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<document ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" approvalRequired="59" approvalRequired_for_taxonomicNames="59" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. &amp; Jansa, Sharon A." docDate="2022" docId="03EFDD5DF77A6968DB2BFE741E18FDE4" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.8:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="8" docTitle="Palaeothentes undetermined" docType="treatment" docVersion="5" lastPageNumber="321" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="321" updateTime="1659987007554" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Beck, Robin M. D.</mods:namePart>
<mods:affiliation>School of Science, Engineering and Environment University of Salford, U. K. &amp; School of Biological, Earth &amp; Environmental Sciences University of New South Wales, Australia &amp; Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Voss, Robert S.</mods:namePart>
<mods:affiliation>Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jansa, Sharon A.</mods:namePart>
<mods:affiliation>Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Bulletin of the American Museum of Natural History</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-06-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>2022</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>457</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>353</mods:end>
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<mods:location>
<mods:url>https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1206/0003-0090.457.1.1</mods:identifier>
<mods:identifier type="ISSN">0003-0090</mods:identifier>
<mods:identifier type="Zenodo-Dep">6971356</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6974448" ID-Zenodo-Dep="6974448" LSID="urn:lsid:plazi:treatment:03EFDD5DF77A6968DB2BFE741E18FDE4" httpUri="http://treatment.plazi.org/id/03EFDD5DF77A6968DB2BFE741E18FDE4" lastPageNumber="321" pageId="321" pageNumber="321">
<subSubSection box="[288,456,473,498]" pageId="321" pageNumber="321" type="nomenclature">
<paragraph blockId="321.[288,456,473,498]" box="[288,456,473,498]" pageId="321" pageNumber="321">
<heading box="[288,456,473,498]" centered="true" fontSize="10" level="2" pageId="321" pageNumber="321" reason="4">
<taxonomicName box="[302,456,473,497]" class="Mammalia" family="Palaeothentidae" genus="Palaeothentes" kingdom="Animalia" order="Paucituberculata" pageId="321" pageNumber="321" phylum="Chordata" rank="species">
<emphasis bold="true" box="[302,456,473,497]" italics="true" pageId="321" pageNumber="321">Palaeothentes</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="321" pageNumber="321" type="synonymic_list">
<paragraph blockId="321.[108,640,523,1738]" pageId="321" pageNumber="321">
SPECIES SCORED: †
<taxonomicName baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[349,590,523,547]" class="Mammalia" family="Palaeothentidae" genus="Palaeothentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="321" pageNumber="321" phylum="Chordata" rank="species">
<emphasis box="[349,590,523,547]" italics="true" pageId="321" pageNumber="321">Palaeothentes minutus</emphasis>
</taxonomicName>
, †
<emphasis italics="true" pageId="321" pageNumber="321">P. lemoinei</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection pageId="321" pageNumber="321" type="materials_examined">
<paragraph blockId="321.[108,640,523,1738]" pageId="321" pageNumber="321">
GEOLOGICAL PROVENANCE OF SCORED
<collectionCode box="[108,167,623,647]" pageId="321" pageNumber="321">SPECIMENS</collectionCode>
: Santa Cruz Formation,
<collectingRegion country="Argentina" name="Santa Cruz" pageId="321" pageNumber="321">Santa Cruz Province</collectingRegion>
,
<collectingCountry box="[167,280,655,679]" name="Argentina" pageId="321" pageNumber="321">Argentina</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="321" pageNumber="321" type="description">
<paragraph blockId="321.[108,640,523,1738]" pageId="321" pageNumber="321">
AGE OF SCORED SPECIMENS: the
<collectingRegion box="[512,635,689,713]" country="Bolivia" name="Santa Cruz" pageId="321" pageNumber="321">Santa Cruz</collectingRegion>
Formation spans a maximum of 14 to 19 Mya (see †
<taxonomicName authorityName="Ameghino" authorityYear="1887" box="[172,304,755,779]" class="Mammalia" family="Caenolestidae" genus="Stilotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="321" pageNumber="321" phylum="Chordata" rank="genus">
<emphasis box="[172,304,755,779]" italics="true" pageId="321" pageNumber="321">Stilotherium</emphasis>
</taxonomicName>
above).
</paragraph>
<paragraph blockId="321.[108,640,523,1738]" box="[140,621,788,812]" pageId="321" pageNumber="321">
ASSIGNED AGE
<collectionCode box="[311,388,788,812]" pageId="321" pageNumber="321">RANGE</collectionCode>
:
<geoCoordinate box="[402,474,788,812]" degrees="19.000" direction="north" orientation="latitude" pageId="321" pageNumber="321" precision="55" value="19.0">19.000</geoCoordinate>
<geoCoordinate box="[473,559,788,812]" degrees="14.000" direction="west" orientation="longitude" pageId="321" pageNumber="321" precision="55" value="-14.0">14.000</geoCoordinate>
Mya.
</paragraph>
</subSubSection>
<subSubSection pageId="321" pageNumber="321" type="discussion">
<paragraph blockId="321.[108,640,523,1738]" pageId="321" pageNumber="321">
REMARKS: We scored †
<taxonomicName baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[395,636,821,845]" class="Mammalia" family="Palaeothentidae" genus="Palaeothentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="321" pageNumber="321" phylum="Chordata" rank="species">
<emphasis box="[395,636,821,845]" italics="true" pageId="321" pageNumber="321">Palaeothentes minutus</emphasis>
</taxonomicName>
based on abundant but mostly fragmentary specimens collected by Carlos Ameghino from several localities in the early Miocene Santa Cruz Formation; this material has been revised by Marshall (1980) and Abello (2007). Bown and Fleagle (1993) subsequently collected considerable additional dental material of †
<taxonomicName baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[517,636,1053,1076]" class="Mammalia" family="Palaeothentidae" genus="Palaeothentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="321" pageNumber="321" phylum="Chordata" rank="species">
<emphasis box="[517,636,1053,1076]" italics="true" pageId="321" pageNumber="321">P. minutus</emphasis>
</taxonomicName>
from localities in the Santa Cruz and Pinturas formations, and Marshall (1990) identified this species from the?middle Miocene (Friasian
<collectionCode box="[108,200,1185,1209]" pageId="321" pageNumber="321">SALMA</collectionCode>
) Río Frias Formation in
<collectingCountry box="[468,529,1184,1208]" name="Chile" pageId="321" pageNumber="321">Chile</collectingCountry>
; however, we did not use these additional specimens for scoring purposes. Croft (2007: table 3) reported †
<taxonomicName baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[121,234,1284,1307]" class="Mammalia" family="Palaeothentidae" genus="Palaeothentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="321" pageNumber="321" phylum="Chordata" rank="species">
<emphasis box="[121,234,1284,1307]" italics="true" pageId="321" pageNumber="321">P. minutus</emphasis>
</taxonomicName>
from the middle Miocene (Laventan
<collectionCode box="[108,201,1317,1341]" pageId="321" pageNumber="321">SALMA</collectionCode>
) Quebrada Honda Fauna in southern
<collectingCountry box="[108,187,1350,1374]" name="Bolivia" pageId="321" pageNumber="321">Bolivia</collectingCountry>
, but Engelman et al. (2016) subsequently referred this material to two new species (†
<emphasis italics="true" pageId="321" pageNumber="321">P. serratus</emphasis>
and †
<emphasis box="[235,338,1416,1440]" italics="true" pageId="321" pageNumber="321">P. relictus</emphasis>
), and we did not use it to score our †
<taxonomicName box="[231,376,1449,1473]" class="Mammalia" family="Palaeothentidae" genus="Palaeothentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="321" pageNumber="321" phylum="Chordata" rank="genus">
<emphasis box="[231,376,1449,1473]" italics="true" pageId="321" pageNumber="321">Palaeothentes</emphasis>
</taxonomicName>
terminal.
</paragraph>
<paragraph blockId="321.[108,640,523,1738]" lastBlockId="321.[684,1212,226,547]" pageId="321" pageNumber="321">
Well-preserved cranial material of †
<taxonomicName baseAuthorityName="Ameghino" baseAuthorityYear="1887" class="Mammalia" family="Palaeothentidae" genus="Palaeothentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="321" pageNumber="321" phylum="Chordata" rank="species">
<emphasis italics="true" pageId="321" pageNumber="321">Palaeothentes minutus</emphasis>
</taxonomicName>
is unavailable, but Forasiepi et al. (2014b) described a well-preserved skull and partial skeleton (MPM-PV 3566) of a congeneric species,
<emphasis box="[201,334,1614,1638]" italics="true" pageId="321" pageNumber="321">†P. lemoinei</emphasis>
, also from the
<collectingRegion box="[511,636,1615,1639]" country="Bolivia" name="Santa Cruz" pageId="321" pageNumber="321">Santa Cruz</collectingRegion>
Formation. Monophyly of †
<taxonomicName box="[479,636,1647,1671]" class="Mammalia" family="Palaeothentidae" genus="Palaeothentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="321" pageNumber="321" phylum="Chordata" rank="genus">
<emphasis box="[479,636,1647,1671]" italics="true" pageId="321" pageNumber="321">Palaeothentes</emphasis>
</taxonomicName>
(excluding Ҡ
<taxonomicName box="[263,412,1680,1704]" class="Mammalia" family="Palaeothentidae" genus="Palaeothentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="321" pageNumber="321" phylum="Chordata" rank="genus">
<emphasis box="[263,412,1680,1704]" italics="true" pageId="321" pageNumber="321">Palaeothentes</emphasis>
</taxonomicName>
<emphasis box="[435,511,1681,1704]" italics="true" pageId="321" pageNumber="321">primus</emphasis>
) has been found in all recent large-scale phylogenetic analyses of
<taxonomicName authorityName="Ameghino" authorityYear="1894" box="[762,944,226,250]" class="Mammalia" kingdom="Animalia" order="Paucituberculata" pageId="321" pageNumber="321" phylum="Chordata" rank="order">Paucituberculata</taxonomicName>
(Abello, 2013; Rincón et al., 2015; Engelman et al., 2016; Abello et al., 2020), so we used Forasiepi et al.s (2014b) description of the skull of
<emphasis box="[984,1114,325,349]" italics="true" pageId="321" pageNumber="321">†P. lemoinei</emphasis>
to score cranial characters for our †
<taxonomicName box="[989,1134,358,382]" class="Mammalia" family="Palaeothentidae" genus="Palaeothentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="321" pageNumber="321" phylum="Chordata" rank="genus">
<emphasis box="[989,1134,358,382]" italics="true" pageId="321" pageNumber="321">Palaeothentes</emphasis>
</taxonomicName>
terminal. †
<taxonomicName box="[743,887,391,415]" class="Mammalia" family="Palaeothentidae" genus="Palaeothentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="321" pageNumber="321" phylum="Chordata" rank="genus">
<emphasis box="[743,887,391,415]" italics="true" pageId="321" pageNumber="321">Palaeothentes</emphasis>
</taxonomicName>
is the
<typeStatus box="[955,1002,391,415]" pageId="321" pageNumber="321">type</typeStatus>
genus of the family †
<taxonomicName baseAuthorityName="Sinclair" baseAuthorityYear="1906" box="[695,878,424,448]" class="Mammalia" family="Palaeothentidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="321" pageNumber="321" phylum="Chordata" rank="family">Palaeothentidae</taxonomicName>
, which is a member of the superfamily †Palaeothentoidea (Abello, 2007, 2013; Goin et al., 2009a; Rincón et al., 2015; Engelman et al., 2016; Abello et al., 2020).
</paragraph>
</subSubSection>
</treatment>
</document>