 
<taxonomicName id="4C574D5B6202FFB5A7B554D92117FCBD" authorityName="Dana" authorityYear="1852" box="[392,514,763,788]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B923EACA6202FFB5A7B554D92117FCBD" box="[392,514,763,788]" italics="true" pageId="7" pageNumber="434">P. tenuipes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C574D5B6202FFB5A40654D9218AFCBC" authorityName="Bruce" authorityYear="2002" box="[571,671,763,789]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="B923EACA6202FFB5A40654D9218AFCBC" box="[571,671,763,789]" italics="true" pageId="7" pageNumber="434">P. foresti</emphasis>
</taxonomicName>
by the rostral configuration, and from 
<taxonomicName id="4C574D5B6202FFB5A26154D927C3FCBD" authorityName="Dana" authorityYear="1852" box="[1116,1238,763,788]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B923EACA6202FFB5A26154D927C3FCBD" box="[1116,1238,763,788]" italics="true" pageId="7" pageNumber="434">P. tenuipes</emphasis>
</taxonomicName>
also by the more ventral and more posterior position of the hepatic tooth (cf. 
<bibRefCitation id="EFC64B296202FFB5A50D553D209EFC90" author="Li, X." box="[816,907,799,825]" pageId="7" pageNumber="434" refId="ref5995" refString="Li, X. (2000) Catalog of the genera and species of Pontoniinae Kingsley, 1878. Xueyuan Press, Beijing, 319 pp." type="book" year="2000">Li 2000</bibRefCitation>
: fig. 117; 
<bibRefCitation id="EFC64B296202FFB5A5C4553D2799FC90" author="Bruce, A. J." box="[1017,1164,799,825]" pageId="7" pageNumber="434" pagination="277 - 298" refId="ref5091" refString="Bruce, A. J. (2002 a) Notes on some Indo-Pacific Pontoniinae, XLVI. Palaemonella foresti sp. nov., a new pontoniine shrimp from western Australia (Decapoda, Palaemonidae), with a review of the Indo-West Pacific species of the genus Palaemonella Dana, 1852. Crustaceana, 75, 277 - 298. https: // doi. org / 10.1163 / 156854002760095390" type="journal article" year="2002">Bruce 2002a</bibRefCitation>
: fig. 1B). The remaining species of the genus present more numerous differences with the new species (e.g., 
<bibRefCitation id="EFC64B296202FFB5A26355612616FCF4" author="Holthuis, L. B." box="[1118,1283,835,861]" pageId="7" pageNumber="434" pagination="1 - 48" refId="ref5792" refString="Holthuis, L. B. (1973) Caridean shrimps found in land-locked saltwater pools at four Indo-West Pacific localities (Sinai Peninsula, Funafuti Atoll, Maui and Hawaii Islands), with the description of one new genus and four new species. Zoologische Verhandelingen, Leiden, 128, 1 - 48, pls. 1 - 7." type="journal article" year="1973">Holthuis 1973</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A32C5561268DFCF4" author="Bruce, A. J." box="[1297,1432,834,861]" pageId="7" pageNumber="434" pagination="169 - 185" refId="ref4879" refString="Bruce, A. J. (1975) Further observations on the Indo-West Pacific species of the genus Palaemonella Dana, 1852 (Decapoda Natantia, Pontoniinae). Crustaceana, 29, 169 - 185. https: // doi. org / 10.1163 / 156854075 X 00180" type="journal article" year="1975">Bruce 1975</bibRefCitation>
, 
<bibRefCitation id="EFC64B296202FFB5A6AA554523C4FC28" author="Bruce, A. J." box="[151,209,871,897]" pageId="7" pageNumber="434" refId="ref5028" refString="Bruce, A. J. (1991) Shallow water palaemonoid shrimps from New Caledonia (Crustacea: Decapoda). In: Richer de Forges, B. (Ed.), Le Benthos des fonds meubles des lagons de Nouvelle-Caledonie. Vol. 1. Etudes et Theses. ORSTOM, Paris, pp. 221 - 279." type="book" year="1991">1991</bibRefCitation>
, 
<bibRefCitation id="EFC64B296202FFB5A6E25545223DFC28" author="Bruce, A. J." box="[223,296,871,897]" pageId="7" pageNumber="434" pagination="15 - 18" refId="ref5163" refString="Bruce, A. J. (2002 b) A new species of Palaemonella (Crustacea: Decapoda: Pontoniinae) from East Africa. The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 18, 15 - 18. https: // doi. org / 10.5962 / p. 286294" type="journal article" year="2002">2002b</bibRefCitation>
, 
<bibRefCitation id="EFC64B296202FFB5A70855452265FC28" author="Bruce, A. J." box="[309,368,871,897]" pageId="7" pageNumber="434" pagination="63 - 68" refId="ref5309" refString="Bruce, A. J. (2008) A new species of Palaemonella Dana, 1852, (Crustacea: Decapoda; Pontoniinae) from the Red Sea. Zootaxa, 1844 (1), 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1844.1.6" type="journal article" year="2008">2008</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A74355452153FC28" author="Li, X. &amp; Bruce, A. J." box="[382,582,871,897]" pageId="7" pageNumber="434" pagination="611 - 738" refId="ref6022" refString="Li, X. &amp; Bruce, A. J. (2006) Further Indo-West Pacific palaemonoid shrimps (Crustacea: Decapoda: Palaemonoidea), principally from the New Caledonian region. Journal of Natural History, 40 (11 - 12), 611 - 738. https: // doi. org / 10.1080 / 00222930600763627" type="journal article" year="2006">Li &amp; Bruce 2006</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A46E554521CEFC28" author="Marin, I." box="[595,731,871,897]" pageId="7" pageNumber="434" pagination="375 - 390" refId="ref6082" refString="Marin, I. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Vir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, Leiden, 82, 375 - 390." type="journal article" year="2008">Marin 2008</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A4D455452085FC28" author="Hayashi, K. I." box="[745,912,871,897]" pageId="7" pageNumber="434" pagination="89 - 103" refId="ref5682" refString="Hayashi, K. I. (2009) Redescription of Palaemonella spinulata Yokoya with designation of a neotype and description of a new species of Palaemonella from Japan (Decapoda: Caridea: Palaemonidae). Bulletin of the National Museum of Nature and Science (A), Zoology, Supplement 3, 89 - 103." type="journal article" year="2009">Hayashi, 2009</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A5A355452768FC28" author="Fransen, C. H. J. M. &amp; van der Veer, E. &amp; Frolova, P." box="[926,1149,871,897]" pageId="7" pageNumber="434" pagination="557 - 580" refId="ref5542" refString="Fransen, C. H. J. M., van der Veer, E. &amp; Frolova, P. (2022) A new species of scleractinian associated shrimp of the genus Palaemonella (Crustacea, Decapoda, Palaemonidae) with a redescription of Palaemonella orientalis Dana, 1852. Zootaxa, 5214 (4), 557 - 580. https: // doi. org / 10.11646 / zootaxa. 5214.4.5" type="journal article" year="2022">
Fransen 
<emphasis id="B923EACA6202FFB5A5C25545272CFC28" box="[1023,1081,871,897]" italics="true" pageId="7" pageNumber="434">et al.</emphasis>
2022
</bibRefCitation>
, 
<bibRefCitation id="EFC64B296202FFB5A2B7554527D0FC28" author="Fransen, C. H. J. M. &amp; van der Veer, E. &amp; Duris, Z." box="[1162,1221,871,897]" pageId="7" pageNumber="434" pagination="345 - 381" refId="ref5471" refString="Fransen, C. H. J. M., van der Veer, E. &amp; Duris, Z. (2023) A new species of Palaemonella (Decapoda, Caridea, Palaemonidae) associated with scleractinian corals of the genus Euphyllia Dana. Crustaceana, 96 (4), 345 - 381. https: // doi. org / 10.1163 / 15685403 - bja 10289" type="journal article" year="2023">2023</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A2EC554523C1FC0C" author="Anker, A. &amp; Benzoni, F." pageId="7" pageNumber="434" pagination="78 - 102" refId="ref4589" refString="Anker, A. &amp; Benzoni, F. (2023) Hidden in plain sight: two new species of decapod crustaceans (Palaemonidae and Porcellanidae) discovered in the Red Sea on a university campus in Saudi Arabia. Zootaxa, 5278 (1), 78 - 102. https: // doi. org / 10.11646 / zootaxa. 5278.1.3" type="journal article" year="2023">Anker &amp; Benzoni 2023</bibRefCitation>
). At least nine of them have different colour patterns, including all species associated with scleractinian corals, up to recently assigned to the genus 
<taxonomicName id="4C574D5B6202FFB5A411558D21EBFC60" authority="Holthuis, 1952" authorityName="Holthuis" authorityYear="1952" box="[556,766,942,969]" class="Malacostraca" family="Palaemonidae" genus="Vir" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B923EACA6202FFB5A411558D215BFC61" box="[556,590,943,968]" italics="true" pageId="7" pageNumber="434">Vir</emphasis>
<bibRefCitation id="EFC64B296202FFB5A468558D21EBFC60" author="Holthuis, L. B." box="[597,766,942,969]" pageId="7" pageNumber="434" pagination="1 - 254" refId="ref5741" refString="Holthuis, L. B. (1952) The Decapoda of the Siboga Expedition. Part XI. The Palaemonidae collected by the Siboga and Snellius Expeditions with remarks on other species. II. Subfamily Pontoniinae. Siboga Expedition Monograph, 39 a 10, 1 - 254." type="journal article" year="1952">Holthuis, 1952</bibRefCitation>
</taxonomicName>
, which is now considered to be a synonym of 
<taxonomicName id="4C574D5B6202FFB5A33C558D2688FC60" authorityName="Dana" authorityYear="1852" box="[1281,1437,943,969]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B923EACA6202FFB5A33C558D2688FC60" box="[1281,1437,943,969]" italics="true" pageId="7" pageNumber="434">Palaemonella</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC64B296202FFB5A6A255F12256FC44" author="Holthuis, L. B." box="[159,323,979,1005]" pageId="7" pageNumber="434" pagination="1 - 48" refId="ref5792" refString="Holthuis, L. B. (1973) Caridean shrimps found in land-locked saltwater pools at four Indo-West Pacific localities (Sinai Peninsula, Funafuti Atoll, Maui and Hawaii Islands), with the description of one new genus and four new species. Zoologische Verhandelingen, Leiden, 128, 1 - 48, pls. 1 - 7." type="journal article" year="1973">Holthuis 1973</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A77255F122C0FC44" author="Marin, I." box="[335,469,979,1005]" pageId="7" pageNumber="434" pagination="375 - 390" refId="ref6082" refString="Marin, I. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Vir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, Leiden, 82, 375 - 390." type="journal article" year="2008">Marin 2008</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A7DC55F121E7FC44" author="Kuiter, R. &amp; Debelius, H." box="[481,754,979,1005]" pageId="7" pageNumber="434" refId="ref5970" refString="Kuiter, R. &amp; Debelius, H. (2009) World atlas of marine fauna. IKAN, Frankfurt, 725 pp." type="book" year="2009">Kuiter &amp; Debelius 2009</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A4C055F12700FC44" author="Anker, A. &amp; De Grave, S." box="[765,1045,979,1005]" pageId="7" pageNumber="434" pagination="145 - 150" refId="ref4696" refString="Anker, A. &amp; De Grave, S. (2019) Further records of burrow-associated palaemonid shrimps (Decapoda: Palaemonidae). Zootaxa, 4612 (1), 145 - 150. https: // doi. org / 10.11646 / zootaxa. 4612.1.13" type="journal article" year="2019">Anker &amp; De Grave 2019</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A21D55F127EFFC44" author="Frolova, P. &amp; Horka, I. &amp; Duris, Z." box="[1056,1274,979,1005]" pageId="7" pageNumber="434" pagination="1 - 11" refId="ref5618" refString="Frolova, P., Horka, I. &amp; Duris, Z. (2022) Molecular phylogeny and historical biogeography of marine palaemonid shrimps (Palaemonidae: Palaemonella - Cuapetes group). Scientific Reports, 12 (15237), 1 - 11. https: // doi. org / 10.1038 / s 41598 - 022 - 19372 - 5" type="journal article" year="2022">
Frolová 
<emphasis id="B923EACA6202FFB5A24255F127A2FC44" box="[1151,1207,979,1005]" italics="true" pageId="7" pageNumber="434">et al.</emphasis>
2022
</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A33955F123C7FBB8" author="Fransen, C. H. J. M. &amp; van der Veer, E. &amp; Frolova, P." pageId="7" pageNumber="434" pagination="557 - 580" refId="ref5542" refString="Fransen, C. H. J. M., van der Veer, E. &amp; Frolova, P. (2022) A new species of scleractinian associated shrimp of the genus Palaemonella (Crustacea, Decapoda, Palaemonidae) with a redescription of Palaemonella orientalis Dana, 1852. Zootaxa, 5214 (4), 557 - 580. https: // doi. org / 10.11646 / zootaxa. 5214.4.5" type="journal article" year="2022">
Fransen 
<emphasis id="B923EACA6202FFB5A35955F12688FC44" box="[1380,1437,979,1005]" italics="true" pageId="7" pageNumber="434">et al.</emphasis>
2022
</bibRefCitation>
, 
<bibRefCitation id="EFC64B296202FFB5A6E055D5220DFBB8" author="Fransen, C. H. J. M. &amp; van der Veer, E. &amp; Duris, Z." box="[221,280,1015,1041]" pageId="7" pageNumber="434" pagination="345 - 381" refId="ref5471" refString="Fransen, C. H. J. M., van der Veer, E. &amp; Duris, Z. (2023) A new species of Palaemonella (Decapoda, Caridea, Palaemonidae) associated with scleractinian corals of the genus Euphyllia Dana. Crustaceana, 96 (4), 345 - 381. https: // doi. org / 10.1163 / 15685403 - bja 10289" type="journal article" year="2023">2023</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A71F55D52139FBB8" author="Anker, A. &amp; Benzoni, F." box="[290,556,1015,1041]" pageId="7" pageNumber="434" pagination="78 - 102" refId="ref4589" refString="Anker, A. &amp; Benzoni, F. (2023) Hidden in plain sight: two new species of decapod crustaceans (Palaemonidae and Porcellanidae) discovered in the Red Sea on a university campus in Saudi Arabia. Zootaxa, 5278 (1), 78 - 102. https: // doi. org / 10.11646 / zootaxa. 5278.1.3" type="journal article" year="2023">Anker &amp; Benzoni 2023</bibRefCitation>
), whereas some are deep-water forms (
<bibRefCitation id="EFC64B296202FFB5A5E255D52771FBB8" author="Bruce, A. J." box="[991,1124,1015,1041]" pageId="7" pageNumber="434" refId="ref5028" refString="Bruce, A. J. (1991) Shallow water palaemonoid shrimps from New Caledonia (Crustacea: Decapoda). In: Richer de Forges, B. (Ed.), Le Benthos des fonds meubles des lagons de Nouvelle-Caledonie. Vol. 1. Etudes et Theses. ORSTOM, Paris, pp. 221 - 279." type="book" year="1991">Bruce 1991</bibRefCitation>
, 
<bibRefCitation id="EFC64B296202FFB5A24D55D527BEFBB8" author="Bruce, A. J." box="[1136,1195,1015,1041]" pageId="7" pageNumber="434" pagination="63 - 68" refId="ref5309" refString="Bruce, A. J. (2008) A new species of Palaemonella Dana, 1852, (Crustacea: Decapoda; Pontoniinae) from the Red Sea. Zootaxa, 1844 (1), 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1844.1.6" type="journal article" year="2008">2008</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A28A55D52662FBB8" author="Li, X. &amp; Bruce, A. J." box="[1207,1399,1015,1041]" pageId="7" pageNumber="434" pagination="611 - 738" refId="ref6022" refString="Li, X. &amp; Bruce, A. J. (2006) Further Indo-West Pacific palaemonoid shrimps (Crustacea: Decapoda: Palaemonoidea), principally from the New Caledonian region. Journal of Natural History, 40 (11 - 12), 611 - 738. https: // doi. org / 10.1080 / 00222930600763627" type="journal article" year="2006">Li &amp; Bruce 2006</bibRefCitation>
) or inhabitants of land-locked lava pools (
<bibRefCitation id="EFC64B296202FFB5A47E523921FCFB9C" author="Holthuis, L. B." box="[579,745,1051,1077]" pageId="7" pageNumber="434" pagination="1 - 48" refId="ref5792" refString="Holthuis, L. B. (1973) Caridean shrimps found in land-locked saltwater pools at four Indo-West Pacific localities (Sinai Peninsula, Funafuti Atoll, Maui and Hawaii Islands), with the description of one new genus and four new species. Zoologische Verhandelingen, Leiden, 128, 1 - 48, pls. 1 - 7." type="journal article" year="1973">Holthuis 1973</bibRefCitation>
).
</paragraph>
<paragraph id="8BE836D86202FFB5A6FA521D27A4FA34" blockId="7.[151,1437,151,1581]" pageId="7" pageNumber="434">
The identity of 
<taxonomicName id="4C574D5B6202FFB5A740521D2157FBF0" box="[381,578,1087,1113]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B923EACA6202FFB5A740521D210CFBF0" box="[381,537,1087,1113]" italics="true" pageId="7" pageNumber="434">Palaemonella</emphasis>
sp.
</taxonomicName>
<emphasis id="B923EACA6202FFB5A471521D219FFBF1" box="[588,650,1087,1112]" italics="true" pageId="7" pageNumber="434">sensu</emphasis>
<bibRefCitation id="EFC64B296202FFB5A4AE521D203EFBF0" author="Bruce, A. J." box="[659,811,1087,1113]" pageId="7" pageNumber="434" pagination="209 - 257" refId="ref5222" refString="Bruce, A. J. (2003) The pontoniine shrimp fauna of Hong Kong and the South China Sea (Crustacea: Decapoda: Palaemonidae). In: Morton, B. (Ed.), Perspectives on marine environment change in Hong Kong and southern China, 1977 - 2001. Proceedings of an International Workshop Reunion Conference, Hong Kong 21 - 26 October 2001. Hong Hong Kong University Press, Hong Kong, pp. 209 - 257." type="book chapter" year="2003">Bruce (2003)</bibRefCitation>
from 
<collectingCountry id="F34076486202FFB5A548521D20EBFBF0" box="[885,1022,1087,1113]" name="Hong Kong" pageId="7" pageNumber="434">Hong Kong</collectingCountry>
, originally reported as 
<taxonomicName id="4C574D5B6202FFB5A337521D2293FBD4" authority="Kemp, 1922" authorityName="Kemp" authorityYear="1922" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="digitalis">
<emphasis id="B923EACA6202FFB5A337521D23E6FBD4" italics="true" pageId="7" pageNumber="434">Periclimenes digitalis</emphasis>
<bibRefCitation id="EFC64B296202FFB5A6C652412293FBD4" author="Kemp, S." box="[251,390,1123,1149]" pageId="7" pageNumber="434" pagination="113 - 228" refId="ref5855" refString="Kemp, S. (1922) Notes on Crustacea Decapoda in the Indian Museum. XV. Pontoniinae. Records of the Indian Museum, 24, 113 - 228. https: // doi. org / 10.26515 / rzsi / v 24 / i 2 / 1922 / 163464" type="journal article" year="1922">Kemp, 1922</bibRefCitation>
</taxonomicName>
by 
<bibRefCitation id="EFC64B296202FFB5A78F52412154FBD4" author="Bruce, A. J." box="[434,577,1123,1149]" pageId="7" pageNumber="434" pagination="215 - 248" refId="ref4929" refString="Bruce, A. J. (1979) Records of some pontoniinid shrimps from the South China Sea. Cahiers de l'Indo-Pacifique, 1 (2), 215 - 248." type="journal article" year="1979">Bruce (1979</bibRefCitation>
, 
<bibRefCitation id="EFC64B296202FFB5A4705241219CFBD4" author="Bruce, A. J." box="[589,649,1123,1149]" pageId="7" pageNumber="434" pagination="233 - 284" refId="ref4962" refString="Bruce, A. J. (1982) The pontoniine shrimp fauna or Hong Kong. In: Morton, B. &amp; Tseng, C. K. (Eds.), Proceedings of the First International Marine Biological Workshop: The marine Fauna of Hong Kong and southern China, Hong Kong 1980, I, pp. 233 - 284." type="journal article" year="1982">1982</bibRefCitation>
), remains unknown. The illustrations of a single male specimen (pocl 
<quantity id="4CAF9B3D6202FFB5A6AA52A523FBFB08" box="[151,238,1159,1185]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="7" pageNumber="434" unit="mm" value="3.2">3.2 mm</quantity>
) provided by 
<bibRefCitation id="EFC64B296202FFB5A7B752A5210CFB08" author="Bruce, A. J." box="[394,537,1159,1185]" pageId="7" pageNumber="434" pagination="233 - 284" refId="ref4962" refString="Bruce, A. J. (1982) The pontoniine shrimp fauna or Hong Kong. In: Morton, B. &amp; Tseng, C. K. (Eds.), Proceedings of the First International Marine Biological Workshop: The marine Fauna of Hong Kong and southern China, Hong Kong 1980, I, pp. 233 - 284." type="journal article" year="1982">Bruce (1982</bibRefCitation>
: figs. 4, 5) suggest that this form may be very close, if not identical with 
<taxonomicName id="4C574D5B6202FFB5A37452A52689FB08" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[1353,1436,1159,1185]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6202FFB5A37452A52689FB08" box="[1353,1436,1159,1185]" italics="true" pageId="7" pageNumber="434">P. yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6202FFB5A6AA528923E7FB6C" bold="true" box="[151,242,1195,1221]" pageId="7" pageNumber="434">
<taxonomicNameLabel id="A21057B16202FFB5A6AA528923E7FB6C" box="[151,242,1195,1221]" pageId="7" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, despite their wide geographic separation. However, two morphological differences between the 
<typeStatus id="54EC887A6202FFB5A30752892689FB6C" box="[1338,1436,1195,1221]" pageId="7" pageNumber="434" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C574D5B6202FFB5A68B52ED221CFB40" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[182,265,1231,1257]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6202FFB5A68B52ED221CFB40" box="[182,265,1231,1257]" italics="true" pageId="7" pageNumber="434">P. yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6202FFB5A72D52ED227FFB40" bold="true" box="[272,362,1231,1257]" pageId="7" pageNumber="434">
<taxonomicNameLabel id="A21057B16202FFB5A72D52ED227FFB40" box="[272,362,1231,1257]" pageId="7" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C574D5B6202FFB5A79D52ED2176FB40" box="[416,611,1231,1257]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B923EACA6202FFB5A79D52ED2129FB40" box="[416,572,1231,1257]" italics="true" pageId="7" pageNumber="434">Palaemonella</emphasis>
sp.
</taxonomicName>
<emphasis id="B923EACA6202FFB5A45452ED21B2FB41" box="[617,679,1231,1256]" italics="true" pageId="7" pageNumber="434">sensu</emphasis>
<bibRefCitation id="EFC64B296202FFB5A49252ED2056FB40" author="Bruce, A. J." box="[687,835,1231,1257]" pageId="7" pageNumber="434" pagination="209 - 257" refId="ref5222" refString="Bruce, A. J. (2003) The pontoniine shrimp fauna of Hong Kong and the South China Sea (Crustacea: Decapoda: Palaemonidae). In: Morton, B. (Ed.), Perspectives on marine environment change in Hong Kong and southern China, 1977 - 2001. Proceedings of an International Workshop Reunion Conference, Hong Kong 21 - 26 October 2001. Hong Hong Kong University Press, Hong Kong, pp. 209 - 257." type="book chapter" year="2003">Bruce (2003)</bibRefCitation>
prevent the inclusion of the 
<collectingCountry id="F34076486202FFB5A2B952ED2612FB40" box="[1156,1287,1231,1257]" name="Hong Kong" pageId="7" pageNumber="434">Hong Kong</collectingCountry>
record under the synonymy of the new species. These are the presence of only one tooth on the ventral margin of the rostrum in the male from 
<collectingCountry id="F34076486202FFB5A707533522A8FA98" box="[314,445,1303,1329]" name="Hong Kong" pageId="7" pageNumber="434">Hong Kong</collectingCountry>
<emphasis id="B923EACA6202FFB5A7FE533522F7FA99" box="[451,482,1303,1328]" italics="true" pageId="7" pageNumber="434">vs.</emphasis>
two in 
<taxonomicName id="4C574D5B6202FFB5A4085335219DFA98" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[565,648,1303,1329]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6202FFB5A4085335219DFA98" box="[565,648,1303,1329]" italics="true" pageId="7" pageNumber="434">P. yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6202FFB5A4B2533521FDFA98" bold="true" box="[655,744,1303,1329]" pageId="7" pageNumber="434">
<taxonomicNameLabel id="A21057B16202FFB5A4B2533521FDFA98" box="[655,744,1303,1329]" pageId="7" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(cf. 
<figureCitation id="136C2A5D6202FFB5A5265335209BFA98" box="[795,910,1303,1329]" captionStart="FIGURE 1" captionStartId="3.[151,250,150,174]" captionTargetId="figure-21@2.[151,1436,179,1873]" captionTargetPageId="2" captionText="FIGURE 1. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10185395" httpUri="https://zenodo.org/record/10185395/files/figure.png" pageId="7" pageNumber="434">Fig. 1B, P</figureCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A5A75335270AFA98" author="Bruce, A. J." box="[922,1055,1303,1329]" pageId="7" pageNumber="434" pagination="233 - 284" refId="ref4962" refString="Bruce, A. J. (1982) The pontoniine shrimp fauna or Hong Kong. In: Morton, B. &amp; Tseng, C. K. (Eds.), Proceedings of the First International Marine Biological Workshop: The marine Fauna of Hong Kong and southern China, Hong Kong 1980, I, pp. 233 - 284." type="journal article" year="1982">Bruce 1982</bibRefCitation>
: fig. 4A), and the presence of four spiniform setae on the antepenultimate article of the third maxilliped 
<emphasis id="B923EACA6202FFB5A59E531920D7FAFD" box="[931,962,1339,1364]" italics="true" pageId="7" pageNumber="434">vs.</emphasis>
seven in 
<taxonomicName id="4C574D5B6202FFB5A21053192794FAFC" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[1069,1153,1339,1365]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6202FFB5A21053192794FAFC" box="[1069,1153,1339,1365]" italics="true" pageId="7" pageNumber="434">P. yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6202FFB5A2B5531927F7FAFC" bold="true" box="[1160,1250,1339,1365]" pageId="7" pageNumber="434">
<taxonomicNameLabel id="A21057B16202FFB5A2B5531927F7FAFC" box="[1160,1250,1339,1365]" pageId="7" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(cf. 
<figureCitation id="136C2A5D6202FFB5A32A53192682FAFC" box="[1303,1431,1339,1365]" captionStart="FIGURE 1" captionStartId="3.[151,250,150,174]" captionTargetId="figure-21@2.[151,1436,179,1873]" captionTargetPageId="2" captionText="FIGURE 1. Palaemonella yalla sp. nov., holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446 [A–O]: A, anterior region of carapace and cephalic appendages, dorsal; B, same, lateral; C, carapace, lateral; D, pleon, posterior pleonites, lateral; E, telson, dorsal; F, same, posterior margin, dorsal; G, right mandible, mesial (molar process broken); H, right maxillule, lateral; I, right maxilla, lateral; J, right first maxilliped, lateral; K, right second maxilliped, lateral; L, same, detail of epipod and podobranch, mesial; M, right third maxilliped, ventrolateral; N, same, detail of antepenultimate article, lateral; O, right uropod, dorsal; non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445 [P]: rostrum, lateral. Scales are in mm." figureDoi="http://doi.org/10.5281/zenodo.10185395" httpUri="https://zenodo.org/record/10185395/files/figure.png" pageId="7" pageNumber="434">Fig. 1M, N</figureCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A6AA537D2208FAD0" author="Bruce, A. J." box="[151,285,1375,1401]" pageId="7" pageNumber="434" pagination="233 - 284" refId="ref4962" refString="Bruce, A. J. (1982) The pontoniine shrimp fauna or Hong Kong. In: Morton, B. &amp; Tseng, C. K. (Eds.), Proceedings of the First International Marine Biological Workshop: The marine Fauna of Hong Kong and southern China, Hong Kong 1980, I, pp. 233 - 284." type="journal article" year="1982">Bruce 1982</bibRefCitation>
: fig. 5E). For the time being, it seems most reasonable to leave 
<taxonomicName id="4C574D5B6202FFB5A5E0537D278AFAD0" box="[989,1183,1375,1401]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B923EACA6202FFB5A5E0537D276CFAD0" box="[989,1145,1375,1401]" italics="true" pageId="7" pageNumber="434">Palaemonella</emphasis>
sp.
</taxonomicName>
<emphasis id="B923EACA6202FFB5A29B537D27F1FAD1" box="[1190,1252,1375,1400]" italics="true" pageId="7" pageNumber="434">sensu</emphasis>
<bibRefCitation id="EFC64B296202FFB5A2D7537D266AFAD0" author="Bruce, A. J." box="[1258,1407,1375,1401]" pageId="7" pageNumber="434" pagination="209 - 257" refId="ref5222" refString="Bruce, A. J. (2003) The pontoniine shrimp fauna of Hong Kong and the South China Sea (Crustacea: Decapoda: Palaemonidae). In: Morton, B. (Ed.), Perspectives on marine environment change in Hong Kong and southern China, 1977 - 2001. Proceedings of an International Workshop Reunion Conference, Hong Kong 21 - 26 October 2001. Hong Hong Kong University Press, Hong Kong, pp. 209 - 257." type="book chapter" year="2003">Bruce (2003)</bibRefCitation>
as a presumably undescribed species with affinities to 
<taxonomicName id="4C574D5B6202FFB5A4E753A12038FA34" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[730,813,1411,1437]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6202FFB5A4E753A12038FA34" box="[730,813,1411,1437]" italics="true" pageId="7" pageNumber="434">P. yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6202FFB5A50953A1209BFA34" bold="true" box="[820,910,1411,1437]" pageId="7" pageNumber="434">
<taxonomicNameLabel id="A21057B16202FFB5A50953A1209BFA34" box="[820,910,1411,1437]" pageId="7" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="4C574D5B6202FFB5A5A153A12716FA34" authorityName="Komai &amp; Yamada" authorityYear="2015" box="[924,1027,1411,1437]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="okunoi">
<emphasis id="B923EACA6202FFB5A5A153A12716FA34" box="[924,1027,1411,1437]" italics="true" pageId="7" pageNumber="434">P. okunoi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C574D5B6202FFB5A20753A127BFFA34" authorityName="Okuno" authorityYear="1999" box="[1082,1194,1411,1437]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="hachijo">
<emphasis id="B923EACA6202FFB5A20753A127BFFA34" box="[1082,1194,1411,1437]" italics="true" pageId="7" pageNumber="434">P. hachijo</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE836D86202FFB5A6FA53852109F984" blockId="7.[151,1437,151,1581]" pageId="7" pageNumber="434">
<taxonomicName id="4C574D5B6202FFB5A6FA538522B5FA68" authority="Anker &amp; Assayie, 2023" authorityName="Anker &amp; Assayie" authorityYear="2023" box="[199,416,1447,1473]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="yalla" status="sp. nov.">
<emphasis id="B923EACA6202FFB5A6FA538522B5FA68" box="[199,416,1447,1473]" italics="true" pageId="7" pageNumber="434">Palaemonella yalla</emphasis>
</taxonomicName>
<emphasis id="B923EACA6202FFB5A79B538522EAFA68" bold="true" box="[422,511,1447,1473]" pageId="7" pageNumber="434">
<taxonomicNameLabel id="A21057B16202FFB5A79B538522EAFA68" box="[422,511,1447,1473]" pageId="7" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the fifth species within the family 
<taxonomicName id="4C574D5B6202FFB5A5A853852727FA68" authorityName="Rafinesque" authorityYear="1815" box="[917,1074,1447,1473]" class="Malacostraca" family="Palaemonidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="434" phylum="Arthropoda" rank="family">Palaemonidae</taxonomicName>
associated, at least facultatively, with larger burrowing animals (
<bibRefCitation id="EFC64B296202FFB5A7CB53E92168FA4C" author="Marin, I." box="[502,637,1483,1509]" pageId="7" pageNumber="434" pagination="375 - 390" refId="ref6082" refString="Marin, I. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Vir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, Leiden, 82, 375 - 390." type="journal article" year="2008">Marin 2008</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A4B553E92082FA4C" author="Kuiter, R. &amp; Debelius, H." box="[648,919,1483,1509]" pageId="7" pageNumber="434" refId="ref5970" refString="Kuiter, R. &amp; Debelius, H. (2009) World atlas of marine fauna. IKAN, Frankfurt, 725 pp." type="book" year="2009">Kuiter &amp; Debelius 2009</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A59F53E9273AFA4C" author="Okuno, J." box="[930,1071,1483,1509]" pageId="7" pageNumber="434" pagination="521 - 528" refId="ref6171" refString="Okuno, J. (2017) Palaemonella shirakawai, a new species of shrimp (Crustacea: Decapoda: Palaemonidae), associated with burrow dwelling opistognathid fish from the Ryukyu Islands, Japan. Zootaxa, 4299 (4), 521 - 528. https: // doi. org / 10.11646 / zootaxa. 4299.4.3" type="journal article" year="2017">Okuno 2017</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A20553E92658FA4C" author="Anker, A. &amp; De Grave, S." box="[1080,1357,1483,1509]" pageId="7" pageNumber="434" pagination="145 - 150" refId="ref4696" refString="Anker, A. &amp; De Grave, S. (2019) Further records of burrow-associated palaemonid shrimps (Decapoda: Palaemonidae). Zootaxa, 4612 (1), 145 - 150. https: // doi. org / 10.11646 / zootaxa. 4612.1.13" type="journal article" year="2019">Anker &amp; De Grave 2019</bibRefCitation>
; 
<bibRefCitation id="EFC64B296202FFB5A36B53E92242F9A0" author="Anker, A. &amp; Benzoni, F." pageId="7" pageNumber="434" pagination="78 - 102" refId="ref4589" refString="Anker, A. &amp; Benzoni, F. (2023) Hidden in plain sight: two new species of decapod crustaceans (Palaemonidae and Porcellanidae) discovered in the Red Sea on a university campus in Saudi Arabia. Zootaxa, 5278 (1), 78 - 102. https: // doi. org / 10.11646 / zootaxa. 5278.1.3" type="journal article" year="2023">Anker &amp; Benzoni 2023</bibRefCitation>
). However, the elusive burrowing host or hosts of the new species remain to be determined, as is the consistency of this association.
</paragraph>
</subSubSection>
</treatment>
</document>