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<document ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" approvalRequired="59" approvalRequired_for_taxonomicNames="59" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. &amp; Jansa, Sharon A." docDate="2022" docId="03EFDD5DF7766964DB22FAA419DAFB92" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.7:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="7" docTitle="Onirocuscus reidi" docType="treatment" docVersion="7" lastPageNumber="333" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="333" updateTime="1661891901480" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Beck, Robin M. D.</mods:namePart>
<mods:affiliation>School of Science, Engineering and Environment University of Salford, U. K. &amp; School of Biological, Earth &amp; Environmental Sciences University of New South Wales, Australia &amp; Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Voss, Robert S.</mods:namePart>
<mods:affiliation>Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jansa, Sharon A.</mods:namePart>
<mods:affiliation>Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-06-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>2022</mods:number>
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<mods:detail type="issue">
<mods:number>457</mods:number>
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<mods:end>353</mods:end>
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<mods:url>https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1206/0003-0090.457.1.1</mods:identifier>
<mods:identifier type="ISSN">0003-0090</mods:identifier>
<mods:identifier type="Zenodo-Dep">6971356</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6974507" ID-Zenodo-Dep="6974507" LSID="urn:lsid:plazi:treatment:03EFDD5DF7766964DB22FAA419DAFB92" httpUri="http://treatment.plazi.org/id/03EFDD5DF7766964DB22FAA419DAFB92" lastPageNumber="333" pageId="333" pageNumber="333">
<subSubSection box="[297,448,1290,1314]" pageId="333" pageNumber="333" type="multiple">
<paragraph blockId="333.[297,448,1290,1314]" box="[297,448,1290,1314]" pageId="333" pageNumber="333">
<heading box="[297,448,1290,1314]" centered="true" fontSize="10" level="2" pageId="333" pageNumber="333" reason="4">
<emphasis bold="true" box="[297,448,1290,1314]" pageId="333" pageNumber="333">
<taxonomicName box="[310,448,1290,1314]" class="Mammalia" family="Phalangeridae" genus="Onirocuscus" kingdom="Animalia" order="Diprotodontia" pageId="333" pageNumber="333" phylum="Chordata" rank="species" species="undetermined">
<emphasis bold="true" box="[310,448,1290,1314]" italics="true" pageId="333" pageNumber="333">Onirocuscus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="333" pageNumber="333" type="nomenclature">
<paragraph blockId="333.[108,637,1348,1736]" pageId="333" pageNumber="333">
SPECIES SCORED: †
<taxonomicName box="[365,564,1348,1372]" class="Mammalia" family="Phalangeridae" genus="Onirocuscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="333" pageNumber="333" phylum="Chordata" rank="species" species="reidi">
<emphasis box="[365,564,1348,1372]" italics="true" pageId="333" pageNumber="333">Onirocuscus reidi</emphasis>
</taxonomicName>
(
<typeStatus box="[586,636,1348,1372]" pageId="333" pageNumber="333">type</typeStatus>
species).
</paragraph>
</subSubSection>
<subSubSection pageId="333" pageNumber="333" type="materials_examined">
<paragraph blockId="333.[108,637,1348,1736]" pageId="333" pageNumber="333">
GEOLOGICAL PROVENANCE OF SCORED
<collectionCode box="[108,168,1448,1472]" pageId="333" pageNumber="333">SPECIMENS</collectionCode>
: Riversleigh Faunal Zone C, Riversleigh World Heritage Area,
<collectingRegion box="[350,482,1480,1504]" country="Australia" name="Queensland" pageId="333" pageNumber="333">Queensland</collectingRegion>
,
<collectingCountry box="[494,596,1480,1504]" name="Australia" pageId="333" pageNumber="333">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="333" pageNumber="333" type="description">
<paragraph blockId="333.[108,637,1348,1736]" pageId="333" pageNumber="333">AGE OF SCORED SPECIMENS: Riversleigh Faunal Zone C is interpreted to be middle Miocene based on biostratigraphy (see above). In the absence of radiometric dates, we have assumed the entire span of the middle Miocene (Langhian to Serravallian; Cohen et al., 2013 [updated]) for this terminal.</paragraph>
<paragraph blockId="333.[684,1214,225,1109]" box="[716,1197,225,250]" pageId="333" pageNumber="333">
<heading box="[716,1197,225,250]" centered="true" fontSize="10" level="2" pageId="333" pageNumber="333" reason="4">
ASSIGNED AGE
<collectionCode box="[887,964,226,250]" pageId="333" pageNumber="333">RANGE</collectionCode>
:
<geoCoordinate box="[978,1050,225,249]" degrees="15.970" direction="north" orientation="latitude" pageId="333" pageNumber="333" precision="55" value="15.97">15.970</geoCoordinate>
<geoCoordinate box="[1049,1135,225,249]" degrees="11.630" direction="west" orientation="longitude" pageId="333" pageNumber="333" precision="55" value="-11.63">11.630</geoCoordinate>
Mya.
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="333" pageNumber="333" type="discussion">
<paragraph blockId="333.[684,1214,225,1109]" pageId="333" pageNumber="333">
REMARKS: †
<taxonomicName box="[853,1045,259,283]" class="Mammalia" family="Phalangeridae" genus="Onirocuscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="333" pageNumber="333" phylum="Chordata" rank="species" species="reidi">
<emphasis box="[853,1045,259,283]" italics="true" pageId="333" pageNumber="333">Onirocuscus reidi</emphasis>
</taxonomicName>
was originally described as a fossil species of
<taxonomicName box="[1043,1175,292,315]" class="Mammalia" family="Phalangeridae" genus="Strigocuscus" kingdom="Animalia" order="Diprotodontia" pageId="333" pageNumber="333" phylum="Chordata" rank="genus">
<emphasis box="[1043,1175,292,315]" italics="true" pageId="333" pageNumber="333">Strigocuscus</emphasis>
</taxonomicName>
by Flannery and Archer (1987c), who highlighted similarities with the ground cuscus
<taxonomicName authorityName="Peters &amp; Doria" authorityYear="1875" class="Mammalia" family="Phalangeridae" genus="Phalanger" kingdom="Animalia" order="Diprotodontia" pageId="333" pageNumber="333" phylum="Chordata" rank="species" species="gymnotis">
<emphasis italics="true" pageId="333" pageNumber="333">Phalanger gymnotis</emphasis>
</taxonomicName>
(formerly also classified in the genus
<taxonomicName authority="sensu Flannery et al., 1987" authorityName="Flannery" authorityYear="1987" box="[684,1129,424,448]" class="Mammalia" family="Phalangeridae" genus="Strigocuscus" kingdom="Animalia" order="Diprotodontia" pageId="333" pageNumber="333" phylum="Chordata" rank="genus">
<emphasis box="[684,815,424,447]" italics="true" pageId="333" pageNumber="333">Strigocuscus</emphasis>
sensu Flannery et al., 1987
</taxonomicName>
). However, based on evidence from a subsequently discovered and substantially complete cranium, Crosby (2007) referred
<taxonomicName box="[962,1012,523,547]" class="Mammalia" family="Phalangeridae" genus="Onirocuscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="333" pageNumber="333" phylum="Chordata" rank="species" species="reidi">
<emphasis box="[962,1012,523,547]" italics="true" pageId="333" pageNumber="333">reidi</emphasis>
</taxonomicName>
to a new genus, †
<taxonomicName authorityName="Crosby" authorityYear="2007" box="[697,829,557,580]" class="Mammalia" family="Phalangeridae" genus="Onirocuscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="333" pageNumber="333" phylum="Chordata" rank="genus">
<emphasis box="[697,829,557,580]" italics="true" pageId="333" pageNumber="333">Onirocuscus</emphasis>
</taxonomicName>
, which she placed in the phalangerid tribe
<taxonomicName authorityName="Flynn" authorityYear="1911" box="[824,968,589,613]" class="Mammalia" family="Phalangeridae" kingdom="Animalia" order="Diprotodontia" pageId="333" pageNumber="333" phylum="Chordata" rank="tribe" tribe="Trichosurini">Trichosurini</taxonomicName>
. Crosby (2007) also described four other species of †
<taxonomicName authorityName="Crosby" authorityYear="2007" box="[1034,1164,623,646]" class="Mammalia" family="Phalangeridae" genus="Onirocuscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="333" pageNumber="333" phylum="Chordata" rank="genus">
<emphasis box="[1034,1164,623,646]" italics="true" pageId="333" pageNumber="333">Onirocuscus</emphasis>
</taxonomicName>
, but we did not use them for scoring purposes. Crosbys (2007: fig. 7) phylogenetic hypothesis suggests that †
<taxonomicName authorityName="Crosby" authorityYear="2007" box="[825,960,722,745]" class="Mammalia" family="Phalangeridae" genus="Onirocuscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="333" pageNumber="333" phylum="Chordata" rank="genus">
<emphasis box="[825,960,722,745]" italics="true" pageId="333" pageNumber="333">Onirocuscus</emphasis>
</taxonomicName>
is closely related to “
<taxonomicName authorityName="Lesson" authorityYear="1828" box="[695,822,755,779]" class="Mammalia" family="Phalangeridae" genus="Trichosurus" kingdom="Animalia" order="Diprotodontia" pageId="333" pageNumber="333" phylum="Chordata" rank="genus">
<emphasis box="[695,822,755,779]" italics="true" pageId="333" pageNumber="333">Trichosurus</emphasis>
</taxonomicName>
” †
<taxonomicName authorityName="Muirhead &amp; Archer" authorityYear="1990" box="[858,946,755,779]" class="Mammalia" family="Thylacinidae" genus="Nimbacinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="333" pageNumber="333" phylum="Chordata" rank="species" species="dicksoni">
<emphasis box="[858,946,755,779]" italics="true" pageId="333" pageNumber="333">dicksoni</emphasis>
</taxonomicName>
, another fossil phalangerid from Riversleigh (see below). However, her phylogeny was not the result of analyzing a published data matrix and it endorsed some relationships—such as a basal position for
<taxonomicName box="[1120,1212,887,911]" class="Mammalia" family="Phalangeridae" genus="Ailurops" kingdom="Animalia" order="Diprotodontia" pageId="333" pageNumber="333" phylum="Chordata" rank="genus">
<emphasis box="[1120,1212,887,911]" italics="true" pageId="333" pageNumber="333">Ailurops</emphasis>
</taxonomicName>
within
<taxonomicName authorityName="Thomas" authorityYear="1888" box="[762,917,920,944]" class="Mammalia" family="Phalangeridae" kingdom="Animalia" order="Diprotodontia" pageId="333" pageNumber="333" phylum="Chordata" rank="family">Phalangeridae</taxonomicName>
and
<taxonomicName baseAuthorityName="Gray" baseAuthorityYear="1858" box="[974,1214,920,944]" class="Mammalia" family="Phalangeridae" genus="Strigocuscus" kingdom="Animalia" order="Diprotodontia" pageId="333" pageNumber="333" phylum="Chordata" rank="species" species="celebensis">
<emphasis box="[974,1214,920,944]" italics="true" pageId="333" pageNumber="333">Strigocuscus celebensis</emphasis>
</taxonomicName>
within
<taxonomicName authorityName="Flynn" authorityYear="1911" box="[762,898,953,977]" class="Mammalia" family="Phalangeridae" kingdom="Animalia" order="Diprotodontia" pageId="333" pageNumber="333" phylum="Chordata" rank="tribe" tribe="Trichosurini">Trichosurini</taxonomicName>
(see also Crosby and Norris, 2003)—that are contradicted by recent molecular analyses (Ruedas and Morales, 2005; Raterman et al., 2006; Meredith et al., 2009b; Mitchell et al., 2014; Kealy et al., 2019).
</paragraph>
</subSubSection>
</treatment>
</document>