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<document id="7FED6F5713221C22E63965296C964310" ID-CLB-Dataset="31353" ID-DOI="10.11646/zootaxa.4353.2.1" ID-GBIF-Dataset="d93b2bd2-75e1-467c-89d3-e2faee3b10a4" ID-ISSN="1175-5326" ID-Zenodo-Dep="1065124" ID-ZooBank="D2902BF9-0213-40F3-91A4-EC4D4FDB3B27" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511423577451" checkinUser="plazi" docAuthor="Serrano, Artur R. M., Capela, Rúben A. &amp; Santos, Carmen Van-Dúnem Neto" docDate="2017" docId="03EFF252F3227F4514BFA683FEF1F92C" docLanguage="en" docName="zootaxa.4353.2.1.pdf" docOrigin="Zootaxa 4353 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tetragonoderus quadrimaculatus Gory 1833" docType="treatment" docVersion="9" lastPageNumber="239" masterDocId="FFD68A2AF3047F631428A273FFF3FFBF" masterDocTitle="Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae)" masterLastPageNumber="256" masterPageNumber="201" pageNumber="239" updateTime="1698503882738" updateUser="ExternalLinkService">
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<mods:title id="033DC7194A44AA8EF11E69AE5EF261FC">Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="C5DFB60807D1F7DC737A583073763581">Serrano, Artur R. M.</mods:namePart>
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<mods:namePart id="0E52C443790D10BF6D0B9F8918AE15D2">Capela, Rúben A.</mods:namePart>
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<mods:namePart id="28357364A1F34530276731C31B1E0325">Santos, Carmen Van-Dúnem Neto</mods:namePart>
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<treatment id="03EFF252F3227F4514BFA683FEF1F92C" ID-DOI="http://doi.org/10.5281/zenodo.6489800" ID-GBIF-Taxon="137541427" ID-Zenodo-Dep="6489800" LSID="urn:lsid:plazi:treatment:03EFF252F3227F4514BFA683FEF1F92C" httpUri="http://treatment.plazi.org/id/03EFF252F3227F4514BFA683FEF1F92C" lastPageNumber="239" pageId="38" pageNumber="239">
<subSubSection id="C35C10CFF3227F4514BFA683FD4AFAB5" box="[151,697,1264,1290]" pageId="38" pageNumber="239" type="nomenclature">
<paragraph id="8BF94344F3227F4514BFA683FD4AFAB5" blockId="38.[151,697,1264,1290]" box="[151,697,1264,1290]" pageId="38" pageNumber="239">
<heading id="D0B1F428F3227F4514BFA683FD4AFAB5" bold="true" box="[151,697,1264,1290]" fontSize="11" level="1" pageId="38" pageNumber="239" reason="1">
<taxonomicName id="4C4638C7F3227F4514BFA683FD4AFAB5" ID-CoL="55N26" authority="Gory, 1833" authorityName="Gory" authorityYear="1833" box="[151,697,1264,1290]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="239" phylum="Arthropoda" rank="species" species="quadrimaculatus">
<emphasis id="B9329F56F3227F4514BFA683FD4AFAB5" bold="true" box="[151,697,1264,1290]" pageId="38" pageNumber="239">
<emphasis id="B9329F56F3227F4514BFA683FDDAFAB5" bold="true" box="[151,553,1264,1290]" italics="true" pageId="38" pageNumber="239">Tetragonoderus quadrimaculatus</emphasis>
Gory, 1833
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C10CFF3227F4514BFA745FCAEFAF0" box="[151,861,1334,1359]" pageId="38" pageNumber="239" type="distribution">
<paragraph id="8BF94344F3227F4514BFA745FCAEFAF0" blockId="38.[151,1437,1334,1683]" box="[151,861,1334,1359]" pageId="38" pageNumber="239">
<emphasis id="B9329F56F3227F4514BFA745FDC8FAF0" bold="true" box="[151,571,1334,1359]" pageId="38" pageNumber="239">
Distribution in
<collectingCountry id="F35103D4F3227F451566A745FE50FAF0" box="[334,419,1334,1359]" name="Angola" pageId="38" pageNumber="239">Angola</collectingCountry>
(Provinces):
</emphasis>
1)
<collectingRegion id="49828DA6F3227F451648A745FD1CFAF1" box="[608,751,1334,1359]" country="Angola" name="Lunda Norte" pageId="38" pageNumber="239">
Lunda
<collectingRegion id="49828DA6F3227F451687A744FD1CFAF1" box="[687,751,1335,1358]" country="Sudan" name="Ash Shimaliyya" pageId="38" pageNumber="239">Norte</collectingRegion>
</collectingRegion>
?; 2)
<collectingRegion id="49828DA6F3227F451707A745FCAAFAF0" box="[815,857,1334,1359]" country="Angola" name="Bie" pageId="38" pageNumber="239">Bié</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C10CFF3227F4514EFA729FCF6FA28" pageId="38" pageNumber="239" type="materials_examined">
<paragraph id="8BF94344F3227F4514EFA729FCF6FA28" blockId="38.[151,1437,1334,1683]" pageId="38" pageNumber="239">
<materialsCitation id="3B2E4919F3227F4514EFA729FCF2FA28" ID-GBIF-Occurrence="1701951654" collectingDate="2014-11-01" collectionCode="ASC" collectorName="Serrano &amp; R. Capela" elevation="1666" latitude="-13.377995" location="BIE" longLatPrecision="1" longitude="16.699463" pageId="38" pageNumber="239" specimenCount="1" specimenCount-male="1" stateProvince="BIE">
<emphasis id="B9329F56F3227F4514EFA729FE40FACC" bold="true" box="[199,435,1370,1395]" pageId="38" pageNumber="239">Material examined.</emphasis>
Chitembo (
<quantity id="4CBEEEA1F3227F451617A728FD79FACC" box="[575,650,1370,1395]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="38" pageNumber="239" unit="km" value="10.0">10 km</quantity>
N) (
<geoCoordinate id="EE722583F3227F4516ECA728FC61FACC" box="[708,914,1371,1395]" direction="south" orientation="latitude" pageId="38" pageNumber="239" precision="1" value="-13.377995">13° 22´40.78´´ S</geoCoordinate>
,
<geoCoordinate id="EE722583F3227F45178AA729FB81FACD" box="[930,1138,1370,1395]" direction="east" orientation="longitude" pageId="38" pageNumber="239" precision="1" value="16.699463">16° 41´58.07´´ E</geoCoordinate>
,
<quantity id="4CBEEEA1F3227F4510A9A729FB29FACD" box="[1153,1242,1370,1395]" metricMagnitude="3" metricUnit="m" metricValue="1.666" pageId="38" pageNumber="239" unit="m" value="1666.0">
<elevation id="006BA477F3227F4510A9A729FB29FACD" box="[1153,1242,1370,1395]" metricMagnitude="3" metricUnit="m" metricValue="1.666" pageId="38" pageNumber="239" unit="m" value="1666.0">1666 m</elevation>
</quantity>
alt., 282) (
<collectingRegion id="49828DA6F3227F451175A729FA7CFACC" box="[1373,1423,1370,1395]" country="Angola" name="Bie" pageId="38" pageNumber="239">BIÉ</collectingRegion>
),
<date id="FFF86584F3227F4514BFA70CFEF8FA28" box="[151,267,1407,1431]" pageId="38" pageNumber="239" value="2014-11-01">
<collectingDate id="EFBC9C6CF3227F4514BFA70CFEF8FA28" box="[151,267,1407,1431]" pageId="38" pageNumber="239" value="2014-11-01">1.XI.2014</collectingDate>
</date>
,
<specimenCount id="9D4088CDF3227F45153EA70CFEC8FA28" box="[278,315,1407,1431]" pageId="38" pageNumber="239" type="male">1♂</specimenCount>
, DO,
<collectingRegion id="49828DA6F3227F451556A70CFE66FA28" box="[382,405,1407,1431]" country="Gambia" name="Upper River" pageId="38" pageNumber="239">A</collectingRegion>
.
<collectorName id="26B32692F3227F451588A70CFE04FA28" box="[416,503,1407,1431]" pageId="38" pageNumber="239">Serrano</collectorName>
&amp;
<collectorName id="26B32692F3227F451633A70CFD79FA28" box="[539,650,1406,1431]" pageId="38" pageNumber="239">R. Capela</collectorName>
leg.,
<collectionCode id="ED57DB81F3227F4516EFA70CFCF2FA28" box="[711,769,1407,1431]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15857" name="Northern Arizona University" pageId="38" pageNumber="239">ASC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C10CFF3227F4514EFA7D1FEF1F92C" pageId="38" pageNumber="239" type="discussion">
<paragraph id="8BF94344F3227F4514EFA7D1FEF1F92C" blockId="38.[151,1437,1334,1683]" pageId="38" pageNumber="239">
<emphasis id="B9329F56F3227F4514EFA7D1FEC8FA04" bold="true" box="[199,315,1442,1467]" pageId="38" pageNumber="239">Remarks.</emphasis>
<collectingRegion id="49828DA6F3227F451561A7D0FEAEFA04" box="[329,349,1443,1467]" country="Gambia" name="Upper River" pageId="38" pageNumber="239">A</collectingRegion>
species widespread throughout western, central and southern Africa (
<collectingCountry id="F35103D4F3227F451083A7D1FAD8FA04" box="[1195,1323,1442,1467]" name="Mauritania" pageId="38" pageNumber="239">Mauritania</collectingCountry>
,
<collectingCountry id="F35103D4F3227F451114A7D1FA64FA04" box="[1340,1431,1442,1467]" name="Senegal" pageId="38" pageNumber="239">Senegal</collectingCountry>
,
<collectingCountry id="F35103D4F3227F4514BFA7B5FF00FA60" box="[151,243,1478,1503]" name="Gambia" pageId="38" pageNumber="239">Gambia</collectingCountry>
,
<collectingCountry id="F35103D4F3227F4514D6A7B5FE55FA60" box="[254,422,1478,1503]" name="Guinea-Bissau" pageId="38" pageNumber="239">Guinea-Bissau</collectingCountry>
,
<collectingCountry id="F35103D4F3227F45159AA7B5FDF5FA60" box="[434,518,1478,1503]" name="Guinea-Bissau" pageId="38" pageNumber="239">Guinea</collectingCountry>
,
<collectingCountry id="F35103D4F3227F451639A7B5FDBCFA60" box="[529,591,1478,1503]" name="Chad" pageId="38" pageNumber="239">Chad</collectingCountry>
,
<collectingCountry id="F35103D4F3227F451673A7B4FD11FA60" box="[603,738,1479,1503]" name="Ivory Coast" pageId="38" pageNumber="239">
Ivory
<collectingRegion id="49828DA6F3227F4516B7A7B4FD11FA60" box="[671,738,1479,1503]" country="Tanzania" name="Pwani" pageId="38" pageNumber="239">Coast</collectingRegion>
</collectingCountry>
,
<collectingCountry id="F35103D4F3227F4516C6A7B5FCD5FA60" box="[750,806,1478,1503]" name="Serbia" pageId="38" pageNumber="239">Mali</collectingCountry>
,
<collectingCountry id="F35103D4F3227F451719A7B5FC75FA60" box="[817,902,1478,1503]" name="Angola" pageId="38" pageNumber="239">Angola</collectingCountry>
, D. R. of the
<collectingRegion id="49828DA6F3227F451008A7B4FB9EFA60" box="[1056,1133,1479,1503]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="38" pageNumber="239">
<collectingCountry id="F35103D4F3227F451008A7B4FB9EFA60" box="[1056,1133,1479,1503]" name="Democratic Republic of the Congo" pageId="38" pageNumber="239">Congo</collectingCountry>
</collectingRegion>
,
<collectingCountry id="F35103D4F3227F451051A7B5FAF9FA60" box="[1145,1290,1478,1503]" name="South Africa" pageId="38" pageNumber="239">
<collectingRegion id="49828DA6F3227F451051A7B5FB49FA60" box="[1145,1210,1478,1503]" country="Tanzania" name="Kusini Unguja" pageId="38" pageNumber="239">South</collectingRegion>
Africa
</collectingCountry>
R.) (
<bibRefCitation id="EFD73EB5F3227F45116BA7B4FF21F9BC" author="Serrano" pageId="38" pageNumber="239" refString="Serrano, A. R. M. (2009) Biodiversity and notes on Carabid beetles from Guinea-Bissau. VI. Lebiinae (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 26 (1), 27 - 46." type="journal article" year="2009">Serrano 2009</bibRefCitation>
, Insectoid.info 2017). It is with some reserve that we ascribe the singleton male adult specimen to this species. The apex of median lobe of aedeagus is slightly different of
<taxonomicName id="4C4638C7F3227F451785A47CFB7AF998" authorityName="Gory" authorityYear="1833" box="[941,1161,1550,1575]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="239" phylum="Arthropoda" rank="species" species="quadrimaculatus">
<emphasis id="B9329F56F3227F451785A47CFB7AF998" box="[941,1161,1550,1575]" italics="true" pageId="38" pageNumber="239">T. quadrimaculatus</emphasis>
</taxonomicName>
and more similar to the same organ of
<taxonomicName id="4C4638C7F3227F451516A447FD7FF9F4" authority="(Fabricius, 1792)" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[318,652,1586,1611]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="239" phylum="Arthropoda" rank="species" species="quadrum">
<emphasis id="B9329F56F3227F451516A447FE4CF9F4" box="[318,447,1586,1611]" italics="true" pageId="38" pageNumber="239">T. quadrum</emphasis>
(Fabricius, 1792)
</taxonomicName>
. However, this species is distributed on northern territories of Africa. The adult specimen was found in a country road within an open secondary forest. It is a new record for the
<collectingRegion id="49828DA6F3227F45115DA425FF0CF92C" country="Angola" name="Bie" pageId="38" pageNumber="239">Bié Province</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>