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<mods:title id="DD86C594F0FC3936A10A32394724DD81">Revision of Hestiasulini Giglio-Tos, 1915 stat. rev. (Insecta: Mantodea: Hymenopodidae) of Borneo, with description of new taxa and comments on the taxonomy of the tribe</mods:title>
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<treatment id="03DC81241812A16AFF6BFA68FA736661" ID-DOI="http://doi.org/10.5281/zenodo.6050966" ID-GBIF-Taxon="132549688" ID-Zenodo-Dep="6050966" LSID="urn:lsid:plazi:treatment:03DC81241812A16AFF6BFA68FA736661" httpUri="http://treatment.plazi.org/id/03DC81241812A16AFF6BFA68FA736661" lastPageId="26" lastPageNumber="269" pageId="25" pageNumber="268">
<subSubSection id="C36F63B91812A169FF6BFA68FE51655E" pageId="25" pageNumber="268" type="nomenclature">
<paragraph id="8BCA30321812A169FF6BFA68FE966695" blockId="25.[151,1436,1497,1592]" box="[151,377,1497,1523]" pageId="25" pageNumber="268">
<heading id="D082875E1812A169FF6BFA68FE966695" bold="true" box="[151,377,1497,1523]" fontSize="11" level="1" pageId="25" pageNumber="268" reason="1">
<emphasis id="B901EC201812A169FF6BFA68FE966695" bold="true" box="[151,377,1497,1523]" pageId="25" pageNumber="268">
Key to
<taxonomicName id="4C754BB11812A169FF0CFA68FE966695" box="[240,377,1497,1523]" class="Insecta" family="Oxypilinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="25" pageNumber="268" phylum="Arthropoda" rank="family">Oxypilinae</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCA30321812A169FF6BFA4AFE51655E" blockId="25.[151,1436,1497,1592]" pageId="25" pageNumber="268">
(after
<bibRefCitation id="EFE44DC31812A169FF21FA4AFEB06572" author="Beier" box="[221,351,1531,1556]" pageId="25" pageNumber="268" refString="Beier, M. (1934) Mantodea. Fam. Mantidae. Subfam. Hymenopodinae. Genera Insectorum, Fascicule 196, 1 - 37, pls. 1 - 2 [Nachtrage 1937, 3 pp.]" type="book chapter" year="1934">Beier 1934</bibRefCitation>
,
<bibRefCitation id="EFE44DC31812A169FE90FA4DFD876572" author="Roy" box="[364,616,1532,1556]" pageId="25" pageNumber="268" refString="Roy, R. &amp; Svenson, G. J. (2007) Revision of the genus Ceratomantis Wood-Mason, 1876 (Dictyoptera, Mantodea). Bulletin de la Societe entomologique de France, 112 (4), 433 - 444." type="journal article" year="2007">Roy &amp; Svenson 2007</bibRefCitation>
,
<bibRefCitation id="EFE44DC31812A169FD8AFA4DFD056572" author="Roy" box="[630,746,1532,1556]" pageId="25" pageNumber="268" refString="Roy, R. (2013) Revision du genre Pachymantis Saussure, 1871 (Mantodea, Hymenopodidae). Bulletin de la Societe entomologique de France, 118 (2), 145 - 154." type="journal article" year="2013">Roy 2013</bibRefCitation>
, and
<bibRefCitation id="EFE44DC31812A169FCD6FA4DFBFD6572" author="Svenson" box="[810,1042,1531,1556]" pageId="25" pageNumber="268" refString="Svenson, G. J., Hardy, N. B., Cahill Wightman, H. M. &amp; Wieland, F. (2015) Of flowers and twigs: phylogenetic revision of the plant-mimicking praying mantises (Mantodea: Empusidae and Hymenopodidae) with a new suprageneric classification. Systematic Entomology, 40 (4), 789 - 834." type="journal article" year="2015">
Svenson
<emphasis id="B901EC201812A169FC6FFA4CFC286572" box="[915,967,1531,1556]" italics="true" pageId="25" pageNumber="268">et al</emphasis>
. 2015
</bibRefCitation>
, modified); Bornean members are keyed out to species level.
</paragraph>
</subSubSection>
<subSubSection id="C36F63B91812A16AFF6BF9D6FA736661" lastPageId="26" lastPageNumber="269" pageId="25" pageNumber="268" type="key">
<keyStep id="308128971812A169FF6BF9D6FA7365B2" pageId="25" pageNumber="268">
<paragraph id="8BCA30321812A169FF6BF9D6FA7465FF" blockId="25.[151,1436,1639,2012]" pageId="25" pageNumber="268">
<keyLead id="308493071812A169FF6BF9D6FA7465FF" pageId="25" pageNumber="268">1. Pronotum with 3–4 pairs of conical or spine-like tubercles; dilated part of femur about half as wide as non-dilated part; pos- tero-ventral margin of fore femur not or very slightly denticulate (tribus Oxypilini)................................. 2</keyLead>
</paragraph>
<paragraph id="8BCA30321812A169FF6BF913FA7365B2" blockId="25.[151,1436,1639,2012]" pageId="25" pageNumber="268">
<keyLead id="308493071812A169FF6BF913FA7365B2" pageId="25" pageNumber="268">- Pronotum smooth or slightly tuberculate, but never with conical or spine-like tubercles; dilated part of femur about as wide as non-dilated part; postero-ventral margin of fore femur distinctly denticulate (tribus Hestiasulini).......................7</keyLead>
</paragraph>
</keyStep>
<keyStep id="308128971812A169FF6BF96DFA736469" pageId="25" pageNumber="268">
<paragraph id="8BCA30321812A169FF6BF96DFA746597" blockId="25.[151,1436,1639,2012]" box="[151,1435,1756,1777]" pageId="25" pageNumber="268">
<keyLead id="308493071812A169FF6BF96DFA746597" box="[151,1435,1756,1777]" pageId="25" pageNumber="268">2. Prozona with two pairs of conical or spine-like tubercles (Afrotropical)...........................................3</keyLead>
</paragraph>
<paragraph id="8BCA30321812A169FF6BF94BFA736469" blockId="25.[151,1436,1639,2012]" box="[151,1436,1786,1807]" pageId="25" pageNumber="268">
<keyLead id="308493071812A169FF6BF94BFA736469" box="[151,1436,1786,1807]" pageId="25" pageNumber="268">- Prozona with one pair of spine-like tubercles (Oriental).......................................................4</keyLead>
</paragraph>
</keyStep>
<keyStep id="308128971812A169FF6BF8A6FA736401" pageId="25" pageNumber="268">
<paragraph id="8BCA30321812A169FF6BF8A6FA73644A" blockId="25.[151,1436,1639,2012]" box="[151,1436,1814,1836]" pageId="25" pageNumber="268">
<keyLead id="308493071812A169FF6BF8A6FA73644A" box="[151,1436,1814,1836]" pageId="25" pageNumber="268">
3. Walking leg femora without lobes; females apterous.................................
<taxonomicName id="4C754BB11812A169FB98F8A7FA73644A" ID-CoL="8NWJC" authority="Audinet-Serville, 1831" authorityName="Audinet-Serville" authorityYear="1831" box="[1124,1436,1814,1836]" class="Insecta" family="Hymenopodidae" genus="Oxypilus" kingdom="Animalia" order="Mantodea" pageId="25" pageNumber="268" phylum="Arthropoda" rank="genus">
<emphasis id="B901EC201812A169FB98F8A7FB56644A" box="[1124,1209,1814,1836]" italics="true" pageId="25" pageNumber="268">Oxypilus</emphasis>
<bibRefCitation id="EFE44DC31812A169FB3CF8A6FA73644A" author="Audinet-Serville" box="[1216,1436,1815,1836]" pageId="25" pageNumber="268" refString="Audinet-Serville, J. - G. (1831) Revue methodique des insectes de l'ordre des Orthopteres. Annales des Sciences Naturelles, 22, 28 - 65." type="journal article" year="1831">Audinet-Serville, 1831</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCA30321812A169FF6BF885FA736401" blockId="25.[151,1436,1639,2012]" pageId="25" pageNumber="268">
<keyLead id="308493071812A169FF6BF885FA736401" pageId="25" pageNumber="268">
- Ventral lamina of walking leg femora produced into 3 small lobes; females brachypterous.....................................................................................................
<taxonomicName id="4C754BB11812A169FBBEF8E0FA736401" ID-CoL="8NT3H" authority="Schulthess-Rechberg, 1899" authorityName="Schulthess-Rechberg" authorityYear="1899" box="[1090,1436,1873,1895]" class="Insecta" family="Hymenopodidae" genus="Junodia" kingdom="Animalia" order="Mantodea" pageId="25" pageNumber="268" phylum="Arthropoda" rank="genus">
<emphasis id="B901EC201812A169FBBEF8E0FB606401" box="[1090,1167,1873,1895]" italics="true" pageId="25" pageNumber="268">Junodia</emphasis>
Schulthess-Rechberg, 1899
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308128971812A169FF6BF8DEFA7364BA" pageId="25" pageNumber="268">
<paragraph id="8BCA30321812A169FF6BF8DEFA7464C7" blockId="25.[151,1436,1639,2012]" pageId="25" pageNumber="268">
<keyLead id="308493071812A169FF6BF8DEFA7464C7" pageId="25" pageNumber="268">
4. Metazona with two spine-like processes; process on the vertex long (over 75% width of head), bifid, with two pairs of smaller lateral processes; frontal shield with a median keel (genus
<taxonomicName id="4C754BB11812A169FCEEF83DFBB664C7" ID-CoL="8NP7N" authority="Wood-Mason, 1876" authorityName="Wood-Mason" authorityYear="1876" box="[786,1113,1932,1954]" class="Insecta" family="Hymenopodidae" genus="Ceratomantis" kingdom="Animalia" order="Mantodea" pageId="25" pageNumber="268" phylum="Arthropoda" rank="genus">
<emphasis id="B901EC201812A169FCEEF83DFC7A64C4" box="[786,917,1932,1954]" italics="true" pageId="25" pageNumber="268">Ceratomantis</emphasis>
<bibRefCitation id="EFE44DC31812A169FC67F83DFBB664C7" author="Wood-Mason" box="[923,1113,1932,1953]" pageId="25" pageNumber="268" refString="Wood-Mason, J. (1876) On some remarkable species of Mantidae. Proceedings of the Asiatic Society of Bengal, 1876, 175 - 176." type="proceedings paper" year="1876">Wood-Mason, 1876</bibRefCitation>
</taxonomicName>
).........................5
</keyLead>
</paragraph>
<paragraph id="8BCA30321812A169FF6BF81BFA7364BA" blockId="25.[151,1436,1639,2012]" pageId="25" pageNumber="268">
<keyLead id="308493071812A169FF6BF81BFA7364BA" pageId="25" pageNumber="268">
- Metazona with four spine-like processes, one paramedian and one lateral set; process on vertex short (less than 25% width of head), simple; frontal shield with two flattened, paramedian keels (genus
<taxonomicName id="4C754BB11812A169FC5DF877FB5A64BA" ID-CoL="8NWK2" authority="Saussure, 1871" authorityName="Saussure" authorityYear="1871" box="[929,1205,1990,2012]" class="Insecta" family="Hymenopodidae" genus="Pachymantis" kingdom="Animalia" order="Mantodea" pageId="25" pageNumber="268" phylum="Arthropoda" rank="genus">
<emphasis id="B901EC201812A169FC5DF877FBF264BA" box="[929,1053,1990,2012]" italics="true" pageId="25" pageNumber="268">Pachymantis</emphasis>
<bibRefCitation id="EFE44DC31812A169FBDFF876FB5A64BA" author="Saussure" box="[1059,1205,1991,2012]" pageId="25" pageNumber="268" refString="Saussure, H. de (1871) Melanges orthopterologiques. Supplement au III me Fascicule. Mantides. Memoires de la Societe de Physique et d'Histoire naturelle de Geneve, 21 (1), 239 - 337, pl. 7." type="journal article" year="1871">Saussure, 1871</bibRefCitation>
</taxonomicName>
).................6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308128971811A16AFF6BFF26FA736262" pageId="26" pageNumber="269">
<paragraph id="8BCA30321811A16AFF6BFF26FA7363AC" blockId="26.[151,1436,151,1287]" pageId="26" pageNumber="269">
<keyLead id="308493071811A16AFF6BFF26FA7363AC" pageId="26" pageNumber="269">
5. Anterior side of fore coxa with a black median stripe extending from base to apex. Pronotum short, about 1.2 times as long as wide..........................................................................
<taxonomicName id="4C754BB11811A16AFB71FF05FA7363AC" ID-CoL="SND4" authority="Svenson, 2007" authorityName="Svenson" authorityYear="2007" box="[1165,1436,180,202]" class="Insecta" family="Hymenopodidae" genus="Ceratomantis" kingdom="Animalia" order="Mantodea" pageId="26" pageNumber="269" phylum="Arthropoda" rank="species" species="kimberlae">
<emphasis id="B901EC201811A16AFB71FF05FAE763AC" box="[1165,1288,180,202]" italics="true" pageId="26" pageNumber="269">C. kimberlae</emphasis>
<bibRefCitation id="EFE44DC31811A16AFAF2FF04FA7363AC" author="Svenson" box="[1294,1436,181,202]" pageId="26" pageNumber="269" refString="Roy, R. &amp; Svenson, G. J. (2007) Revision of the genus Ceratomantis Wood-Mason, 1876 (Dictyoptera, Mantodea). Bulletin de la Societe entomologique de France, 112 (4), 433 - 444." type="journal article" year="2007">Svenson, 2007</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCA30321811A16AFF6BFF63FA736262" blockId="26.[151,1436,151,1287]" pageId="26" pageNumber="269">
<keyLead id="308493071811A16AFF6BFF63FA736262" pageId="26" pageNumber="269">
- Anterior side of fore coxa with a subapical black spot only; pronotum longer, about 1.4 times as long as wide...............................................................................................
<taxonomicName id="4C754BB11811A16AFB40FF5EFA736262" ID-CoL="SND3" authority="Roy, 2007" authorityName="Roy" authorityYear="2007" box="[1212,1436,239,261]" class="Insecta" family="Hymenopodidae" genus="Ceratomantis" kingdom="Animalia" order="Mantodea" pageId="26" pageNumber="269" phylum="Arthropoda" rank="species" species="gigliotosi">
<emphasis id="B901EC201811A16AFB40FF5EFADC6263" box="[1212,1331,239,261]" italics="true" pageId="26" pageNumber="269">C. gigliotosi</emphasis>
<bibRefCitation id="EFE44DC31811A16AFAC5FF5EFA736262" author="Roy" box="[1337,1436,239,260]" pageId="26" pageNumber="269" refString="Roy, R. &amp; Svenson, G. J. (2007) Revision of the genus Ceratomantis Wood-Mason, 1876 (Dictyoptera, Mantodea). Bulletin de la Societe entomologique de France, 112 (4), 433 - 444." type="journal article" year="2007">Roy, 2007</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308128971811A16AFF6BFEBCFA73621C" pageId="26" pageNumber="269">
<paragraph id="8BCA30321811A16AFF6BFEBCFA736259" blockId="26.[151,1436,151,1287]" pageId="26" pageNumber="269">
<keyLead id="308493071811A16AFF6BFEBCFA736259" pageId="26" pageNumber="269">
6. Anterior side of fore coxa yellowish-brown with black apex and two large round spots; antero-ventral side of fore trochanter black; dilated part of fore femur black, ventral part reddish-brown............................
<taxonomicName id="4C754BB11811A16AFB51FE9BFA736259" ID-CoL="4BSQH" authority="Roy, 2013" authorityName="Roy" authorityYear="2013" box="[1197,1436,297,319]" class="Insecta" family="Hymenopodidae" genus="Pachymantis" kingdom="Animalia" order="Mantodea" pageId="26" pageNumber="269" phylum="Arthropoda" rank="species" species="maculicoxa">
<emphasis id="B901EC201811A16AFB51FE9BFADC6259" box="[1197,1331,297,319]" italics="true" pageId="26" pageNumber="269">P. maculicoxa</emphasis>
<bibRefCitation id="EFE44DC31811A16AFAC5FE9BFA736259" author="Roy" box="[1337,1436,298,319]" pageId="26" pageNumber="269" refString="Roy, R. (2013) Revision du genre Pachymantis Saussure, 1871 (Mantodea, Hymenopodidae). Bulletin de la Societe entomologique de France, 118 (2), 145 - 154." type="journal article" year="2013">Roy, 2013</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCA30321811A16AFF6BFEF6FA73621C" blockId="26.[151,1436,151,1287]" pageId="26" pageNumber="269">
<keyLead id="308493071811A16AFF6BFEF6FA73621C" pageId="26" pageNumber="269">
- Anterior side of fore coxa yellowish-brown at base, then largely black; antero-ventral side of fore trochanter yellowish-brown; anterior side of fore femur totally black..................................................
<taxonomicName id="4C754BB11811A16AFB4CFED4FA73621C" ID-CoL="4BSQJ" authority="Roy, 2013" authorityName="Roy" authorityYear="2013" box="[1200,1436,356,378]" class="Insecta" family="Hymenopodidae" genus="Pachymantis" kingdom="Animalia" order="Mantodea" pageId="26" pageNumber="269" phylum="Arthropoda" rank="species" species="piceifemur">
<emphasis id="B901EC201811A16AFB4CFED4FADD621C" box="[1200,1330,356,378]" italics="true" pageId="26" pageNumber="269">P. piceifemur</emphasis>
<bibRefCitation id="EFE44DC31811A16AFAC5FED4FA73621C" author="Roy" box="[1337,1436,357,378]" pageId="26" pageNumber="269" refString="Roy, R. (2013) Revision du genre Pachymantis Saussure, 1871 (Mantodea, Hymenopodidae). Bulletin de la Societe entomologique de France, 118 (2), 145 - 154." type="journal article" year="2013">Roy, 2013</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308128971811A16AFF6BFE33FA746121" pageId="26" pageNumber="269">
<paragraph id="8BCA30321811A16AFF6BFE33FA7362B4" blockId="26.[151,1436,151,1287]" pageId="26" pageNumber="269">
<keyLead id="308493071811A16AFF6BFE33FA7362B4" pageId="26" pageNumber="269">
7. Male subgenital plate with styli (
<figureCitation id="134E2CB71811A16AFDDDFE33FD7D62F1" box="[545,658,386,407]" captionStart="FIGURES 21 – 22" captionStartId="9.[151,264,1104,1127]" captionTargetBox="[180,1396,199,1063]" captionTargetId="figure@9.[159,1428,173,1083]" captionTargetPageId="9" captionText="FIGURES 21 – 22. Subgenital plate and genitalia of 21. H. brunneriana from India (ES) and 22. C. moultoni from Sabah (ES). Arrows indicate weakly sclerotized membrane at base of sclerotized left margin of ventral phallomere (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829537/files/figure.png" pageId="26" pageNumber="269">Figs 21–22</figureCitation>
); ventral phallomere with membraneous gap along left margin (
<figureCitation id="134E2CB71811A16AFB01FE33FA8362F1" box="[1277,1388,386,407]" captionStart="FIGURES 21 – 22" captionStartId="9.[151,264,1104,1127]" captionTargetBox="[180,1396,199,1063]" captionTargetId="figure@9.[159,1428,173,1083]" captionTargetPageId="9" captionText="FIGURES 21 – 22. Subgenital plate and genitalia of 21. H. brunneriana from India (ES) and 22. C. moultoni from Sabah (ES). Arrows indicate weakly sclerotized membrane at base of sclerotized left margin of ventral phallomere (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829537/files/figure.png" pageId="26" pageNumber="269">Figs 21–22</figureCitation>
,
<figureCitation id="134E2CB71811A16AFA84FE33FF1362D2" captionStart="FIGURES 43 – 50" captionStartId="22.[151,264,1570,1593]" captionTargetBox="[151,1436,467,1550]" captionTargetId="figure@22.[151,1436,467,1550]" captionTargetPageId="22" captionText="FIGURES 43 – 50. Genitalia of various Hestiasulini. 43. A. phyllopus from Sabah (CS). 44. A. basinigra from Myanmar (SMNK). 45. C. nitida from Myanmar (SMNK). 46. C. nitida from Sumatra (SMNK). 47. C. sp. from Kerala (SMNK). 48. Ephestiasula amoena from Nepal (ES). 49. E. pictipes from India (CS). 50. H. gyldenstolpei from Thailand (ES) (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829549/files/figure.png" pageId="26" pageNumber="269">45– 50</figureCitation>
); dorsal lamina of left phallomere not folded longitudinally (
<figureCitation id="134E2CB71811A16AFCBEFE2EFC5F62D2" box="[834,944,415,436]" captionStart="FIGURES 21 – 22" captionStartId="9.[151,264,1104,1127]" captionTargetBox="[180,1396,199,1063]" captionTargetId="figure@9.[159,1428,173,1083]" captionTargetPageId="9" captionText="FIGURES 21 – 22. Subgenital plate and genitalia of 21. H. brunneriana from India (ES) and 22. C. moultoni from Sabah (ES). Arrows indicate weakly sclerotized membrane at base of sclerotized left margin of ventral phallomere (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829537/files/figure.png" pageId="26" pageNumber="269">Figs 21–22</figureCitation>
,
<figureCitation id="134E2CB71811A16AFC46FE2EFC1562D2" box="[954,1018,415,436]" captionStart="FIGURES 43 – 50" captionStartId="22.[151,264,1570,1593]" captionTargetBox="[151,1436,467,1550]" captionTargetId="figure@22.[151,1436,467,1550]" captionTargetPageId="22" captionText="FIGURES 43 – 50. Genitalia of various Hestiasulini. 43. A. phyllopus from Sabah (CS). 44. A. basinigra from Myanmar (SMNK). 45. C. nitida from Myanmar (SMNK). 46. C. nitida from Sumatra (SMNK). 47. C. sp. from Kerala (SMNK). 48. Ephestiasula amoena from Nepal (ES). 49. E. pictipes from India (CS). 50. H. gyldenstolpei from Thailand (ES) (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829549/files/figure.png" pageId="26" pageNumber="269">45–50</figureCitation>
); head with a process or tubercle (
<figureCitation id="134E2CB71811A16AFABDFE2EFA7762D2" box="[1345,1432,415,436]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[323,1273,195,1896]" captionTargetId="figure@6.[323,1273,195,1896]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Dorsal habitus of Hestiasulini. 1. Male of Hestiasula brunneriana (Sri Lanka) (CS). 2. Female of H. brunneriana (Sri Lanka) (CS). 3. Male of Astyliasula phyllopus from Sabah (CS). 4. Female of A. phyllopus from Brunei (CS). 5. Male of A. sarawaca from Brunei (SMNK). 6. Female of A. sarawaca from Kalimantan (RMNH). 7. Pseudohestiasula borneana, holotype (SMNK). 8. Male of Catestiasula moultoni from Sarawak (CS) (scale bar: 10 mm)." httpUri="https://zenodo.org/record/829533/files/figure.png" pageId="26" pageNumber="269">Figs 1–2</figureCitation>
,
<figureCitation id="134E2CB71811A16AFF1DFE0CFECF62B4" box="[225,288,445,466]" captionStart="FIGURES 51 – 52" captionStartId="23.[151,264,1634,1657]" captionTargetBox="[359,1219,469,1613]" captionTargetId="figure@23.[340,1247,469,1613]" captionTargetPageId="23" captionText="FIGURES 51 – 52. Life aspect of Catestiasula. 51. Male of C. moultoni, Mt. Trus Madi, Sabah, Malaysia; Photo © Alexey Yakovlev. 52. Male of unidentified Catestiasula specimen from Ulu Gombak NP, Selangor, Malaysia; Photo © Bernhard Jacobi." httpUri="https://zenodo.org/record/829551/files/figure.png" pageId="26" pageNumber="269">51–52</figureCitation>
); ventral side of meso femora without lobe; alae of female hyaline or subhyaline (
<figureCitation id="134E2CB71811A16AFBCDFE0CFB8462B4" box="[1073,1131,445,466]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[323,1273,195,1896]" captionTargetId="figure@6.[323,1273,195,1896]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Dorsal habitus of Hestiasulini. 1. Male of Hestiasula brunneriana (Sri Lanka) (CS). 2. Female of H. brunneriana (Sri Lanka) (CS). 3. Male of Astyliasula phyllopus from Sabah (CS). 4. Female of A. phyllopus from Brunei (CS). 5. Male of A. sarawaca from Brunei (SMNK). 6. Female of A. sarawaca from Kalimantan (RMNH). 7. Pseudohestiasula borneana, holotype (SMNK). 8. Male of Catestiasula moultoni from Sarawak (CS) (scale bar: 10 mm)." httpUri="https://zenodo.org/record/829533/files/figure.png" pageId="26" pageNumber="269">Fig. 2</figureCitation>
).......................8
</keyLead>
</paragraph>
<paragraph id="8BCA30321811A16AFF6BFE6BFA746121" blockId="26.[151,1436,151,1287]" pageId="26" pageNumber="269">
<keyLead id="308493071811A16AFF6BFE6BFA746121" pageId="26" pageNumber="269">
- Male subgenital plate without styli (
<figureCitation id="134E2CB71811A16AFDC4FE6BFD496289" box="[568,678,474,495]" captionStart="FIGURES 23 – 25" captionStartId="15.[151,264,998,1021]" captionTargetBox="[289,1298,202,960]" captionTargetId="figure@15.[251,1335,193,977]" captionTargetPageId="15" captionText="FIGURES 23 – 25. Terminalia of A. sarawaca. 23. Supra-anal plate and cerci. 24. Subgenital plate, dorsal view. 25. Subgenital plate, posterior view (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829539/files/figure.png" pageId="26" pageNumber="269">Figs 24–25</figureCitation>
,
<figureCitation id="134E2CB71811A16AFD53FE6BFD256289" box="[687,714,474,495]" captionStart="FIGURES 40 – 42" captionStartId="21.[151,264,1095,1118]" captionTargetBox="[215,1378,208,1059]" captionTargetId="figure@21.[192,1395,193,1075]" captionTargetPageId="21" captionText="FIGURES 40 – 42. Genitalia of P. borneana. 40. Supra-anal plate and cerci (pilosity omitted). 41. Subgenital plate in dorsal view. 42. Genitalia in ventral view. Arrow indicates fold in dorsal lamina (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829547/files/figure.png" pageId="26" pageNumber="269">41</figureCitation>
); ventral phallomere without membraneous gap along left margin (
<figureCitation id="134E2CB71811A16AFAB4FE6BFF14616A" captionStart="FIGURES 26 – 32" captionStartId="16.[151,264,1780,1803]" captionTargetBox="[240,1366,494,1751]" captionTargetId="figure@16.[195,1392,481,1759]" captionTargetPageId="16" captionText="FIGURES 26 – 32. Genitalia of A. sarawaca. 26. Genitalia of Trus Madi male in ventral view. 27 – 28. Ventral processes of two Brunei males. 29 – 30. Phalloid apophysises of two Brunei males. 31. Right phallomere in antero-sinistral view. 32. Left phallomere in dorsal view. Arrow indicates fold in dorsal lamina (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829541/files/figure.png" pageId="26" pageNumber="269">Figs 26– 28</figureCitation>
,
<figureCitation id="134E2CB71811A16AFEFAFE46FEAA616A" box="[262,325,503,524]" captionStart-0="FIGURES 40 – 42" captionStart-1="FIGURES 43 – 50" captionStartId-0="21.[151,264,1095,1118]" captionStartId-1="22.[151,264,1570,1593]" captionTargetBox-0="[215,1378,208,1059]" captionTargetBox-1="[151,1436,467,1550]" captionTargetId-0="figure@21.[192,1395,193,1075]" captionTargetId-1="figure@22.[151,1436,467,1550]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURES 40 – 42. Genitalia of P. borneana. 40. Supra-anal plate and cerci (pilosity omitted). 41. Subgenital plate in dorsal view. 42. Genitalia in ventral view. Arrow indicates fold in dorsal lamina (scale bar: 1 mm)." captionText-1="FIGURES 43 – 50. Genitalia of various Hestiasulini. 43. A. phyllopus from Sabah (CS). 44. A. basinigra from Myanmar (SMNK). 45. C. nitida from Myanmar (SMNK). 46. C. nitida from Sumatra (SMNK). 47. C. sp. from Kerala (SMNK). 48. Ephestiasula amoena from Nepal (ES). 49. E. pictipes from India (CS). 50. H. gyldenstolpei from Thailand (ES) (scale bar: 1 mm)." httpUri-0="https://zenodo.org/record/829547/files/figure.png" httpUri-1="https://zenodo.org/record/829549/files/figure.png" pageId="26" pageNumber="269">42–44</figureCitation>
); dorsal lamina of left phallomere folded longitudinally (
<figureCitation id="134E2CB71811A16AFC93FE46FC54616A" box="[879,955,503,524]" captionStart="FIGURES 26 – 32" captionStartId="16.[151,264,1780,1803]" captionTargetBox="[240,1366,494,1751]" captionTargetId="figure@16.[195,1392,481,1759]" captionTargetPageId="16" captionText="FIGURES 26 – 32. Genitalia of A. sarawaca. 26. Genitalia of Trus Madi male in ventral view. 27 – 28. Ventral processes of two Brunei males. 29 – 30. Phalloid apophysises of two Brunei males. 31. Right phallomere in antero-sinistral view. 32. Left phallomere in dorsal view. Arrow indicates fold in dorsal lamina (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829541/files/figure.png" pageId="26" pageNumber="269">Figs 26</figureCitation>
,
<figureCitation id="134E2CB71811A16AFC39FE46FBEA616A" box="[965,1029,503,524]" captionStart-0="FIGURES 40 – 42" captionStart-1="FIGURES 43 – 50" captionStartId-0="21.[151,264,1095,1118]" captionStartId-1="22.[151,264,1570,1593]" captionTargetBox-0="[215,1378,208,1059]" captionTargetBox-1="[151,1436,467,1550]" captionTargetId-0="figure@21.[192,1395,193,1075]" captionTargetId-1="figure@22.[151,1436,467,1550]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURES 40 – 42. Genitalia of P. borneana. 40. Supra-anal plate and cerci (pilosity omitted). 41. Subgenital plate in dorsal view. 42. Genitalia in ventral view. Arrow indicates fold in dorsal lamina (scale bar: 1 mm)." captionText-1="FIGURES 43 – 50. Genitalia of various Hestiasulini. 43. A. phyllopus from Sabah (CS). 44. A. basinigra from Myanmar (SMNK). 45. C. nitida from Myanmar (SMNK). 46. C. nitida from Sumatra (SMNK). 47. C. sp. from Kerala (SMNK). 48. Ephestiasula amoena from Nepal (ES). 49. E. pictipes from India (CS). 50. H. gyldenstolpei from Thailand (ES) (scale bar: 1 mm)." httpUri-0="https://zenodo.org/record/829547/files/figure.png" httpUri-1="https://zenodo.org/record/829549/files/figure.png" pageId="26" pageNumber="269">42–44</figureCitation>
); head without tubercle (
<figureCitation id="134E2CB71811A16AFB0BFE46FAA1616A" box="[1271,1358,503,524]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[323,1273,195,1896]" captionTargetId="figure@6.[323,1273,195,1896]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Dorsal habitus of Hestiasulini. 1. Male of Hestiasula brunneriana (Sri Lanka) (CS). 2. Female of H. brunneriana (Sri Lanka) (CS). 3. Male of Astyliasula phyllopus from Sabah (CS). 4. Female of A. phyllopus from Brunei (CS). 5. Male of A. sarawaca from Brunei (SMNK). 6. Female of A. sarawaca from Kalimantan (RMNH). 7. Pseudohestiasula borneana, holotype (SMNK). 8. Male of Catestiasula moultoni from Sarawak (CS) (scale bar: 10 mm)." httpUri="https://zenodo.org/record/829533/files/figure.png" pageId="26" pageNumber="269">Figs 3–6</figureCitation>
,
<figureCitation id="134E2CB71811A16AFAA6FE46FA77616A" box="[1370,1432,503,524]" captionStart="FIGURES 33 – 36" captionStartId="17.[151,264,1946,1969]" captionTargetBox="[151,1436,193,1925]" captionTargetId="figure@17.[151,1436,193,1925]" captionTargetPageId="17" captionText="FIGURES 33 – 36. Holotype of A. sarawaca; Photos Katherine Child © OUMNH. 33. Dorsal view and labels. 34. Ventral view (scale bar: 10 mm). 35. Pronotum in dorsal view (scale bar: 1 mm). 36. Right femur in anterior view (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829543/files/figure.png" pageId="26" pageNumber="269">33–34</figureCitation>
,
<figureCitation id="134E2CB71811A16AFF1DFDA4FECF614C" box="[225,288,533,554]" captionStart-0="FIGURES 53 – 57" captionStart-1="FIGURES 58 – 59" captionStartId-0="24.[151,264,1839,1862]" captionStartId-1="25.[151,264,864,887]" captionTargetBox-0="[151,1436,375,1818]" captionTargetBox-1="[348,1232,193,843]" captionTargetId-0="figure@24.[151,1436,375,1818]" captionTargetId-1="figure@25.[340,1247,193,843]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURES 53 – 57. Life aspect of Bornean Astyliasula. 53. Male of A. sarawaca, Kubah NP, Sarawak, Malaysia; Photo © Fu- Sheng Huang. 54. Female of A. phyllopus in dorsal view, Mulu NP, Borneo, Sarawak, Malaysia; Photo © Paul Bertner. 55. Captive raised female of A. phyllopus in cryptic resting position; Photo © Daniel Döpfner. 56. A. phyllopus, displaying female, Mulu NP, Borneo, Sarawak, Malaysia; Photo © Paul Bertner. 57. A. phyllopus, displaying male, Mt. Trus Madi, Borneo, Sabah, Malaysia; Photo © Evgeny Shcherbakov." captionText-1="FIGURES 58 – 59. A. phyllopus. 58. Ootheca with hatching nymphs. 59. Displaying first instar nymph. Photos © Jürg Sommerhalder." httpUri-0="https://zenodo.org/record/829553/files/figure.png" httpUri-1="https://zenodo.org/record/829555/files/figure.png" pageId="26" pageNumber="269">53–59</figureCitation>
); if with tubercle (
<figureCitation id="134E2CB71811A16AFE29FDA4FDF4614C" box="[469,539,533,554]" captionStart="FIGURES 37 – 39" captionStartId="20.[151,264,1980,2003]" captionTargetBox="[243,1341,718,1941]" captionTargetId="figure@20.[216,1371,697,1960]" captionTargetPageId="20" captionText="FIGURES 37 – 39. P. borneana. 37. Head in anterior view. 38. Pronotum. 39. Left fore leg in anterior view (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829545/files/figure.png" pageId="26" pageNumber="269">Fig. 37</figureCitation>
) then ventral side of meso femora with a proximal lobe; alae of female (where known) subopaque (
<figureCitation id="134E2CB71811A16AFECFFD83FE666121" box="[307,393,562,583]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[323,1273,195,1896]" captionTargetId="figure@6.[323,1273,195,1896]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Dorsal habitus of Hestiasulini. 1. Male of Hestiasula brunneriana (Sri Lanka) (CS). 2. Female of H. brunneriana (Sri Lanka) (CS). 3. Male of Astyliasula phyllopus from Sabah (CS). 4. Female of A. phyllopus from Brunei (CS). 5. Male of A. sarawaca from Brunei (SMNK). 6. Female of A. sarawaca from Kalimantan (RMNH). 7. Pseudohestiasula borneana, holotype (SMNK). 8. Male of Catestiasula moultoni from Sarawak (CS) (scale bar: 10 mm)." httpUri="https://zenodo.org/record/829533/files/figure.png" pageId="26" pageNumber="269">Figs 4, 6</figureCitation>
,
<figureCitation id="134E2CB71811A16AFE6FFD83FE3E6121" box="[403,465,562,583]" captionStart="FIGURES 33 – 36" captionStartId="17.[151,264,1946,1969]" captionTargetBox="[151,1436,193,1925]" captionTargetId="figure@17.[151,1436,193,1925]" captionTargetPageId="17" captionText="FIGURES 33 – 36. Holotype of A. sarawaca; Photos Katherine Child © OUMNH. 33. Dorsal view and labels. 34. Ventral view (scale bar: 10 mm). 35. Pronotum in dorsal view (scale bar: 1 mm). 36. Right femur in anterior view (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829543/files/figure.png" pageId="26" pageNumber="269">33–34</figureCitation>
,
<figureCitation id="134E2CB71811A16AFE27FD83FE1A6121" box="[475,501,562,583]" captionStart="FIGURES 53 – 57" captionStartId="24.[151,264,1839,1862]" captionTargetBox="[151,1436,375,1818]" captionTargetId="figure@24.[151,1436,375,1818]" captionTargetPageId="24" captionText="FIGURES 53 – 57. Life aspect of Bornean Astyliasula. 53. Male of A. sarawaca, Kubah NP, Sarawak, Malaysia; Photo © Fu- Sheng Huang. 54. Female of A. phyllopus in dorsal view, Mulu NP, Borneo, Sarawak, Malaysia; Photo © Paul Bertner. 55. Captive raised female of A. phyllopus in cryptic resting position; Photo © Daniel Döpfner. 56. A. phyllopus, displaying female, Mulu NP, Borneo, Sarawak, Malaysia; Photo © Paul Bertner. 57. A. phyllopus, displaying male, Mt. Trus Madi, Borneo, Sabah, Malaysia; Photo © Evgeny Shcherbakov." httpUri="https://zenodo.org/record/829553/files/figure.png" pageId="26" pageNumber="269">56</figureCitation>
)........................................................................... 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308128971811A16AFF6BFDFEFA746191" pageId="26" pageNumber="269">
<paragraph id="8BCA30321811A16AFF6BFDFEFA7461F9" blockId="26.[151,1436,151,1287]" pageId="26" pageNumber="269">
<keyLead id="308493071811A16AFF6BFDFEFA7461F9" pageId="26" pageNumber="269">
8. Discoidal field of tegmen in both sexes iridescent, without false crossveins, hyaline or very slightly infumate, without dark markings (
<figureCitation id="134E2CB71811A16AFEBBFDDCFE6961E4" box="[327,390,621,642]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[323,1273,195,1896]" captionTargetId="figure@6.[323,1273,195,1896]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Dorsal habitus of Hestiasulini. 1. Male of Hestiasula brunneriana (Sri Lanka) (CS). 2. Female of H. brunneriana (Sri Lanka) (CS). 3. Male of Astyliasula phyllopus from Sabah (CS). 4. Female of A. phyllopus from Brunei (CS). 5. Male of A. sarawaca from Brunei (SMNK). 6. Female of A. sarawaca from Kalimantan (RMNH). 7. Pseudohestiasula borneana, holotype (SMNK). 8. Male of Catestiasula moultoni from Sarawak (CS) (scale bar: 10 mm)." httpUri="https://zenodo.org/record/829533/files/figure.png" pageId="26" pageNumber="269">Figs 8</figureCitation>
,
<figureCitation id="134E2CB71811A16AFE6EFDDCFE3E61E4" box="[402,465,621,642]" captionStart="FIGURES 51 – 52" captionStartId="23.[151,264,1634,1657]" captionTargetBox="[359,1219,469,1613]" captionTargetId="figure@23.[340,1247,469,1613]" captionTargetPageId="23" captionText="FIGURES 51 – 52. Life aspect of Catestiasula. 51. Male of C. moultoni, Mt. Trus Madi, Sabah, Malaysia; Photo © Alexey Yakovlev. 52. Male of unidentified Catestiasula specimen from Ulu Gombak NP, Selangor, Malaysia; Photo © Bernhard Jacobi." httpUri="https://zenodo.org/record/829551/files/figure.png" pageId="26" pageNumber="269">51–52</figureCitation>
); dorsal and ventral laminae of left phallomere fused along left margin (
<figureCitation id="134E2CB71811A16AFB70FDDCFB3861E4" box="[1164,1239,621,642]" captionStart="FIGURES 21 – 22" captionStartId="9.[151,264,1104,1127]" captionTargetBox="[180,1396,199,1063]" captionTargetId="figure@9.[159,1428,173,1083]" captionTargetPageId="9" captionText="FIGURES 21 – 22. Subgenital plate and genitalia of 21. H. brunneriana from India (ES) and 22. C. moultoni from Sabah (ES). Arrows indicate weakly sclerotized membrane at base of sclerotized left margin of ventral phallomere (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829537/files/figure.png" pageId="26" pageNumber="269">Figs 22</figureCitation>
,
<figureCitation id="134E2CB71811A16AFB1EFDDCFACD61E4" box="[1250,1314,621,642]" captionStart="FIGURES 43 – 50" captionStartId="22.[151,264,1570,1593]" captionTargetBox="[151,1436,467,1550]" captionTargetId="figure@22.[151,1436,467,1550]" captionTargetPageId="22" captionText="FIGURES 43 – 50. Genitalia of various Hestiasulini. 43. A. phyllopus from Sabah (CS). 44. A. basinigra from Myanmar (SMNK). 45. C. nitida from Myanmar (SMNK). 46. C. nitida from Sumatra (SMNK). 47. C. sp. from Kerala (SMNK). 48. Ephestiasula amoena from Nepal (ES). 49. E. pictipes from India (CS). 50. H. gyldenstolpei from Thailand (ES) (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829549/files/figure.png" pageId="26" pageNumber="269">45–47</figureCitation>
); apical process missing (Indo-Malayan) (genus
<taxonomicName id="4C754BB11811A16AFDBFFD38FC8A61F9" ID-CoL="8NP4F" authority="Giglio-Tos, 1915" authorityName="Giglio-Tos" authorityYear="1915" box="[579,869,649,671]" class="Insecta" family="Hymenopodidae" genus="Catestiasula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="26" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="B901EC201811A16AFDBFFD38FD5461F9" box="[579,699,649,671]" italics="true" pageId="26" pageNumber="269">Catestiasula</emphasis>
<bibRefCitation id="EFE44DC31811A16AFD3CFD3BFC8A61F9" author="Giglio-Tos" box="[704,869,650,671]" pageId="26" pageNumber="269" refString="Giglio-Tos, E. (1915) Mantidi esotici. Generi e specie nuove. Bollettino della Societa Entomologica Italiana, 46, 31 - 108." type="journal article" year="1915">Giglio-Tos, 1915</bibRefCitation>
</taxonomicName>
).............................................9
</keyLead>
</paragraph>
<paragraph id="8BCA30321811A16AFF6BFD16FA746191" blockId="26.[151,1436,151,1287]" pageId="26" pageNumber="269">
<keyLead id="308493071811A16AFF6BFD16FA746191" pageId="26" pageNumber="269">
- Discoidal field of tegmen in both sexes with false crossveins, slightly infumate to subopaque, with dark markings (
<figureCitation id="134E2CB71811A16AFAC4FD16FA6061DA" box="[1336,1423,679,700]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[323,1273,195,1896]" captionTargetId="figure@6.[323,1273,195,1896]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Dorsal habitus of Hestiasulini. 1. Male of Hestiasula brunneriana (Sri Lanka) (CS). 2. Female of H. brunneriana (Sri Lanka) (CS). 3. Male of Astyliasula phyllopus from Sabah (CS). 4. Female of A. phyllopus from Brunei (CS). 5. Male of A. sarawaca from Brunei (SMNK). 6. Female of A. sarawaca from Kalimantan (RMNH). 7. Pseudohestiasula borneana, holotype (SMNK). 8. Male of Catestiasula moultoni from Sarawak (CS) (scale bar: 10 mm)." httpUri="https://zenodo.org/record/829533/files/figure.png" pageId="26" pageNumber="269">Figs 1–2</figureCitation>
); dorsal and ventral laminae of left phallomere not fused along left margin; apical process broad and short (
<figureCitation id="134E2CB71811A16AFB02FD74FAA461BC" box="[1278,1355,709,730]" captionStart="FIGURES 21 – 22" captionStartId="9.[151,264,1104,1127]" captionTargetBox="[180,1396,199,1063]" captionTargetId="figure@9.[159,1428,173,1083]" captionTargetPageId="9" captionText="FIGURES 21 – 22. Subgenital plate and genitalia of 21. H. brunneriana from India (ES) and 22. C. moultoni from Sabah (ES). Arrows indicate weakly sclerotized membrane at base of sclerotized left margin of ventral phallomere (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829537/files/figure.png" pageId="26" pageNumber="269">Figs 21</figureCitation>
,
<figureCitation id="134E2CB71811A16AFAA4FD74FA7B61BC" box="[1368,1428,709,730]" captionStart="FIGURES 43 – 50" captionStartId="22.[151,264,1570,1593]" captionTargetBox="[151,1436,467,1550]" captionTargetId="figure@22.[151,1436,467,1550]" captionTargetPageId="22" captionText="FIGURES 43 – 50. Genitalia of various Hestiasulini. 43. A. phyllopus from Sabah (CS). 44. A. basinigra from Myanmar (SMNK). 45. C. nitida from Myanmar (SMNK). 46. C. nitida from Sumatra (SMNK). 47. C. sp. from Kerala (SMNK). 48. Ephestiasula amoena from Nepal (ES). 49. E. pictipes from India (CS). 50. H. gyldenstolpei from Thailand (ES) (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829549/files/figure.png" pageId="26" pageNumber="269">48–50</figureCitation>
) (Indo-Singhalese)....................................................................................10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308128971811A16AFF6BFD4EFA73600A" pageId="26" pageNumber="269">
<paragraph id="8BCA30321811A16AFF6BFD4EFA736054" blockId="26.[151,1436,151,1287]" pageId="26" pageNumber="269">
<keyLead id="308493071811A16AFF6BFD4EFA736054" pageId="26" pageNumber="269">
9. Anterior side of fore femora with only one small whitish spot in the claw groove, distal spots, if present, not clearly white (
<figureCitation id="134E2CB71811A16AFA8DFD4EFEF26054" captionStart="FIGURES 9 – 20" captionStartId="7.[151,264,1281,1304]" captionTargetBox="[206,1384,223,1244]" captionTargetId="figure@7.[182,1404,195,1257]" captionTargetPageId="7" captionText="FIGURES 9 – 20. Fore legs of Hestiasulini treated in this paper, anterior view. 9. A. phyllopus, male from the Malay Peninsula (ES). 10. A. phyllopus, male from Brunei, mirrored (CS). 11. A. phyllopus, female from Brunei (CS). 12. A. sarawaca, male from Brunei (CS). 13. P. borneana, holotype (SMNK). 14. H. brunneriana, male from Sri Lanka (CS). 15. H. brunneriana, female from Sri Lanka (CS). 16. A. sarawaca, female from Kalimantan; Photo C. J. Schwarz © RMNH (RMNH). 17. C. moultoni, male from Sarawak (CS). 18. C. moultoni, female from Brunei, mirrored (CS). 19. C. nitida, male from Myanmar (SMNK). 20. C. nitida, male from Sumatra (SMNK) (not to scale)." httpUri="https://zenodo.org/record/829535/files/figure.png" pageId="26" pageNumber="269">Figs 17–18</figureCitation>
).......................................................................
<taxonomicName id="4C754BB11811A16AFB7DFCADFA736054" authority="Giglio-Tos, 1915" authorityName="Giglio-Tos" authorityYear="1915" box="[1153,1436,796,818]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="26" pageNumber="269" phylum="Arthropoda" rank="species" species="moultoni">
<emphasis id="B901EC201811A16AFB7DFCADFB1D6054" box="[1153,1266,796,818]" italics="true" pageId="26" pageNumber="269">C. moultoni</emphasis>
<bibRefCitation id="EFE44DC31811A16AFB04FCACFA736054" author="Giglio-Tos" box="[1272,1436,797,818]" pageId="26" pageNumber="269" refString="Giglio-Tos, E. (1915) Mantidi esotici. Generi e specie nuove. Bollettino della Societa Entomologica Italiana, 46, 31 - 108." type="journal article" year="1915">Giglio-Tos, 1915</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCA30321811A16AFF6BFC8BFA73600A" blockId="26.[151,1436,151,1287]" pageId="26" pageNumber="269">
<keyLead id="308493071811A16AFF6BFC8BFA73600A" pageId="26" pageNumber="269">
- Anterior side of fore femora with two to three whitish spots along ventral margin (
<figureCitation id="134E2CB71811A16AFC09FC8BFB8A6029" box="[1013,1125,826,847]" captionStart="FIGURES 9 – 20" captionStartId="7.[151,264,1281,1304]" captionTargetBox="[206,1384,223,1244]" captionTargetId="figure@7.[182,1404,195,1257]" captionTargetPageId="7" captionText="FIGURES 9 – 20. Fore legs of Hestiasulini treated in this paper, anterior view. 9. A. phyllopus, male from the Malay Peninsula (ES). 10. A. phyllopus, male from Brunei, mirrored (CS). 11. A. phyllopus, female from Brunei (CS). 12. A. sarawaca, male from Brunei (CS). 13. P. borneana, holotype (SMNK). 14. H. brunneriana, male from Sri Lanka (CS). 15. H. brunneriana, female from Sri Lanka (CS). 16. A. sarawaca, female from Kalimantan; Photo C. J. Schwarz © RMNH (RMNH). 17. C. moultoni, male from Sarawak (CS). 18. C. moultoni, female from Brunei, mirrored (CS). 19. C. nitida, male from Myanmar (SMNK). 20. C. nitida, male from Sumatra (SMNK) (not to scale)." httpUri="https://zenodo.org/record/829535/files/figure.png" pageId="26" pageNumber="269">Figs 19–20</figureCitation>
)..............................................................................................
<taxonomicName id="4C754BB11811A16AFBDEFCE6FB99600B" box="[1058,1142,855,877]" class="Insecta" family="Hymenopodidae" genus="Pachymantis" kingdom="Animalia" order="Mantodea" pageId="26" pageNumber="269" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="B901EC201811A16AFBDEFCE6FB99600B" box="[1058,1142,855,877]" italics="true" pageId="26" pageNumber="269">C. nitida</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE44DC31811A16AFB7FFCE6FA7A600A" author="Brunner" box="[1155,1429,855,876]" pageId="26" pageNumber="269" refString="Brunner de Wattenwyl, K. (1893) Revision du systeme des Orthopteres et description des especes rapportees par M. Leonardo Fea de Birmanie. Annali del Museo Civico di Storia Naturale di Genova, Serie 2 a, 13 (33), 1 - 230." type="journal article" year="1893">Brunner d. Wattenwyl, 1893</bibRefCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308128971811A16AFF6BFCC4FA73677A" pageId="26" pageNumber="269">
<paragraph id="8BCA30321811A16AFF6BFCC4FA7360A2" blockId="26.[151,1436,151,1287]" pageId="26" pageNumber="269">
<keyLead id="308493071811A16AFF6BFCC4FA7360A2" pageId="26" pageNumber="269">
10. Process on vertex with pointed apex in both sexes, and with a pair of accessory conical tubercles at base at least in females; paramedian carinae on frontal shield well-developed; denticulation at postero-ventral margin of fore femur heavier; discoidal field of tegmen at most with numerous dark spots....................................
<taxonomicName id="4C754BB11811A16AFB8AFC1EFA7360A2" ID-CoL="8NRDP" authority="Giglio-Tos, 1915" authorityName="Giglio-Tos" authorityYear="1915" box="[1142,1436,943,965]" class="Insecta" family="Hymenopodidae" genus="Ephestiasula" kingdom="Animalia" order="Mantodea" pageId="26" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="B901EC201811A16AFB8AFC1EFB1D60A3" box="[1142,1266,943,965]" italics="true" pageId="26" pageNumber="269">Ephestiasula</emphasis>
<bibRefCitation id="EFE44DC31811A16AFB04FC1EFA7360A2" author="Giglio-Tos" box="[1272,1436,943,964]" pageId="26" pageNumber="269" refString="Giglio-Tos, E. (1915) Mantidi esotici. Generi e specie nuove. Bollettino della Societa Entomologica Italiana, 46, 31 - 108." type="journal article" year="1915">Giglio-Tos, 1915</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCA30321811A16AFF6BFC7CFA73677A" blockId="26.[151,1436,151,1287]" pageId="26" pageNumber="269">
<keyLead id="308493071811A16AFF6BFC7CFA73677A" pageId="26" pageNumber="269">
- Process on vertex with bifid apex in females (
<figureCitation id="134E2CB71811A16AFD6BFC7CFD3D6084" box="[663,722,973,994]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[323,1273,195,1896]" captionTargetId="figure@6.[323,1273,195,1896]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Dorsal habitus of Hestiasulini. 1. Male of Hestiasula brunneriana (Sri Lanka) (CS). 2. Female of H. brunneriana (Sri Lanka) (CS). 3. Male of Astyliasula phyllopus from Sabah (CS). 4. Female of A. phyllopus from Brunei (CS). 5. Male of A. sarawaca from Brunei (SMNK). 6. Female of A. sarawaca from Kalimantan (RMNH). 7. Pseudohestiasula borneana, holotype (SMNK). 8. Male of Catestiasula moultoni from Sarawak (CS) (scale bar: 10 mm)." httpUri="https://zenodo.org/record/829533/files/figure.png" pageId="26" pageNumber="269">Fig. 2</figureCitation>
), without accessory conical tubercles at base; paramedian carinae on frontal shield indistinct; denticulation at postero-ventral margin of fore femur fainter; discoidal field of tegmen with darker apex and with a large smoky patch in proximal half (may be indistinct in males)....................
<taxonomicName id="4C754BB11811A16AFB64FBB6FA73677A" ID-CoL="8NSHN" authority="Saussure, 1871" authorityName="Saussure" authorityYear="1871" box="[1176,1436,1031,1053]" class="Insecta" family="Hymenopodidae" genus="Hestiasula" kingdom="Animalia" order="Mantodea" pageId="26" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="B901EC201811A16AFB64FBB6FAEB677B" box="[1176,1284,1031,1053]" italics="true" pageId="26" pageNumber="269">Hestiasula</emphasis>
<bibRefCitation id="EFE44DC31811A16AFAF7FBB6FA73677A" author="Saussure" box="[1291,1436,1031,1052]" pageId="26" pageNumber="269" refString="Saussure, H. de (1871) Melanges orthopterologiques. Supplement au III me Fascicule. Mantides. Memoires de la Societe de Physique et d'Histoire naturelle de Geneve, 21 (1), 239 - 337, pl. 7." type="journal article" year="1871">Saussure, 1871</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308128971811A16AFF6BFB94FA7567F4" pageId="26" pageNumber="269">
<paragraph id="8BCA30321811A16AFF6BFB94FA73673E" blockId="26.[151,1436,151,1287]" pageId="26" pageNumber="269">
<keyLead id="308493071811A16AFF6BFB94FA73673E" pageId="26" pageNumber="269">
11. Vertex with a tubercle (
<figureCitation id="134E2CB71811A16AFE38FB94FDE2675C" box="[452,525,1061,1082]" captionStart="FIGURES 37 – 39" captionStartId="20.[151,264,1980,2003]" captionTargetBox="[243,1341,718,1941]" captionTargetId="figure@20.[216,1371,697,1960]" captionTargetPageId="20" captionText="FIGURES 37 – 39. P. borneana. 37. Head in anterior view. 38. Pronotum. 39. Left fore leg in anterior view (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829545/files/figure.png" pageId="26" pageNumber="269">Fig. 37</figureCitation>
); ventral side of meso femora with a proximal lobe. Fore coxae dichromatic (
<figureCitation id="134E2CB71811A16AFB03FB94FAA8675C" box="[1279,1351,1061,1082]" captionStart="FIGURES 9 – 20" captionStartId="7.[151,264,1281,1304]" captionTargetBox="[206,1384,223,1244]" captionTargetId="figure@7.[182,1404,195,1257]" captionTargetPageId="7" captionText="FIGURES 9 – 20. Fore legs of Hestiasulini treated in this paper, anterior view. 9. A. phyllopus, male from the Malay Peninsula (ES). 10. A. phyllopus, male from Brunei, mirrored (CS). 11. A. phyllopus, female from Brunei (CS). 12. A. sarawaca, male from Brunei (CS). 13. P. borneana, holotype (SMNK). 14. H. brunneriana, male from Sri Lanka (CS). 15. H. brunneriana, female from Sri Lanka (CS). 16. A. sarawaca, female from Kalimantan; Photo C. J. Schwarz © RMNH (RMNH). 17. C. moultoni, male from Sarawak (CS). 18. C. moultoni, female from Brunei, mirrored (CS). 19. C. nitida, male from Myanmar (SMNK). 20. C. nitida, male from Sumatra (SMNK) (not to scale)." httpUri="https://zenodo.org/record/829535/files/figure.png" pageId="26" pageNumber="269">Fig. 13</figureCitation>
); dilated part of fore femur less wide than non-dilated part (
<figureCitation id="134E2CB71811A16AFD39FBF3FCE16731" box="[709,782,1090,1111]" captionStart="FIGURES 9 – 20" captionStartId="7.[151,264,1281,1304]" captionTargetBox="[206,1384,223,1244]" captionTargetId="figure@7.[182,1404,195,1257]" captionTargetPageId="7" captionText="FIGURES 9 – 20. Fore legs of Hestiasulini treated in this paper, anterior view. 9. A. phyllopus, male from the Malay Peninsula (ES). 10. A. phyllopus, male from Brunei, mirrored (CS). 11. A. phyllopus, female from Brunei (CS). 12. A. sarawaca, male from Brunei (CS). 13. P. borneana, holotype (SMNK). 14. H. brunneriana, male from Sri Lanka (CS). 15. H. brunneriana, female from Sri Lanka (CS). 16. A. sarawaca, female from Kalimantan; Photo C. J. Schwarz © RMNH (RMNH). 17. C. moultoni, male from Sarawak (CS). 18. C. moultoni, female from Brunei, mirrored (CS). 19. C. nitida, male from Myanmar (SMNK). 20. C. nitida, male from Sumatra (SMNK) (not to scale)." httpUri="https://zenodo.org/record/829535/files/figure.png" pageId="26" pageNumber="269">Figs 13</figureCitation>
,
<figureCitation id="134E2CB71811A16AFCE5FBF3FCDD6731" box="[793,818,1090,1111]" captionStart="FIGURES 37 – 39" captionStartId="20.[151,264,1980,2003]" captionTargetBox="[243,1341,718,1941]" captionTargetId="figure@20.[216,1371,697,1960]" captionTargetPageId="20" captionText="FIGURES 37 – 39. P. borneana. 37. Head in anterior view. 38. Pronotum. 39. Left fore leg in anterior view (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829545/files/figure.png" pageId="26" pageNumber="269">39</figureCitation>
) (genus
<taxonomicName id="4C754BB11811A16AFC79FBF0FBC26731" box="[901,1069,1089,1111]" class="Insecta" family="Empusidae" genus="Pseudohestiasula" kingdom="Animalia" order="Mantodea" pageId="26" pageNumber="269" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B901EC201811A16AFC79FBF0FBC26731" box="[901,1069,1089,1111]" italics="true" pageId="26" pageNumber="269">Pseudohestiasula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A232515B1811A16AFBCFFBF3FB636731" box="[1075,1164,1090,1111]" pageId="26" pageNumber="269" rank="genus">gen. nov.</taxonomicNameLabel>
).....
<taxonomicName id="4C754BB11811A16AFB2AFBF3FAA66731" box="[1238,1353,1089,1111]" class="Insecta" family="Empusidae" genus="Pseudohestiasula" kingdom="Animalia" order="Mantodea" pageId="26" pageNumber="269" phylum="Arthropoda" rank="species" species="borneana" status="sp. nov.">
<emphasis id="B901EC201811A16AFB2AFBF3FAA66731" box="[1238,1353,1089,1111]" italics="true" pageId="26" pageNumber="269">P. borneana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A232515B1811A16AFAB3FBF3FA73673E" box="[1359,1436,1090,1112]" pageId="26" pageNumber="269" rank="species">
<emphasis id="B901EC201811A16AFAB3FBF3FA73673E" bold="true" box="[1359,1436,1090,1112]" pageId="26" pageNumber="269">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BCA30321811A16AFF6BFBEEFA7567F4" blockId="26.[151,1436,151,1287]" pageId="26" pageNumber="269">
<keyLead id="308493071811A16AFF6BFBEEFA7567F4" pageId="26" pageNumber="269">
- Vertex without tubercle (
<figureCitation id="134E2CB71811A16AFE2FFBEEFDC46712" box="[467,555,1119,1140]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[323,1273,195,1896]" captionTargetId="figure@6.[323,1273,195,1896]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Dorsal habitus of Hestiasulini. 1. Male of Hestiasula brunneriana (Sri Lanka) (CS). 2. Female of H. brunneriana (Sri Lanka) (CS). 3. Male of Astyliasula phyllopus from Sabah (CS). 4. Female of A. phyllopus from Brunei (CS). 5. Male of A. sarawaca from Brunei (SMNK). 6. Female of A. sarawaca from Kalimantan (RMNH). 7. Pseudohestiasula borneana, holotype (SMNK). 8. Male of Catestiasula moultoni from Sarawak (CS) (scale bar: 10 mm)." httpUri="https://zenodo.org/record/829533/files/figure.png" pageId="26" pageNumber="269">Figs 3–6</figureCitation>
,
<figureCitation id="134E2CB71811A16AFDCAFBEEFDBF6712" box="[566,592,1119,1140]" captionStart="FIGURES 33 – 36" captionStartId="17.[151,264,1946,1969]" captionTargetBox="[151,1436,193,1925]" captionTargetId="figure@17.[151,1436,193,1925]" captionTargetPageId="17" captionText="FIGURES 33 – 36. Holotype of A. sarawaca; Photos Katherine Child © OUMNH. 33. Dorsal view and labels. 34. Ventral view (scale bar: 10 mm). 35. Pronotum in dorsal view (scale bar: 1 mm). 36. Right femur in anterior view (scale bar: 1 mm)." httpUri="https://zenodo.org/record/829543/files/figure.png" pageId="26" pageNumber="269">33</figureCitation>
,
<figureCitation id="134E2CB71811A16AFDA0FBEEFD746712" box="[604,667,1119,1140]" captionStart-0="FIGURES 53 – 57" captionStart-1="FIGURES 58 – 59" captionStartId-0="24.[151,264,1839,1862]" captionStartId-1="25.[151,264,864,887]" captionTargetBox-0="[151,1436,375,1818]" captionTargetBox-1="[348,1232,193,843]" captionTargetId-0="figure@24.[151,1436,375,1818]" captionTargetId-1="figure@25.[340,1247,193,843]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURES 53 – 57. Life aspect of Bornean Astyliasula. 53. Male of A. sarawaca, Kubah NP, Sarawak, Malaysia; Photo © Fu- Sheng Huang. 54. Female of A. phyllopus in dorsal view, Mulu NP, Borneo, Sarawak, Malaysia; Photo © Paul Bertner. 55. Captive raised female of A. phyllopus in cryptic resting position; Photo © Daniel Döpfner. 56. A. phyllopus, displaying female, Mulu NP, Borneo, Sarawak, Malaysia; Photo © Paul Bertner. 57. A. phyllopus, displaying male, Mt. Trus Madi, Borneo, Sabah, Malaysia; Photo © Evgeny Shcherbakov." captionText-1="FIGURES 58 – 59. A. phyllopus. 58. Ootheca with hatching nymphs. 59. Displaying first instar nymph. Photos © Jürg Sommerhalder." httpUri-0="https://zenodo.org/record/829553/files/figure.png" httpUri-1="https://zenodo.org/record/829555/files/figure.png" pageId="26" pageNumber="269">53–59</figureCitation>
); meso femora without lobes; dilated part of fore femur much wider than nondilated part (
<figureCitation id="134E2CB71811A16AFEA6FBCCFE3067F4" box="[346,479,1149,1170]" captionStart="FIGURES 9 – 20" captionStartId="7.[151,264,1281,1304]" captionTargetBox="[206,1384,223,1244]" captionTargetId="figure@7.[182,1404,195,1257]" captionTargetPageId="7" captionText="FIGURES 9 – 20. Fore legs of Hestiasulini treated in this paper, anterior view. 9. A. phyllopus, male from the Malay Peninsula (ES). 10. A. phyllopus, male from Brunei, mirrored (CS). 11. A. phyllopus, female from Brunei (CS). 12. A. sarawaca, male from Brunei (CS). 13. P. borneana, holotype (SMNK). 14. H. brunneriana, male from Sri Lanka (CS). 15. H. brunneriana, female from Sri Lanka (CS). 16. A. sarawaca, female from Kalimantan; Photo C. J. Schwarz © RMNH (RMNH). 17. C. moultoni, male from Sarawak (CS). 18. C. moultoni, female from Brunei, mirrored (CS). 19. C. nitida, male from Myanmar (SMNK). 20. C. nitida, male from Sumatra (SMNK) (not to scale)." httpUri="https://zenodo.org/record/829535/files/figure.png" pageId="26" pageNumber="269">Figs 9–12, 16</figureCitation>
) (genus
<taxonomicName id="4C754BB11811A16AFDCEFBCDFD7267F4" ID-CoL="8NMSH" box="[562,669,1148,1170]" class="Insecta" family="Hymenopodidae" genus="Astyliasula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mantodea" pageId="26" pageNumber="269" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B901EC201811A16AFDCEFBCDFD7267F4" box="[562,669,1148,1170]" italics="true" pageId="26" pageNumber="269">Astyliasula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A232515B1811A16AFD5FFBCDFD1267F4" box="[675,765,1148,1170]" pageId="26" pageNumber="269" rank="genus">
<emphasis id="B901EC201811A16AFD5FFBCDFD1267F4" bold="true" box="[675,765,1148,1170]" pageId="26" pageNumber="269">gen. nov.</emphasis>
</taxonomicNameLabel>
).....................................................12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308128971811A16AFF6BFB2BFA736661" pageId="26" pageNumber="269">
<paragraph id="8BCA30321811A16AFF6BFB2BFA7467AA" blockId="26.[151,1436,151,1287]" pageId="26" pageNumber="269">
<keyLead id="308493071811A16AFF6BFB2BFA7467AA" pageId="26" pageNumber="269">
12. Both pairs of wings of males with an extensive dark pattern (
<figureCitation id="134E2CB71811A16AFCD4FB2BFC8967C9" box="[808,870,1178,1199]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[323,1273,195,1896]" captionTargetId="figure@6.[323,1273,195,1896]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Dorsal habitus of Hestiasulini. 1. Male of Hestiasula brunneriana (Sri Lanka) (CS). 2. Female of H. brunneriana (Sri Lanka) (CS). 3. Male of Astyliasula phyllopus from Sabah (CS). 4. Female of A. phyllopus from Brunei (CS). 5. Male of A. sarawaca from Brunei (SMNK). 6. Female of A. sarawaca from Kalimantan (RMNH). 7. Pseudohestiasula borneana, holotype (SMNK). 8. Male of Catestiasula moultoni from Sarawak (CS) (scale bar: 10 mm)." httpUri="https://zenodo.org/record/829533/files/figure.png" pageId="26" pageNumber="269">Figs 3</figureCitation>
,
<figureCitation id="134E2CB71811A16AFC8DFB2BFC6367C9" box="[881,908,1178,1199]" captionStart="FIGURES 53 – 57" captionStartId="24.[151,264,1839,1862]" captionTargetBox="[151,1436,375,1818]" captionTargetId="figure@24.[151,1436,375,1818]" captionTargetPageId="24" captionText="FIGURES 53 – 57. Life aspect of Bornean Astyliasula. 53. Male of A. sarawaca, Kubah NP, Sarawak, Malaysia; Photo © Fu- Sheng Huang. 54. Female of A. phyllopus in dorsal view, Mulu NP, Borneo, Sarawak, Malaysia; Photo © Paul Bertner. 55. Captive raised female of A. phyllopus in cryptic resting position; Photo © Daniel Döpfner. 56. A. phyllopus, displaying female, Mulu NP, Borneo, Sarawak, Malaysia; Photo © Paul Bertner. 57. A. phyllopus, displaying male, Mt. Trus Madi, Borneo, Sabah, Malaysia; Photo © Evgeny Shcherbakov." httpUri="https://zenodo.org/record/829553/files/figure.png" pageId="26" pageNumber="269">57</figureCitation>
); pronotal ratio of female&gt; 1.25...............................................................................................
<taxonomicName id="4C754BB11811A16AFB81FB06FA7467AA" authority="De Haan, 1842" authorityName="De Haan" authorityYear="1842" box="[1149,1435,1207,1229]" class="Insecta" family="Mantidae" genus="Mantis" kingdom="Animalia" order="Mantodea" pageId="26" pageNumber="269" phylum="Arthropoda" rank="species" species="phyllopus">
<emphasis id="B901EC201811A16AFB81FB06FB1A67AB" box="[1149,1269,1207,1229]" italics="true" pageId="26" pageNumber="269">A. phyllopus</emphasis>
(
<bibRefCitation id="EFE44DC31811A16AFAFEFB06FA7B67AA" author="De" box="[1282,1428,1207,1228]" pageId="26" pageNumber="269" refString="De Haan, W. (1842) Bijdragen tot de Kennis der Orthoptera Zoologie. Verhandelingen over de Natuurlijke Geschiedenis der Nederlandsche overzeesche Bezittingen door de Leden der Natuurkundige commisie in Indie et andere Schrijvers, 6, 45 - 248, 13 pls." type="journal article" year="1842">De Haan, 1842</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCA30321811A16AFF6BFB64FA736661" blockId="26.[151,1436,151,1287]" pageId="26" pageNumber="269">
<keyLead id="308493071811A16AFF6BFB64FA736661" pageId="26" pageNumber="269">
- Both pairs of wings of males hyaline with dark apices, tegmina with a transverse line of brownish spots, alae with darkened posterior corner (
<figureCitation id="134E2CB71811A16AFE7AFB43FE2C6661" box="[390,451,1266,1287]" captionStart="FIGURES 1 – 8" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[323,1273,195,1896]" captionTargetId="figure@6.[323,1273,195,1896]" captionTargetPageId="6" captionText="FIGURES 1 – 8. Dorsal habitus of Hestiasulini. 1. Male of Hestiasula brunneriana (Sri Lanka) (CS). 2. Female of H. brunneriana (Sri Lanka) (CS). 3. Male of Astyliasula phyllopus from Sabah (CS). 4. Female of A. phyllopus from Brunei (CS). 5. Male of A. sarawaca from Brunei (SMNK). 6. Female of A. sarawaca from Kalimantan (RMNH). 7. Pseudohestiasula borneana, holotype (SMNK). 8. Male of Catestiasula moultoni from Sarawak (CS) (scale bar: 10 mm)." httpUri="https://zenodo.org/record/829533/files/figure.png" pageId="26" pageNumber="269">Figs 5</figureCitation>
,
<figureCitation id="134E2CB71811A16AFE32FB43FE066661" box="[462,489,1266,1287]" captionStart="FIGURES 53 – 57" captionStartId="24.[151,264,1839,1862]" captionTargetBox="[151,1436,375,1818]" captionTargetId="figure@24.[151,1436,375,1818]" captionTargetPageId="24" captionText="FIGURES 53 – 57. Life aspect of Bornean Astyliasula. 53. Male of A. sarawaca, Kubah NP, Sarawak, Malaysia; Photo © Fu- Sheng Huang. 54. Female of A. phyllopus in dorsal view, Mulu NP, Borneo, Sarawak, Malaysia; Photo © Paul Bertner. 55. Captive raised female of A. phyllopus in cryptic resting position; Photo © Daniel Döpfner. 56. A. phyllopus, displaying female, Mulu NP, Borneo, Sarawak, Malaysia; Photo © Paul Bertner. 57. A. phyllopus, displaying male, Mt. Trus Madi, Borneo, Sabah, Malaysia; Photo © Evgeny Shcherbakov." httpUri="https://zenodo.org/record/829553/files/figure.png" pageId="26" pageNumber="269">53</figureCitation>
); pronotal ratio of female &lt;1.2.............................
<taxonomicName id="4C754BB11811A16AFB95FB40FA736661" authority="Westwood, 1889" authorityName="Westwood" authorityYear="1889" box="[1129,1436,1265,1287]" class="Insecta" family="Hymenopodidae" genus="Hestias" kingdom="Animalia" order="Mantodea" pageId="26" pageNumber="269" phylum="Arthropoda" rank="species" species="sarawaca">
<emphasis id="B901EC201811A16AFB95FB40FB0B6661" box="[1129,1252,1265,1287]" italics="true" pageId="26" pageNumber="269">A. sarawaca</emphasis>
(
<bibRefCitation id="EFE44DC31811A16AFB0EFB43FA7A6661" author="Westwood" box="[1266,1429,1266,1287]" pageId="26" pageNumber="269" refString="Westwood, J. O. (1889) Revisio insectorum familiae mantidarum, speciebus novis aut minus cognitis descriptis et delineates. Gurney &amp; Jackson, London, 54 + III pp., 14 pl." type="book" year="1889">Westwood, 1889</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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