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<document id="3CFD7E583945618F3CAF5B0C2D17C463" ID-DOI="10.1111/j.1096-3642.2009.00543.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5443367" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630455436479" checkinUser="felipe" docAuthor="Orrico, Victor G. D. &amp; Kury, Adriano B." docDate="2009" docId="03DA4503FFE4731F0A85FC654C72FDB3" docLanguage="en" docName="j.1096-3642.2009.00543.x.pdf" docOrigin="Zoological Journal of the Linnean Society 157 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00543.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Stygnicranaus alessandroi Orrico &amp; Kury 2009, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="480" masterDocId="FFE33D7BFFE073150A2DFFBC4972FB3A" masterDocTitle="A cladistic analysis of the Stygnicranainae Roewer, 1913 (Arachnida, Opiliones, Cranaidae) - where do longipalp cranaids belong?" masterLastPageNumber="494" masterPageNumber="470" pageNumber="474" updateTime="1699277118945" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="D35B521D0AE46394C791161184612BF7">A cladistic analysis of the Stygnicranainae Roewer, 1913 (Arachnida, Opiliones, Cranaidae) - where do longipalp cranaids belong?</mods:title>
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<mods:namePart id="E4D9C9B2DB7E8939C91D1CC7284540D3">Orrico, Victor G. D.</mods:namePart>
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<mods:namePart id="39A22F3FFA7648D33A06102706C759D5">Kury, Adriano B.</mods:namePart>
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<mods:date id="03E61AA9701B3FE5E6DC2DB5382BF24A">2009</mods:date>
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<treatment id="03DA4503FFE4731F0A85FC654C72FDB3" LSID="urn:lsid:plazi:treatment:03DA4503FFE4731F0A85FC654C72FDB3" httpUri="http://treatment.plazi.org/id/03DA4503FFE4731F0A85FC654C72FDB3" lastPageId="10" lastPageNumber="480" pageId="4" pageNumber="474">
<subSubSection id="C369A79EFFE473110A85FC654A75F8D5" box="[168,775,985,1007]" pageId="4" pageNumber="474" type="nomenclature">
<paragraph id="8BCCF415FFE473110A85FC654A75F8D5" blockId="4.[168,775,985,1007]" box="[168,775,985,1007]" pageId="4" pageNumber="474">
<heading id="D0844379FFE473110A85FC654A75F8D5" box="[168,775,985,1007]" centered="true" fontSize="9" level="2" pageId="4" pageNumber="474" reason="4">
<taxonomicName id="4C738F96FFE473110A85FC654B65F8D4" ID-CoL="7Z529" authority="Orrico &amp; Kury, 2009" authorityName="Orrico &amp; Kury" authorityYear="2009" box="[168,535,985,1006]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="474" phylum="Arthropoda" rank="species" species="alessandroi" status="sp. nov.">STYGNICRANAUS ALESSANDROI</taxonomicName>
<emphasis id="B9072807FFE47311080CFC614BF4F8D5" bold="true" box="[545,646,986,1007]" pageId="4" pageNumber="474">
<taxonomicNameLabel id="A234957CFFE47311080CFC614BF4F8D5" box="[545,646,986,1007]" pageId="4" pageNumber="474" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="1348E890FFE4731108B5FC664B8DF8D5" box="[664,767,985,1007]" captionStart-0="Figures 23" captionStart-1="Figures 45" captionStartId-0="5.[144,234,1348,1367]" captionStartId-1="6.[164,254,866,885]" captionTargetBox-0="[399,1167,196,1317]" captionTargetBox-1="[419,1187,196,837]" captionTargetId-0="figure-239@5.[399,1167,196,1317]" captionTargetId-1="figure-362@6.[419,1187,196,837]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figures 23. Stygnicranaus alessandroi sp. nov., male holotype (UNAL) from Colombia. Habitus. Fig. 2. Dorsal view. Fig. 3. Lateral view, spots omitted. Scale bars = 3 mm." captionText-1="Figures 45. Stygnicranaus alessandroi sp. nov., male holotype (UNAL) from Colombia. Penis, distal portion. Fig. 4. Dorsal view. Fig. 5. Lateral view. Arrows indicate capellum. Scale bars = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5443373" figureDoi-1="http://doi.org/10.5281/zenodo.5443375" httpUri-0="https://zenodo.org/record/5443373/files/figure.png" httpUri-1="https://zenodo.org/record/5443375/files/figure.png" pageId="4" pageNumber="474">FIGS 25</figureCitation>
)
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</paragraph>
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<subSubSection id="C369A79EFFE4731D0A89FBBD4D97FEB6" lastPageId="8" lastPageNumber="478" pageId="4" pageNumber="474" type="materials_examined">
<paragraph id="8BCCF415FFE473110A89FBBD4BF4FF69" blockId="4.[164,779,1024,1107]" pageId="4" pageNumber="474">
<emphasis id="B9072807FFE473110A89FBBD484FFF2F" box="[164,317,1024,1046]" italics="true" pageId="4" pageNumber="474">Type locality:</emphasis>
<materialsCitation id="3B1BFE48FFE473110B65FBBC4BF1FF69" collectionCode="WWF" collectorName="Dagua. El Salto. World Wildlife Fund &amp; Ecoregion NT" country="Colombia" location="Valle Del Cauca" pageId="4" pageNumber="474" specimenCount="1" stateProvince="Valle del Cauca">
<collectingCountry id="F364B485FFE473110B65FBBC48C4FF2C" box="[328,438,1024,1046]" name="Colombia" pageId="4" pageNumber="474">Colombia</collectingCountry>
,
<collectingRegion id="49B73AF7FFE473110BE8FBBC4BF7FF2F" box="[453,645,1024,1046]" country="Colombia" name="Valle del Cauca" pageId="4" pageNumber="474">Valle Del Cauca</collectingRegion>
,
<collectorName id="268691C3FFE4731108B9FBBC48A5FF0F" pageId="4" pageNumber="474">Dagua. El Salto. World Wildlife Fund</collectorName>
(
<collectionCode id="ED626CD0FFE473110BC8FBA34B5AFF0F" box="[485,552,1055,1077]" country="USA" lsid="urn:lsid:biocol.org:col:13930" name="Rob and Bessie Welder Wildlife Foundation" pageId="4" pageNumber="474" type="Herbarium">WWF</collectionCode>
)
<collectorName id="268691C3FFE47311081AFBA34BA6FF0E" box="[567,724,1054,1077]" pageId="4" pageNumber="474">Ecoregion NT</collectorName>
0145 (North-western Andean montane forests)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCCF415FFE473110A89FBC7498DFD06" blockId="4.[164,779,1146,1596]" pageId="4" pageNumber="474">
<emphasis id="B9072807FFE473110A89FBC74B25FFB5" box="[164,599,1146,1168]" italics="true" pageId="4" pageNumber="474">
Diagnosis:
<taxonomicName id="4C738F96FFE473110B08FBC74B25FFB5" authorityName="Orrico &amp; Kury" authorityYear="2009" box="[293,599,1146,1168]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="474" phylum="Arthropoda" rank="species" species="alessandroi">Stygnicranaus alessandroi</taxonomicName>
</emphasis>
differs from
<taxonomicName id="4C738F96FFE4731108DEFBC74873FF94" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="474" phylum="Arthropoda" rank="species" species="concolor">
<emphasis id="B9072807FFE4731108DEFBC74873FF94" italics="true" pageId="4" pageNumber="474">S. concolor</emphasis>
</taxonomicName>
by the presence of white granulation on the scutum, the straight, upward, stout spines in the ocularium (
<taxonomicName id="4C738F96FFE473110B00FB6B48DDFFD1" box="[301,431,1238,1260]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="474" phylum="Arthropoda" rank="species" species="concolor">
<emphasis id="B9072807FFE473110B00FB6B48DDFFD1" box="[301,431,1238,1260]" italics="true" pageId="4" pageNumber="474">S. concolor</emphasis>
</taxonomicName>
has curved spines), and for having two small spines in the frontal border of the scute (
<taxonomicName id="4C738F96FFE473110ADEFAA84800FE12" box="[243,370,1299,1321]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="474" phylum="Arthropoda" rank="species" species="concolor">
<emphasis id="B9072807FFE473110ADEFAA84800FE12" box="[243,370,1299,1321]" italics="true" pageId="4" pageNumber="474">S. concolor</emphasis>
</taxonomicName>
has none). It differs from both
<taxonomicName id="4C738F96FFE4731108DEFAA8487DFE7D" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="474" phylum="Arthropoda" rank="species" species="abnormis">
<emphasis id="B9072807FFE4731108DEFAA8487DFE7D" italics="true" pageId="4" pageNumber="474">S. abnormis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C738F96FFE473110B61FA8E48B8FE7D" box="[332,458,1330,1351]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="474" phylum="Arthropoda" rank="species" species="concolor">
<emphasis id="B9072807FFE473110B61FA8E48B8FE7D" box="[332,458,1330,1351]" italics="true" pageId="4" pageNumber="474">S. concolor</emphasis>
</taxonomicName>
in the absence of spines in area I, whereas
<taxonomicName id="4C738F96FFE473110B74FAED48A6FE5C" box="[345,468,1361,1382]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="474" phylum="Arthropoda" rank="species" species="concolor">
<emphasis id="B9072807FFE473110B74FAED48A6FE5C" box="[345,468,1361,1382]" italics="true" pageId="4" pageNumber="474">S. concolor</emphasis>
</taxonomicName>
and
<taxonomicName id="4C738F96FFE473110821FAED4BE7FE5C" box="[524,661,1361,1382]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="474" phylum="Arthropoda" rank="species" species="abnormis">
<emphasis id="B9072807FFE473110821FAED4BE7FE5C" box="[524,661,1361,1382]" italics="true" pageId="4" pageNumber="474">S. abnormis</emphasis>
</taxonomicName>
both have two small spines. It differs from
<taxonomicName id="4C738F96FFE47311083EFACC4BB3FEBE" authorityName="Orrico &amp; Kury" authorityYear="2009" box="[531,705,1391,1413]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="474" phylum="Arthropoda" rank="species" species="poncedeleoni">
<emphasis id="B9072807FFE47311083EFACC4BB3FEBE" box="[531,705,1391,1413]" italics="true" pageId="4" pageNumber="474">S. poncedeleoni</emphasis>
</taxonomicName>
in the presence of small spines instead of the pearly tubercles in coxa IV and by having two and not three teeth at each side of the anterolateral corners of the carapace. It also differs from all other members of Stygnicranainae by the numerous small dark spots on scutum.
</paragraph>
<paragraph id="8BCCF415FFE473130A89F9D94837FC55" blockId="4.[164,779,1636,1903]" lastBlockId="6.[164,779,1024,1903]" lastPageId="6" lastPageNumber="476" pageId="4" pageNumber="474">
<emphasis id="B9072807FFE473110A89F9D94884FD43" box="[164,502,1636,1658]" italics="true" pageId="4" pageNumber="474">
Description: Male
<typeStatus id="54C84AB7FFE473110BB5F9D84884FD43" box="[408,502,1636,1657]" pageId="4" pageNumber="474" type="holotype">holotype</typeStatus>
</emphasis>
: Dorsum. (
<figureCitation id="1348E890FFE4731108B7F9D84B8DFD40" box="[666,767,1636,1658]" captionStart="Figures 23" captionStartId="5.[144,234,1348,1367]" captionTargetBox="[399,1167,196,1317]" captionTargetId="figure-239@5.[399,1167,196,1317]" captionTargetPageId="5" captionText="Figures 23. Stygnicranaus alessandroi sp. nov., male holotype (UNAL) from Colombia. Habitus. Fig. 2. Dorsal view. Fig. 3. Lateral view, spots omitted. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5443373" httpUri="https://zenodo.org/record/5443373/files/figure.png" pageId="4" pageNumber="474">Figs 23</figureCitation>
). Dorsal scutum attenuate pyriform, with carapace only a little smaller than abdominal scutum. Posterior border of scutum straight. Cheliceral sockets shallow with a median tooth. Anterolateral corners of carapace with two teeth each side, one at pedipalpus, the other between it and leg I. Cheliceral sockets with a small pearly tubercle on each side. Posterior corners of scutum with a small tooth on each side. Ocularium elliptical, low, with a pair of stout, paramedian erect (lateral view), slightly divergent (frontal view) spines with blunt point. Surface of ocularium with seven pearly granules concentrated around the base of the spines. Carapace behind ocularium with a pair of rounded pearly setiferous granules. Median third of lateral area with a group of such granules. Posterior part of lateral area of scutum each side with an coalescent island of such tubercles. Grooves of scutum well marked, defining a mesotergum with three areas; original area IV completely fused with area
<collectionCode id="ED626CD0FFE473110F73FB824C0DFF69" box="[1374,1407,1086,1107]" pageId="4" pageNumber="474">III</collectionCode>
. A single shallow longitudinal medial groove dividing areas
<collectionCode id="ED626CD0FFE473110952FBC74AE1FFAA" box="[895,915,1147,1168]" pageId="4" pageNumber="474">II</collectionCode>
and
<collectionCode id="ED626CD0FFE4731109E6FBC74A9BFFAA" box="[971,1001,1147,1168]" pageId="4" pageNumber="474">III</collectionCode>
in two halves each. All areas unarmed except for area
<collectionCode id="ED626CD0FFE4731109D7FB254D6AFF94" box="[1018,1048,1177,1198]" pageId="4" pageNumber="474">III</collectionCode>
armed with a pair of very high acuminate spiniform processes. Scutal area
<collectionCode id="ED626CD0FFE473110F1DFB044C36FFF7" box="[1328,1348,1208,1229]" pageId="4" pageNumber="474">II</collectionCode>
projecting anteriorly into area I dividing it into left and right halves. Areas I and
<collectionCode id="ED626CD0FFE473110E39FB494D5AFE30" box="[1044,1064,1269,1290]" pageId="4" pageNumber="474">II</collectionCode>
each with a transverse row of five to six setiferous tubercles. Area
<collectionCode id="ED626CD0FFE473110E93FAA84DAEFE13" box="[1214,1244,1300,1321]" pageId="4" pageNumber="474">III</collectionCode>
with clusters of setiferous tubercles anterior to the large spines and reaching their basal part. In the place of area IV a pair of paramedian setiferous tubercles. Single ozopore opening laterally at tip of a dorsolateral low protuberance. Free tergites I
<collectionCode id="ED626CD0FFE473110EB9FA114DC3FEF8" box="[1172,1201,1453,1474]" pageId="4" pageNumber="474">III</collectionCode>
smooth, except for a pair of paramedian granules each. Corners of tergites I
<collectionCode id="ED626CD0FFE47311097EFA564A02FEC5" box="[851,880,1514,1535]" pageId="4" pageNumber="474">III</collectionCode>
each bearing an small acuminate tooth. Venter. Coxae I
<collectionCode id="ED626CD0FFE473110988F9B54AB0FD24" box="[933,962,1545,1566]" pageId="4" pageNumber="474">III</collectionCode>
smooth, except for a transverse row of tubercles each. Intercoxal bridges weak. Coxae I
<collectionCode id="ED626CD0FFE473110FA9F99B4CD3FD06" box="[1412,1441,1575,1596]" pageId="4" pageNumber="474">III</collectionCode>
subrectangular, transverse. Coxa IV the largest, equalling the other three taken together, shaped elongate pentagonal, divergent posteriorly. Coxa IV with one ventral subdistal small tubercle. Stigmatic area long, Y-shaped. Stigmata large, elongate elliptic, clearly apparent, not hidden by any bridges or tubercles, each occupying one third of the area. Maxillary lobes of coxae I
<collectionCode id="ED626CD0FFE4731109BAF8A04AD8FC0B" box="[919,938,1820,1841]" pageId="4" pageNumber="474">II</collectionCode>
clearly visible. Free sternites
<collectionCode id="ED626CD0FFE473110ED3F8A04C60FC0B" box="[1278,1298,1820,1841]" pageId="4" pageNumber="474">II</collectionCode>
and
<collectionCode id="ED626CD0FFE473110F64F8A04C15FC0B" box="[1353,1383,1820,1841]" pageId="4" pageNumber="474">III</collectionCode>
with their corners each bearing an small acuminate process. Chelicera. Basichelicerite long, with bulla attenuate and a medial belt of granules and a mesal subdistal small acuminate tubercle. Hand not swollen, with two frontal rows of setiferous tubercles. Pedipalpus. Much elongate and smooth, with tibia and tarsus forming a subchela. Coxa large, conical, smooth. Trochanter cylindrical with typical sub-basal constriction, and apical dorsomesal tubercles and one ventral setiferous tubercle. Femur longer than dorsal scutum, thicker at distal third, with dorsoapical tubercles and a ventral row of setiferous tubercles. Patella longer than abdominal scutum, thicker at distal third. Tibia as long as patella, cylindrical with four ectal (IiIi) and four mesal (IiIi) spines. Tarsus fusiform, slightly shorter than tibia, with four ectal (IiIi) and six mesal (IiiiIi) spines and two ventral rows of setiferous tubercles. Tarsal claw almost as long as tarsus. Legs. All segments slender and unarmed. Relative length I
<emphasis id="B9072807FFE57310091AFA564A37FEC5" box="[823,837,1514,1535]" italics="true" pageId="5" pageNumber="475">&lt;</emphasis>
<collectionCode id="ED626CD0FFE573100966FA564A1BFEC5" box="[843,873,1514,1535]" pageId="5" pageNumber="475">III</collectionCode>
<emphasis id="B9072807FFE57310095DFA564A0CFEC5" box="[880,894,1514,1535]" italics="true" pageId="5" pageNumber="475">&lt;</emphasis>
<collectionCode id="ED626CD0FFE5731009A8FA564AD3FEC5" box="[901,929,1514,1535]" pageId="5" pageNumber="475">IV</collectionCode>
<emphasis id="B9072807FFE57310098AFA564AC7FEC5" box="[935,949,1514,1535]" italics="true" pageId="5" pageNumber="475">&lt;</emphasis>
<collectionCode id="ED626CD0FFE573100991FA564AA1FEC5" box="[956,979,1514,1535]" pageId="5" pageNumber="475">II</collectionCode>
. Dorsobasal apophyses of coxae I
<collectionCode id="ED626CD0FFE573100F57FA564CFFFEC5" box="[1402,1421,1514,1535]" pageId="5" pageNumber="475">II</collectionCode>
close to the ozopore: coxa I anterior spiniform, posterior small, directed backwards; coxa
<collectionCode id="ED626CD0FFE573100EE1F99B4D92FD06" box="[1228,1248,1575,1596]" pageId="5" pageNumber="475">II</collectionCode>
anterior spiniform, posterior as a lobe joining coxa
<collectionCode id="ED626CD0FFE573100ED4F9FA4C68FD61" box="[1273,1306,1606,1627]" pageId="5" pageNumber="475">III</collectionCode>
. Coxa IV hidden under dorsal scutum in dorsal view, with prolateral dorsoapical spiniform apophysis and one pearly median prolateral dorsal granule. Trochanters IIV with basal ventral row of tubercles, interlocking with corresponding tubercles at respective coxae. Trochanter
<collectionCode id="ED626CD0FFE5731009A7F9414ADAFC28" box="[906,936,1789,1810]" pageId="5" pageNumber="475">III</collectionCode>
with a few ventral setiferous tubercles. Trochanter IV with a row of small retrolateral apophyses and a small subdistal prolateral spiniform apophysis. All femora straight. Femora
<collectionCode id="ED626CD0FFE573100EDEF8E54C7DFC54" box="[1267,1295,1881,1902]" pageId="5" pageNumber="475">III</collectionCode>
IV with a pair of small dorsoapical spurs. Femur IV with two rows of granules decreasing in size towards apex. Tarsi
<collectionCode id="ED626CD0FFE673130ADDFB82487EFF69" box="[240,268,1086,1107]" pageId="6" pageNumber="476">III</collectionCode>
IV with unpectinated double claws and well-developed tarsal process (pseudonychium). Colour. Body background dorsal Brilliant Orange Yellow with numerous Moderate Red small spots. All granules of scutum contrasting Pale Greenish Yellow surrounded by a circular islet of the same colour. In the clusters of lateral margins the islets are coalesced into a larger irregular spot. All appendages more or less the same colour as the scutum, but glossy, darker, and more saturated (background Strong Orange with Deep Orange reticule). Legs growing darker towards the apex. Free tergite
<collectionCode id="ED626CD0FFE673130AD6FA11486BFEF8" box="[251,281,1453,1474]" pageId="6" pageNumber="476">III</collectionCode>
completely Pale Greenish Yellow. Genitalia. (
<figureCitation id="1348E890FFE673130ACFFA774834FEDB" box="[226,326,1483,1505]" captionStart="Figures 45" captionStartId="6.[164,254,866,885]" captionTargetBox="[419,1187,196,837]" captionTargetId="figure-362@6.[419,1187,196,837]" captionTargetPageId="6" captionText="Figures 45. Stygnicranaus alessandroi sp. nov., male holotype (UNAL) from Colombia. Penis, distal portion. Fig. 4. Dorsal view. Fig. 5. Lateral view. Arrows indicate capellum. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5443375" httpUri="https://zenodo.org/record/5443375/files/figure.png" pageId="6" pageNumber="476">Figs 45</figureCitation>
). Ventral plate subrectangular with shallow lateromedial and anterior constrictions in dorsal view; setae of ventral plate divided in two groups, distal and medial, on each side. Distal group with four helicoidal setae, the innermost hooked and directed distally; the two apical ones directed laterodistally, and the remaining, transverse. Medial group with two substraight, subequal setae directed lateroposteriorly. Follis measuring about two-thirds of the ventral plate maximum length, extending from truncus tip to the base of the distal helicoidal setae with shallow folds in the basal portion. Stylus straight with capellum shovel-like, presenting lateral beaks.
</paragraph>
<caption id="DF0CA49DFFE573100ABDFAF84BA4FE4E" ID-DOI="http://doi.org/10.5281/zenodo.5443373" ID-Zenodo-Dep="5443373" httpUri="https://zenodo.org/record/5443373/files/figure.png" pageId="5" pageNumber="475" startId="5.[144,234,1348,1367]" targetBox="[399,1167,196,1317]" targetPageId="5">
<paragraph id="8BCCF415FFE573100ABDFAF84BA4FE4E" blockId="5.[144,1422,1347,1396]" pageId="5" pageNumber="475">
<emphasis id="B9072807FFE573100ABDFAF84BCCFE6D" bold="true" box="[144,702,1347,1367]" pageId="5" pageNumber="475">
Figures 23.
<taxonomicName id="4C738F96FFE573100B05FAF84B10FE6C" authority="Orrico &amp; Kury, 2009" authorityName="Orrico &amp; Kury" authorityYear="2009" box="[296,610,1347,1367]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="475" phylum="Arthropoda" rank="species" species="alessandroi" status="sp. nov.">
<emphasis id="B9072807FFE573100B05FAF84B10FE6C" bold="true" box="[296,610,1347,1367]" italics="true" pageId="5" pageNumber="475">Stygnicranaus alessandroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234957CFFE573100845FAF84BCCFE6D" box="[616,702,1348,1367]" pageId="5" pageNumber="475" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (UNAL) from Colombia. Habitus. Fig. 2. Dorsal view. Fig. 3. Lateral view, spots omitted. Scale bars = 3 mm.
</paragraph>
</caption>
<caption id="DF0CA49DFFE673130A89FCDE4D7BF8A9" ID-DOI="http://doi.org/10.5281/zenodo.5443375" ID-Zenodo-Dep="5443375" httpUri="https://zenodo.org/record/5443375/files/figure.png" pageId="6" pageNumber="476" startId="6.[164,254,866,885]" targetBox="[419,1187,196,837]" targetPageId="6">
<paragraph id="8BCCF415FFE673130A89FCDE4D7BF8A9" blockId="6.[164,1441,866,915]" pageId="6" pageNumber="476">
<emphasis id="B9072807FFE673130A89FCDE4BA6F84F" bold="true" box="[164,724,866,885]" pageId="6" pageNumber="476">
Figures 45.
<taxonomicName id="4C738F96FFE673130B11FCDE4B05F84F" authority="Orrico &amp; Kury, 2009" authorityName="Orrico &amp; Kury" authorityYear="2009" box="[316,631,866,885]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="476" phylum="Arthropoda" rank="species" species="alessandroi" status="sp. nov.">
<emphasis id="B9072807FFE673130B11FCDE4B05F84F" bold="true" box="[316,631,866,885]" italics="true" pageId="6" pageNumber="476">Stygnicranaus alessandroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234957CFFE673130850FCDE4BA6F84F" box="[637,724,866,885]" pageId="6" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (UNAL) from Colombia. Penis, distal portion. Fig. 4. Dorsal view. Fig. 5. Lateral view. Arrows indicate capellum. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BCCF415FFE673130917FBBC4D66FF2C" blockId="6.[826,1044,1024,1046]" box="[826,1044,1024,1046]" pageId="6" pageNumber="476">
<emphasis id="B9072807FFE673130917FBBC4AE6FF2F" box="[826,916,1024,1045]" italics="true" pageId="6" pageNumber="476">Female:</emphasis>
Unknown.
</paragraph>
<paragraph id="8BCCF415FFE673130917FB834DEDFFA8" blockId="6.[826,1441,1087,1170]" pageId="6" pageNumber="476">
<emphasis id="B9072807FFE673130917FB834A97FF6E" box="[826,997,1087,1108]" italics="true" pageId="6" pageNumber="476">Type material:</emphasis>
<specimenCount id="9D753F9CFFE6731309D9FBFE4D7FFF6E" box="[1012,1037,1090,1108]" pageId="6" pageNumber="476" type="male"></specimenCount>
<typeStatus id="54C84AB7FFE673130E31FB834D0DFF6F" box="[1052,1151,1087,1109]" pageId="6" pageNumber="476" type="holotype">holotype</typeStatus>
,
<materialsCitation id="3B1BFE48FFE673130EBFFB834DEDFFA8" collectingDate="2006-08" collectionCode="UNAL" collectorName="Dagua. El Salto. &amp; Giupponi, A." country="Colombia" location="Valle Del Cauca" pageId="6" pageNumber="476" specimenCount="1" specimenCount-juvenile="1" stateProvince="Valle del Cauca" typeStatus="paratype">
<specimenCount id="9D753F9CFFE673130EBFFB834C55FF6F" box="[1170,1319,1087,1109]" pageId="6" pageNumber="476" type="juvenile">one juvenile</specimenCount>
.
<typeStatus id="54C84AB7FFE673130F14FB834CD3FF6E" box="[1337,1441,1087,1108]" pageId="6" pageNumber="476" type="paratype">Paratype</typeStatus>
(
<collectionCode id="ED626CD0FFE67313096CFBE14AFFFF49" box="[833,909,1117,1139]" pageId="6" pageNumber="476">UNAL</collectionCode>
)
<collectingCountry id="F364B485FFE6731309B3FBE14D7EFF49" box="[926,1036,1117,1139]" name="Colombia" pageId="6" pageNumber="476">Colombia</collectingCountry>
,
<collectingRegion id="49B73AF7FFE673130E34FBE14DA7FF48" box="[1049,1237,1117,1139]" country="Colombia" name="Valle del Cauca" pageId="6" pageNumber="476">Valle Del Cauca</collectingRegion>
,
<collectorName id="268691C3FFE673130ECFFBE24CD3FF49" box="[1250,1441,1117,1139]" pageId="6" pageNumber="476">Dagua. El Salto.</collectorName>
<date id="FFCDD2D5FFE673130917FBC04ABDFFA8" box="[826,975,1148,1170]" pageId="6" pageNumber="476" value="2006-08">
<collectingDate id="EF892B3DFFE673130917FBC04ABDFFA8" box="[826,975,1148,1170]" pageId="6" pageNumber="476" value="2006-08">August 2006</collectingDate>
</date>
,
<collectorName id="268691C3FFE6731309F0FBC04D1DFFAB" box="[989,1135,1148,1170]" pageId="6" pageNumber="476">Giupponi, A.</collectorName>
col.
</materialsCitation>
</paragraph>
<paragraph id="8BCCF415FFE673130917FB074D2FFE34" blockId="6.[826,1441,1211,1294]" pageId="6" pageNumber="476">
<emphasis id="B9072807FFE673130917FB074ACAFFEA" box="[826,952,1211,1232]" italics="true" pageId="6" pageNumber="476">Etymology:</emphasis>
The species name honours the collector of the
<typeStatus id="54C84AB7FFE673130947FB664AE9FFD5" box="[874,923,1242,1263]" pageId="6" pageNumber="476">type</typeStatus>
series, our colleague and friend Alessandro Ponce de Leão Giupponi.
</paragraph>
<paragraph id="8BCCF415FFE673130958FAF84C15FE63" blockId="6.[885,1383,1348,1402]" box="[885,1383,1348,1369]" pageId="6" pageNumber="476">
<taxonomicName id="4C738F96FFE673130958FAF84D85FE63" authority="Orrico &amp; Kury, 2009" authorityName="Orrico &amp; Kury" authorityYear="2009" box="[885,1271,1348,1369]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="476" phylum="Arthropoda" rank="species" species="poncedeleoni" status="sp. nov.">STYGNICRANAUS PONCEDELEONI</taxonomicName>
<emphasis id="B9072807FFE673130F2FFAF44C15FE63" bold="true" box="[1282,1383,1348,1369]" pageId="6" pageNumber="476">
<taxonomicNameLabel id="A234957CFFE673130F2FFAF44C15FE63" box="[1282,1383,1348,1369]" pageId="6" pageNumber="476" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BCCF415FFE673130E01FAD84DDDFE40" blockId="6.[885,1383,1348,1402]" box="[1068,1199,1380,1402]" pageId="6" pageNumber="476">
(
<figureCitation id="1348E890FFE673130E19FAD84DD5FE40" box="[1076,1191,1380,1402]" captionStart-0="Figures 67" captionStart-1="Figures 89" captionStart-2="Figures 1011" captionStartId-0="7.[144,234,1318,1337]" captionStartId-1="8.[164,254,698,717]" captionStartId-2="9.[144,234,807,826]" captionTargetBox-0="[399,1167,195,1289]" captionTargetBox-1="[164,782,196,669]" captionTargetBox-2="[399,1167,196,777]" captionTargetId-0="figure-278@7.[399,1167,195,1289]" captionTargetId-1="figure-657@8.[164,782,196,669]" captionTargetId-2="figure-507@9.[399,1167,196,777]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="Figures 67. Stygnicranaus poncedeleoni sp. nov., male holotype (UNAL) from Colombia. Habitus. Fig. 6. Dorsal view. Fig. 7. Lateral view. Scale bars = 3 mm." captionText-1="Figures 89. Stygnicranaus poncedeleoni sp. nov., male holotype (UNAL) from Colombia. Left pedipalpus. Fig. 8. Mesal view. Fig. 9. Ectal view. Scale bars = 1 mm." captionText-2="Figures 1011. Stygnicranaus poncedeleoni sp. nov., male holotype (UNAL) from Colombia. Penis, distal portion. Fig. 10. Dorsal view. Fig. 11. Lateral view. Arrows indicates capellum. Scale bars = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5443377" figureDoi-1="http://doi.org/10.5281/zenodo.5443379" figureDoi-2="http://doi.org/10.5281/zenodo.5443381" httpUri-0="https://zenodo.org/record/5443377/files/figure.png" httpUri-1="https://zenodo.org/record/5443379/files/figure.png" httpUri-2="https://zenodo.org/record/5443381/files/figure.png" pageId="6" pageNumber="476">FIGS 611</figureCitation>
)
</paragraph>
<paragraph id="8BCCF415FFE673130917FA374D17FEE4" blockId="6.[826,1441,1419,1502]" pageId="6" pageNumber="476">
<emphasis id="B9072807FFE673130917FA374AA1FE9A" box="[826,979,1419,1440]" italics="true" pageId="6" pageNumber="476">Type locality:</emphasis>
<collectingCountry id="F364B485FFE6731309F3FA374D3EFE9B" box="[990,1100,1419,1441]" name="Colombia" pageId="6" pageNumber="476">Colombia</collectingCountry>
,
<collectingRegion id="49B73AF7FFE673130E76FA374C6EFE9A" box="[1115,1308,1419,1441]" country="Colombia" name="Valle del Cauca" pageId="6" pageNumber="476">Valle Del Cauca</collectingRegion>
, Dagua. El Salto. WWF Ecoregion NT0145 (North-western Andean montane forests).
</paragraph>
<paragraph id="8BCCF415FFE673130917F9BB4D74FD60" blockId="6.[826,1441,1543,1626]" pageId="6" pageNumber="476">
<emphasis id="B9072807FFE673130917F9BB4AC3FD26" box="[826,945,1543,1564]" italics="true" pageId="6" pageNumber="476">Diagnosis:</emphasis>
It is easily distinguished from all other species by the presence of pearly tubercles in coxa IV instead of spines.
</paragraph>
<paragraph id="8BCCF415FFE6731D0917F93F4C6EFF0F" blockId="6.[826,1441,1667,1903]" lastBlockId="8.[826,1442,198,1077]" lastPageId="8" lastPageNumber="478" pageId="6" pageNumber="476">
<emphasis id="B9072807FFE673130917F93F4D15FDA2" box="[826,1127,1667,1688]" italics="true" pageId="6" pageNumber="476">
Description: Male
<typeStatus id="54C84AB7FFE673130E24F93F4D15FDA2" box="[1033,1127,1667,1688]" pageId="6" pageNumber="476" type="holotype">holotype</typeStatus>
</emphasis>
: Dorsum (
<figureCitation id="1348E890FFE673130EF3F93F4C31FDA3" box="[1246,1347,1667,1689]" captionStart="Figures 67" captionStartId="7.[144,234,1318,1337]" captionTargetBox="[399,1167,195,1289]" captionTargetId="figure-278@7.[399,1167,195,1289]" captionTargetPageId="7" captionText="Figures 67. Stygnicranaus poncedeleoni sp. nov., male holotype (UNAL) from Colombia. Habitus. Fig. 6. Dorsal view. Fig. 7. Lateral view. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5443377" httpUri="https://zenodo.org/record/5443377/files/figure.png" pageId="6" pageNumber="476">Figs 67</figureCitation>
). Dorsal scutum attenuate pyriform, with carapace smaller than abdominal scutum. Posterior border of scutum straight. Cheliceral sockets very shallow with a median tooth. Anterolateral corners of carapace with three teeth on each side, two by the pedipalpus, ectal larger, the third at leg I. Frontal hump with one tubercle each side. Posterior corners of scutum with two teeth on each side. Ocularium elliptical, low, with a pair of stout, paramedian erect (lateral view), slightly divergent (frontal view) spines with blunt point. Surface of ocularium with 16 coarse scattered granules. Posterior part of lateral area of scutum each side with an island of such granules. Posterior border of dorsal scutum with two paramedian pearly setiferous tubercles. Carapace behind ocularium with a pair of rounded setiferous tubercles. Anterior half of lateral area with a group of such tubercles. Lateral border of scutum with a irregular row of small scarce tubercles. Grooves of scutum well marked, defining a mesotergum with three areas; original area IV completely fused with area
<collectionCode id="ED626CD0FFE773120B8AF8A048BAFC0B" box="[423,456,1820,1841]" pageId="7" pageNumber="477">III</collectionCode>
. A single shallow longitudinal medial groove dividing areas
<collectionCode id="ED626CD0FFE77312081DF8874B36FC6A" box="[560,580,1851,1872]" pageId="7" pageNumber="477">II</collectionCode>
and
<collectionCode id="ED626CD0FFE7731208A9F8874BD0FC6A" box="[644,674,1851,1872]" pageId="7" pageNumber="477">III</collectionCode>
in two halves each. All areas unarmed except for area
<collectionCode id="ED626CD0FFE7731208F5F8E54B84FC54" box="[728,758,1881,1902]" pageId="7" pageNumber="477">III</collectionCode>
armed with a pair of very high acuminate spiniform processes. Scutal area
<collectionCode id="ED626CD0FFE773120E13FA114D20FEF8" box="[1086,1106,1453,1474]" pageId="7" pageNumber="477">II</collectionCode>
projecting anteriorly into area I, dividing it in left and right halves. Areas I and
<collectionCode id="ED626CD0FFE77312090BFA564A48FEC5" box="[806,826,1514,1535]" pageId="7" pageNumber="477">II</collectionCode>
each with a transverse row of six to eight setiferous tubercles. Area
<collectionCode id="ED626CD0FFE7731209D1F9B54D68FD24" box="[1020,1050,1545,1566]" pageId="7" pageNumber="477">III</collectionCode>
with clusters of setiferous tubercles anterior to the large spines and reaching their basal part. Single V-shaped ozopore opening laterally at the tip of a dorsolateral low protuberance. Free tergites I
<collectionCode id="ED626CD0FFE7731209F2F93F4A80FDA2" box="[991,1010,1667,1688]" pageId="7" pageNumber="477">II</collectionCode>
smooth, except for a pair of paramedian pearly granules each, at the same level as the tubercles in the posterior border of the scutum. Free tergite
<collectionCode id="ED626CD0FFE773120951F9634AE8FDCE" box="[892,922,1759,1780]" pageId="7" pageNumber="477">III</collectionCode>
bearing a paramedian pair of acuminated spiniform tubercules, a lot larger than those in tergites I
<collectionCode id="ED626CD0FFE77312095BF8A04AF9FC0B" box="[886,907,1820,1841]" pageId="7" pageNumber="477">II</collectionCode>
. Their corners each bearing an acuminate small process tooth. Venter. Coxae I
<collectionCode id="ED626CD0FFE773120F29F8874C53FC6A" box="[1284,1313,1851,1872]" pageId="7" pageNumber="477">III</collectionCode>
smooth, except for a transverse row of tubercles in coxa I, two rows in
<collectionCode id="ED626CD0FFE8731D0B2CFCF54867F864" box="[257,277,841,862]" pageId="8" pageNumber="478">II</collectionCode>
<collectionCode id="ED626CD0FFE8731D0B08FCF54836F864" box="[293,324,841,862]" pageId="8" pageNumber="478">III</collectionCode>
. Three irregular rows in coxa IV. Intercoxal bridges weak. Coxae I
<collectionCode id="ED626CD0FFE8731D0BC7FCDB4B75F846" box="[490,519,871,892]" pageId="8" pageNumber="478">III</collectionCode>
subrectangular, transverse. Coxa IV the largest, almost equalling the other three taken together, shaped lanceolate and pentagonal, divergent posteriorly. Stigmatic area long, Y-shaped. Stigmata large, elongate elliptic, clearly apparent, not hidden by any bridges or tubercles, each occupying one third of the area. Maxillary lobes of coxae
<collectionCode id="ED626CD0FFE8731D0AC5FB82498EFF69" box="[232,252,1086,1107]" pageId="8" pageNumber="478">II</collectionCode>
clearly visible. Chelicera. Basichelicerite long, with bulla attenuate, with a medial belt of granules and a mesal subdistal small acuminate tubercle. Hand not swollen, with a frontal medial hooked spine. Pedipalpus (
<figureCitation id="1348E890FFE8731D0AD0FB044812FFF4" box="[253,352,1208,1230]" captionStart="Figures 89" captionStartId="8.[164,254,698,717]" captionTargetBox="[164,782,196,669]" captionTargetId="figure-657@8.[164,782,196,669]" captionTargetPageId="8" captionText="Figures 89. Stygnicranaus poncedeleoni sp. nov., male holotype (UNAL) from Colombia. Left pedipalpus. Fig. 8. Mesal view. Fig. 9. Ectal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5443379" httpUri="https://zenodo.org/record/5443379/files/figure.png" pageId="8" pageNumber="478">Figs 89</figureCitation>
). Very elongate and smooth, with tibia and tarsus forming a subchela. Coxa large, conical, with a ventral medial row of three setiferous tubercles. Trochanter cylindrical with typical sub-basal constriction, with apical dorsomesal tubercles and a ventral pair of setiferous tubercles, mesal larger than ectal. Femur longer than dorsal scutum, thicker at distal third, with dorsoapical tubercles and a ventral row of setiferous tubercles. Patella longer than abdominal scutum, thicker at distal third. Tibia as long as patella, cylindrical with four ectal (IiIi) and four mesal (IiIi) spines. Tarsus fusiform, slightly shorter than tibia, with 11 ectal (iiIiiiiiIii) and ten mesal (iiIiiiiIii) spines. Tarsal claw almost as long as tarsus. Legs. All segments slender and unarmed. Relative length I
<emphasis id="B9072807FFE8731D0A98F93F49B1FDA2" box="[181,195,1667,1688]" italics="true" pageId="8" pageNumber="478">&lt;</emphasis>
<collectionCode id="ED626CD0FFE8731D0AE4F93F4995FDA2" box="[201,231,1667,1688]" pageId="8" pageNumber="478">III</collectionCode>
<emphasis id="B9072807FFE8731D0AC3F93F498EFDA2" box="[238,252,1667,1688]" italics="true" pageId="8" pageNumber="478">&lt;</emphasis>
<collectionCode id="ED626CD0FFE8731D0B2FF93F486CFDA2" box="[258,286,1667,1688]" pageId="8" pageNumber="478">IV</collectionCode>
<emphasis id="B9072807FFE8731D0B08F93F4841FDA2" box="[293,307,1667,1688]" italics="true" pageId="8" pageNumber="478">&lt;</emphasis>
<collectionCode id="ED626CD0FFE8731D0B14F93F4822FDA2" box="[313,336,1667,1688]" pageId="8" pageNumber="478">II</collectionCode>
. Dorsobasal apophyses of coxae I
<collectionCode id="ED626CD0FFE8731D08DAF93F4A78FDA2" box="[759,778,1667,1688]" pageId="8" pageNumber="478">II</collectionCode>
close to the ozopore: coxa I anterior spiniform, posterior small, directed backwards; coxa
<collectionCode id="ED626CD0FFE8731D0867F97C4B2CFDEF" box="[586,606,1728,1749]" pageId="8" pageNumber="478">II</collectionCode>
anterior spiniform, posterior as a lobe joining coxa
<collectionCode id="ED626CD0FFE8731D085AF9634BEAFDCE" box="[631,664,1759,1780]" pageId="8" pageNumber="478">III</collectionCode>
. Coxa IV hidden under dorsal scutum in dorsal view, with prolateral dorsoapical spiniform apophysis and dorsal granules. Trochanters IIV with ventrobasal tubercles interlocking with respective coxae. Trochanter
<collectionCode id="ED626CD0FFE8731D08E6F8E54BACFC54" box="[715,734,1881,1902]" pageId="8" pageNumber="478">II</collectionCode>
<collectionCode id="ED626CD0FFE8731D08C0F8E54A78FC54" box="[749,778,1881,1902]" pageId="8" pageNumber="478">III</collectionCode>
with a few prolateral and retrolateral setiferous tubercles. Trochanter IV with a row of small retrolateral apophyses and small subdistal prolateral and retrolateral spiniform apophyses. All femora straight. Femora
<collectionCode id="ED626CD0FFE8731D09B4FEFD4AC7FA6C" box="[921,949,321,342]" pageId="8" pageNumber="478">III</collectionCode>
IV with a pair of small dorsoapical spurs, retrolateral larger than prolateral. Femur IV with two rows of granules decreasing in size towards apex. Tarsi
<collectionCode id="ED626CD0FFE8731D0917FE204A24FA8B" box="[826,854,412,433]" pageId="8" pageNumber="478">III</collectionCode>
IV with unpectinated double claws and welldeveloped tarsal process (pseudonychium). Colour. Body and appendage background colour Deep Reddish Orange with strong mottling and spotting in Strong Reddish Orange, especially at scutal grooves. Chelicera by contrast has conspicuous darker reticulation. All granules are Pale Greenish Yellow as well as the inverted Y (from behind the ocularium to lateral of area
<collectionCode id="ED626CD0FFE8731D0956FD2D4AFDF99C" box="[891,911,657,678]" pageId="8" pageNumber="478">II</collectionCode>
) formed by the spots of scutum. Genitalia (
<figureCitation id="1348E890FFE8731D096FFD134AB3F9FC" box="[834,961,687,710]" captionStart="Figures 1011" captionStartId="9.[144,234,807,826]" captionTargetBox="[399,1167,196,777]" captionTargetId="figure-507@9.[399,1167,196,777]" captionTargetPageId="9" captionText="Figures 1011. Stygnicranaus poncedeleoni sp. nov., male holotype (UNAL) from Colombia. Penis, distal portion. Fig. 10. Dorsal view. Fig. 11. Lateral view. Arrows indicates capellum. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5443381" httpUri="https://zenodo.org/record/5443381/files/figure.png" pageId="8" pageNumber="478">Figs 1011</figureCitation>
). Ventral plate subrectangular with shallow lateromedial and marked anterior constrictions in dorsal view; setae of ventral plate divided into two groups, distal and medial, on each side. Distal group with four helicoidal setae, the innermost hooked and directed distally; two apical oblique, directed laterodistally, and the remaining one somewhat transverse. Medial group with two substraight, subequal setae directed lateroposteriorly. Follis measuring about two-thirds of the ventral plate maximum length, extending from truncus tip to the base of the distal helicoidal setae with shallow folds in the basal portion. Stylus straight with capellum shovel-like.
</paragraph>
<caption id="DF0CA49DFFE773120ABDFA9A4B07FE6D" ID-DOI="http://doi.org/10.5281/zenodo.5443377" ID-Zenodo-Dep="5443377" httpUri="https://zenodo.org/record/5443377/files/figure.png" pageId="7" pageNumber="477" startId="7.[144,234,1318,1337]" targetBox="[399,1167,195,1289]" targetPageId="7">
<paragraph id="8BCCF415FFE773120ABDFA9A4B07FE6D" blockId="7.[144,1421,1318,1367]" pageId="7" pageNumber="477">
<emphasis id="B9072807FFE773120ABDFA9A4BABFE00" bold="true" box="[144,729,1318,1338]" pageId="7" pageNumber="477">
Figures 67.
<taxonomicName id="4C738F96FFE773120B05FA9A4B07FE03" authority="Orrico &amp; Kury, 2009" authorityName="Orrico &amp; Kury" authorityYear="2009" box="[296,629,1318,1337]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="477" phylum="Arthropoda" rank="species" species="poncedeleoni" status="sp. nov.">
<emphasis id="B9072807FFE773120B05FA9A4B07FE03" bold="true" box="[296,629,1318,1337]" italics="true" pageId="7" pageNumber="477">Stygnicranaus poncedeleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234957CFFE773120852FA9B4BABFE00" box="[639,729,1319,1338]" pageId="7" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (UNAL) from Colombia. Habitus. Fig. 6. Dorsal view. Fig. 7. Lateral view. Scale bars = 3 mm.
</paragraph>
</caption>
<caption id="DF0CA49DFFE8731D0A89FD064A77F833" ID-DOI="http://doi.org/10.5281/zenodo.5443379" ID-Zenodo-Dep="5443379" httpUri="https://zenodo.org/record/5443379/files/figure.png" pageId="8" pageNumber="478" startId="8.[164,254,698,717]" targetBox="[164,782,196,669]" targetPageId="8">
<paragraph id="8BCCF415FFE8731D0A89FD064A77F833" blockId="8.[164,779,698,777]" pageId="8" pageNumber="478">
<emphasis id="B9072807FFE8731D0A89FD064A77F9F4" bold="true" box="[164,773,698,718]" pageId="8" pageNumber="478">
Figures 89.
<taxonomicName id="4C738F96FFE8731D0B11FD064BE3F9F7" authority="Orrico &amp; Kury, 2009" authorityName="Orrico &amp; Kury" authorityYear="2009" box="[316,657,698,717]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="478" phylum="Arthropoda" rank="species" species="poncedeleoni" status="sp. nov.">
<emphasis id="B9072807FFE8731D0B11FD064BE3F9F7" bold="true" box="[316,657,698,717]" italics="true" pageId="8" pageNumber="478">Stygnicranaus poncedeleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234957CFFE8731D088EFD074A77F9F4" box="[675,773,699,718]" pageId="8" pageNumber="478" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (UNAL) from Colombia. Left pedipalpus. Fig. 8. Mesal view. Fig. 9. Ectal view. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8BCCF415FFE8731D0917FBE84D91FF9D" blockId="8.[826,1441,1108,1191]" pageId="8" pageNumber="478">
<emphasis id="B9072807FFE8731D0917FBE84AE6FF53" box="[826,916,1108,1129]" italics="true" pageId="8" pageNumber="478">Female:</emphasis>
Very similar to male, except for the slightly larger size and the absence of the frontal medial hooked spine in the cheliceral hand.
</paragraph>
<paragraph id="8BCCF415FFE8731D0917FB7B4C1AFE23" blockId="8.[826,1441,1222,1305]" pageId="8" pageNumber="478">
<emphasis id="B9072807FFE8731D0917FB7B4A91FFE1" box="[826,995,1222,1244]" italics="true" pageId="8" pageNumber="478">Type material:</emphasis>
<materialsCitation id="3B1BFE48FFE8731D09DCFB754D91FFE6" box="[1009,1251,1222,1244]" collectionCode="UNAL" pageId="8" pageNumber="478" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="9D753F9CFFE8731D09DCFB754D78FFE1" box="[1009,1034,1225,1243]" pageId="8" pageNumber="478" type="male"></specimenCount>
<typeStatus id="54C84AB7FFE8731D0E35FB7A4D0BFFE6" box="[1048,1145,1222,1244]" pageId="8" pageNumber="478" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED626CD0FFE8731D0EA0FB7A4DAEFFE6" box="[1165,1244,1222,1244]" pageId="8" pageNumber="478">UNAL</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B1BFE48FFE8731D0ED8FB7B4C1AFE23" collectingDate="2006-08" collectionCode="MNRJ" collectorName="Dagua. El Salto. &amp; Giupponi, A." country="Colombia" location="Valle Del Cauca" pageId="8" pageNumber="478" specimenCode="MNRJ 17906" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Valle del Cauca" typeStatus="paratype">
<specimenCount id="9D753F9CFFE8731D0ED8FB7B4C31FFE1" box="[1269,1347,1223,1244]" pageId="8" pageNumber="478" type="male">one ♂</specimenCount>
,
<specimenCount id="9D753F9CFFE8731D0F7AFB7B4CD3FFE1" box="[1367,1441,1223,1244]" pageId="8" pageNumber="478" type="female">one ♀</specimenCount>
<typeStatus id="54C84AB7FFE8731D0917FB594AD9FFC0" box="[826,939,1253,1274]" pageId="8" pageNumber="478" type="paratype">paratypes</typeStatus>
(
<specimenCode id="DBD55C6EFFE8731D0991FB594D27FFC1" box="[956,1109,1253,1275]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="8" pageNumber="478">MNRJ 17906</specimenCode>
)
<collectingCountry id="F364B485FFE8731D0E4BFB594DA6FFC1" box="[1126,1236,1253,1275]" name="Colombia" pageId="8" pageNumber="478">Colombia</collectingCountry>
,
<collectingRegion id="49B73AF7FFE8731D0ECCFB594CEFFFC0" box="[1249,1437,1253,1275]" country="Colombia" name="Valle del Cauca" pageId="8" pageNumber="478">Valle Del Cauca</collectingRegion>
,
<collectorName id="268691C3FFE8731D0917FAB84A89FE23" box="[826,1019,1283,1305]" pageId="8" pageNumber="478">Dagua. El Salto.</collectorName>
<date id="FFCDD2D5FFE8731D0E2EFAB84DEAFE23" box="[1027,1176,1283,1305]" pageId="8" pageNumber="478" value="2006-08">
<collectingDate id="EF892B3DFFE8731D0E2EFAB84DEAFE23" box="[1027,1176,1283,1305]" pageId="8" pageNumber="478" value="2006-08">August 2006</collectingDate>
</date>
,
<collectorName id="268691C3FFE8731D0E8BFABF4C4AFE23" box="[1190,1336,1283,1305]" pageId="8" pageNumber="478">Giupponi, A.</collectorName>
col.
</materialsCitation>
</paragraph>
<paragraph id="8BCCF415FFE8731D0917FA854D97FEB6" blockId="8.[826,1441,1337,1420]" pageId="8" pageNumber="478">
<emphasis id="B9072807FFE8731D0917FA854ACAFE74" box="[826,952,1337,1358]" italics="true" pageId="8" pageNumber="478">Etymology:</emphasis>
The species name also honours the collector of the
<typeStatus id="54C84AB7FFE8731D09E1FAE44A8FFE57" box="[972,1021,1368,1389]" pageId="8" pageNumber="478">type</typeStatus>
series, our colleague and friend Alessandro Ponce de Leão Giupponi.
</paragraph>
</subSubSection>
<subSubSection id="C369A79EFFE8731D095AFA6D4DADFD71" pageId="8" pageNumber="478" type="reference_group">
<paragraph id="8BCCF415FFE8731D095AFA6D4C16FEDF" blockId="8.[887,1380,1488,1542]" box="[887,1380,1488,1510]" pageId="8" pageNumber="478">
<taxonomicName id="4C738F96FFE8731D095AFA6D4C16FEDF" ID-CoL="7Z528" authority="ROEWER, 1913" authorityName="Roewer" authorityYear="1913" box="[887,1380,1488,1510]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="478" phylum="Arthropoda" rank="species" species="abnormis">
<emphasis id="B9072807FFE8731D095AFA6D4DC2FEDE" box="[887,1200,1489,1510]" italics="true" pageId="8" pageNumber="478">STYGNICRANAUS ABNORMIS</emphasis>
<bibRefCitation id="EFE289E4FFE8731D0E97FA6D4C16FEDF" author="Roewer C-F" box="[1210,1380,1488,1510]" pageId="8" pageNumber="478" pagination="257 - 472" refId="ref11247" refString="Roewer C-F. 1913. Die Familie der Gonyleptiden de Opiliones - Laniatores. [part 2]. Archiv fur Naturgeschichte 79: 257 - 472 + Taf. 2." type="journal article" year="1913">ROEWER, 1913</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BCCF415FFE8731D0E09FA4C4DC5FD3C" blockId="8.[887,1380,1488,1542]" box="[1060,1207,1520,1542]" pageId="8" pageNumber="478">
(
<figureCitation id="1348E890FFE8731D0E01FA4C4DDCFD3C" box="[1068,1198,1520,1542]" captionStart="Figures 1213" captionStartId="10.[164,254,1265,1284]" captionTargetBox="[419,1187,195,1235]" captionTargetId="figure-233@10.[419,1187,195,1235]" captionTargetPageId="10" captionText="Figures 1213. Stygnicranaus abnormis Roewer, 1913, female holotype (SMF 9800868) from Colombia. Habitus. Fig. 12. Dorsal view. Fig. 13. Lateral view. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5443383" httpUri="https://zenodo.org/record/5443383/files/figure.png" pageId="8" pageNumber="478">FIGS 1213</figureCitation>
)
</paragraph>
<paragraph id="8BCCF415FFE8731D0917F9AB4DADFD71" blockId="8.[826,1441,1559,1611]" pageId="8" pageNumber="478">
<treatmentCitationGroup id="AB63D33BFFE8731D0917F9AB4DADFD71" pageId="8" pageNumber="478">
<treatmentCitation id="0AD2D204FFE8731D0917F9AB4C4BFD16" author="Roewer C-F" box="[826,1337,1559,1580]" page="423" pageId="8" pageNumber="478" year="1913">
<taxonomicName id="4C738F96FFE8731D0917F9AB4C4BFD16" ID-CoL="7Z528" authority="Roewer, 1913: 423" authorityName="Roewer" authorityPageNumber="423" authorityYear="1913" box="[826,1337,1559,1580]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="478" phylum="Arthropoda" rank="species" species="abnormis">
<emphasis id="B9072807FFE8731D0917F9AB4D27FD16" box="[826,1109,1559,1580]" italics="true" pageId="8" pageNumber="478">Stygnicranaus abnormis</emphasis>
<bibRefCitation id="EFE289E4FFE8731D0E4DF9AB4C4BFD16" author="Roewer C-F" box="[1120,1337,1559,1580]" pageId="8" pageNumber="478" pagination="257 - 472" refId="ref11247" refString="Roewer C-F. 1913. Die Familie der Gonyleptiden de Opiliones - Laniatores. [part 2]. Archiv fur Naturgeschichte 79: 257 - 472 + Taf. 2." type="journal article" year="1913">Roewer, 1913: 423</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 167; 1923: 570, fig. 716;
<treatmentCitation id="0AD2D204FFE8731D0E32F98A4DA9FD71" author="Kury AB" box="[1055,1243,1590,1611]" page="102" pageId="8" pageNumber="478" year="2003">
<bibRefCitation id="EFE289E4FFE8731D0E32F98A4DA9FD71" author="Kury AB" box="[1055,1243,1590,1611]" pageId="8" pageNumber="478" pagination="1 - 337" refId="ref10890" refString="Kury AB. 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia vol. especial monografico 1: 1 - 337." type="journal article" year="2003">Kury, 2003: 102</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C369A79EFFE8731F0917F9D74C72FDB3" lastPageId="10" lastPageNumber="480" pageId="8" pageNumber="478" type="materials_examined">
<paragraph id="8BCCF415FFE8731D0917F9D74CF0FD84" blockId="8.[826,1441,1643,1726]" pageId="8" pageNumber="478">
<emphasis id="B9072807FFE8731D0917F9D74AA8FDBA" box="[826,986,1643,1664]" italics="true" pageId="8" pageNumber="478">Type locality:</emphasis>
<collectingCountry id="F364B485FFE8731D09C0F9D74D29FDBB" box="[1005,1115,1643,1665]" name="Colombia" pageId="8" pageNumber="478">Colombia</collectingCountry>
, Maracaibo (
<collectingCountry id="F364B485FFE8731D0F2EF9D74C0AFDBB" box="[1283,1400,1643,1665]" name="Venezuela" pageId="8" pageNumber="478">Venezuela</collectingCountry>
or
<collectingCountry id="F364B485FFE8731D0917F9354AD4FDA5" box="[826,934,1673,1695]" name="Colombia" pageId="8" pageNumber="478">Colombia</collectingCountry>
? See discussion below). WWF Ecoregion NT0145 (North-western Andean montane forests).
</paragraph>
<paragraph id="8BCCF415FFE8731D0917F9624AD8FC57" blockId="8.[826,1441,1757,1901]" pageId="8" pageNumber="478">
<emphasis id="B9072807FFE8731D0917F9624AC3FDC9" box="[826,945,1758,1779]" italics="true" pageId="8" pageNumber="478">Diagnosis:</emphasis>
It is very similar to
<taxonomicName id="4C738F96FFE8731D0E9BF9614C1AFDC8" authorityName="Orrico &amp; Kury" authorityYear="2009" box="[1206,1384,1757,1778]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="478" phylum="Arthropoda" rank="species" species="poncedeleoni">
<emphasis id="B9072807FFE8731D0E9BF9614C1AFDC8" box="[1206,1384,1757,1778]" italics="true" pageId="8" pageNumber="478">S. poncedeleoni</emphasis>
</taxonomicName>
, but does not present pearly rounded granules in femora IV. Also, it is the only species in the genus with a dorsomedial basal small curved spine in the basis of the bulla.
</paragraph>
<caption id="DF0CA49DFFE9731C0ABDFC9B4D10F862" ID-DOI="http://doi.org/10.5281/zenodo.5443381" ID-Zenodo-Dep="5443381" httpUri="https://zenodo.org/record/5443381/files/figure.png" pageId="9" pageNumber="479" startId="9.[144,234,807,826]" targetBox="[399,1167,196,777]" targetPageId="9">
<paragraph id="8BCCF415FFE9731C0ABDFC9B4D10F862" blockId="9.[144,1422,807,856]" pageId="9" pageNumber="479">
<emphasis id="B9072807FFE9731C0ABDFC9B4B83F801" bold="true" box="[144,753,807,827]" pageId="9" pageNumber="479">
Figures 1011.
<taxonomicName id="4C738F96FFE9731C0B6CFC944BFFF800" authority="Orrico &amp; Kury, 2009" authorityName="Orrico &amp; Kury" authorityYear="2009" box="[321,653,807,827]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="479" phylum="Arthropoda" rank="species" species="poncedeleoni" status="sp. nov.">
<emphasis id="B9072807FFE9731C0B6CFC944BFFF800" bold="true" box="[321,653,807,827]" italics="true" pageId="9" pageNumber="479">Stygnicranaus poncedeleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234957CFFE9731C08BAFC944B83F801" box="[663,753,808,827]" pageId="9" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (UNAL) from Colombia. Penis, distal portion. Fig. 10. Dorsal view. Fig. 11. Lateral view. Arrows indicates capellum. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BCCF415FFE9731C0ABDFC194890F880" blockId="9.[144,482,932,954]" box="[144,482,932,954]" pageId="9" pageNumber="479">
<emphasis id="B9072807FFE9731C0ABDFC194828F883" box="[144,346,932,954]" italics="true" pageId="9" pageNumber="479">Description: Male</emphasis>
: Unknown.
</paragraph>
<paragraph id="8BCCF415FFE9731F0ABDFC5D4D1DFD24" blockId="9.[144,759,993,1903]" lastBlockId="10.[826,1441,1391,1566]" lastPageId="10" lastPageNumber="480" pageId="9" pageNumber="479">
<emphasis id="B9072807FFE9731C0ABDFC5D483CF8CC" box="[144,334,993,1014]" italics="true" pageId="9" pageNumber="479">
Female
<typeStatus id="54C84AB7FFE9731C0AC4FC5D483AF8CC" box="[233,328,993,1014]" pageId="9" pageNumber="479" type="holotype">holotype</typeStatus>
:
</emphasis>
Dorsum (
<figureCitation id="1348E890FFE9731C0B93FC5D4B4CF8C2" box="[446,574,993,1016]" captionStart="Figures 1213" captionStartId="10.[164,254,1265,1284]" captionTargetBox="[419,1187,195,1235]" captionTargetId="figure-233@10.[419,1187,195,1235]" captionTargetPageId="10" captionText="Figures 1213. Stygnicranaus abnormis Roewer, 1913, female holotype (SMF 9800868) from Colombia. Habitus. Fig. 12. Dorsal view. Fig. 13. Lateral view. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5443383" httpUri="https://zenodo.org/record/5443383/files/figure.png" pageId="9" pageNumber="479">Figs 1213</figureCitation>
). Dorsal scutum attenuate pyriform, with carapace smaller than abdominal scutum. Posterior border of scutum straight. Cheliceral sockets very shallow with a median tooth. Anterolateral corners of carapace with three to four small teeth on each side, all at pedipalpus level. Posterior corners of scutum with a setiferous tubercle on each side. Ocularium low, with a pair of stout, paramedian erect (lateral view), slightly divergent (frontal view) spines with blunt point. Surface of ocularium with 14 (seven + seven) coarse scattered granules. Carapace behind ocularium with a few small tubercles. Posterior part of lateral area of scutum with an island of such tubercles on each side. Grooves of scutum well marked, defining a mesotergum with three areas; original area IV completely fused with area
<collectionCode id="ED626CD0FFE9731C0B4DFA7748F3FEDA" box="[352,385,1483,1504]" pageId="9" pageNumber="479">III</collectionCode>
. A single shallow longitudinal medial groove dividing areas
<collectionCode id="ED626CD0FFE9731C082EFA564B65FEC5" box="[515,535,1514,1535]" pageId="9" pageNumber="479">II</collectionCode>
and
<collectionCode id="ED626CD0FFE9731C084FFA564BF2FEC5" box="[610,640,1514,1535]" pageId="9" pageNumber="479">III</collectionCode>
into two halves each. All areas armed with a pair of spines except for area
<collectionCode id="ED626CD0FFE9731C0B64F99B482DFD06" box="[329,351,1575,1596]" pageId="9" pageNumber="479">II</collectionCode>
, which is unarmed. Scutal area
<collectionCode id="ED626CD0FFE9731C08CFF99B4B84FD06" box="[738,758,1575,1596]" pageId="9" pageNumber="479">II</collectionCode>
projecting anteriorly into area I dividing it partially into left and right halves. Areas I and
<collectionCode id="ED626CD0FFE9731C0848F9D84B0BFD43" box="[613,633,1636,1657]" pageId="9" pageNumber="479">II</collectionCode>
each with islands of small white setiferous tubercles. Area
<collectionCode id="ED626CD0FFE9731C08F5F93F4B84FDA2" box="[728,758,1667,1688]" pageId="9" pageNumber="479">III</collectionCode>
with clusters of setiferous tubercles anterior to the large spines and reaching their basal part. In the place of area IV, a pair of paramedian setiferous tubercles. Single ozopore opening laterally at the tip of a dorsolateral low protuberance. Free tergites I
<collectionCode id="ED626CD0FFE9731C08CEF8A04B84FC0B" box="[739,758,1820,1841]" pageId="9" pageNumber="479">II</collectionCode>
smooth, except for a pair of paramedian granules each. Free tergite
<collectionCode id="ED626CD0FFE9731C0B57F8E548EAFC54" box="[378,408,1881,1902]" pageId="9" pageNumber="479">III</collectionCode>
with a pair of paramedian strong spines Their corners each bear an acuminate small tooth. Venter. Coxae I
<collectionCode id="ED626CD0FFE9731C0E56FC7F4DEAF8E2" box="[1147,1176,963,984]" pageId="9" pageNumber="479">III</collectionCode>
smooth, except for a transverse row of small tubercles in coxae I and
<collectionCode id="ED626CD0FFE9731C0F5FFC5E4CFBF8CD" box="[1394,1417,994,1015]" pageId="9" pageNumber="479">II</collectionCode>
. Intercoxal bridges weak. Coxae I
<collectionCode id="ED626CD0FFE9731C0E99FBBC4DA3FF2F" box="[1204,1233,1024,1045]" pageId="9" pageNumber="479">III</collectionCode>
subrectangular, transverse. Coxa IV the largest, almost equalling the other three taken together, shape lanceolate and pentagonal, divergent posteriorly. Stigmatic area long, Y-shaped. Stigmata large, elongate elliptic, oblique, clearly apparent, not hidden by any bridges or tubercles, each occupying one-quarter of the area. Maxillary lobes of coxae
<collectionCode id="ED626CD0FFE9731C0E67FB6B4D2CFFD6" box="[1098,1118,1239,1260]" pageId="9" pageNumber="479">II</collectionCode>
clearly visible. Chelicera. Basichelicerite long, with bulla well marked, and a mesal subdistal small acuminate setiferous tubercle. Median tubercle at posterior rim of bulla. Hand not swollen, with one frontal row of granules. Pedipalpus. Very elongate and smooth, with tibia and tarsus forming a subchela. Coxa large, conical, with a row of ventromedial tubercles. Trochanter cylindrical with typical sub-basal constriction, and apical dorsomesal tubercles and one ventral row of setiferous tubercles. Femur as long as dorsal scutum, slightly thicker at distal third, smooth. Patella longer than abdominal scutum, thicker at distal third. Tibia as long as patella, cylindrical with three ectal (IIi) and three mesal (IIi) spines, dorsally covered with coarse tubercles. Tarsus fusiform, slightly shorter than tibia, with seven ectal (iIiiiIi) and 11 mesal (iiIiiiiiIii) spines and two ventral rows of setiferous tubercles. Tarsal claw almost as long as tarsus. Legs. All segments slender and unarmed. Relative length I
<emphasis id="B9072807FFE9731C0EC3F8A04D8EFC0B" box="[1262,1276,1820,1841]" italics="true" pageId="9" pageNumber="479">&lt;</emphasis>
<collectionCode id="ED626CD0FFE9731C0F2FF8A04C52FC0B" box="[1282,1312,1820,1841]" pageId="9" pageNumber="479">III</collectionCode>
<emphasis id="B9072807FFE9731C0F0AF8A04C47FC0B" box="[1319,1333,1820,1841]" italics="true" pageId="9" pageNumber="479">&lt;</emphasis>
<collectionCode id="ED626CD0FFE9731C0F16F8A04C25FC0B" box="[1339,1367,1820,1841]" pageId="9" pageNumber="479">IV</collectionCode>
<emphasis id="B9072807FFE9731C0F73F8A04C1EFC0B" box="[1374,1388,1820,1841]" italics="true" pageId="9" pageNumber="479">&lt;</emphasis>
<collectionCode id="ED626CD0FFE9731C0F5FF8A04CFBFC0B" box="[1394,1417,1820,1841]" pageId="9" pageNumber="479">II</collectionCode>
. Dorsobasal apophyses of coxae I
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close to the ozopore: coxa I anterior spiniform, posterior small, directed backwards; coxa
<collectionCode id="ED626CD0FFEA731F0BF8FACC489BFEBF" box="[469,489,1392,1413]" pageId="10" pageNumber="480">II</collectionCode>
anterior spiniform, posterior apophysis absent. Coxa IV hidden under dorsal scutum in dorsal view, with two prolateral dorsoapical spiniform apophyses and ventral granules. Trochanters IIV with ventrobasal tubercles connecting with respective coxae and scattered small setiferous tubercles all over the ventral surface. Trochanter
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with a few prolateral and retrolateral setiferous tubercles and distally with anterior and posterior spiniform processes. Trochanter IV with row of retrolateral tubercles, increasing distally and two posterior dorsal subequal spurs. All femora straight (even though the
<typeStatus id="54C84AB7FFEA731F0B67F96248D9FDCE" box="[330,427,1758,1780]" pageId="10" pageNumber="480" type="holotype">holotype</typeStatus>
specimen presents curved femora, it is surely a result of the small vial used). Femora
<collectionCode id="ED626CD0FFEA731F0B2EF8A0486DFC0B" box="[259,287,1820,1841]" pageId="10" pageNumber="480">III</collectionCode>
IV with a pair of small dorsoapical spurs. Femora IV also bear a small basal dorsoretrolateral row of five spines. Tarsi
<collectionCode id="ED626CD0FFEA731F0BF0F8E5488BFC54" box="[477,505,1881,1902]" pageId="10" pageNumber="480">III</collectionCode>
IV with unpectinated double claws and well-developed tarsal process (pseudonychium). For tarsal counts see
<tableCitation id="C6F1C1AEFFEA731F0F6AFA324CE9FE99" box="[1351,1435,1422,1444]" captionStart="Table 1" captionStartId="2.[164,228,870,889]" captionText="Table 1. Tarsal counts for the species of Stygnicranainae" pageId="10" pageNumber="480">Table 1</tableCitation>
. Colour. Somewhat faded because of conservation in alcohol. Body and appendages background Deep Orange with lighter reticulations. Granules and spots of the scutum Pale Yellow.
</paragraph>
<caption id="DF0CA49DFFEA731F0A89FB4D4B96FE18" ID-DOI="http://doi.org/10.5281/zenodo.5443383" ID-Zenodo-Dep="5443383" httpUri="https://zenodo.org/record/5443383/files/figure.png" pageId="10" pageNumber="480" startId="10.[164,254,1265,1284]" targetBox="[419,1187,195,1235]" targetPageId="10">
<paragraph id="8BCCF415FFEA731F0A89FB4D4B96FE18" blockId="10.[164,1441,1265,1314]" pageId="10" pageNumber="480">
<emphasis id="B9072807FFEA731F0A89FB4D4838FE3E" bold="true" box="[164,330,1265,1284]" pageId="10" pageNumber="480">Figures 1213.</emphasis>
<taxonomicName id="4C738F96FFEA731F0B7BFB4E4B9EFE3E" authority="Roewer, 1913" authorityName="Roewer" authorityYear="1913" box="[342,748,1265,1285]" class="Arachnida" family="Cranaidae" genus="Stygnicranaus" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="480" phylum="Arthropoda" rank="species" species="abnormis">
<emphasis id="B9072807FFEA731F0B7BFB4E4B25FE3E" box="[342,599,1265,1285]" italics="true" pageId="10" pageNumber="480">Stygnicranaus abnormis</emphasis>
<bibRefCitation id="EFE289E4FFEA731F0873FB4E4B9EFE3E" author="Roewer C-F" box="[606,748,1265,1285]" pageId="10" pageNumber="480" pagination="257 - 472" refId="ref11247" refString="Roewer C-F. 1913. Die Familie der Gonyleptiden de Opiliones - Laniatores. [part 2]. Archiv fur Naturgeschichte 79: 257 - 472 + Taf. 2." type="journal article" year="1913">Roewer, 1913</bibRefCitation>
</taxonomicName>
, female holotype (SMF 9800868) from Colombia. Habitus. Fig. 12. Dorsal view. Fig. 13. Lateral view. Scale bars = 3 mm.
</paragraph>
</caption>
<paragraph id="8BCCF415FFEA731F0917F98B4C72FDB3" blockId="10.[826,1441,1590,1673]" pageId="10" pageNumber="480">
<emphasis id="B9072807FFEA731F0917F98B4D15FD71" box="[826,1127,1590,1612]" italics="true" pageId="10" pageNumber="480">Type material (examined):</emphasis>
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<typeStatus id="54C84AB7FFEA731F0EA2F98A4D82FD76" box="[1167,1264,1590,1612]" pageId="10" pageNumber="480" type="holotype">holotype</typeStatus>
(and not
<specimenCount id="9D753F9CFFEA731F0F48F9854C0CFD71" box="[1381,1406,1593,1611]" pageId="10" pageNumber="480" type="male"></specimenCount>
as in
<bibRefCitation id="EFE289E4FFEA731F0977F9E94A84FD50" author="Roewer C-F" box="[858,1014,1621,1642]" pageId="10" pageNumber="480" pagination="257 - 472" refId="ref11247" refString="Roewer C-F. 1913. Die Familie der Gonyleptiden de Opiliones - Laniatores. [part 2]. Archiv fur Naturgeschichte 79: 257 - 472 + Taf. 2." type="journal article" year="1913">Roewer, 1913</bibRefCitation>
,
<bibRefCitation id="EFE289E4FFEA731F0E2EF9E94D48FD51" author="Roewer C-F" box="[1027,1082,1621,1643]" pageId="10" pageNumber="480" pagination="1 - 1212" refId="ref11279" refString="Roewer C-F. 1923. Die Weberknechte der Erde. Systematische Bearbeitung der bisher bekannten Opiliones. Jena: Gustav Fischer. 1116 pp., figs 1 - 1212." type="book chapter" year="1923">1923</bibRefCitation>
) (
<collectionCode id="ED626CD0FFEA731F0E44F9E94DD3FD50" box="[1129,1185,1621,1642]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="10" pageNumber="480">SMF</collectionCode>
9800868 RI/868-43)
<collectingCountry id="F364B485FFEA731F097FF9CF4ACFFDB3" box="[850,957,1651,1673]" name="Colombia" pageId="10" pageNumber="480">Colombia</collectingCountry>
[or
<collectingCountry id="F364B485FFEA731F09DDF9CF4D15FDB3" box="[1008,1127,1651,1673]" name="Venezuela" pageId="10" pageNumber="480">Venezuela</collectingCountry>
?] Maracaibo.
</paragraph>
</subSubSection>
</treatment>
</document>