231 lines
31 KiB
XML
231 lines
31 KiB
XML
<document id="A6A9C8B0C8E851216A3D499553EB320D" ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. & Jansa, Sharon A." docDate="2022" docId="03EFDD5DF6E268F3DAD5FAF81860FD40" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Dasyuromorphia Gill 1872" docType="treatment" docVersion="15" lastPageNumber="217" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="217" updateTime="1699350301078" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="733B9B00F4839F7E6493FBF1D3E60FDF">Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
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<mods:namePart id="4589CE1DF3A089C624D78BA928FE916B">Beck, Robin M. D.</mods:namePart>
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<mods:affiliation id="DA6A19158487F779AD51482F4B1D5BE0">School of Science, Engineering and Environment University of Salford, U. K. & School of Biological, Earth & Environmental Sciences University of New South Wales, Australia & Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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<mods:namePart id="8782A25756605E74B87ADA34F6789F58">Voss, Robert S.</mods:namePart>
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<mods:affiliation id="AC4E026C38B62A8A5B016413F7B2D07D">Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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<mods:namePart id="6FD7766A3887E6C4BD1F4A291001517D">Jansa, Sharon A.</mods:namePart>
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<mods:affiliation id="921003D2DE7F52DBFCAFACCFBB5E4D36">Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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<mods:date id="374AFA92EA98B02BE67E6BEDBCCE2CFA">2022</mods:date>
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<subSubSection id="C35C3FC0F6E268F0DAD5FAF81868FAA8" box="[222,522,1366,1391]" pageId="217" pageNumber="217" type="nomenclature">
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<paragraph id="8BF96C4BF6E268F0DAD5FAF81868FAA8" blockId="217.[222,522,1366,1391]" box="[222,522,1366,1391]" pageId="217" pageNumber="217">
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<heading id="D0B1DB27F6E268F0DAD5FAF81868FAA8" box="[222,522,1366,1391]" centered="true" fontSize="9" level="2" pageId="217" pageNumber="217" reason="2">
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<taxonomicName id="4C4617C8F6E268F0DAD5FAF81868FAA8" ID-CoL="33L" authority="Gill, 1872" authorityName="Gill" authorityYear="1872" box="[222,522,1366,1391]" class="Mammalia" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="order">
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Dasyuromorphia
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<bibRefCitation id="EFD711BAF6E268F0DBABFAF81868FAA8" author="Gill, T." box="[416,522,1366,1391]" pageId="217" pageNumber="217" pagination="1 - 98" refId="ref205030" refString="Gill, T. 1872. Arrangement of the families of mammals with analytical tables. Smithsonian Miscellaneous Collections 11 (1): 1 - 98." type="journal article" year="1872">Gill, 1872</bibRefCitation>
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</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C35C3FC0F6E268F0DA87FA26181FFA06" pageId="217" pageNumber="217" type="synonymic_list">
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<paragraph id="8BF96C4BF6E268F0DA87FA26181FFA06" blockId="217.[108,637,1416,1738]" pageId="217" pageNumber="217">
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CONTENTS: †
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<taxonomicName id="4C4617C8F6E268F0DB11FA261BE0FA67" baseAuthorityName="Muirhead and Wroe" baseAuthorityYear="1998" box="[282,386,1416,1440]" class="Mammalia" family="Thylacinidae" genus="Badjcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E268F0DB11FA261BE0FA67" box="[282,386,1416,1440]" italics="true" pageId="217" pageNumber="217">Badjcinus</emphasis>
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</taxonomicName>
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, †
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<taxonomicName id="4C4617C8F6E268F0DB90FA271B8DFA67" baseAuthorityName="Wroe" baseAuthorityYear="1999" box="[411,495,1417,1440]" class="Mammalia" family="Dasyuridae" genus="Barinya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E268F0DB90FA271B8DFA67" box="[411,495,1417,1440]" italics="true" pageId="217" pageNumber="217">Barinya</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C4617C8F6E268F0DBF0FA26181AFA67" authorityName="Goldfuss" authorityYear="1820" box="[507,632,1416,1440]" class="Mammalia" family="Dasyuridae" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="family">Dasyuridae</taxonomicName>
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, †
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<taxonomicName id="4C4617C8F6E268F0DA72FA041B6DFA06" authorityName="Murray & Megirian" authorityYear="2000" box="[121,271,1450,1473]" class="Mammalia" family="Thylacinidae" genus="Mutpuracinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E268F0DA72FA041B6DFA06" box="[121,271,1450,1473]" italics="true" pageId="217" pageNumber="217">Mutpuracinus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C4617C8F6E268F0DB16FA071BCEFA06" baseAuthorityName="Flower" baseAuthorityYear="1867" box="[285,428,1449,1473]" class="Mammalia" family="Myrmecobiidae" genus="Myrmecobius" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E268F0DB16FA071BCEFA06" box="[285,428,1449,1473]" italics="true" pageId="217" pageNumber="217">Myrmecobius</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C4617C8F6E268F0DBE0FA07181BFA06" authorityName="Bonaparte" authorityYear="1838" box="[491,633,1449,1473]" class="Mammalia" family="Thylacinidae" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="family">Thylacinidae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C3FC0F6E268F0DA87FA64194AFE26" pageId="217" pageNumber="217" type="description">
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<paragraph id="8BF96C4BF6E268F0DA87FA641ACEF9C4" blockId="217.[108,637,1416,1738]" pageId="217" pageNumber="217">STEM AGE: 43.7 Mya (95% HPD: 39.6–47.4 Mya).</paragraph>
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<footnote id="E85D7045F6E268F0D8C7FA6B1EC5F90E" pageId="217" pageNumber="217">
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<paragraph id="8BF96C4BF6E268F0D8C7FA6B1EC5F90E" blockId="217.[684,1213,1477,1737]" pageId="217" pageNumber="217">
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||
<superScript id="7C33C103F6E268F0D8C7FA6B18B8FA16" attach="right" box="[716,730,1477,1489]" fontSize="5" pageId="217" pageNumber="217">32</superScript>
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†
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||
<taxonomicName id="4C4617C8F6E268F0D8E5FA681923FA1E" baseAuthorityName="Muirhead and Wroe" baseAuthorityYear="1998" box="[750,833,1478,1497]" class="Mammalia" family="Thylacinidae" genus="Badjcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E268F0D8E5FA681923FA1E" box="[750,833,1478,1497]" italics="true" pageId="217" pageNumber="217">Badjcinus</emphasis>
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</taxonomicName>
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exhibits a carnivorously adapted dentition that most closely resembles those of other taxa currently placed in
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<taxonomicName id="4C4617C8F6E268F0D90BFA58190FF9CE" box="[768,877,1526,1545]" class="Mammalia" family="Thylacinidae" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="family">Thylacinidae</taxonomicName>
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(see
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<bibRefCitation id="EFD711BAF6E268F0D991FA581E1DF9CE" author="Muirhead, J. & S. Wroe" box="[922,1151,1526,1545]" pageId="217" pageNumber="217" pagination="612 - 626" refId="ref217610" refString="Muirhead, J., and S. Wroe. 1998. A new genus and species, Badjcinus turnbulli (Thylacinidae: Marsupialia), from the late Oligocene of Riversleigh, northern Australia, and an investigation of thylacinid phylogeny. Journal of Vertebrate Paleontology 18: 612 - 626." type="journal article" year="1998">Muirhead and Wroe, 1998</bibRefCitation>
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), but it has some unusual features—most notably, fusion between stB and the paracone on M1 (char. 136), which is not seen in definitive thylacinids, but which is seen in several dasyurids (e.g.,
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<taxonomicName id="4C4617C8F6E268F0D8EEF9F81924F9AF" box="[741,838,1622,1640]" class="Mammalia" family="Dasyuridae" genus="Dasycercus" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E268F0D8EEF9F81924F9AF" box="[741,838,1622,1640]" italics="true" pageId="217" pageNumber="217">Dasycercus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C4617C8F6E268F0D953F9F819A1F9AE" authorityName="Spencer" authorityYear="1896" box="[856,963,1622,1641]" class="Mammalia" family="Dasyuridae" genus="Dasyuroides" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E268F0D953F9F819A1F9AE" box="[856,963,1622,1641]" italics="true" pageId="217" pageNumber="217">Dasyuroides</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C4617C8F6E268F0D9DDF9F81E4BF9AF" authorityName="E. Geoffroy Saint-Hilaire" authorityYear="1796" box="[982,1065,1622,1640]" class="Mammalia" family="Dasyuridae" genus="Dasyurus" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E268F0D9DDF9F81E4BF9AF" box="[982,1065,1622,1640]" italics="true" pageId="217" pageNumber="217">Dasyurus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C4617C8F6E268F0DE37F9F81EC2F9AE" authorityName="F. G. Cuvier" authorityYear="1837" box="[1084,1184,1622,1641]" class="Mammalia" family="Dasyuridae" genus="Sarcophilus" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E268F0DE37F9F81EC2F9AE" box="[1084,1184,1622,1641]" italics="true" pageId="217" pageNumber="217">Sarcophilus</emphasis>
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</taxonomicName>
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)— together with a seemingly plesiomorphic basicranium. Most strikingly, it lacks a posterior squamosal epitympanic sinus (char. 84), a structure that is present in all other known dasyuromorphians (
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<bibRefCitation id="EFD711BAF6E268F0D939F91819CCF90E" author="Archer, M." box="[818,942,1718,1737]" pageId="217" pageNumber="217" pagination="217 - 322" refId="ref189612" refString="Archer, M. 1976 b. The basicranial region of marsupicarnivores (Marsupialia), inter-relationships of carnivorous marsupials, and affinities of the insectivorous marsupial peramelids. Zoological Journal of the Linnean Society 59 (3): 217 - 322." type="journal article" year="1976">Archer, 1976b</bibRefCitation>
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;
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<bibRefCitation id="EFD711BAF6E268F0D9BCF9181EFEF90E" author="Muirhead, J. & S. Wroe" box="[951,1180,1718,1737]" pageId="217" pageNumber="217" pagination="612 - 626" refId="ref217610" refString="Muirhead, J., and S. Wroe. 1998. A new genus and species, Badjcinus turnbulli (Thylacinidae: Marsupialia), from the late Oligocene of Riversleigh, northern Australia, and an investigation of thylacinid phylogeny. Journal of Vertebrate Paleontology 18: 612 - 626." type="journal article" year="1998">Muirhead and Wroe, 1998</bibRefCitation>
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).
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</paragraph>
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</footnote>
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<paragraph id="8BF96C4BF6E268F0DA87F9A21ACFF981" blockId="217.[108,637,1416,1738]" pageId="217" pageNumber="217">CROWN AGE: 31.2 Mya (95% HPD: 26.6–36.5 Mya).</paragraph>
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<paragraph id="8BF96C4BF6E268F0DA87F9E1194AFE26" blockId="217.[108,637,1416,1738]" lastBlockId="217.[684,1213,226,1440]" pageId="217" pageNumber="217">UNAMBIGUOUS CRANIODENTAL SYNAPOMORPHIES: Nasals not produced beyond premaxillary facial processes, incisive foramina exposed in dorsal view (char. 1: 0→1; ci = 1.000); postorbital processes present (char. 18: 0→1; ci = 0.042); posterolateral palatal foramina absent, posterior margins incomplete (char. 40: 0→1; ci = 0.200); stD taller than stB on M2 (char. 134: 0→1; ci = 0.250); m3 hypoconid lingual to salient protoconid (char. 173: 0→1; ci = 0.045); and lower molars with distinct posterior cingulid (char. 180: 0→1; ci = 0.333).</paragraph>
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</subSubSection>
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<subSubSection id="C35C3FC0F6E268F3D8C7FE441860FD40" lastPageId="218" lastPageNumber="218" pageId="217" pageNumber="217" type="discussion">
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<paragraph id="8BF96C4BF6E268F0D8C7FE441E2EFC48" blockId="217.[684,1213,226,1440]" pageId="217" pageNumber="217">
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COMMENTS: Monophyly of
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<taxonomicName id="4C4617C8F6E268F0DE0AFE4419BCFDE4" authority="sensu Kealy and Beck (2017)" authorityName="sensu Kealy and Beck" authorityYear="2017" class="Mammalia" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="order">
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Dasyuromorphia sensu
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<bibRefCitation id="EFD711BAF6E268F0D8E5FDA519BCFDE4" author="Kealy, S. & R. M. D. Beck" box="[750,990,523,547]" pageId="217" pageNumber="217" pagination="240" refId="ref210569" refString="Kealy, S., and R. M. D. Beck. 2017. Total evidence phylogeny and evolutionary timescale for Australian faunivorous marsupials (Dasyuromorphia). BMC Evolutionary Biology 17 (1): 240." type="journal article" year="2017">Kealy and Beck (2017)</bibRefCitation>
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</taxonomicName>
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—the most inclusive clade including
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<taxonomicName id="4C4617C8F6E268F0D950FD831E51FD83" baseAuthorityName="Shaw" baseAuthorityYear="1800" box="[859,1075,557,580]" class="Mammalia" family="Dasyuridae" genus="Dasyurus" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="species" species="viverrinus">
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<emphasis id="B932B059F6E268F0D950FD831E51FD83" box="[859,1075,557,580]" italics="true" pageId="217" pageNumber="217">Dasyurus viverrinus</emphasis>
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</taxonomicName>
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, but excluding
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<taxonomicName id="4C4617C8F6E268F0D8D3FDE319F7FDA2" authorityName="E. Geoffroy Saint-Hilaire" authorityYear="1804" box="[728,917,589,613]" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="species" species="nasuta">
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<emphasis id="B932B059F6E268F0D8D3FDE319F7FDA2" box="[728,917,589,613]" italics="true" pageId="217" pageNumber="217">Perameles nasuta</emphasis>
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</taxonomicName>
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,
|
||
<taxonomicName id="4C4617C8F6E268F0D9AEFDE01E1AFDA2" baseAuthorityName="Stirling" baseAuthorityYear="1889" box="[933,1144,589,613]" class="Mammalia" family="Notoryctidae" genus="Notoryctes" kingdom="Animalia" order="Notoryctemorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="species" species="typhlops">
|
||
<emphasis id="B932B059F6E268F0D9AEFDE01E1AFDA2" box="[933,1144,589,613]" italics="true" pageId="217" pageNumber="217">Notoryctes typhlops</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4617C8F6E268F0DE82FDE31901FD41" class="Mammalia" family="Phalangeridae" genus="Phalanger" kingdom="Animalia" order="Diprotodontia" pageId="217" pageNumber="217" phylum="Chordata" rank="species" species="orientalis">
|
||
<emphasis id="B932B059F6E268F0DE82FDE31901FD41" italics="true" pageId="217" pageNumber="217">Phalanger orientalis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C4617C8F6E268F0D9A7FDC11EF2FD41" authorityName="Thomas" authorityYear="1894" box="[940,1168,622,646]" class="Mammalia" family="Microbiotheriidae" genus="Dromiciops" kingdom="Animalia" order="Microbiotheria" pageId="217" pageNumber="217" phylum="Chordata" rank="species" species="gliroides">
|
||
<emphasis id="B932B059F6E268F0D9A7FDC11EF2FD41" box="[940,1168,622,646]" italics="true" pageId="217" pageNumber="217">Dromiciops gliroides</emphasis>
|
||
</taxonomicName>
|
||
—is strongly supported in our total-evidence analyses (figs. 32,
|
||
<figureCitation id="137D70CEF6E268F0D915FD1F195EFD0E" box="[798,828,689,713]" captionStart="FIG" captionStartId="181.[108,150,938,959]" captionTargetId="figure-12@181.[241,1216,224,1485]" captionTargetPageId="181" captionText="FIG. 33. (opposite page and at right) Fifty-per- cent majority rule consensus of post-burn-in trees that results from dated Bayesian analysis (using combined tip-and-node dating and separate Independent Gamma Rates [IGR] clock models for the molecular and morpho- logical partitions) of our total evidence data- set. Black dots at nodes indicate ≥0.95 Bayesian posterior probability (“strong support”); dark gray dots indicate 0.75–0.94 Bayesian poste- rior probability (“moderate support”); light gray dots indicate 0.50–0.74 Bayesian poste- rior probability (“weak support”). Nodes with- out dots were constrained a priori so that their ages could be calibrated (see tables 6, 13). Orange bars represent 95% Highest Posterior Density (HPD) intervals on the ages of nodes. For clarity, 95% HPD intervals are not shown for the ages of fossil terminals." pageId="217" pageNumber="217">33</figureCitation>
|
||
). This clade is supported by six unambiguous craniodental synapomorphies, of which four—nasals not produced beyond premaxillary facial processes, posterolateral palatal foramina absent, stD taller than stB on M2, and lower molars with distinct posterior cingulid present—show little or no homoplasy.
|
||
</paragraph>
|
||
<paragraph id="8BF96C4BF6E268F3D8C6FC361860FD40" blockId="217.[684,1213,226,1440]" lastBlockId="218.[108,637,226,647]" lastPageId="218" lastPageNumber="218" pageId="217" pageNumber="217">
|
||
†
|
||
<taxonomicName id="4C4617C8F6E268F0D8D2FC3619C8FC77" authorityName="Muirhead & Wroe" authorityYear="1998" box="[729,938,920,944]" class="Mammalia" family="Thylacinidae" genus="Badjcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="species" species="turnbulli">
|
||
<emphasis id="B932B059F6E268F0D8D2FC3619C8FC77" box="[729,938,920,944]" italics="true" pageId="217" pageNumber="217">Badjcinus turnbulli</emphasis>
|
||
</taxonomicName>
|
||
from the late Oligocene (Faunal Zone A) of Riversleigh World Heritage area was originally described as a thylacinid (see
|
||
<bibRefCitation id="EFD711BAF6E268F0D8A6FC5519BBFBD4" author="Muirhead, J. & S. Wroe" box="[685,985,1019,1043]" pageId="217" pageNumber="217" pagination="612 - 626" refId="ref217610" refString="Muirhead, J., and S. Wroe. 1998. A new genus and species, Badjcinus turnbulli (Thylacinidae: Marsupialia), from the late Oligocene of Riversleigh, northern Australia, and an investigation of thylacinid phylogeny. Journal of Vertebrate Paleontology 18: 612 - 626." type="journal article" year="1998">Muirhead and Wroe, 1998</bibRefCitation>
|
||
), and it is placed in
|
||
<taxonomicName id="4C4617C8F6E268F0D8A7FBB21E1EFBF3" authority="sensu Kealy and Beck" authorityName="Kealy and Beck" box="[684,1148,1052,1076]" class="Mammalia" family="Thylacinidae" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="family">
|
||
Thylacinidae sensu
|
||
<bibRefCitation id="EFD711BAF6E268F0D98DFBB21E1EFBF3" author="Kealy, S. & R. M. D. Beck" box="[902,1148,1052,1076]" pageId="217" pageNumber="217" pagination="240" refId="ref210569" refString="Kealy, S., and R. M. D. Beck. 2017. Total evidence phylogeny and evolutionary timescale for Australian faunivorous marsupials (Dasyuromorphia). BMC Evolutionary Biology 17 (1): 240." type="journal article" year="2017">Kealy and Beck (2017</bibRefCitation>
|
||
</taxonomicName>
|
||
:
|
||
<tableCitation id="C6C459F0F6E268F0DE83FBB218DAFB92" captionStart="TABLE 1" captionStartId="7.[611,691,226,250]" captionTargetBox="[119,1110,327,1337]" captionTargetPageId="7" captionText="TABLE 1 Higher Classification and Geographic Distribution of Recent Marsupialsa" httpUri="http://table.plazi.org/id/DF393CC3F63C682ED868FF4C1E65FED9" pageId="217" pageNumber="217" tableUuid="DF393CC3F63C682ED868FF4C1E65FED9">table 1</tableCitation>
|
||
) in our undated total-evidence analysis (fig. 32). However, in our dated total-evidence analysis, †
|
||
<taxonomicName id="4C4617C8F6E268F0D8ECFBD11930FB50" baseAuthorityName="Muirhead and Wroe" baseAuthorityYear="1998" box="[743,850,1151,1175]" class="Mammalia" family="Thylacinidae" genus="Badjcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932B059F6E268F0D8ECFBD11930FB50" box="[743,850,1151,1175]" italics="true" pageId="217" pageNumber="217">Badjcinus</emphasis>
|
||
</taxonomicName>
|
||
is placed in a trichotomy with
|
||
<taxonomicName id="4C4617C8F6E268F0D8A7FB0E1955FB7F" box="[684,823,1184,1208]" class="Mammalia" family="Thylacinidae" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="family">Thylacinidae</taxonomicName>
|
||
(†
|
||
<taxonomicName id="4C4617C8F6E268F0D95CFB0E19B5FB7F" authorityName="Muirhead & Archer" authorityYear="1990" box="[855,983,1184,1208]" class="Mammalia" family="Thylacinidae" genus="Nimbacinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932B059F6E268F0D95CFB0E19B5FB7F" box="[855,983,1184,1208]" italics="true" pageId="217" pageNumber="217">Nimbacinus</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C4617C8F6E268F0D9F2FB0E1E09FB7F" authorityName="Temminck" authorityYear="1824" box="[1017,1131,1184,1208]" class="Mammalia" family="Thylacinidae" genus="Thylacinus" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932B059F6E268F0D9F2FB0E1E09FB7F" box="[1017,1131,1184,1208]" italics="true" pageId="217" pageNumber="217">Thylacinus</emphasis>
|
||
</taxonomicName>
|
||
) and a clade comprising our remaining dasyuromorphian terminals (
|
||
<figureCitation id="137D70CEF6E268F0D963FB4D19D3FB3C" box="[872,945,1251,1275]" captionStart="FIG" captionStartId="181.[108,150,938,959]" captionTargetId="figure-12@181.[241,1216,224,1485]" captionTargetPageId="181" captionText="FIG. 33. (opposite page and at right) Fifty-per- cent majority rule consensus of post-burn-in trees that results from dated Bayesian analysis (using combined tip-and-node dating and separate Independent Gamma Rates [IGR] clock models for the molecular and morpho- logical partitions) of our total evidence data- set. Black dots at nodes indicate ≥0.95 Bayesian posterior probability (“strong support”); dark gray dots indicate 0.75–0.94 Bayesian poste- rior probability (“moderate support”); light gray dots indicate 0.50–0.74 Bayesian poste- rior probability (“weak support”). Nodes with- out dots were constrained a priori so that their ages could be calibrated (see tables 6, 13). Orange bars represent 95% Highest Posterior Density (HPD) intervals on the ages of nodes. For clarity, 95% HPD intervals are not shown for the ages of fossil terminals." pageId="217" pageNumber="217">fig. 33</figureCitation>
|
||
). It is, therefore, unclear whether †
|
||
<taxonomicName id="4C4617C8F6E268F0D912FAAA19E0FADB" baseAuthorityName="Muirhead and Wroe" baseAuthorityYear="1998" box="[793,898,1284,1308]" class="Mammalia" family="Thylacinidae" genus="Badjcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="217" pageNumber="217" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932B059F6E268F0D912FAAA19E0FADB" box="[793,898,1284,1308]" italics="true" pageId="217" pageNumber="217">Badjcinus</emphasis>
|
||
</taxonomicName>
|
||
is a member of the dasyuromorphian crown clade (= Dasyuroidea;
|
||
<bibRefCitation id="EFD711BAF6E268F0DE75FA8B1935FA99" author="Kealy, S. & R. M. D. Beck" pageId="217" pageNumber="217" pagination="240" refId="ref210569" refString="Kealy, S., and R. M. D. Beck. 2017. Total evidence phylogeny and evolutionary timescale for Australian faunivorous marsupials (Dasyuromorphia). BMC Evolutionary Biology 17 (1): 240." type="journal article" year="2017">Kealy and Beck, 2017</bibRefCitation>
|
||
).
|
||
<superScript id="7C33C103F6E268F0D96AFAED1910FA93" attach="left" box="[865,882,1347,1364]" fontSize="7" pageId="217" pageNumber="217">32</superScript>
|
||
In this respect, our results are broadly similar to those of
|
||
<bibRefCitation id="EFD711BAF6E268F0DE0AFAC91897FA67" author="Kealy, S. & R. M. D. Beck" pageId="217" pageNumber="217" pagination="240" refId="ref210569" refString="Kealy, S., and R. M. D. Beck. 2017. Total evidence phylogeny and evolutionary timescale for Australian faunivorous marsupials (Dasyuromorphia). BMC Evolutionary Biology 17 (1): 240." type="journal article" year="2017">Kealy and Beck (2017)</bibRefCitation>
|
||
, who also failed to unambiguously place †
|
||
<taxonomicName id="4C4617C8F6E168F3DA72FF4C1A80FF3D" baseAuthorityName="Muirhead and Wroe" baseAuthorityYear="1998" box="[121,226,226,250]" class="Mammalia" family="Thylacinidae" genus="Badjcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932B059F6E168F3DA72FF4C1A80FF3D" box="[121,226,226,250]" italics="true" pageId="218" pageNumber="218">Badjcinus</emphasis>
|
||
</taxonomicName>
|
||
within
|
||
<taxonomicName id="4C4617C8F6E168F3DB31FF4C1BA5FF3D" box="[314,455,226,250]" class="Mammalia" family="Thylacinidae" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="family">Thylacinidae</taxonomicName>
|
||
, despite a much denser sampling of dasyuromorphians than that used here; instead, it was placed as sister to all other dasyuromorphians in some analyses, congruent with its antiquity (late Oligocene) and putative basicranial plesiomorphies.
|
||
<bibRefCitation id="EFD711BAF6E168F3D800FE291A92FE07" author="Kealy, S. & R. M. D. Beck" pageId="218" pageNumber="218" pagination="240" refId="ref210569" refString="Kealy, S., and R. M. D. Beck. 2017. Total evidence phylogeny and evolutionary timescale for Australian faunivorous marsupials (Dasyuromorphia). BMC Evolutionary Biology 17 (1): 240." type="journal article" year="2017">Kealy and Beck (2017)</bibRefCitation>
|
||
argued that †
|
||
<taxonomicName id="4C4617C8F6E168F3DB87FE061B97FE07" baseAuthorityName="Muirhead and Wroe" baseAuthorityYear="1998" box="[396,501,424,448]" class="Mammalia" family="Thylacinidae" genus="Badjcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932B059F6E168F3DB87FE061B97FE07" box="[396,501,424,448]" italics="true" pageId="218" pageNumber="218">Badjcinus</emphasis>
|
||
</taxonomicName>
|
||
is best considered?
|
||
<taxonomicName id="4C4617C8F6E168F3DADEFE671B05FE26" box="[213,359,457,481]" class="Mammalia" family="Thylacinidae" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="family">Thylacinidae</taxonomicName>
|
||
, based on current evidence. By contrast, the phylogenetic analyses of
|
||
<bibRefCitation id="EFD711BAF6E168F3DA67FDA51B3EFDE4" author="Rovinsky" box="[108,348,523,547]" etAl="et al." firstAuthor="Rovinsky" pageId="218" pageNumber="218" pagination="7457" refId="ref225576" refString="Rovinsky, D. S., A. R. Evans, and J. W. Adams. 2019. The pre-Pleistocene fossil thylacinids (Dasyuromorphia: Thylacinidae) and the evolutionary context of the modern thylacine. PeerJ 7: e 7457." type="journal article" year="2019">Rovinsky et al. (2019)</bibRefCitation>
|
||
consistently placed †
|
||
<taxonomicName id="4C4617C8F6E168F3D843FDA51AC8FD83" baseAuthorityName="Muirhead and Wroe" baseAuthorityYear="1998" class="Mammalia" family="Thylacinidae" genus="Badjcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932B059F6E168F3D843FDA51AC8FD83" italics="true" pageId="218" pageNumber="218">Badjcinus</emphasis>
|
||
</taxonomicName>
|
||
within
|
||
<taxonomicName id="4C4617C8F6E168F3DAF5FD821BE9FD83" box="[254,395,556,580]" class="Mammalia" family="Thylacinidae" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="family">Thylacinidae</taxonomicName>
|
||
, but these employed a less dense sampling of dasyuromorphian taxa than those of
|
||
<bibRefCitation id="EFD711BAF6E168F3DB0FFDC01B9DFD40" author="Kealy, S. & R. M. D. Beck" box="[260,511,622,647]" pageId="218" pageNumber="218" pagination="240" refId="ref210569" refString="Kealy, S., and R. M. D. Beck. 2017. Total evidence phylogeny and evolutionary timescale for Australian faunivorous marsupials (Dasyuromorphia). BMC Evolutionary Biology 17 (1): 240." type="journal article" year="2017">Kealy and Beck (2017)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |