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<document id="476C6707EFF35E6BB460764CEFD5A3D9" ID-ISSN="1536-9307" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638260192403" checkinUser="felipe" docAuthor="Teruel, Rolando, Kovařík, František &amp; Fet, Victor" docDate="2018" docId="03D887E8FB76FFACFCCEF452FAFEED7F" docLanguage="en" docName="Euscorpius.270-1-45.pdf" docOrigin="Euscorpius 270" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Anomalobuthus rickmersi Kraepelin 1900" docType="treatment" docVersion="2" lastPageNumber="11" masterDocId="FFE1FF90FB72FFA0FFA1F66FFFB6EB54" masterDocTitle="Revision of the Central Asian scorpion genus Anomalobuthus Kraepelin, 1900, with descriptions of three new species and a generic synonymy (Scorpiones: Buthidae)" masterLastPageNumber="45" masterPageNumber="1" pageNumber="3" updateTime="1699436338496" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="ECB3728AAF0208F96ECE054624BB2335">Revision of the Central Asian scorpion genus Anomalobuthus Kraepelin, 1900, with descriptions of three new species and a generic synonymy (Scorpiones: Buthidae)</mods:title>
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<mods:namePart id="D8578DB631DA3E16F98068EB038DF736">Teruel, Rolando</mods:namePart>
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<mods:namePart id="5FD0082CD42E979E82053A96E719F053">Kovařík, František</mods:namePart>
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<mods:title id="0D5E53B73102E0D72D0DD491AE98C4BB">Euscorpius</mods:title>
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<mods:date id="812710B0E1C248B717C2664E9EBA41C3">2018</mods:date>
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<taxonomicName id="4C714D7DFB76FFA4FCCEF452FAD9E90C" ID-CoL="5V74N" authority="Kraepelin, 1900" authorityName="Kraepelin" authorityYear="1900" box="[879,1391,573,600]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="rickmersi">
<emphasis id="B905EAECFB76FFA4FCCEF452FB1CE90C" bold="true" box="[879,1194,573,600]" italics="true" pageId="4" pageNumber="3">Anomalobuthus rickmersi</emphasis>
<bibRefCitation id="EFE04B0FFB76FFA4FB13F451FAD9E90C" author="KRAEPELIN" box="[1202,1391,574,600]" pageId="4" pageNumber="3" pagination="1 - 17" refId="ref25373" refString="KRAEPELIN, K. 1900. Uber einige neue Gliederspinnen. Abhandlungen aus dem Gebiete der Naturwissenschaften. Herausgegeben vom Naturwissenschaftlichen Verein in Hamburg, 16, 1 (4): 1 - 17." type="journal article" year="1900">Kraepelin, 1900</bibRefCitation>
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<figureCitation id="134A2A7BFB76FFA4FCEBF430FC67E922" box="[842,977,607,630]" captionStart-0="Figures 1–3" captionStart-1="Figures 4–7" captionStart-2="Figures 8–13" captionStart-3="Figures 14–26" captionStart-4="Figures 27–33" captionStartId-0="5.[192,278,1579,1602]" captionStartId-1="6.[192,278,1734,1757]" captionStartId-2="7.[192,278,1543,1566]" captionStartId-3="9.[192,278,1609,1632]" captionStartId-4="10.[192,278,1394,1417]" captionTargetBox-0="[191,1440,191,1550]" captionTargetBox-1="[207,1403,201,1681]" captionTargetBox-2="[191,1437,191,1514]" captionTargetBox-3="[196,1409,215,1573]" captionTargetBox-4="[192,1437,192,1364]" captionTargetId-0="figure-6@5.[192,1439,192,1549]" captionTargetId-1="figure-12@6.[192,1439,188,1704]" captionTargetId-2="figure-6@7.[192,1437,192,1513]" captionTargetId-3="figure-6@9.[192,1439,192,1580]" captionTargetId-4="figure-12@10.[192,1437,192,1364]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="9" captionTargetPageId-4="10" captionText-0="Figures 1–3: Anomalobuthus rickmersi, adult male holotype: full-body views, dorsal (1) and ventral (2), plus original labels (3). Scale bar = 10 mm (1–2). The red inset at bottom right shows the label of the specimen examined by Lourenço (2001), demonstrating it was not the actual Rickmers's type specimen." captionText-1="Figures 4–7: Anomalobuthus rickmersi from Turkmenistan: adult male from Repetek (4–5) and adult female from Chainury (6– 7), full-body views, dorsal (4, 6) and ventral (5, 7). Scale bar = 10 mm." captionText-2="Figures 8–13: Anomalobuthus rickmersi. Figures 8–9. Juvenile male from Repetek, Turkmenistan: full-body views, dorsal (8) and ventral (9). Scale bar = 10 mm. Figures 10–13. Adult male holotype: chelicerae, carapace and tergites I–III, dorsal (10), sternopectinal region and sternites III–V, ventral (11), distal part of left leg III, internal (12), distal part of left leg IV, internal (13)." captionText-3="Figures 14–26: Anomalobuthus rickmersi, adult male holotype: right pedipalp trochanter and femur, dorsal (14) and ventral (15), right pedipalp patella, dorsal (16), external (17) and ventral (18), right pedipalp chela, dorsal (19–20), external (21) and ventrointernal (22), metasoma and telson, lateral (23), dorsal (24) and ventral (25), telson, lateral (26)." captionText-4="Figures 27–33: Anomalobuthus rickmersi from Turkmenistan, adult male from Repetek (27–30) and adult female from Chainury (31–33): metasoma and telson, lateral (27, 31), dorsal (28, 32) and ventral (29, 33), telson, lateral (30)." pageId="4" pageNumber="3">Figures 1–33</figureCitation>
,
<figureCitation id="134A2A7BFB76FFA4FC7AF430FB80E922" box="[987,1078,607,630]" captionStart="Figures 107–119" captionStartId="33.[192,278,1734,1757]" captionTargetBox="[191,1440,216,1705]" captionTargetId="figure-6@33.[192,1439,217,1704]" captionTargetPageId="33" captionText="Figures 107–119: Anomalobuthus spp., from ZISP historical collection: adult male topotypes of A. pavlovskyi sp. n. (107– 112), full-body dorsal views (107, 110), labels (108, 111) and bottles (109, 112); adult male (113) and adult female (114) A. pavlovskyi sp. n. from Kunya-Urgench, Turkmenistan, full-body dorsal views (113, 114), labels (115) and bottle (116); adult male A. rickmersi from Tejen, Turkmenistan (117–119), full-body dorsal view (117), labels (118) and bottle (119). Photographs courtesy Alexander Koval." pageId="4" pageNumber="3">117–119</figureCitation>
,
<figureCitation id="134A2A7BFB76FFA4FBE1F430FB2DE922" box="[1088,1179,607,630]" captionStart="Figures 126–133" captionStartId="36.[192,278,1688,1711]" captionTargetBox="[191,1440,167,1659]" captionTargetId="figure-12@36.[192,1439,168,1658]" captionTargetPageId="36" captionText="Figures 126–133: Anomalobuthus spp., SEM views of right pedipalp of adult male A. rickmersi from Repetek (126–127) and adult male paratopotype A. krivochatskyi sp. n. (128–133): chela (126, 128), close-up of manus (127, 129), three ventrointernal magnified views of mid part of both fingers in consecutive figures (130–132, movable finger on top), ventrointernal close-up of fingertips (133, movable finger on top). See trichobothria highlighted in red." pageId="4" pageNumber="3">126–127</figureCitation>
,
<figureCitation id="134A2A7BFB76FFA4FB07F430FB78E922" box="[1190,1230,607,630]" captionStart="Figures 134–140" captionStartId="37.[192,278,1748,1771]" captionTargetBox="[191,1440,191,1719]" captionTargetId="figure-6@37.[192,1439,192,1718]" captionTargetPageId="37" captionText="Figures 134–140: Anomalobuthus spp., SEM views showing size variation of leg IV tibial spur (highlighted in red): large in paratopotype female A. krivochatskyi sp. n. (134), moderately large in paratype female A. krivochatskyi sp. n. from Bukhara (135), small in paratopotype female A. krivochatskyi sp. n. (136), absent in male A. rickmersi from Repetek (137), close-up of a medium-sized leg IV tibial spur of a paratopotype female A. krivochatskyi sp. n. from Zarafshan (138), see spinulate and striate sculpture. Anomalobuthus krivochatskyi sp. n., adult female paratopotype: SEM views of right pecten and adjacent parts of leg coxae and sternite III (139), peg sensilla of tooth from same pecten (140)." pageId="4" pageNumber="3">137</figureCitation>
,
<figureCitation id="134A2A7BFB76FFA4FB79F430FAB6E922" box="[1240,1280,607,630]" captionStart="Figure 141" captionStartId="40.[192,268,1144,1167]" captionTargetBox="[191,1440,191,1114]" captionTargetId="figure-12@40.[192,1439,192,1114]" captionTargetPageId="40" captionText="Figure 141: Distribution of the genus Anomalobuthus: A. krivochatskyi sp. n. (1), A. lowei sp. n. (2), A. pavlovskyi sp. n. (3), A. rickmersi (4), A. talebii (5), A. zarudnyi comb. n. (6), and literature records whose identification is pending (black squares). Data compiled from specimens studied herein and the following supplementary sources: Rickmer Rickmers (1913), Fet (1989, 1994), Gromov &amp; Kopdykbaev (1994), Gromov (2006), Mityaev et al. (2008), and Graham et al. (2012)." pageId="4" pageNumber="3">141</figureCitation>
;
<tableCitation id="C6F30345FB76FFA4FAADF430FAD4E922" box="[1292,1378,607,630]" captionStart="Table 1" captionStartId="12.[192,257,1118,1141]" captionText="Table 1: Measurements of three adults of Anomalobuthus rickmersi. Abbreviations: length (L), width (W), posterior width (Wp), depth (D)." pageId="4" pageNumber="3">Tables 1</tableCitation>
,
<tableCitation id="C6F30345FB76FFA4FACEF430FA22E922" box="[1391,1428,607,630]" captionStart-0="Table 4" captionStart-1="Table 5" captionStartId-0="35.[192,257,1030,1053]" captionStartId-1="38.[192,257,680,703]" captionText-0="Table 4: Comparison between five species of Anomalobuthus based upon selected morphometric ratios. Abbreviations: length (L)," captionText-1="Table 5: Pectinal tooth count variation in all known species of Anomalobuthus. Abbreviations: number of pectines (N), average (Ave.), standard deviation (SD). This includes counts taken from sources with precise and reliable geographic data (Birula, 1905, 1910)." pageId="4" pageNumber="3">4–5</tableCitation>
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<treatmentCitation id="0AD010EFFB76FFA4FCCFF4CDFA2AE9ED" author="KRAEPELIN" box="[878,1436,674,697]" page="11" pageId="4" pageNumber="3" year="1900">
<taxonomicName id="4C714D7DFB76FFA4FCCFF4CDFA2AE9ED" ID-CoL="5V74N" authority="Kraepelin, 1900: 1 - 11" authorityName="Kraepelin" authorityPageNumber="1 - 11" authorityYear="1900" box="[878,1436,674,697]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="rickmersi">
<emphasis id="B905EAECFB76FFA4FCCFF4CDFB3CE9ED" box="[878,1162,674,697]" italics="true" pageId="4" pageNumber="3">Anomalobuthus rickmersi</emphasis>
<bibRefCitation id="EFE04B0FFB76FFA4FB38F4CDFA2AE9ED" author="KRAEPELIN" box="[1177,1436,674,697]" pageId="4" pageNumber="3" pagination="1 - 17" refId="ref25373" refString="KRAEPELIN, K. 1900. Uber einige neue Gliederspinnen. Abhandlungen aus dem Gebiete der Naturwissenschaften. Herausgegeben vom Naturwissenschaftlichen Verein in Hamburg, 16, 1 (4): 1 - 17." type="journal article" year="1900">Kraepelin, 1900: 1–11</bibRefCitation>
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, figs. 4–7; Birula, 1904: 32; Birula, 1905: 449 (in part); Birula, 1911a: 73–74 Birula, 1911b: 171 (in part); Rickmer Rickmers, 1913: 442; Birula, 1917: 112 (in part);
<treatmentCitation id="0AD010EFFB76FFA4FC9FF572FB9DE860" author="PAVLOVSKY" box="[830,1067,797,820]" page="201" pageId="4" pageNumber="3" year="1934">
<bibRefCitation id="EFE04B0FFB76FFA4FC9FF572FB9DE860" author="PAVLOVSKY" box="[830,1067,797,820]" pageId="4" pageNumber="3" pagination="191 - 204" refId="ref25791" refString="PAVLOVSKY, E. N. 1934. [Some data on the poisonous animals of Turkmenia]. Pp. 191 - 204 in: Trudy Kara-Kalinskoi i Kzyl-Atrekskoi parazitologicheskoi ekspeditsii 1931 goda i materialy po faune Turkmenii [Transactions of the 1931 Kara- Kala and Kzyl-Atrek Parasitological Expedition, with contributions to the fauna of Turkmenia]. Leningrad: Ed. Acad. Sci. USSR and Narkomzdrav of Turkmenia (in Russian)." type="book chapter" year="1934">Pavlovsky, 1934: 201</bibRefCitation>
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(in part).
<treatmentCitation id="0AD010EFFB76FFA4FB3CF572FAD8E860" author="WERNER" box="[1181,1390,797,820]" page="272" pageId="4" pageNumber="3" year="1934">
<bibRefCitation id="EFE04B0FFB76FFA4FB3CF572FAD8E860" author="WERNER" box="[1181,1390,797,820]" pageId="4" pageNumber="3" pagination="8" refId="ref26500" refString="WERNER, F. 1934. Scorpiones, Pedipalpi. Pp. 1 - 316 in: Bronns, H. G. (ed.). Klassen und Ordnungen des Tierreichs. Akademische Verlag Gesellschaft, Leipzig. 5, IV, 8, Lief. 1 - 2 (Scorpiones), 316 pp." type="book chapter" year="1934">Werner, 1934: 272</bibRefCitation>
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; fig. 341;
<treatmentCitation id="0AD010EFFB76FFA4FCD8F553FBFAE807" author="WEIDNER" box="[889,1100,828,851]" page="98" pageId="4" pageNumber="3" year="1959">
<bibRefCitation id="EFE04B0FFB76FFA4FCD8F553FBFAE807" author="WEIDNER" box="[889,1100,828,851]" pageId="4" pageNumber="3" pagination="89 - 142" refId="ref26459" refString="WEIDNER, H. 1959. Die Entomologischen Sammlungen des Zoologischen Staatsinstituts und Zoologischen Museums Hamburg, I. Teil, Pararthropoda und Chelicerata I. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 57: 89 - 142." type="journal article" year="1959">Weidner, 1959: 98</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AD010EFFB76FFA4FBFDF553FA85E807" author="STAHNKE" box="[1116,1331,828,851]" page="122" pageId="4" pageNumber="3" year="1972">
<bibRefCitation id="EFE04B0FFB76FFA4FBFDF553FA85E807" author="STAHNKE" box="[1116,1331,828,851]" pageId="4" pageNumber="3" pagination="121 - 133" refId="ref26140" refString="STAHNKE, H. L. 1972. A key to the genera of Buthidae (Scorpionida). Entomological News, 83 (5): 121 - 133." type="journal article" year="1972">Stahnke, 1972: 122</bibRefCitation>
</treatmentCitation>
(in part), fig. 3;
<treatmentCitation id="0AD010EFFB76FFA4FC2FF535FBD0E825" author="VACHON" box="[910,1126,858,881]" page="908" pageId="4" pageNumber="3" year="1974">
<bibRefCitation id="EFE04B0FFB76FFA4FC2FF535FBD0E825" author="VACHON" box="[910,1126,858,881]" pageId="4" pageNumber="3" pagination="857 - 958" refId="ref26267" refString="VACHON, M. 1974. Etudes des caracteres utilises pour classer les familles et les genres des scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, 3 e serie, 140 (Zoologie, 104): 857 - 958." type="journal article" year="1974">Vachon, 1974: 908</bibRefCitation>
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, fig. 37; Fet, 1980: 224 (in part); Fet, 1989: 80 (in part);
<treatmentCitation id="0AD010EFFB76FFA4FB22F516FAF7E8C4" author="SISSOM" box="[1155,1345,889,912]" page="37" pageId="4" pageNumber="3" year="1990">
<bibRefCitation id="EFE04B0FFB76FFA4FB22F516FAF7E8C4" author="SISSOM" box="[1155,1345,889,912]" pageId="4" pageNumber="3" pagination="64 - 160" refId="ref25919" refString="SISSOM, W. D. 1990. Systematics, biogeography and paleontology. Pp. 64 - 160 in: Polis, G. A. (ed.). Biology of Scorpions. Stanford, California: Stanford University Press, 587 pp." type="book chapter" year="1990">Sissom, 1990: 37</bibRefCitation>
</treatmentCitation>
, 46, 48– 51, 96, 101, 120, figs. 3.17h, m; Fet, 1994: 531 (in part); Gromov &amp; Kopdykbaev, 1994: 20 (in part); Fet et al., 1998: 613 (in part), fig. 6 [specimen from Repetek]; Fet &amp; Lowe, 2000: 75 (in part, includes full list of references before 1998); Fet et al., 2001: 183–185, tab. 1 (in part). Fet et al., 2003: 4 (in part); Fet et al., 2005: 2– 3, 6–7, 10–11, 13, 22, 24–25, figs. 10, 23–25, tab. 1 (in part);
<treatmentCitation id="0AD010EFFB76FFA4FC24F201FB09EFD1" author="GRAHAM" box="[901,1215,1134,1157]" page="95" pageId="4" pageNumber="3" year="2012">
<bibRefCitation id="EFE04B0FFB76FFA4FC24F201FB09EFD1" author="GRAHAM" box="[901,1215,1134,1157]" pageId="4" pageNumber="3" pagination="95 - 110" refId="ref24752" refString="GRAHAM, M. R., V. OLAH-HEMMINGS &amp; V. FET. 2012. Phylogeography of co-distributed dune scorpions identifies the Amu Darya River as a longstanding component of Central Asian biogeography (Scorpiones: Buthidae). Zoology in the Middle East, 55: 95 - 110." type="journal article" year="2012">Graham et al., 2012: 95–96</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0AD010EFFB76FFA4FA9BF201FC17EFF0" author="WARBURG" page="96" pageId="4" pageNumber="3" year="2013">
<bibRefCitation id="EFE04B0FFB76FFA4FA9BF201FC17EFF0" author="WARBURG" pageId="4" pageNumber="3" pagination="95 - 104" refId="ref26429" refString="WARBURG, M. R. 2013. The locomotory rhythmic activity in scorpions: with a review. Arthropods, 2 (3): 95 - 104." type="journal article" year="2013">Warburg, 2013: 96</bibRefCitation>
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, 102 (in part);
<treatmentCitation id="0AD010EFFB76FFA4FBE4F2E2FA9CEFF0" author="TERUEL" box="[1093,1322,1165,1188]" page="1" pageId="4" pageNumber="3" year="2014">
<bibRefCitation id="EFE04B0FFB76FFA4FBE4F2E2FA9CEFF0" author="TERUEL" box="[1093,1322,1165,1188]" pageId="4" pageNumber="3" pagination="1 - 10" refId="ref26171" refString="TERUEL, R., F. KOVARIK, S. NAVIDPOUR &amp; V. FET. 2014. The first record of the genus Anomalobuthus Kraepelin, 1900 from Iran, with description of a new species (Scorpiones: Buthidae). Euscorpius, 192: 1 - 10." type="journal article" year="2014">Teruel et al., 2014: 1</bibRefCitation>
</treatmentCitation>
, 4, 6–7, 9, fig. 20, tabs. 1–2 (in part).
<emphasis id="B905EAECFB76FFA4FBCFF2C3FB21EF97" box="[1134,1175,1196,1219]" italics="true" pageId="4" pageNumber="3">Nec</emphasis>
<treatmentCitation id="0AD010EFFB76FFA4FB00F2C3FA16EF97" author="LOURENCO" box="[1185,1440,1196,1219]" page="15" pageId="4" pageNumber="3" year="2001">
<bibRefCitation id="EFE04B0FFB76FFA4FB00F2C3FA16EF97" author="LOURENCO" box="[1185,1440,1196,1219]" pageId="4" pageNumber="3" pagination="15 - 20" refId="ref25502" refString="LOURENCO, W. R. 2001. Un nouveau genre de Buthidae, probable vicariant geographique d'Anomalobuthus Kraepelin (Chelicerata, Scorpiones). Biogeographica, 77 (1): 15 - 20." type="journal article" year="2001">Lourenço, 2001: 15–20</bibRefCitation>
</treatmentCitation>
[misidentification, see below in Remarks section].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36B6575FB76FFA4FCCFF361FC15EC25" pageId="4" pageNumber="3" type="materials_examined">
<paragraph id="8BCE36FEFB76FFA4FCCFF361FC24EECB" blockId="4.[830,1441,1294,1439]" pageId="4" pageNumber="3">
<materialsCitation id="3B193CA3FB76FFA4FCCFF361FC24EECB" collectionCode="ZMUH" collectorName="W. Rickmer Rickmers. See" country="Tajikistan" location="Kyzylsu River valley" pageId="4" pageNumber="3" specimenCount="1" specimenCount-male="1" stateProvince="Khatlon" typeStatus="holotype">
<typeStatus id="54CA885CFB76FFA4FCCFF361FC55EE70" box="[878,995,1294,1317]" pageId="4" pageNumber="3" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D77FD77FB76FFA4FC58F361FBBBEE71" box="[1017,1037,1294,1317]" pageId="4" pageNumber="3" type="male">♂</specimenCount>
(
<collectionCode id="ED60AE3BFB76FFA4FB8BF361FBCBEE71" box="[1066,1149,1294,1317]" country="Viet Nam" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoological Museum, University of Hanoi" pageId="4" pageNumber="3">ZMUH</collectionCode>
, examined).
<emphasis id="B905EAECFB76FFA4FA85F361FA2FEE71" bold="true" box="[1316,1433,1294,1317]" pageId="4" pageNumber="3">
<collectingCountry id="F366766EFB76FFA4FA85F361FA2FEE71" box="[1316,1433,1294,1317]" name="Tajikistan" pageId="4" pageNumber="3">Tajikistan</collectingCountry>
</emphasis>
,
<collectingRegion id="49B5F81CFB76FFA4FC9FF343FBB6EE17" box="[830,1024,1324,1347]" country="Tajikistan" name="Khatlon" pageId="4" pageNumber="3">Khatlon Province</collectingRegion>
,
<location id="8EAE6025FB76FFA4FBACF343FB4FEE17" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB76FFA4FBACF343FB4FEE17" box="[1037,1273,1324,1347]" country="Tajikistan" name="Kyzylsu River valley" pageId="4" pageNumber="3" stateProvince="Khatlon">Kyzylsu River valley</location>
, near Baljuvon (= Bal'dzhuvon or Baljuan), early summer 1896, leg.
<collectorName id="26845328FB76FFA4FA21F324FBF0EED5" pageId="4" pageNumber="3">W. Rickmer Rickmers. See</collectorName>
discussion below, in Remarks section.
</materialsCitation>
</paragraph>
<paragraph id="8BCE36FEFB76FFA4FCCFF3A3FC15EC25" blockId="4.[830,1441,1484,1905]" pageId="4" pageNumber="3">
<materialsCitation id="3B193CA3FB76FFA4FCCFF3A3FBA4ED6B" collectingDate="1985-07-21" collectionCode="OTHER, MATERIAL, EXAMINED" country="Turkmenistan" county="Karakum Desert" elevation="201" latitude="38.565" location="Province" longLatPrecision="19" longitude="63.180836" municipality="Chardzhou" pageId="4" pageNumber="3" specimenCount="1" stateProvince="Lebap Welayaty">
<collectionCode id="ED60AE3BFB76FFA4FCCFF3A3FB13EEB6" box="[878,1189,1484,1507]" pageId="4" pageNumber="3">OTHER MATERIAL EXAMINED</collectionCode>
.
<emphasis id="B905EAECFB76FFA4FB12F3A3FAE0EEB7" bold="true" box="[1203,1366,1484,1507]" pageId="4" pageNumber="3">
<collectingCountry id="F366766EFB76FFA4FB12F3A3FAE0EEB7" box="[1203,1366,1484,1507]" name="Turkmenistan" pageId="4" pageNumber="3">Turkmenistan</collectingCountry>
</emphasis>
,
<collectingCounty id="62AF4E72FB76FFA4FAC2F3A3FC71ED55" pageId="4" pageNumber="3">Karakum Desert</collectingCounty>
,
<collectingRegion id="49B5F81CFB76FFA4FC7BF385FBABED55" box="[986,1053,1514,1537]" country="Turkmenistan" name="Lebap Welayaty" pageId="4" pageNumber="3">Lebap</collectingRegion>
(=
<collectingMunicipality id="6BAAAC84FB76FFA4FBE2F385FB0AED55" box="[1091,1212,1514,1537]" pageId="4" pageNumber="3">Chardzhou</collectingMunicipality>
)
<location id="8EAE6025FB76FFA4FB72F385FA83ED55" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB76FFA4FB72F385FA83ED55" box="[1235,1333,1514,1537]" country="Turkmenistan" county="Karakum Desert" latitude="38.565" longLatPrecision="19" longitude="63.180836" municipality="Chardzhou" name="Province" pageId="4" pageNumber="3" stateProvince="Lebap Welayaty">Province</location>
,
<location id="8EAE6025FB76FFA4FAE8F385FC5AED74" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB76FFA4FAE8F385FC5AED74" country="Turkmenistan" county="Karakum Desert" latitude="38.565" longLatPrecision="19" longitude="63.180836" municipality="Chardzhou" name="Repetek Nature Reserve" pageId="4" pageNumber="3" stateProvince="Lebap Welayaty">Repetek Nature Reserve</location>
,
<geoCoordinate id="EE455039FB76FFA4FC5DF066FBCFED74" box="[1020,1145,1545,1568]" degrees="38" direction="north" minutes="33" orientation="latitude" pageId="4" pageNumber="3" precision="15" seconds="54" value="38.565">38°33'54&quot;N</geoCoordinate>
<geoCoordinate id="EE455039FB76FFA4FB27F066FAB4ED74" box="[1158,1282,1545,1568]" degrees="63" direction="east" minutes="10" orientation="longitude" pageId="4" pageNumber="3" precision="15" seconds="51" value="63.180836">63°10'51&quot;E</geoCoordinate>
,
<elevation id="005CD1CDFB76FFA4FAB2F066FA2CED74" box="[1299,1434,1545,1568]" metricMagnitude="2" metricUnit="m" metricValue="2.01" pageId="4" pageNumber="3" unit="m" value="201.0">
<quantity id="4C899B1BFB76FFA4FAB2F066FAEDED74" box="[1299,1371,1545,1568]" metricMagnitude="2" metricUnit="m" metricValue="2.01" pageId="4" pageNumber="3" unit="m" value="201.0">201 m</quantity>
a.s.l.
</elevation>
,
<date id="FFCF103EFB76FFA4FC9FF047FC6DED6B" box="[830,987,1576,1599]" pageId="4" pageNumber="3" value="1985-07-21">
<collectingDate id="EF8BE9D6FB76FFA4FC9FF047FC6DED6B" box="[830,987,1576,1599]" pageId="4" pageNumber="3" value="1985-07-21">21 July 1985</collectingDate>
</date>
, leg
</materialsCitation>
.
<collectionCode id="ED60AE3BFB76FFA4FB84F047FB8DED6B" box="[1061,1083,1576,1599]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="3" type="Museum">V</collectionCode>
.
<materialsCitation id="3B193CA3FB76FFA4FBECF047FA2BED09" collectingDate="1967-04-29" collectionCode="NHMW, VFPC" collectorName="Fet" country="Turkmenistan" county="Karakum Desert" elevation="201" latitude="38.565" location="Province" longLatPrecision="19" longitude="63.180836" municipality="Chardzhou" pageId="4" pageNumber="3" specimenCount="8" specimenCount-juvenile="1" specimenCount-male="7" stateProvince="Lebap Welayaty">
<collectorName id="26845328FB76FFA4FBECF047FBC4ED6B" box="[1101,1138,1576,1599]" pageId="4" pageNumber="3">Fet</collectorName>
,
<specimenCount id="9D77FD77FB76FFA4FB24F047FB0CED6B" box="[1157,1210,1576,1599]" pageId="4" pageNumber="3" type="male">2♂♂</specimenCount>
(
<collectionCode id="ED60AE3BFB76FFA4FB71F047FA9BED6B" box="[1232,1325,1576,1599]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="3" type="Museum">NHMW</collectionCode>
),
<specimenCount id="9D77FD77FB76FFA4FAE9F047FACBED6B" box="[1352,1405,1576,1599]" pageId="4" pageNumber="3" type="male">5♂♂</specimenCount>
,
<specimenCount id="9D77FD77FB76FFA4FA32F047FC22ED09" pageId="4" pageNumber="3" type="juvenile">1 juvenile</specimenCount>
(
<collectionCode id="ED60AE3BFB76FFA4FC07F029FC59ED09" box="[934,1007,1606,1629]" pageId="4" pageNumber="3">VFPC</collectionCode>
); same data except
<date id="FFCF103EFB76FFA4FB6DF029FADCED09" box="[1228,1386,1606,1629]" pageId="4" pageNumber="3" value="1967-04-29">
<collectingDate id="EF8BE9D6FB76FFA4FB6DF029FADCED09" box="[1228,1386,1606,1629]" pageId="4" pageNumber="3" value="1967-04-29">29 April 1967</collectingDate>
</date>
, leg
</materialsCitation>
.
<collectionCode id="ED60AE3BFB76FFA4FC9FF00AFCE2ED28" box="[830,852,1637,1660]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="3" type="Museum">V</collectionCode>
.
<materialsCitation id="3B193CA3FB76FFA4FCC1F00AFBBFEDCF" collectingDate="2002-04-15" collectionCode="ZSRO" collectorName="I. Kuznetsov" country="Turkmenistan" county="Karakum Desert" elevation="201" latitude="38.565" location="Province" longLatPrecision="19" longitude="63.180836" municipality="Chardzhou" pageId="4" pageNumber="3" specimenCount="1" specimenCount-male="1" stateProvince="Lebap Welayaty">
<collectorName id="26845328FB76FFA4FCC1F00AFC5BED28" box="[864,1005,1637,1660]" pageId="4" pageNumber="3">I. Kuznetsov</collectorName>
,
<specimenCount id="9D77FD77FB76FFA4FC59F00AFBACED28" box="[1016,1050,1637,1660]" pageId="4" pageNumber="3" type="male">1♂</specimenCount>
(
<collectionCode id="ED60AE3BFB76FFA4FB88F00AFBD9ED28" box="[1065,1135,1637,1660]" pageId="4" pageNumber="3">ZSRO</collectionCode>
Sc-1249); same data except
<date id="FFCF103EFB76FFA4FC9FF0EBFC6FEDCF" box="[830,985,1668,1691]" pageId="4" pageNumber="3" value="2002-04-15">
<collectingDate id="EF8BE9D6FB76FFA4FC9FF0EBFC6FEDCF" box="[830,985,1668,1691]" pageId="4" pageNumber="3" value="2002-04-15">15 April 2002</collectingDate>
</date>
, leg
</materialsCitation>
.
<collectionCode id="ED60AE3BFB76FFA4FBB5F0EBFB9CEDCF" box="[1044,1066,1668,1691]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="3" type="Museum">V</collectionCode>
.
<materialsCitation id="3B193CA3FB76FFA4FB97F0EBFC1DEC07" collectingDate="1904-07-10" collectingDateMax="2002-04-06" collectingDateMin="1904-07-10" collectionCode="VFPC" collectorName="Fet &amp; K. Aris" country="Turkmenistan" county="Karakum Desert" elevation="188" latitude="35.681114" location="Tejen" longLatPrecision="19" longitude="62.02611" municipality="Akhal Province" pageId="4" pageNumber="3" specimenCode="ZISP 809" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Mary Welayaty">
<collectorName id="26845328FB76FFA4FB97F0EBFBEDEDCF" box="[1078,1115,1668,1691]" pageId="4" pageNumber="3">Fet</collectorName>
,
<specimenCount id="9D77FD77FB76FFA4FBC7F0EBFB67EDCF" box="[1126,1233,1668,1691]" pageId="4" pageNumber="3" type="juvenile">1 juvenile</specimenCount>
(
<collectionCode id="ED60AE3BFB76FFA4FB40F0EBFA9CEDCF" box="[1249,1322,1668,1691]" pageId="4" pageNumber="3">VFPC</collectionCode>
);
<collectingCounty id="62AF4E72FB76FFA4FA9CF0EBFC30EDED" pageId="4" pageNumber="3">Karakum Desert</collectingCounty>
,
<collectingMunicipality id="6BAAAC84FB76FFA4FC37F0CDFBF2EDED" box="[918,1092,1698,1721]" pageId="4" pageNumber="3">Akhal Province</collectingMunicipality>
,
<location id="8EAE6025FB76FFA4FBF2F0CDFB26EDED" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB76FFA4FBF2F0CDFB26EDED" box="[1107,1168,1698,1721]" country="Turkmenistan" county="Karakum Desert" latitude="35.681114" longLatPrecision="19" longitude="62.02611" municipality="Akhal Province" name="Tejen" pageId="4" pageNumber="3" stateProvince="Mary Welayaty">Tejen</location>
(=
<location id="8EAE6025FB76FFA4FB1CF0CDFAADEDED" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB76FFA4FB1CF0CDFAADEDED" box="[1213,1307,1698,1721]" country="Turkmenistan" county="Karakum Desert" latitude="35.681114" longLatPrecision="19" longitude="62.02611" municipality="Akhal Province" name="Tedzhen" pageId="4" pageNumber="3" stateProvince="Mary Welayaty">Tedzhen</location>
,
<location id="8EAE6025FB76FFA4FA8BF0CDFA23EDED" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB76FFA4FA8BF0CDFA23EDED" box="[1322,1429,1698,1721]" country="Turkmenistan" county="Karakum Desert" latitude="35.681114" longLatPrecision="19" longitude="62.02611" municipality="Akhal Province" name="Tedschen" pageId="4" pageNumber="3" stateProvince="Mary Welayaty">Tedschen</location>
),
<geoCoordinate id="EE455039FB76FFA4FC9FF0AEFC0AED8C" box="[830,956,1729,1752]" degrees="37" direction="north" minutes="22" orientation="latitude" pageId="4" pageNumber="3" precision="15" seconds="39" value="37.3775">37°22'39&quot;N</geoCoordinate>
<geoCoordinate id="EE455039FB76FFA4FC66F0AEFBF2ED8C" box="[967,1092,1729,1752]" degrees="60" direction="east" minutes="29" orientation="longitude" pageId="4" pageNumber="3" precision="15" seconds="52" value="60.49778">60°29'52&quot;E</geoCoordinate>
,
<elevation id="005CD1CDFB76FFA4FBF3F0AEFB60ED8C" box="[1106,1238,1729,1752]" metricMagnitude="2" metricUnit="m" metricValue="1.88" pageId="4" pageNumber="3" unit="m" value="188.0">
<quantity id="4C899B1BFB76FFA4FBF3F0AEFB2CED8C" box="[1106,1178,1729,1752]" metricMagnitude="2" metricUnit="m" metricValue="1.88" pageId="4" pageNumber="3" unit="m" value="188.0">188 m</quantity>
a.s.l.
</elevation>
,
<date id="FFCF103EFB76FFA4FB47F0AEFA17ED8C" box="[1254,1441,1729,1752]" pageId="4" pageNumber="3" value="1904-07-10" valueMax="1904-07-24" valueMin="1904-07-10">
<collectingDate id="EF8BE9D6FB76FFA4FB47F0AEFA17ED8C" box="[1254,1441,1729,1752]" pageId="4" pageNumber="3" value="1904-07-10" valueMax="1904-07-24" valueMin="1904-07-10">10–24 July 1904</collectingDate>
</date>
[23 July–6 August], leg.
<collectorName id="26845328FB76FFA4FBF0F08FFB12EDA3" box="[1105,1188,1760,1783]" pageId="4" pageNumber="3">K. Aris</collectorName>
,
<specimenCount id="9D77FD77FB76FFA4FB13F08FFB65EDA3" box="[1202,1235,1760,1783]" pageId="4" pageNumber="3" type="male">1♂</specimenCount>
(
<specimenCode id="DBD79E85FB76FFA4FB45F08FFAE7EDA3" box="[1252,1361,1760,1783]" collectionCode="ZISP" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="4" pageNumber="3">ZISP 809</specimenCode>
);
<collectingRegion id="49B5F81CFB76FFA4FAC6F08FFC16EC41" country="Turkmenistan" name="Mary Welayaty" pageId="4" pageNumber="3">Mary Province</collectingRegion>
,
<location id="8EAE6025FB76FFA4FC0FF091FB39EC41" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB76FFA4FC0FF091FB39EC41" box="[942,1167,1790,1813]" country="Turkmenistan" county="Karakum Desert" latitude="35.681114" longLatPrecision="19" longitude="62.02611" municipality="Akhal Province" name="Serkhetabad District" pageId="4" pageNumber="3" stateProvince="Mary Welayaty">Serkhetabad District</location>
,
<location id="8EAE6025FB76FFA4FB3CF091FAE1EC41" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB76FFA4FB3CF091FAE1EC41" box="[1181,1367,1790,1813]" country="Turkmenistan" county="Karakum Desert" latitude="35.681114" longLatPrecision="19" longitude="62.02611" municipality="Akhal Province" name="Badghyz Plateau" pageId="4" pageNumber="3" stateProvince="Mary Welayaty">Badghyz Plateau</location>
,
<location id="8EAE6025FB76FFA4FAC4F091FCC4EC60" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB76FFA4FAC4F091FCC4EC60" country="Turkmenistan" county="Karakum Desert" latitude="35.681114" longLatPrecision="19" longitude="62.02611" municipality="Akhal Province" name="Chainury" pageId="4" pageNumber="3" stateProvince="Mary Welayaty">Chainury</location>
,
<geoCoordinate id="EE455039FB76FFA4FC27F172FBB2EC60" box="[902,1028,1821,1844]" degrees="35" direction="north" minutes="40" orientation="latitude" pageId="4" pageNumber="3" precision="15" seconds="52" value="35.681114">35°40'52&quot;N</geoCoordinate>
<geoCoordinate id="EE455039FB76FFA4FBB5F172FB27EC60" box="[1044,1169,1821,1844]" degrees="62" direction="east" minutes="01" orientation="longitude" pageId="4" pageNumber="3" precision="15" seconds="34" value="62.02611">62°01'34&quot;E</geoCoordinate>
,
<elevation id="005CD1CDFB76FFA4FB04F172FA82EC60" box="[1189,1332,1821,1844]" metricMagnitude="2" metricUnit="m" metricValue="4.58" pageId="4" pageNumber="3" unit="m" value="458.0">
<quantity id="4C899B1BFB76FFA4FB04F172FB47EC60" box="[1189,1265,1821,1844]" metricMagnitude="2" metricUnit="m" metricValue="4.58" pageId="4" pageNumber="3" unit="m" value="458.0">458 m</quantity>
a.s.l.
</elevation>
,
<date id="FFCF103EFB76FFA4FAEAF172FCC0EC07" pageId="4" pageNumber="3" value="2002-04-06">
<collectingDate id="EF8BE9D6FB76FFA4FAEAF172FCC0EC07" pageId="4" pageNumber="3" value="2002-04-06">6 April 2002</collectingDate>
</date>
, leg
</materialsCitation>
.
<materialsCitation id="3B193CA3FB76FFA4FC1AF153FBF1EC07" box="[955,1095,1852,1875]" collectionCode="V" country="Turkmenistan" location="Fet" pageId="4" pageNumber="3" specimenCount="1">
<collectionCode id="ED60AE3BFB76FFA4FC1AF153FC67EC07" box="[955,977,1852,1875]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="3" type="Museum">V</collectionCode>
.
<location id="8EAE6025FB76FFA4FC40F153FBB5EC07" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB76FFA4FC40F153FBB5EC07" box="[993,1027,1852,1875]" country="Turkmenistan" name="Fet" pageId="4" pageNumber="3">Fet</location>
&amp; A
</materialsCitation>
.
<collectionCode id="ED60AE3BFB76FFA4FBF6F153FBDBEC07" box="[1111,1133,1852,1875]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="3" type="Museum">V</collectionCode>
.
<materialsCitation id="3B193CA3FB76FFA4FBDFF153FC29EC25" collectionCode="FKCP, NMPC" collectorName="Gromov" country="Turkmenistan" pageId="4" pageNumber="3" specimenCount="2" specimenCount-female="2">
<collectorName id="26845328FB76FFA4FBDFF153FB6CEC07" box="[1150,1242,1852,1875]" pageId="4" pageNumber="3">Gromov</collectorName>
,
<specimenCount id="9D77FD77FB76FFA4FB48F153FABDEC07" box="[1257,1291,1852,1875]" pageId="4" pageNumber="3" type="female">1♀</specimenCount>
(
<collectionCode id="ED60AE3BFB76FFA4FABEF153FAD1EC07" box="[1311,1383,1852,1875]" pageId="4" pageNumber="3">FKCP</collectionCode>
),
<specimenCount id="9D77FD77FB76FFA4FADEF153FA16EC07" box="[1407,1440,1852,1875]" pageId="4" pageNumber="3" type="female">1♀</specimenCount>
(
<collectionCode id="ED60AE3BFB76FFA4FCE7F135FC2EEC25" box="[838,920,1882,1905]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="3">NMPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF0E6676FB77FFA5FF61F044FCA0ED27" pageId="5" pageNumber="4" startId="5.[192,278,1579,1602]" targetBox="[191,1440,191,1550]" targetPageId="5">
<paragraph id="8BCE36FEFB77FFA5FF61F044FCA0ED27" blockId="5.[192,1441,1579,1651]" pageId="5" pageNumber="4">
<emphasis id="B905EAECFB77FFA5FF61F044FEFBED16" bold="true" box="[192,333,1579,1602]" pageId="5" pageNumber="4">Figures 1–3:</emphasis>
<taxonomicName id="4C714D7DFB77FFA5FEF5F042FDFDED16" authorityName="Kraepelin" authorityYear="1900" box="[340,587,1581,1602]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="rickmersi">
<emphasis id="B905EAECFB77FFA5FEF5F042FDFDED16" box="[340,587,1581,1602]" italics="true" pageId="5" pageNumber="4">Anomalobuthus rickmersi</emphasis>
</taxonomicName>
, adult male holotype: full-body views, dorsal (1) and ventral (2), plus original labels (3). Scale bar = 10 mm (1–2). The red inset at bottom right shows the label of the specimen examined by
<bibRefCitation id="EFE04B0FFB77FFA5FB52F029FA2AED0F" author="LOURENCO" box="[1267,1436,1606,1627]" pageId="5" pageNumber="4" pagination="15 - 20" refId="ref25502" refString="LOURENCO, W. R. 2001. Un nouveau genre de Buthidae, probable vicariant geographique d'Anomalobuthus Kraepelin (Chelicerata, Scorpiones). Biogeographica, 77 (1): 15 - 20." type="journal article" year="2001">Lourenço (2001)</bibRefCitation>
, demonstrating it was not the actual Rickmers's type specimen.
</paragraph>
</caption>
<subSubSection id="C36B6575FB77FFA7FF51F0C7FD7CEDA6" lastPageId="7" lastPageNumber="6" pageId="5" pageNumber="4" type="diagnosis">
<paragraph id="8BCE36FEFB77FFA7FF51F0C7FD7CEDA6" blockId="5.[192,802,1704,1911]" lastBlockId="7.[192,802,1693,1778]" lastPageId="7" lastPageNumber="6" pageId="5" pageNumber="4">
DIAGNOSIS (emended). Adult size
<quantity id="4C899B1BFB77FFA5FD22F0C7FD4BEDEB" box="[643,765,1704,1727]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" metricValueMax="3.4" metricValueMin="3.0" pageId="5" pageNumber="4" unit="mm" value="32.0" valueMax="34.0" valueMin="30.0">30–34 mm</quantity>
in males,
<quantity id="4C899B1BFB77FFA5FEB1F0A8FE3EED8A" box="[272,392,1735,1758]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="3.7" metricValueMin="3.2" pageId="5" pageNumber="4" unit="mm" value="34.5" valueMax="37.0" valueMin="32.0">32–37 mm</quantity>
in females. Coloration light yellow, essentially immaculate, only with some regular blackish spots on pedipalps and metasoma (darker and more extended in female); metasomal segment V and telson reddish brown (male) to black (female). Pedipalp fingers with 9–10 principal rows of denticles and 7–9 internal accessory denticles. Pectines with 23–27 teeth in males and
<quantity id="4C899B1BFB77FFA5FCCEF0A8FC66ED8A" box="[879,976,1735,1758]" metricMagnitude="-1" metricUnit="m" metricValue="5.461" metricValueMax="5.842" metricValueMin="5.08" pageId="5" pageNumber="4" unit="in" value="21.5" valueMax="23.0" valueMin="20.0">20–23 in</quantity>
females. Tibial spur variable from absent (specimens from
<collectingCountry id="F366766EFB77FFA5FBADF089FB14EDA9" box="[1036,1186,1766,1789]" name="Turkmenistan" pageId="5" pageNumber="4">Turkmenistan</collectingCountry>
) to present but very small on leg IV only (male
<typeStatus id="54CA885CFB77FFA5FBDFF16BFB6DEC4F" box="[1150,1243,1796,1819]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
from
<collectingCountry id="F366766EFB77FFA5FA87F16BFA27EC4F" box="[1318,1425,1796,1819]" name="Tajikistan" pageId="5" pageNumber="4">Tajikistan</collectingCountry>
). Metasoma conspicuously attenuate, with most carinae obsolete to vestigial; ventral lateral carinae of segment V composed of denticles narrowly conical (male) to narrowly lobate (female), dorsal lateral and lateral supramedian carinae of segments I–IV with terminal denticle enlarged; intercarinal spaces essentially smooth on segments I–III, sparsely granulose on ventral and lateral areas of IV and very densely granulose on ventral and lateral areas of V (granulation sparser in female). Telson vesicle elongate oval and sparsely setose.
</paragraph>
</subSubSection>
<caption id="DF0E6676FB74FFA6FF61F0A9FCDAEDA1" pageId="6" pageNumber="5" startId="6.[192,278,1734,1757]" targetBox="[207,1403,201,1681]" targetPageId="6">
<paragraph id="8BCE36FEFB74FFA6FF61F0A9FCDAEDA1" blockId="6.[192,1440,1734,1781]" pageId="6" pageNumber="5">
<emphasis id="B905EAECFB74FFA6FF61F0A9FEFBED89" bold="true" box="[192,333,1734,1757]" pageId="6" pageNumber="5">Figures 4–7:</emphasis>
<taxonomicName id="4C714D7DFB74FFA6FEF5F0A8FDFDED88" authorityName="Kraepelin" authorityYear="1900" box="[340,587,1735,1756]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="rickmersi">
<emphasis id="B905EAECFB74FFA6FEF5F0A8FDFDED88" box="[340,587,1735,1756]" italics="true" pageId="6" pageNumber="5">Anomalobuthus rickmersi</emphasis>
</taxonomicName>
from Turkmenistan: adult male from Repetek (4–5) and adult female from Chainury (6– 7), full-body views, dorsal (4, 6) and ventral (5, 7). Scale bar = 10 mm.
</paragraph>
</caption>
<caption id="DF0E6676FB75FFA7FF61F068FF59ED3C" pageId="7" pageNumber="6" startId="7.[192,278,1543,1566]" targetBox="[191,1437,191,1514]" targetPageId="7">
<paragraph id="8BCE36FEFB75FFA7FF61F068FF59ED3C" blockId="7.[192,1440,1543,1640]" pageId="7" pageNumber="6">
<emphasis id="B905EAECFB75FFA7FF61F068FEEDED4A" bold="true" box="[192,347,1543,1566]" pageId="7" pageNumber="6">Figures 8–13:</emphasis>
<taxonomicName id="4C714D7DFB75FFA7FEC3F066FDD6ED4A" authority="rickmersi." authorityName="Rickmersi." box="[354,608,1545,1566]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAECFB75FFA7FEC3F066FDECED4A" box="[354,602,1545,1566]" italics="true" pageId="7" pageNumber="6">Anomalobuthus rickmersi</emphasis>
.
</taxonomicName>
<emphasis id="B905EAECFB75FFA7FDCCF066FD5DED4A" bold="true" box="[621,747,1545,1566]" pageId="7" pageNumber="6">Figures 8–9.</emphasis>
Juvenile male from Repetek, Turkmenistan: full-body views, dorsal (8) and ventral (9). Scale bar = 10 mm.
<emphasis id="B905EAECFB75FFA7FD90F04DFD7FED63" bold="true" box="[561,713,1570,1591]" pageId="7" pageNumber="6">Figures 10–13.</emphasis>
Adult male holotype: chelicerae, carapace and tergites I–III, dorsal (10), sternopectinal region and sternites III–V, ventral (11), distal part of left leg III, internal (12), distal part of left leg IV, internal (13).
</paragraph>
</caption>
<subSubSection id="C36B6575FB75FFABFF51F177FEA5EA87" lastPageId="11" lastPageNumber="10" pageId="7" pageNumber="6" type="description">
<paragraph id="8BCE36FEFB75FFA8FF51F177FEF4E91A" blockId="7.[192,802,1816,1900]" lastBlockId="8.[192,802,199,1847]" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
REDESCRIPTION (adult male
<typeStatus id="54CA885CFB75FFA7FD8FF177FD3DEC7B" box="[558,651,1816,1839]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
).
<emphasis id="B905EAECFB75FFA7FD06F177FC94EC7B" bold="true" box="[679,802,1816,1839]" pageId="7" pageNumber="6">Coloration</emphasis>
(
<figureCitation id="134A2A7BFB75FFA7FF66F159FE9BEC19" box="[199,301,1846,1869]" captionStart="Figures 1–3" captionStartId="5.[192,278,1579,1602]" captionTargetBox="[191,1440,191,1550]" captionTargetId="figure-6@5.[192,1439,192,1549]" captionTargetPageId="5" captionText="Figures 1–3: Anomalobuthus rickmersi, adult male holotype: full-body views, dorsal (1) and ventral (2), plus original labels (3). Scale bar = 10 mm (1–2). The red inset at bottom right shows the label of the specimen examined by Lourenço (2001), demonstrating it was not the actual Rickmers's type specimen." pageId="7" pageNumber="6">Figs. 1–2</figureCitation>
) moderately faded and translucent due to long preservation, but still reliably traceable. Base light yellowish; in general, the base color is paler on pedipalp chelae, legs, and pectines. Chelicerae immaculate, except for dark brown finger teeth. Pedipalp femur and patella with internal surface infuscate; chela immaculate, only with finger denticles dark brown. Carapace immaculate, only with a blackish spot under every ocular group. Tergites immaculate. Coxosternal region and genital operculum immaculate. Pectines pale yellowish, immaculate. Sternites immaculate. Legs essentially immaculate; claws with distal half dark brown. Metasoma not conspicuously bicolor, only becoming progressively darker and redder distally, segments I–IV with a diffusely annulated appearance (basal and distal parts of each segment faintly infuscate as thin brown rings, which become larger and more diffuse dorsally) and segment V faintly infuscate, becoming progressively darker and redder distally; carinae not infuscate nor underlined with dark pigment. Telson vesicle reddish brown, aculeus with basal half yellowish and distal half dark brown.
</paragraph>
<paragraph id="8BCE36FEFB7AFFA8FF51F439FEA7E9FE" blockId="8.[192,802,199,1847]" pageId="8" pageNumber="7">
<emphasis id="B905EAECFB7AFFA8FF51F439FEDCE939" bold="true" box="[240,362,598,621]" pageId="8" pageNumber="7">Chelicerae</emphasis>
(
<figureCitation id="134A2A7BFB7AFFA8FEDBF439FE7AE939" box="[378,460,598,621]" captionStart="Figures 8–13" captionStartId="7.[192,278,1543,1566]" captionTargetBox="[191,1437,191,1514]" captionTargetId="figure-6@7.[192,1437,192,1513]" captionTargetPageId="7" captionText="Figures 8–13: Anomalobuthus rickmersi. Figures 8–9. Juvenile male from Repetek, Turkmenistan: full-body views, dorsal (8) and ventral (9). Scale bar = 10 mm. Figures 10–13. Adult male holotype: chelicerae, carapace and tergites I–III, dorsal (10), sternopectinal region and sternites III–V, ventral (11), distal part of left leg III, internal (12), distal part of left leg IV, internal (13)." pageId="8" pageNumber="7">Fig. 10</figureCitation>
). With dentition typical for the genus, as described for
<taxonomicName id="4C714D7DFB7AFFA8FE7FF41BFD25E9DF" authority="Teruel &amp; Kovařík &amp; Fet, 2018" authorityName="Teruel &amp; Kovařík &amp; Fet" authorityYear="2018" box="[478,659,628,651]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="krivochatskyi" status="sp. nov.">
<emphasis id="B905EAECFB7AFFA8FE7FF41BFD25E9DF" box="[478,659,628,651]" italics="true" pageId="8" pageNumber="7">A. krivochatskyi</emphasis>
</taxonomicName>
<emphasis id="B905EAECFB7AFFA8FD03F41BFD51E9DF" bold="true" box="[674,743,628,651]" pageId="8" pageNumber="7">
<taxonomicNameLabel id="A2365797FB7AFFA8FD03F41BFD51E9DF" box="[674,743,628,651]" pageId="8" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(see below).
</paragraph>
<paragraph id="8BCE36FEFB7AFFA8FF51F4DDFEAAEE7A" blockId="8.[192,802,199,1847]" pageId="8" pageNumber="7">
<emphasis id="B905EAECFB7AFFA8FF51F4DDFEE9E99D" bold="true" box="[240,351,690,713]" pageId="8" pageNumber="7">Pedipalps</emphasis>
(
<figureCitation id="134A2A7BFB7AFFA8FED3F4DDFE4EE99D" box="[370,504,690,713]" captionStart="Figures 14–26" captionStartId="9.[192,278,1609,1632]" captionTargetBox="[196,1409,215,1573]" captionTargetId="figure-6@9.[192,1439,192,1580]" captionTargetPageId="9" captionText="Figures 14–26: Anomalobuthus rickmersi, adult male holotype: right pedipalp trochanter and femur, dorsal (14) and ventral (15), right pedipalp patella, dorsal (16), external (17) and ventral (18), right pedipalp chela, dorsal (19–20), external (21) and ventrointernal (22), metasoma and telson, lateral (23), dorsal (24) and ventral (25), telson, lateral (26)." pageId="8" pageNumber="7">Figs. 14–22</figureCitation>
). Relatively short but very slender, essentially bare. Femur subtly curved inwards, with carinae weak, granulose to subdenticulate; intercarinal tegument smooth and glossy. Patella straight, with carinae obsolete to absent, smooth except on internal surface where vestigially denticulate; intercarinal tegument smooth and glossy. Chela elongate and very slender; manus conspicuously narrower than patella (ratio 0.79), subcylindrical (1.80 times longer than wide, 1.07 times wider than deep), with carinae weak to moderate, smooth; intercarinal tegument smooth and glossy; fingers very long (movable finger 2.44 times longer than underhand), only subtly curved and with 10 principal rows of denticles (the two basalmost rows are moderately well-defined), basal lobe/notch combination absent, external accessory denticles absent, internal accessory denticles very large and claw-like (increasing in size distally), numbering eight on both fixed and movable fingers, movable finger with one claw-like accessory denticle basal to the very large terminal denticle.
</paragraph>
<paragraph id="8BCE36FEFB7AFFA8FF51F359FDCBEDCA" blockId="8.[192,802,199,1847]" pageId="8" pageNumber="7">
<emphasis id="B905EAECFB7AFFA8FF51F359FEE8EE19" bold="true" box="[240,350,1334,1357]" pageId="8" pageNumber="7">Carapace</emphasis>
(
<figureCitation id="134A2A7BFB7AFFA8FEDBF359FE61EE19" box="[378,471,1334,1357]" captionStart="Figures 8–13" captionStartId="7.[192,278,1543,1566]" captionTargetBox="[191,1437,191,1514]" captionTargetId="figure-6@7.[192,1437,192,1513]" captionTargetPageId="7" captionText="Figures 8–13: Anomalobuthus rickmersi. Figures 8–9. Juvenile male from Repetek, Turkmenistan: full-body views, dorsal (8) and ventral (9). Scale bar = 10 mm. Figures 10–13. Adult male holotype: chelicerae, carapace and tergites I–III, dorsal (10), sternopectinal region and sternites III–V, ventral (11), distal part of left leg III, internal (12), distal part of left leg IV, internal (13)." pageId="8" pageNumber="7">Fig. 10</figureCitation>
). Very strongly trapezoidal (much narrower anteriorly) and wider than long; anterior margin shallowly convex, with 5–6 pairs of thin macrosetae and some very short microsetae. Carination essentially absent: the only carinae present are the superciliaries, which are smooth. Furrows: anterior median, median ocular, central median, posterior median and posterior marginal fused, wide and moderately deep; lateral oculars, lateral centrals, central transverse, and posterior laterals long, narrow and shallow. Tegument very finely and densely granulose, with a few small to medium-sized granules scattered all over.
</paragraph>
<paragraph id="8BCE36FEFB7AFFA8FF51F0C9FD2CED8F" blockId="8.[192,802,199,1847]" pageId="8" pageNumber="7">
<emphasis id="B905EAECFB7AFFA8FF51F0C9FEE5EDE9" bold="true" box="[240,339,1702,1725]" pageId="8" pageNumber="7">Sternum</emphasis>
(
<figureCitation id="134A2A7BFB7AFFA8FECBF0C9FE77EDE9" box="[362,449,1702,1725]" captionStart="Figures 8–13" captionStartId="7.[192,278,1543,1566]" captionTargetBox="[191,1437,191,1514]" captionTargetId="figure-6@7.[192,1437,192,1513]" captionTargetPageId="7" captionText="Figures 8–13: Anomalobuthus rickmersi. Figures 8–9. Juvenile male from Repetek, Turkmenistan: full-body views, dorsal (8) and ventral (9). Scale bar = 10 mm. Figures 10–13. Adult male holotype: chelicerae, carapace and tergites I–III, dorsal (10), sternopectinal region and sternites III–V, ventral (11), distal part of left leg III, internal (12), distal part of left leg IV, internal (13)." pageId="8" pageNumber="7">Fig. 11</figureCitation>
). Standard for the genus, relatively small and widely triangular in shape.
</paragraph>
<paragraph id="8BCE36FEFB7AFFA8FF51F08CFDB9EC63" blockId="8.[192,802,199,1847]" pageId="8" pageNumber="7">
<emphasis id="B905EAECFB7AFFA8FF51F08CFE78EDAE" bold="true" box="[240,462,1763,1786]" pageId="8" pageNumber="7">Genital operculum</emphasis>
(
<figureCitation id="134A2A7BFB7AFFA8FE44F08CFD8DEDAE" box="[485,571,1763,1786]" captionStart="Figures 8–13" captionStartId="7.[192,278,1543,1566]" captionTargetBox="[191,1437,191,1514]" captionTargetId="figure-6@7.[192,1437,192,1513]" captionTargetPageId="7" captionText="Figures 8–13: Anomalobuthus rickmersi. Figures 8–9. Juvenile male from Repetek, Turkmenistan: full-body views, dorsal (8) and ventral (9). Scale bar = 10 mm. Figures 10–13. Adult male holotype: chelicerae, carapace and tergites I–III, dorsal (10), sternopectinal region and sternites III–V, ventral (11), distal part of left leg III, internal (12), distal part of left leg IV, internal (13)." pageId="8" pageNumber="7">Fig. 11</figureCitation>
). Slightly damaged. Relatively large, each half roundly subtriangular in shape. Genital papillae present.
</paragraph>
<paragraph id="8BCE36FEFB7AFFA8FCCFF6A8FA2CEA23" blockId="8.[830,1441,199,1847]" pageId="8" pageNumber="7">
<emphasis id="B905EAECFB7AFFA8FCCFF6A8FC7DEB8A" bold="true" box="[878,971,199,222]" pageId="8" pageNumber="7">Pectines</emphasis>
(
<figureCitation id="134A2A7BFB7AFFA8FC43F6A8FB8EEB8A" box="[994,1080,199,222]" captionStart="Figures 8–13" captionStartId="7.[192,278,1543,1566]" captionTargetBox="[191,1437,191,1514]" captionTargetId="figure-6@7.[192,1437,192,1513]" captionTargetPageId="7" captionText="Figures 8–13: Anomalobuthus rickmersi. Figures 8–9. Juvenile male from Repetek, Turkmenistan: full-body views, dorsal (8) and ventral (9). Scale bar = 10 mm. Figures 10–13. Adult male holotype: chelicerae, carapace and tergites I–III, dorsal (10), sternopectinal region and sternites III–V, ventral (11), distal part of left leg III, internal (12), distal part of left leg IV, internal (13)." pageId="8" pageNumber="7">Fig. 11</figureCitation>
). Standard-sized for the genus: very long, extending beyond leg IV coxa-trochanter joint), subrectangular and densely setose. Tooth count 25/25. Basal plate heavily sclerotized, much wider than long, anterior margin with a very deep, narrow anteromedian furrow, posterior margin very shallowly convex.
</paragraph>
<paragraph id="8BCE36FEFB7AFFA8FCCFF710FAF8EAA6" blockId="8.[830,1441,199,1847]" pageId="8" pageNumber="7">
<emphasis id="B905EAECFB7AFFA8FCCFF710FC15EAC2" bold="true" box="[878,931,383,406]" pageId="8" pageNumber="7">Legs</emphasis>
(
<figureCitation id="134A2A7BFB7AFFA8FC16F710FB8BEAC2" box="[951,1085,383,406]" captionStart="Figures 8–13" captionStartId="7.[192,278,1543,1566]" captionTargetBox="[191,1437,191,1514]" captionTargetId="figure-6@7.[192,1437,192,1513]" captionTargetPageId="7" captionText="Figures 8–13: Anomalobuthus rickmersi. Figures 8–9. Juvenile male from Repetek, Turkmenistan: full-body views, dorsal (8) and ventral (9). Scale bar = 10 mm. Figures 10–13. Adult male holotype: chelicerae, carapace and tergites I–III, dorsal (10), sternopectinal region and sternites III–V, ventral (11), distal part of left leg III, internal (12), distal part of left leg IV, internal (13)." pageId="8" pageNumber="7">Figs. 12–13</figureCitation>
). Very slender, with all carinae weak and subgranulose to subcostate; intercarinal tegument smooth and glossy. Tibial spurs entirely absent on legs III, reduced to a sinuous remnant on legs IV.
</paragraph>
<paragraph id="8BCE36FEFB7AFFA8FCCFF795FAD7E8AF" blockId="8.[830,1441,199,1847]" pageId="8" pageNumber="7">
<emphasis id="B905EAECFB7AFFA8FCCFF795FC50E945" bold="true" box="[878,998,506,529]" pageId="8" pageNumber="7">Mesosoma</emphasis>
(
<figureCitation id="134A2A7BFB7AFFA8FC5AF795FB32E945" box="[1019,1156,506,529]" captionStart="Figures 8–13" captionStartId="7.[192,278,1543,1566]" captionTargetBox="[191,1437,191,1514]" captionTargetId="figure-6@7.[192,1437,192,1513]" captionTargetPageId="7" captionText="Figures 8–13: Anomalobuthus rickmersi. Figures 8–9. Juvenile male from Repetek, Turkmenistan: full-body views, dorsal (8) and ventral (9). Scale bar = 10 mm. Figures 10–13. Adult male holotype: chelicerae, carapace and tergites I–III, dorsal (10), sternopectinal region and sternites III–V, ventral (11), distal part of left leg III, internal (12), distal part of left leg IV, internal (13)." pageId="8" pageNumber="7">Figs. 10–11</figureCitation>
). Tergites with the same sculpture as on carapace; I–VI irregularly tricarinate: the median longitudinal carina is weak, short, and formed by irregular medium-sized granules that do not project beyond posterior margin, but the submedian carinae are defined only by few coarse granules irregularly aligned; tergite VII with five well-defined carinae: the median carina is very short but strongly subserrate, the submedians and laterals are long, strong and finely serrate to denticulate. Sternites sparsely setose; III–VI glossy and with very subtle vestiges of smooth submedian carinae, spiracles relatively short and slit-like, almost transversely arranged (not strongly oblique), V with smooth patch absent; VII with two pairs of carinae: the submedians are long and finely crenulate to subcostate and the laterals are very short and subcrenulate, intercarinal tegument coriaceous to minutely granulose.
</paragraph>
<paragraph id="8BCE36FEFB7AFFA8FCCFF26CFB02ED35" blockId="8.[830,1441,199,1847]" pageId="8" pageNumber="7">
<emphasis id="B905EAECFB7AFFA8FCCFF26CFC52EF4E" bold="true" box="[878,996,1027,1050]" pageId="8" pageNumber="7">Metasoma</emphasis>
(
<figureCitation id="134A2A7BFB7AFFA8FC5AF26CFB31EF4E" box="[1019,1159,1027,1050]" captionStart="Figures 14–26" captionStartId="9.[192,278,1609,1632]" captionTargetBox="[196,1409,215,1573]" captionTargetId="figure-6@9.[192,1439,192,1580]" captionTargetPageId="9" captionText="Figures 14–26: Anomalobuthus rickmersi, adult male holotype: right pedipalp trochanter and femur, dorsal (14) and ventral (15), right pedipalp patella, dorsal (16), external (17) and ventral (18), right pedipalp chela, dorsal (19–20), external (21) and ventrointernal (22), metasoma and telson, lateral (23), dorsal (24) and ventral (25), telson, lateral (26)." pageId="8" pageNumber="7">Figs. 23–25</figureCitation>
). Elongated and slightly wider subdistally; with 10/10/10/8/5 complete to almost complete carinae, almost all obsolete to vestigial and smooth, with coarse punctations scattered: dorsal laterals very weak on I–III (with one terminal denticle enlarged), obsolete to vestigial as rounded ridges on IV, absent on V; lateral supramedians very weak on I–III (with one terminal denticle enlarged), obsolete to vestigial as rounded ridges on IV–V; lateral inframedians obsolete on I–III, absent on IV–V; ventral laterals obsolete on I– IV, weak to moderate on V, where become progressively stronger and somewhat flared distally, formed by sharply serrate, subequal denticles; ventral submedians obsolete on I–IV, absent on V; ventral median absent on I–IV, weak to moderate on V. Intercarinal tegument smooth and glossy, with minute but sharp granulation on lateral and ventral surfaces, which become much denser towards distal segments. Dorsal furrow obsolete on all segments. Setation sparse, mostly represented by 5–8 dark macrosetae over every carina.
</paragraph>
<paragraph id="8BCE36FEFB7AFFA9FCCFF007FDADEDA8" blockId="8.[830,1441,199,1847]" lastBlockId="9.[192,801,1734,1788]" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">
<emphasis id="B905EAECFB7AFFA8FCCFF007FC0CED2B" bold="true" box="[878,954,1640,1663]" pageId="8" pageNumber="7">Telson</emphasis>
(
<figureCitation id="134A2A7BFB7AFFA8FC6BF007FB90ED2B" box="[970,1062,1640,1663]" captionStart="Figures 14–26" captionStartId="9.[192,278,1609,1632]" captionTargetBox="[196,1409,215,1573]" captionTargetId="figure-6@9.[192,1439,192,1580]" captionTargetPageId="9" captionText="Figures 14–26: Anomalobuthus rickmersi, adult male holotype: right pedipalp trochanter and femur, dorsal (14) and ventral (15), right pedipalp patella, dorsal (16), external (17) and ventral (18), right pedipalp chela, dorsal (19–20), external (21) and ventrointernal (22), metasoma and telson, lateral (23), dorsal (24) and ventral (25), telson, lateral (26)." pageId="8" pageNumber="7">Figs. 26</figureCitation>
). Sparsely setose, with a few setae scattered on dorsal and lateral surfaces. Vesicle elongate oval (2.50 times longer than wide, 0.95 times wider than deep), tegument smooth and glossy, with vestiges of coarse granules arranged into three obsolete longitudinal carinae (ventral median and ventral submedians), and ventrally with some coarse punctations. Subaculear tubercle absent. Aculeus conspicuously shorter than vesicle, thick and shallowly curved.
</paragraph>
<caption id="DF0E6676FB7BFFA9FF61F026FB20EDC6" pageId="9" pageNumber="8" startId="9.[192,278,1609,1632]" targetBox="[196,1409,215,1573]" targetPageId="9">
<paragraph id="8BCE36FEFB7BFFA9FF61F026FB20EDC6" blockId="9.[192,1440,1609,1682]" pageId="9" pageNumber="8">
<emphasis id="B905EAECFB7BFFA9FF61F026FEDCED34" bold="true" box="[192,362,1609,1632]" pageId="9" pageNumber="8">Figures 14–26:</emphasis>
<taxonomicName id="4C714D7DFB7BFFA9FED5F024FDD8ED34" authorityName="Kraepelin" authorityYear="1900" box="[372,622,1611,1632]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="rickmersi">
<emphasis id="B905EAECFB7BFFA9FED5F024FDD8ED34" box="[372,622,1611,1632]" italics="true" pageId="9" pageNumber="8">Anomalobuthus rickmersi</emphasis>
</taxonomicName>
, adult male holotype: right pedipalp trochanter and femur, dorsal (14) and ventral (15), right pedipalp patella, dorsal (16), external (17) and ventral (18), right pedipalp chela, dorsal (19–20), external (21) and ventrointernal (22), metasoma and telson, lateral (23), dorsal (24) and ventral (25), telson, lateral (26).
</paragraph>
</caption>
<paragraph id="8BCE36FEFB7BFFA9FF51F14DFA2AEC23" blockId="9.[192,802,1826,1911]" lastBlockId="9.[830,1441,1734,1911]" pageId="9" pageNumber="8">
FEMALE (Chainury,
<collectingCountry id="F366766EFB7BFFA9FE6FF14DFDD1EC6D" box="[462,615,1826,1849]" name="Turkmenistan" pageId="9" pageNumber="8">Turkmenistan</collectingCountry>
). Very similar to male, sexual dimorphism evident by:
<emphasis id="B905EAECFB7BFFA9FD29F12EFD28EC0C" bold="true" box="[648,670,1857,1880]" pageId="9" pageNumber="8">1)</emphasis>
size comparatively larger inside each size-class;
<emphasis id="B905EAECFB7BFFA9FD31F10FFD11EC23" bold="true" box="[656,679,1888,1911]" pageId="9" pageNumber="8">2)</emphasis>
coloration darker and much more sharply patterned, e.g., pedipalp femur with a conspicuous blackish spot on dorsodistal apex, which diffusely continues as a broad stripe over almost all internal surface, patella with internal surface blackish, carapace with anterior margin deeply infuscate, metasomal segment V and telson black;
<emphasis id="B905EAECFB7BFFA9FAA9F10FFAA8EC23" bold="true" box="[1288,1310,1888,1911]" pageId="9" pageNumber="8">3)</emphasis>
mesosoma
</paragraph>
<paragraph id="8BCE36FEFB78FFAAFE81F606FA16EBD1" blockId="10.[288,1440,103,133]" box="[288,1440,103,133]" pageId="10" pageNumber="9">
<emphasis id="B905EAECFB78FFAAFE81F606FE62EBD1" bold="true" box="[288,468,105,133]" pageId="10" pageNumber="9">Teruel, Kova</emphasis>
ř
<emphasis id="B905EAECFB78FFAAFE43F606FA16EBD1" bold="true" box="[482,1440,103,133]" pageId="10" pageNumber="9">
ík &amp; Fet: Revision of Genus
<taxonomicName id="4C714D7DFB78FFAAFCCCF608FBF7EBD0" box="[877,1089,103,132]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAECFB78FFAAFCCCF608FBF7EBD0" bold="true" box="[877,1089,103,132]" italics="true" pageId="10" pageNumber="9">Anomalobuthus</emphasis>
</taxonomicName>
9
</emphasis>
</paragraph>
<caption id="DF0E6676FB78FFAAFF61F31DFB4FEEF6" pageId="10" pageNumber="9" startId="10.[192,278,1394,1417]" targetBox="[192,1437,192,1364]" targetPageId="10">
<paragraph id="8BCE36FEFB78FFAAFF61F31DFB4FEEF6" blockId="10.[192,1440,1394,1442]" pageId="10" pageNumber="9">
<emphasis id="B905EAECFB78FFAAFF61F31DFED8EEDD" bold="true" box="[192,366,1394,1417]" pageId="10" pageNumber="9">Figures 27–33:</emphasis>
<taxonomicName id="4C714D7DFB78FFAAFEDBF31BFDCEEEDD" authorityName="Kraepelin" authorityYear="1900" box="[378,632,1396,1417]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="rickmersi">
<emphasis id="B905EAECFB78FFAAFEDBF31BFDCEEEDD" box="[378,632,1396,1417]" italics="true" pageId="10" pageNumber="9">Anomalobuthus rickmersi</emphasis>
</taxonomicName>
from Turkmenistan, adult male from Repetek (27–30) and adult female from Chainury (31–33): metasoma and telson, lateral (27, 31), dorsal (28, 32) and ventral (29, 33), telson, lateral (30).
</paragraph>
</caption>
<paragraph id="8BCE36FEFB78FFAAFF61F3B8FEBFED3D" blockId="10.[192,802,1495,1641]" pageId="10" pageNumber="9">
and metasoma slightly less slender;
<emphasis id="B905EAECFB78FFAAFDF9F3B8FDD8EEBA" bold="true" box="[600,622,1495,1518]" pageId="10" pageNumber="9">4)</emphasis>
genital papillae absent;
<emphasis id="B905EAECFB78FFAAFEB2F399FE9FED59" bold="true" box="[275,297,1526,1549]" pageId="10" pageNumber="9">5)</emphasis>
pectines smaller, with consistently lower tooth counts;
<emphasis id="B905EAECFB78FFAAFEB5F07BFE9CED7F" bold="true" box="[276,298,1556,1579]" pageId="10" pageNumber="9">6)</emphasis>
metasomal segment V with lateral intercarinal tegument less granulose. See
<figureCitation id="134A2A7BFB78FFAAFE5FF05CFDD7ED1E" box="[510,609,1587,1610]" captionStart="Figures 4–7" captionStartId="6.[192,278,1734,1757]" captionTargetBox="[207,1403,201,1681]" captionTargetId="figure-12@6.[192,1439,188,1704]" captionTargetPageId="6" captionText="Figures 4–7: Anomalobuthus rickmersi from Turkmenistan: adult male from Repetek (4–5) and adult female from Chainury (6– 7), full-body views, dorsal (4, 6) and ventral (5, 7). Scale bar = 10 mm." pageId="10" pageNumber="9">Figs. 6–7</figureCitation>
,
<figureCitation id="134A2A7BFB78FFAAFDD1F05CFD04ED1E" box="[624,690,1587,1610]" captionStart="Figures 27–33" captionStartId="10.[192,278,1394,1417]" captionTargetBox="[192,1437,192,1364]" captionTargetId="figure-12@10.[192,1437,192,1364]" captionTargetPageId="10" captionText="Figures 27–33: Anomalobuthus rickmersi from Turkmenistan, adult male from Repetek (27–30) and adult female from Chainury (31–33): metasoma and telson, lateral (27, 31), dorsal (28, 32) and ventral (29, 33), telson, lateral (30)." pageId="10" pageNumber="9">31–33</figureCitation>
and
<tableCitation id="C6F30345FB78FFAAFD46F05CFF78ED3D" captionStart="Table 1" captionStartId="12.[192,257,1118,1141]" captionText="Table 1: Measurements of three adults of Anomalobuthus rickmersi. Abbreviations: length (L), width (W), posterior width (Wp), depth (D)." pageId="10" pageNumber="9">Tabs. 1</tableCitation>
,
<tableCitation id="C6F30345FB78FFAAFF7AF03DFEBFED3D" box="[219,265,1618,1641]" captionStart-0="Table 4" captionStart-1="Table 5" captionStartId-0="35.[192,257,1030,1053]" captionStartId-1="38.[192,257,680,703]" captionText-0="Table 4: Comparison between five species of Anomalobuthus based upon selected morphometric ratios. Abbreviations: length (L)," captionText-1="Table 5: Pectinal tooth count variation in all known species of Anomalobuthus. Abbreviations: number of pectines (N), average (Ave.), standard deviation (SD). This includes counts taken from sources with precise and reliable geographic data (Birula, 1905, 1910)." pageId="10" pageNumber="9">4–5.</tableCitation>
</paragraph>
<paragraph id="8BCE36FEFB78FFAAFF51F0E0FDD3ED91" blockId="10.[192,802,1679,1917]" pageId="10" pageNumber="9">
VARIATION. Adult size varied from
<quantity id="4C899B1BFB78FFAAFD22F0E0FC94EDF2" box="[643,802,1679,1702]" metricMagnitude="-2" metricUnit="m" metricValue="3.225" metricValueMax="3.41" metricValueMin="3.04" pageId="10" pageNumber="9" unit="mm" value="32.25" valueMax="34.1" valueMin="30.4">30.4–34.1 mm</quantity>
in males and
<quantity id="4C899B1BFB78FFAAFEECF0C1FE5FED91" box="[333,489,1710,1733]" metricMagnitude="-2" metricUnit="m" metricValue="3.4299999999999997" metricValueMax="3.65" metricValueMin="3.21" pageId="10" pageNumber="9" unit="mm" value="34.3" valueMax="36.5" valueMin="32.1">32.1–36.5 mm</quantity>
in females.
</paragraph>
<paragraph id="8BCE36FEFB78FFAAFF51F0A3FE5EEC75" blockId="10.[192,802,1679,1917]" pageId="10" pageNumber="9">
Pectinal tooth count varied as follows: 23–27 teeth in males and
<quantity id="4C899B1BFB78FFAAFEF7F084FE0EEC56" box="[342,440,1771,1794]" metricMagnitude="-1" metricUnit="m" metricValue="5.461" metricValueMax="5.842" metricValueMin="5.08" pageId="10" pageNumber="9" unit="in" value="21.5" valueMax="23.0" valueMin="20.0">20–23 in</quantity>
females, without clearly defined mode in either sex (
<tableCitation id="C6F30345FB78FFAAFE33F165FE6FEC75" box="[402,473,1802,1825]" captionStart="Table 5" captionStartId="38.[192,257,680,703]" captionText="Table 5: Pectinal tooth count variation in all known species of Anomalobuthus. Abbreviations: number of pectines (N), average (Ave.), standard deviation (SD). This includes counts taken from sources with precise and reliable geographic data (Birula, 1905, 1910)." pageId="10" pageNumber="9">Tab. 5</tableCitation>
).
</paragraph>
<paragraph id="8BCE36FEFB78FFAAFF51F147FB4BED3D" blockId="10.[192,802,1679,1917]" lastBlockId="10.[830,1440,1495,1641]" pageId="10" pageNumber="9">
<materialsCitation id="3B193CA3FB78FFAAFF51F147FB4BED3D" collectionCode="IV" country="Tajikistan" county="Tejen" location="Males" municipality="In" pageId="10" pageNumber="9" specimenCode="ZISP 809" specimenCount="1" typeStatus="holotype">
<location id="8EAE6025FB78FFAAFF51F147FE87EC6B" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB78FFAAFF51F147FE87EC6B" box="[240,305,1832,1855]" country="Tajikistan" county="Tejen" municipality="In" name="Males" pageId="10" pageNumber="9">Males</location>
from
<location id="8EAE6025FB78FFAAFEDFF147FE60EC6B" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB78FFAAFEDFF147FE60EC6B" box="[382,470,1832,1855]" country="Tajikistan" county="Tejen" municipality="In" name="Repetek" pageId="10" pageNumber="9">Repetek</location>
are virtually identical to the
<typeStatus id="54CA885CFB78FFAAFF61F128FEABEC0A" box="[192,285,1863,1886]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
in all diagnostic characters.
<location id="8EAE6025FB78FFAAFDF3F128FDCAEC0A" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB78FFAAFDF3F128FDCAEC0A" box="[594,636,1863,1886]" country="Tajikistan" county="Tejen" municipality="In" name="The" pageId="10" pageNumber="9">The</location>
only exception is that the tibial spurs are usually entirely absent.
<location id="8EAE6025FB78FFAAFD53F109FC97EC29" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB78FFAAFD53F109FC97EC29" box="[754,801,1894,1917]" country="Tajikistan" county="Tejen" municipality="In" name="This" pageId="10" pageNumber="9">This</location>
character alone is inappropriate to separate both populations taxonomically, especially when only a single specimen from
<collectingCountry id="F366766EFB78FFAAFC4FF07BFBEFED7F" box="[1006,1113,1556,1579]" name="Tajikistan" pageId="10" pageNumber="9">Tajikistan</collectingCountry>
is available.
<collectingMunicipality id="6BAAAC84FB78FFAAFB55F07BFABCED7F" box="[1268,1290,1556,1579]" pageId="10" pageNumber="9">In</collectingMunicipality>
addition, the male from
<collectingCounty id="62AF4E72FB78FFAAFC15F05CFC47ED1E" box="[948,1009,1587,1610]" pageId="10" pageNumber="9">Tejen</collectingCounty>
(
<specimenCode id="DBD79E85FB78FFAAFC5EF05CFBD0ED1E" box="[1023,1126,1587,1610]" collectionCode="ZISP" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="10" pageNumber="9">ZISP 809</specimenCode>
) has tibial spurs on both legs
<collectionCode id="ED60AE3BFB78FFAAFC9EF03DFCEDED3D" box="[831,859,1618,1641]" pageId="10" pageNumber="9">IV</collectionCode>
as noted already by Birula (1905: 450)
</materialsCitation>
</paragraph>
<paragraph id="8BCE36FEFB78FFABFCCFF0E0FE37EA4F" blockId="10.[830,1441,1679,1917]" lastBlockId="11.[192,803,199,467]" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9">
COMPARISON. Adults of
<taxonomicName id="4C714D7DFB78FFAAFB27F0E0FAB8EDF2" authorityName="Kraepelin" authorityYear="1900" box="[1158,1294,1679,1702]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="rickmersi">
<emphasis id="B905EAECFB78FFAAFB27F0E0FAB8EDF2" box="[1158,1294,1679,1702]" italics="true" pageId="10" pageNumber="9">A. rickmersi</emphasis>
</taxonomicName>
can be very easily distinguished by the greater slenderness of the metasoma, especially in males (
<figureCitation id="134A2A7BFB78FFAAFB13F0A3FAA9EDB7" box="[1202,1311,1740,1763]" captionStart="Figures 1–3" captionStartId="5.[192,278,1579,1602]" captionTargetBox="[191,1440,191,1550]" captionTargetId="figure-6@5.[192,1439,192,1549]" captionTargetPageId="5" captionText="Figures 1–3: Anomalobuthus rickmersi, adult male holotype: full-body views, dorsal (1) and ventral (2), plus original labels (3). Scale bar = 10 mm (1–2). The red inset at bottom right shows the label of the specimen examined by Lourenço (2001), demonstrating it was not the actual Rickmers's type specimen." pageId="10" pageNumber="9">Figs. 1–2</figureCitation>
,
<figureCitation id="134A2A7BFB78FFAAFA92F0A3FAEDEDB7" box="[1331,1371,1740,1763]" captionStart="Figures 4–7" captionStartId="6.[192,278,1734,1757]" captionTargetBox="[207,1403,201,1681]" captionTargetId="figure-12@6.[192,1439,188,1704]" captionTargetPageId="6" captionText="Figures 4–7: Anomalobuthus rickmersi from Turkmenistan: adult male from Repetek (4–5) and adult female from Chainury (6– 7), full-body views, dorsal (4, 6) and ventral (5, 7). Scale bar = 10 mm." pageId="10" pageNumber="9">4–7</figureCitation>
,
<figureCitation id="134A2A7BFB78FFAAFAD0F0A3FA2FEDB7" box="[1393,1433,1740,1763]" captionStart="Figures 107–119" captionStartId="33.[192,278,1734,1757]" captionTargetBox="[191,1440,216,1705]" captionTargetId="figure-6@33.[192,1439,217,1704]" captionTargetPageId="33" captionText="Figures 107–119: Anomalobuthus spp., from ZISP historical collection: adult male topotypes of A. pavlovskyi sp. n. (107– 112), full-body dorsal views (107, 110), labels (108, 111) and bottles (109, 112); adult male (113) and adult female (114) A. pavlovskyi sp. n. from Kunya-Urgench, Turkmenistan, full-body dorsal views (113, 114), labels (115) and bottle (116); adult male A. rickmersi from Tejen, Turkmenistan (117–119), full-body dorsal view (117), labels (118) and bottle (119). Photographs courtesy Alexander Koval." pageId="10" pageNumber="9">117</figureCitation>
;
<tableCitation id="C6F30345FB78FFAAFC9FF084FC26EC56" box="[830,912,1771,1794]" captionStart="Table 1" captionStartId="12.[192,257,1118,1141]" captionText="Table 1: Measurements of three adults of Anomalobuthus rickmersi. Abbreviations: length (L), width (W), posterior width (Wp), depth (D)." pageId="10" pageNumber="9">Tabs. 1</tableCitation>
,
<tableCitation id="C6F30345FB78FFAAFC01F084FC18EC56" box="[928,942,1771,1794]" captionStart="Table 4" captionStartId="35.[192,257,1030,1053]" captionText="Table 4: Comparison between five species of Anomalobuthus based upon selected morphometric ratios. Abbreviations: length (L)," pageId="10" pageNumber="9">4</tableCitation>
); even to unaided eye, most other congeners are more robust (
<figureCitation id="134A2A7BFB78FFAAFC57F165FBC0EC75" box="[1014,1142,1802,1825]" captionStart-0="Figures 34–37" captionStart-1="Figures 38–41" captionStart-2="Figures 42–45" captionStartId-0="13.[192,278,1630,1653]" captionStartId-1="14.[192,278,1754,1777]" captionStartId-2="16.[192,278,1584,1607]" captionTargetBox-0="[212,1415,153,1571]" captionTargetBox-1="[201,1418,199,1682]" captionTargetBox-2="[212,1422,194,1547]" captionTargetId-0="figure-6@13.[192,1439,144,1601]" captionTargetId-1="figure-12@14.[192,1439,192,1697]" captionTargetId-2="figure-12@16.[192,1440,192,1554]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="16" captionText-0="Figures 34–37: Anomalobuthus krivochatskyi sp. n., adult male holotype (34–35) and adult female paratopotype (36–37): fullbody views, dorsal (34, 36) and ventral (35, 37). Scale bar = 10 mm." captionText-1="Figures 38–41: Anomalobuthus krivochatskyi sp. n., adult male (38–39) and adult female paratypes (40–41) from between Bukhara and Gazli, Uzbekistan: full-body views, dorsal (38, 40) and ventral (39, 41). Scale bar = 10 mm." captionText-2="Figures 42–45: Anomalobuthus krivochatskyi sp. n., adult female (42–43) and juvenile male paratypes (44–45) from Chardara, Kazakhstan: full-body views, dorsal (42, 44) and ventral (43, 45)." pageId="10" pageNumber="9">Figs. 34–45</figureCitation>
,
<figureCitation id="134A2A7BFB78FFAAFB20F165FB71EC75" box="[1153,1223,1802,1825]" captionStart="Figures 65–74" captionStartId="22.[192,278,1590,1613]" captionTargetBox="[191,1437,215,1561]" captionTargetId="figure-12@22.[192,1437,216,1560]" captionTargetPageId="22" captionText="Figures 65–74: Anomalobuthus lowei sp. n., adult female holotype. Figures 65–66. full-body dorsal views, under white light (65) and under UV light (66). Figures 67–68. Mesosoma dorsal (67) and ventral (68) under UV light. Figure 69. Metasoma and telson under UV light, lateral. Figures 70–74. Pedipalp under UV light: right pedipalp trochanter and femur, dorsal (70), right pedipalp patella, dorsal (71), right pedipalp chela, dorsal (72), external (73) and ventral (74). Scale bars = 10 mm (65–66) and 1 mm (67–68, 70–74). Photographs courtesy Graeme Lowe." pageId="10" pageNumber="9">65–66</figureCitation>
,
<figureCitation id="134A2A7BFB78FFAAFB70F165FAA2EC75" box="[1233,1300,1802,1825]" captionStart="Figures 79–82" captionStartId="25.[192,278,1675,1698]" captionTargetBox="[244,1418,176,1638]" captionTargetId="figure-6@25.[192,1439,168,1645]" captionTargetPageId="25" captionText="Figures 79–82: Anomalobuthus pavlovskyi sp. n., adult male (79–80) and adult female paratopotypes (81–82): full-body views, dorsal (79, 81) and ventral (80, 82). Scale bar = 10 mm." pageId="10" pageNumber="9">79–82</figureCitation>
,
<figureCitation id="134A2A7BFB78FFAAFA80F165FAD1EC75" box="[1313,1383,1802,1825]" captionStart="Figures 90–91" captionStartId="30.[192,278,1499,1522]" captionTargetBox="[203,1424,204,1444]" captionTargetId="figure-12@30.[192,1439,192,1469]" captionTargetPageId="30" captionText="Figures 90–91: Anomalobuthus talebii, adult female holotype: full-body views, dorsal (90) and ventral (91)." pageId="10" pageNumber="9">90–91</figureCitation>
,
<figureCitation id="134A2A7BFB78FFAAFAD3F165FC6EEC6B" captionStart="Figures 107–119" captionStartId="33.[192,278,1734,1757]" captionTargetBox="[191,1440,216,1705]" captionTargetId="figure-6@33.[192,1439,217,1704]" captionTargetPageId="33" captionText="Figures 107–119: Anomalobuthus spp., from ZISP historical collection: adult male topotypes of A. pavlovskyi sp. n. (107– 112), full-body dorsal views (107, 110), labels (108, 111) and bottles (109, 112); adult male (113) and adult female (114) A. pavlovskyi sp. n. from Kunya-Urgench, Turkmenistan, full-body dorsal views (113, 114), labels (115) and bottle (116); adult male A. rickmersi from Tejen, Turkmenistan (117–119), full-body dorsal view (117), labels (118) and bottle (119). Photographs courtesy Alexander Koval." pageId="10" pageNumber="9">107, 110, 113–114</figureCitation>
;
<tableCitation id="C6F30345FB78FFAAFC4CF147FBE8EC6B" box="[1005,1118,1832,1855]" captionStart-0="Table 2" captionStart-1="Table 3" captionStart-2="Table 4" captionStartId-0="20.[192,257,1203,1226]" captionStartId-1="28.[192,257,1138,1161]" captionStartId-2="35.[192,257,1030,1053]" captionText-0="Table 2: Measurements of four types of Anomalobuthus krivochatskyi sp. n. Abbreviations: length (L), width (W), posterior width (Wp), depth (D)." captionText-1="Table 3: Measurements of the types of two new species of Anomalobuthus. Abbreviations: length (L), width (W), posterior width (Wp), depth (D)." captionText-2="Table 4: Comparison between five species of Anomalobuthus based upon selected morphometric ratios. Abbreviations: length (L)," pageId="10" pageNumber="9">Tabs. 2–4</tableCitation>
). For example, see the following male ratios: metasoma + telson length / carapace length = 7.76
<emphasis id="B905EAECFB78FFAAFC78F109FC40EC29" box="[985,1014,1894,1917]" italics="true" pageId="10" pageNumber="9">vs.</emphasis>
6.26–7.14, metasomal segments length / width = 1.74
<emphasis id="B905EAECFB79FFABFEC7F6A8FE32EB8A" box="[358,388,199,222]" italics="true" pageId="11" pageNumber="10">vs.</emphasis>
1.02–1.35 (I), 2.13
<emphasis id="B905EAECFB79FFABFDC0F6A8FDCAEB8A" box="[609,636,199,222]" italics="true" pageId="11" pageNumber="10">vs.</emphasis>
1.35–1.82 (II), 2.20
<emphasis id="B905EAECFB79FFABFF57F689FEA5EBA9" box="[246,275,230,253]" italics="true" pageId="11" pageNumber="10">vs.</emphasis>
1.39–1.88 (III), 2.56
<emphasis id="B905EAECFB79FFABFE5AF689FDAEEBA9" box="[507,536,230,253]" italics="true" pageId="11" pageNumber="10">vs.</emphasis>
1.75–2.23 (IV) and 2.71
<emphasis id="B905EAECFB79FFABFF61F76BFF68EA4F" box="[192,222,260,283]" italics="true" pageId="11" pageNumber="10">vs.</emphasis>
1.85–2.38 (V).
</paragraph>
<paragraph id="8BCE36FEFB79FFABFF51F74CFEA5EA87" blockId="11.[192,803,199,467]" pageId="11" pageNumber="10">
Moreover, this species is the only one that has metasomal segment V and telson light reddish-colored in adult males (
<figureCitation id="134A2A7BFB79FFABFEF1F70FFE00EA23" box="[336,438,352,375]" captionStart="Figures 1–3" captionStartId="5.[192,278,1579,1602]" captionTargetBox="[191,1440,191,1550]" captionTargetId="figure-6@5.[192,1439,192,1549]" captionTargetPageId="5" captionText="Figures 1–3: Anomalobuthus rickmersi, adult male holotype: full-body views, dorsal (1) and ventral (2), plus original labels (3). Scale bar = 10 mm (1–2). The red inset at bottom right shows the label of the specimen examined by Lourenço (2001), demonstrating it was not the actual Rickmers's type specimen." pageId="11" pageNumber="10">Figs. 1–2</figureCitation>
,
<figureCitation id="134A2A7BFB79FFABFE66F70FFE59EA23" box="[455,495,352,375]" captionStart="Figures 4–7" captionStartId="6.[192,278,1734,1757]" captionTargetBox="[207,1403,201,1681]" captionTargetId="figure-12@6.[192,1439,188,1704]" captionTargetPageId="6" captionText="Figures 4–7: Anomalobuthus rickmersi from Turkmenistan: adult male from Repetek (4–5) and adult female from Chainury (6– 7), full-body views, dorsal (4, 6) and ventral (5, 7). Scale bar = 10 mm." pageId="11" pageNumber="10">4–5</figureCitation>
,
<figureCitation id="134A2A7BFB79FFABFDA1F70FFDF3EA23" box="[512,581,352,375]" captionStart="Figures 14–26" captionStartId="9.[192,278,1609,1632]" captionTargetBox="[196,1409,215,1573]" captionTargetId="figure-6@9.[192,1439,192,1580]" captionTargetPageId="9" captionText="Figures 14–26: Anomalobuthus rickmersi, adult male holotype: right pedipalp trochanter and femur, dorsal (14) and ventral (15), right pedipalp patella, dorsal (16), external (17) and ventral (18), right pedipalp chela, dorsal (19–20), external (21) and ventrointernal (22), metasoma and telson, lateral (23), dorsal (24) and ventral (25), telson, lateral (26)." pageId="11" pageNumber="10">23–25</figureCitation>
,
<figureCitation id="134A2A7BFB79FFABFDF2F70FFD2EEA23" box="[595,664,352,375]" captionStart="Figures 27–33" captionStartId="10.[192,278,1394,1417]" captionTargetBox="[192,1437,192,1364]" captionTargetId="figure-12@10.[192,1437,192,1364]" captionTargetPageId="10" captionText="Figures 27–33: Anomalobuthus rickmersi from Turkmenistan, adult male from Repetek (27–30) and adult female from Chainury (31–33): metasoma and telson, lateral (27, 31), dorsal (28, 32) and ventral (29, 33), telson, lateral (30)." pageId="11" pageNumber="10">27–29</figureCitation>
,
<figureCitation id="134A2A7BFB79FFABFD07F70FFD79EA23" box="[678,719,352,375]" captionStart="Figures 107–119" captionStartId="33.[192,278,1734,1757]" captionTargetBox="[191,1440,216,1705]" captionTargetId="figure-6@33.[192,1439,217,1704]" captionTargetPageId="33" captionText="Figures 107–119: Anomalobuthus spp., from ZISP historical collection: adult male topotypes of A. pavlovskyi sp. n. (107– 112), full-body dorsal views (107, 110), labels (108, 111) and bottles (109, 112); adult male (113) and adult female (114) A. pavlovskyi sp. n. from Kunya-Urgench, Turkmenistan, full-body dorsal views (113, 114), labels (115) and bottle (116); adult male A. rickmersi from Tejen, Turkmenistan (117–119), full-body dorsal view (117), labels (118) and bottle (119). Photographs courtesy Alexander Koval." pageId="11" pageNumber="10">117</figureCitation>
); in all other
<taxonomicName id="4C714D7DFB79FFABFEA3F710FE69EAC2" box="[258,479,383,406]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B905EAECFB79FFABFEA3F710FE1FEAC2" box="[258,425,383,406]" italics="true" pageId="11" pageNumber="10">Anomalobuthus</emphasis>
spp.
</taxonomicName>
, it is blackish, entirely or at least in its distal half (
<figureCitation id="134A2A7BFB79FFABFE0CF7F1FD9DEAE1" box="[429,555,414,437]" captionStart="Figures 34–37" captionStartId="13.[192,278,1630,1653]" captionTargetBox="[212,1415,153,1571]" captionTargetId="figure-6@13.[192,1439,144,1601]" captionTargetPageId="13" captionText="Figures 34–37: Anomalobuthus krivochatskyi sp. n., adult male holotype (34–35) and adult female paratopotype (36–37): fullbody views, dorsal (34, 36) and ventral (35, 37). Scale bar = 10 mm." pageId="11" pageNumber="10">Figs. 34–35</figureCitation>
,
<figureCitation id="134A2A7BFB79FFABFD99F7F1FDCBEAE1" box="[568,637,414,437]" captionStart="Figures 38–41" captionStartId="14.[192,278,1754,1777]" captionTargetBox="[201,1418,199,1682]" captionTargetId="figure-12@14.[192,1439,192,1697]" captionTargetPageId="14" captionText="Figures 38–41: Anomalobuthus krivochatskyi sp. n., adult male (38–39) and adult female paratypes (40–41) from between Bukhara and Gazli, Uzbekistan: full-body views, dorsal (38, 40) and ventral (39, 41). Scale bar = 10 mm." pageId="11" pageNumber="10">38–39</figureCitation>
,
<figureCitation id="134A2A7BFB79FFABFD29F7F1FD7DEAE1" box="[648,715,414,437]" captionStart="Figures 59–64" captionStartId="19.[192,278,1542,1565]" captionTargetBox="[204,1417,183,1488]" captionTargetId="figure-6@19.[192,1437,168,1512]" captionTargetPageId="19" captionText="Figures 59–64: Anomalobuthus krivochatskyi sp. n., adult male holotype (59–61) and adult female paratopotype (62–64): metasoma and telson, lateral (59, 62), dorsal (60, 63) and ventral (61, 64)." pageId="11" pageNumber="10">59–61</figureCitation>
,
<figureCitation id="134A2A7BFB79FFABFD78F7F1FCA8EAE1" box="[729,798,414,437]" captionStart="Figures 79–82" captionStartId="25.[192,278,1675,1698]" captionTargetBox="[244,1418,176,1638]" captionTargetId="figure-6@25.[192,1439,168,1645]" captionTargetPageId="25" captionText="Figures 79–82: Anomalobuthus pavlovskyi sp. n., adult male (79–80) and adult female paratopotypes (81–82): full-body views, dorsal (79, 81) and ventral (80, 82). Scale bar = 10 mm." pageId="11" pageNumber="10">79–80</figureCitation>
,
<figureCitation id="134A2A7BFB79FFABFF61F7D3FEB2EA87" box="[192,260,444,467]" captionStart="Figures 83–89" captionStartId="27.[192,278,1682,1705]" captionTargetBox="[199,1433,180,1618]" captionTargetId="figure-6@27.[192,1439,168,1653]" captionTargetPageId="27" captionText="Figures 83–89: Anomalobuthus pavlovskyi sp. n., adult male (83–86) and adult female paratopotypes (87–89): right pedipalp (83), metasoma and telson, lateral (84, 87), dorsal (85, 88) and ventral (86, 89)." pageId="11" pageNumber="10">84–86</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B6575FB79FFABFF51F46FFDA3E9C5" pageId="11" pageNumber="10" type="distribution">
<paragraph id="8BCE36FEFB79FFABFF51F46FFDA3E9C5" blockId="11.[192,803,512,657]" pageId="11" pageNumber="10">
DISTRIBUTION (
<figureCitation id="134A2A7BFB79FFABFE3CF46FFE48E943" box="[413,510,512,535]" captionStart="Figure 141" captionStartId="40.[192,268,1144,1167]" captionTargetBox="[191,1440,191,1114]" captionTargetId="figure-12@40.[192,1439,192,1114]" captionTargetPageId="40" captionText="Figure 141: Distribution of the genus Anomalobuthus: A. krivochatskyi sp. n. (1), A. lowei sp. n. (2), A. pavlovskyi sp. n. (3), A. rickmersi (4), A. talebii (5), A. zarudnyi comb. n. (6), and literature records whose identification is pending (black squares). Data compiled from specimens studied herein and the following supplementary sources: Rickmer Rickmers (1913), Fet (1989, 1994), Gromov &amp; Kopdykbaev (1994), Gromov (2006), Mityaev et al. (2008), and Graham et al. (2012)." pageId="11" pageNumber="10">Fig. 141</figureCitation>
). Widespread in southern Central Asia, from the sands of the Karakum Desert (eastern
<collectingCountry id="F366766EFB79FFABFEBDF452FE03E900" box="[284,437,573,596]" name="Turkmenistan" pageId="11" pageNumber="10">Turkmenistan</collectingCountry>
), to the loess soils of the Kyzylsu River valley (southwestern
<collectingCountry id="F366766EFB79FFABFE55F433FDE9E927" box="[500,607,604,627]" name="Tajikistan" pageId="11" pageNumber="10">Tajikistan</collectingCountry>
). See further details below, in Remarks section.
</paragraph>
</subSubSection>
<subSubSection id="C36B6575FB79FFACFF51F4D1FAFEED7F" lastPageId="12" lastPageNumber="11" pageId="11" pageNumber="10" type="biology_ecology">
<paragraph id="8BCE36FEFB79FFABFF51F4D1FD49E89E" blockId="11.[192,802,702,970]" pageId="11" pageNumber="10">
ECOLOGY. According to the personal data of the collector (Rickmer Rickmers, 1913: 442), the
<typeStatus id="54CA885CFB79FFABFD64F4B2FC94E9A0" box="[709,802,733,756]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
was collected in early summer, from crevices or cracks of fine alluvial soil (“
<emphasis id="B905EAECFB79FFABFE0AF575FDF2E865" box="[427,580,794,817]" italics="true" pageId="11" pageNumber="10">loess crannies</emphasis>
”). Populations from the Karakum Desert inhabit sand dunes but tend not to occupy loose sands where two other sympatric ultrapsammophiles are found,
<taxonomicName id="4C714D7DFB79FFABFDADF519FD54E8D9" authorityName="Birula" authorityYear="1898" box="[524,738,886,909]" class="Arachnida" family="Buthidae" genus="Liobuthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="kessleri">
<emphasis id="B905EAECFB79FFABFDADF519FD54E8D9" box="[524,738,886,909]" italics="true" pageId="11" pageNumber="10">Liobuthus kessleri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C714D7DFB79FFABFF61F5FAFDADE8F8" authority="Fet, 1984" authorityName="Fet" authorityYear="1984" box="[192,539,917,940]" class="Arachnida" family="Buthidae" genus="Pectinibuthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="birulai">
<emphasis id="B905EAECFB79FFABFF61F5FAFE10E8F8" box="[192,422,917,940]" italics="true" pageId="11" pageNumber="10">Pectinibuthus birulai</emphasis>
Fet, 1984
</taxonomicName>
. Detailed ecological information from Repetek was published by Fet (1980).
</paragraph>
<paragraph id="8BCE36FEFB79FFABFF51F598FD59EE76" blockId="11.[192,802,1015,1651]" pageId="11" pageNumber="10">
NOTES. The
<typeStatus id="54CA885CFB79FFABFE20F598FE69EF5A" box="[385,479,1015,1038]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C714D7DFB79FFABFDACF598FD23EF5A" authorityName="Kraepelin" authorityYear="1900" box="[525,661,1015,1038]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="rickmersi">
<emphasis id="B905EAECFB79FFABFDACF598FD23EF5A" box="[525,661,1015,1038]" italics="true" pageId="11" pageNumber="10">A. rickmersi</emphasis>
</taxonomicName>
remains the single specimen known from
<collectingCountry id="F366766EFB79FFABFDB2F27AFDC8EF78" box="[531,638,1045,1068]" name="Tajikistan" pageId="11" pageNumber="10">Tajikistan</collectingCountry>
. Its collecting site was reported by
<bibRefCitation id="EFE04B0FFB79FFABFE6BF25BFD3BEF1F" author="KRAEPELIN" box="[458,653,1076,1099]" pageId="11" pageNumber="10" pagination="1 - 17" refId="ref25373" refString="KRAEPELIN, K. 1900. Uber einige neue Gliederspinnen. Abhandlungen aus dem Gebiete der Naturwissenschaften. Herausgegeben vom Naturwissenschaftlichen Verein in Hamburg, 16, 1 (4): 1 - 17." type="journal article" year="1900">Kraepelin (1900)</bibRefCitation>
as Bukhara (“Bucharei”, see the original museum label in
<figureCitation id="134A2A7BFB79FFABFD74F23CFCAEEF3E" box="[725,792,1107,1130]" captionStart="Figures 1–3" captionStartId="5.[192,278,1579,1602]" captionTargetBox="[191,1440,191,1550]" captionTargetId="figure-6@5.[192,1439,192,1549]" captionTargetPageId="5" captionText="Figures 1–3: Anomalobuthus rickmersi, adult male holotype: full-body views, dorsal (1) and ventral (2), plus original labels (3). Scale bar = 10 mm (1–2). The red inset at bottom right shows the label of the specimen examined by Lourenço (2001), demonstrating it was not the actual Rickmers's type specimen." pageId="11" pageNumber="10">Fig. 3</figureCitation>
) and interpreted by all subsequent authors as originating from the city, the capital of the homonymous Bukhara Khanate, located in modern
<collectingCountry id="F366766EFB79FFABFDB1F2C0FD3EEF92" box="[528,648,1199,1222]" name="Uzbekistan" pageId="11" pageNumber="10">Uzbekistan</collectingCountry>
. In addition, Birula (1911b: 171) erroneously stated that the type specimen of
<taxonomicName id="4C714D7DFB79FFABFEF4F283FE6BEE57" authorityName="Kraepelin" authorityYear="1900" box="[341,477,1260,1283]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="rickmersi">
<emphasis id="B905EAECFB79FFABFEF4F283FE6BEE57" box="[341,477,1260,1283]" italics="true" pageId="11" pageNumber="10">A. rickmersi</emphasis>
</taxonomicName>
originated from “westlichen Buchara” (i.e. western part of the Bukhara Emirate).
</paragraph>
<paragraph id="8BCE36FEFB79FFABFF51F346FED8ED27" blockId="11.[192,802,1015,1651]" pageId="11" pageNumber="10">
<materialsCitation id="3B193CA3FB79FFABFF51F346FED1ED27" country="Uzbekistan and Tajikistan" county="Rickmer Rickmers" location="This" municipality="During" pageId="11" pageNumber="10" specimenCount="1" typeStatus="holotype">
<location id="8EAE6025FB79FFABFF51F346FE96EE14" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB79FFABFF51F346FE96EE14" box="[240,288,1321,1344]" country="Uzbekistan and Tajikistan" county="Rickmer Rickmers" municipality="During" name="This" pageId="11" pageNumber="10">This</location>
specimen was collected by the famous
<location id="8EAE6025FB79FFABFD6CF346FE93EE0B" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB79FFABFD6CF346FE93EE0B" country="Uzbekistan and Tajikistan" county="Rickmer Rickmers" municipality="During" name="German mountain" pageId="11" pageNumber="10">German mountain</location>
explorer
<location id="8EAE6025FB79FFABFE2AF327FD27EE0B" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB79FFABFE2AF327FD27EE0B" box="[395,657,1352,1375]" country="Uzbekistan and Tajikistan" county="Rickmer Rickmers" municipality="During" name="Willi Rickmer Rickmers" pageId="11" pageNumber="10">Willi Rickmer Rickmers</location>
(1873–1965) who repeatedly visited the
<location id="8EAE6025FB79FFABFDA2F308FD72EE2A" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB79FFABFDA2F308FD72EE2A" box="[515,708,1383,1406]" country="Uzbekistan and Tajikistan" county="Rickmer Rickmers" municipality="During" name="Bukhara Emirate" pageId="11" pageNumber="10">Bukhara Emirate</location>
, then a protectorate of the Russian Empire, which included part of modern
<collectingCountry id="F366766EFB79FFABFEE5F3CBFDDDEEEF" box="[324,619,1444,1467]" isEnumeration="true" pageId="11" pageNumber="10">Uzbekistan and Tajikistan</collectingCountry>
(Rickmer-Rickmers, 1899; Rickmer-Rickmers, 1913).
<collectingMunicipality id="6BAAAC84FB79FFABFD2CF3ACFD6FEE8E" box="[653,729,1475,1498]" pageId="11" pageNumber="10">During</collectingMunicipality>
preparation of the present revision, we discovered that
<collectingCounty id="62AF4E72FB79FFABFF61F06FFE26ED43" box="[192,400,1536,1559]" pageId="11" pageNumber="10">Rickmer Rickmers</collectingCounty>
(1913: 442) himself actually published unequivocal information on the precise collection locality and date of the
<typeStatus id="54CA885CFB79FFABFE73F052FD99ED00" box="[466,559,1597,1620]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
, which we reproduce below
<emphasis id="B905EAECFB79FFABFEA9F033FED1ED27" box="[264,359,1628,1651]" italics="true" pageId="11" pageNumber="10">verbatim</emphasis>
</materialsCitation>
:
</paragraph>
<paragraph id="8BCE36FEFB79FFABFF51F0F7FD65EC3B" blockId="11.[240,754,1688,1903]" pageId="11" pageNumber="10">
“On the way to Baljuan I noticed that many poplar trees were pitted with wormholes bored by that stately Capricorn beetle the
<taxonomicName id="4C714D7DFB79FFABFDAEF0A0FD44EDB0" authority="Sartus. In" authorityName="Sartus. In" box="[527,754,1743,1764]" class="Insecta" family="Cerambycidae" genus="Pachydissus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAECFB79FFABFDAEF0A0FD7BEDB0" box="[527,717,1743,1764]" italics="true" pageId="11" pageNumber="10">Pachydissus Sartus</emphasis>
. In
</taxonomicName>
the early summer of 1896 the loess crannies around had yielded up to my collection new specimens of poisonous spider and scorpion,
<taxonomicName id="4C714D7DFB79FFABFD90F14DFD44EC63" authorityName="Kraepelin" authorityYear="1899" box="[561,754,1826,1847]" class="Arachnida" family="Gylippidae" genus="Gylippus" kingdom="Animalia" order="Solifugae" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="rickmersi">
<emphasis id="B905EAECFB79FFABFD90F14DFD44EC63" box="[561,754,1826,1847]" italics="true" pageId="11" pageNumber="10">Gylippus Rickmersi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C714D7DFB79FFABFEB8F151FD9DEC07" authority="Rickmersi. I" authorityName="Rickmersi. I" box="[281,555,1854,1875]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAECFB79FFABFEB8F151FDA0EC07" box="[281,534,1854,1875]" italics="true" pageId="11" pageNumber="10">Anomalobuthus Rickmersi</emphasis>
. I
</taxonomicName>
feel highly gratified that such vicious creatures should bear my name.”
</paragraph>
<paragraph id="8BCE36FEFB79FFABFCCFF6A9FCC0EAC6" blockId="11.[830,1441,198,1904]" pageId="11" pageNumber="10">
By the way, the “poisonous spider” mentioned by the impressed collector is a solpugid, also described by
<bibRefCitation id="EFE04B0FFB79FFABFC9FF76DFC4FEA4D" author="KRAEPELIN" box="[830,1017,258,281]" pageId="11" pageNumber="10" pagination="197 - 259" refId="ref25348" refString="KRAEPELIN, K. 1899. Zur Systematik der Solifugen. Mitteilungen aus dem Naturhistorischen Museum in Hamburg, 16: 197 - 259." type="journal article" year="1899">Kraepelin (1899)</bibRefCitation>
. It is a member of the
<taxonomicName id="4C714D7DFB79FFABFB51F76DFAD1EA4D" authorityName="Roewer" authorityYear="1933" box="[1264,1383,258,281]" class="Arachnida" family="Gylippidae" kingdom="Animalia" order="Solifugae" pageId="11" pageNumber="10" phylum="Arthropoda" rank="family">Gylippidae</taxonomicName>
family, currently regarded as a junior synonym of
<taxonomicName id="4C714D7DFB79FFABFAE1F74EFBFCEA02" authority="Birula, 1893" authorityName="Birula" authorityYear="1893" class="Arachnida" family="Gylippidae" genus="Gylippus" kingdom="Animalia" order="Solifugae" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="ferganensis">
<emphasis id="B905EAECFB79FFABFAE1F74EFC0FEA02" italics="true" pageId="11" pageNumber="10">Gylippus ferganensis</emphasis>
Birula, 1893
</taxonomicName>
(see Harms &amp; Dupérré, 2018). As all solpugids, it is indeed “vicious” but not “poisonous”.
</paragraph>
<paragraph id="8BCE36FEFB79FFABFCCFF7F6FB48E990" blockId="11.[830,1441,198,1904]" pageId="11" pageNumber="10">
Baljuan (also spelled Baldzhuan, now Baljuvon or Bal'dzhuvon) is a well-known town in modern
<collectingCountry id="F366766EFB79FFABFAFEF7D7FCC6EAB9" name="Tajikistan" pageId="11" pageNumber="10">Tajikistan</collectingCountry>
, in the valley of Kyzylsu River, a tributary of the larger Panj River (= Pyandzh, Piandj), itself a tributary of the great Amudarya (the ancient Oxus). Baljuvon stands on the Silk Road, not far from the ancient
<collectingRegion id="49B5F81CFB79FFABFAFAF45DFA17E91D" box="[1371,1441,562,585]" country="Afghanistan" name="Balkh" pageId="11" pageNumber="10">Bactra</collectingRegion>
(now
<collectingRegion id="49B5F81CFB79FFABFCDFF43EFC08E93C" box="[894,958,593,616]" country="Afghanistan" name="Balkh" pageId="11" pageNumber="10">Balkh</collectingRegion>
in northern
<collectingCountry id="F366766EFB79FFABFBE6F43EFB7DE93C" box="[1095,1227,593,616]" name="Afghanistan" pageId="11" pageNumber="10">Afghanistan</collectingCountry>
), and was last time noted as the place where Enver Pasha, a rival of the Turkish leader Mustafa Kemal (Ataturk), was killed in
<quantity id="4C899B1BFB79FFABFC9FF4C2FC26E990" box="[830,912,685,708]" metricMagnitude="1" metricUnit="m" metricValue="4.88188" pageId="11" pageNumber="10" unit="in" value="1922.0">1922 in</quantity>
a skirmish with the Soviet troops.
</paragraph>
<paragraph id="8BCE36FEFB79FFABFCCFF4A3FCCEE8A3" blockId="11.[830,1441,198,1904]" pageId="11" pageNumber="10">
In more peaceful times, Rickmer-Rickmers (1899) published a map of his fourth travel to the Bukhara Emirate, conducted in 1898 (the 1896 trip was his third travel). We can see that he followed the same and (then) only route from
<collectingRegion id="49B5F81CFB79FFABFC5EF529FBDCE809" box="[1023,1130,838,861]" country="Tajikistan" name="Dushanbe" pageId="11" pageNumber="10">Dushanbe</collectingRegion>
(now the capital of
<collectingCountry id="F366766EFB79FFABFAFEF529FCC6E828" name="Tajikistan" pageId="11" pageNumber="10">Tajikistan</collectingCountry>
) to Khovaling, via Baljuvon. Baljuvon lies very far to the southeast from the former Emirate's capital city of Bukhara (modern
<collectingCountry id="F366766EFB79FFABFBA5F5CDFBCAE8ED" box="[1028,1148,930,953]" name="Uzbekistan" pageId="11" pageNumber="10">Uzbekistan</collectingCountry>
) and is isolated by several mountain ranges from the lowland deserts of
<collectingCountry id="F366766EFB79FFABFAF3F5AEFCC6E8A3" name="Uzbekistan" pageId="11" pageNumber="10">Uzbekistan</collectingCountry>
.
</paragraph>
<paragraph id="8BCE36FEFB79FFABFCCFF591FC32EE97" blockId="11.[830,1441,198,1904]" pageId="11" pageNumber="10">
Although quite active zoological research was conducted in Central Asia by numerous Russian (and then Soviet) researchers, very few scorpions have been collected in the remote valleys of northern tributaries of Amu Darya in modern
<collectingCountry id="F366766EFB79FFABFBE4F216FB0BEFC4" box="[1093,1213,1145,1168]" name="Uzbekistan" pageId="11" pageNumber="10">Uzbekistan</collectingCountry>
and
<collectingCountry id="F366766EFB79FFABFB59F216FAD0EFC4" box="[1272,1382,1145,1168]" name="Tajikistan" pageId="11" pageNumber="10">Tajikistan</collectingCountry>
. It is not surprising, therefore, that no other specimens of
<taxonomicName id="4C714D7DFB79FFABFA28F2F7FC15EF99" authorityName="Kraepelin" authorityYear="1899" class="Arachnida" family="Gylippidae" genus="Gylippus" kingdom="Animalia" order="Solifugae" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="rickmersi">
<emphasis id="B905EAECFB79FFABFA28F2F7FC15EF99" italics="true" pageId="11" pageNumber="10">A. rickmersi</emphasis>
</taxonomicName>
have been collected in this area since 1896; in fact, no other scorpion collections we know of originate from Baljuvon. Moreover, another, much more amazing scorpion from the same area was not discovered until the 1990s: the relict
<taxonomicName id="4C714D7DFB79FFABFBA1F35EFBFDEE33" authority="Gromov, 1998" authorityName="Gromov" authorityYear="1998" class="Arachnida" family="Pseudochactidae" genus="Pseudochactas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="ovchinnikovi">
<emphasis id="B905EAECFB79FFABFBA1F35EFA82EE1C" box="[1024,1332,1329,1352]" italics="true" pageId="11" pageNumber="10">Pseudochactas ovchinnikovi</emphasis>
<bibRefCitation id="EFE04B0FFB79FFABFAE1F35EFCC5EE33" author="GROMOV" pageId="11" pageNumber="10" pagination="409 - 413" refId="ref24805" refString="GROMOV, A. V. 1998. [A new family, genus, and species of scorpion (Arachnida, Scorpiones) from the southern part of Central Asia]. Zoologicheskii Zhurnal, 77 (9): 1003 - 1008 (in Russian, English summary). English translation (same title): Russian Journal of Zoology, 1998, 2 (3): 409 - 413." type="journal article" year="1998">Gromov, 1998</bibRefCitation>
(Pseudochactidae)
</taxonomicName>
. Its
<typeStatus id="54CA885CFB79FFABFB29F33FFB00EE33" box="[1160,1206,1360,1383]" pageId="11" pageNumber="10">type</typeStatus>
locality lies in the Surkhandarya River valley, which separates modern
<collectingCountry id="F366766EFB79FFABFC9FF3E2FC00EEF0" box="[830,950,1421,1444]" name="Uzbekistan" pageId="11" pageNumber="10">Uzbekistan</collectingCountry>
from
<collectingCountry id="F366766EFB79FFABFBA0F3E2FBDAEEF0" box="[1025,1132,1421,1444]" name="Tajikistan" pageId="11" pageNumber="10">Tajikistan</collectingCountry>
(
<bibRefCitation id="EFE04B0FFB79FFABFB21F3E2FA96EEF0" author="GROMOV" box="[1152,1312,1421,1444]" pageId="11" pageNumber="10" pagination="409 - 413" refId="ref24805" refString="GROMOV, A. V. 1998. [A new family, genus, and species of scorpion (Arachnida, Scorpiones) from the southern part of Central Asia]. Zoologicheskii Zhurnal, 77 (9): 1003 - 1008 (in Russian, English summary). English translation (same title): Russian Journal of Zoology, 1998, 2 (3): 409 - 413." type="journal article" year="1998">Gromov, 1998</bibRefCitation>
; Fet et al., 2004).
</paragraph>
<paragraph id="8BCE36FEFB79FFACFCCFF3A5FE4DEEC6" blockId="11.[830,1441,198,1904]" lastBlockId="12.[192,802,1219,1917]" lastPageId="12" lastPageNumber="11" pageId="11" pageNumber="10">
<materialsCitation id="3B193CA3FB79FFACFCCFF3A5FE41EEC6" collectingDate="1904-02" collectingDateMax="1904-06" collectingDateMin="1904-02" collectionCode="ZISP, ZMMSU, NHMW, R, ZSRO" collectorName="The &amp; Birula &amp; Tejen &amp; Tedzhen &amp; Tedshen &amp; K. Aris &amp; Further &amp; Fet &amp; Repetek" country="Turkmenistan" county="Akhal Province" lastPageId="12" lastPageNumber="11" location="The" municipality="Eduard Nikolaevich Fischer" pageId="11" pageNumber="10" specimenCode="ZISP 808, ZISP 809, ZISP 1699" specimenCount="21" specimenCount-male="1">
<location id="8EAE6025FB79FFABFCCFF3A5FC2EEEB5" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB79FFABFCCFF3A5FC2EEEB5" box="[878,920,1482,1505]" country="Turkmenistan" county="Akhal Province" municipality="Eduard Nikolaevich Fischer" name="The" pageId="11" pageNumber="10">The</location>
first known specimen from
<collectingCountry id="F366766EFB79FFABFB6CF3A5FAD7EEB5" box="[1229,1377,1482,1505]" name="Turkmenistan" pageId="11" pageNumber="10">Turkmenistan</collectingCountry>
(then the
<location id="8EAE6025FB79FFABFCCCF386FBE5ED54" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB79FFABFCCCF386FBE5ED54" box="[877,1107,1513,1536]" country="Turkmenistan" county="Akhal Province" municipality="Eduard Nikolaevich Fischer" name="Transcaspian Region" pageId="11" pageNumber="10">Transcaspian Region</location>
of the Russian Empire) was reported by
<location id="8EAE6025FB79FFABFC70F067FBA2ED4B" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB79FFABFC70F067FBA2ED4B" box="[977,1044,1544,1567]" country="Turkmenistan" county="Akhal Province" municipality="Eduard Nikolaevich Fischer" name="Birula" pageId="11" pageNumber="10">Birula</location>
(1904: 32, 35).
<location id="8EAE6025FB79FFABFB42F067FB45ED4B" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB79FFABFB42F067FB45ED4B" box="[1251,1267,1544,1567]" country="Turkmenistan" county="Akhal Province" municipality="Eduard Nikolaevich Fischer" name="It" pageId="11" pageNumber="10">It</location>
was a female collected next to the railway station of
<location id="8EAE6025FB79FFABFB5CF049FAE2ED69" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB79FFABFB5CF049FAE2ED69" box="[1277,1364,1574,1597]" country="Turkmenistan" county="Akhal Province" municipality="Eduard Nikolaevich Fischer" name="Repetek" pageId="11" pageNumber="10">Repetek</location>
in the
<location id="8EAE6025FB79FFABFC9FF02AFC68ED08" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB79FFABFC9FF02AFC68ED08" box="[830,990,1605,1628]" country="Turkmenistan" county="Akhal Province" municipality="Eduard Nikolaevich Fischer" name="East Karakum" pageId="11" pageNumber="10">East Karakum</location>
, the future site of the famed
<location id="8EAE6025FB79FFABFAE8F02AFC45ED2F" LSID="urn:lsid:plazi:treatment:03D887E8FB76FFACFCCEF452FAFEED7F:8EAE6025FB79FFABFAE8F02AFC45ED2F" country="Turkmenistan" county="Akhal Province" municipality="Eduard Nikolaevich Fischer" name="Repetek Natural Reserve" pageId="11" pageNumber="10">Repetek Natural Reserve</location>
, by the coleopterologist
<collectingMunicipality id="6BAAAC84FB79FFABFAABF00BFC5DEDCD" pageId="11" pageNumber="10">Eduard Nikolaevich Fischer</collectingMunicipality>
in
<date id="FFCF103EFB79FFABFB85F0EDFB7CEDCD" box="[1060,1226,1666,1689]" pageId="11" pageNumber="10" value="1904-02">
<collectingDate id="EF8BE9D6FB79FFABFB85F0EDFB7CEDCD" box="[1060,1226,1666,1689]" pageId="11" pageNumber="10" value="1904-02">February 1904</collectingDate>
</date>
(
<specimenCode id="DBD79E85FB79FFABFB42F0EDFAE0EDCD" box="[1251,1366,1666,1689]" collectionCode="ZISP" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="11" pageNumber="10">ZISP 808</specimenCode>
).
<collectorName id="26845328FB79FFABFAD6F0EDFA17EDCD" box="[1399,1441,1666,1689]" pageId="11" pageNumber="10">The</collectorName>
second specimen was also published by
<collectorName id="26845328FB79FFABFAADF0CEFAF9EDEC" box="[1292,1359,1697,1720]" pageId="11" pageNumber="10">Birula</collectorName>
(1905: 449–450); it was a male from
<collectorName id="26845328FB79FFABFB75F0AFFAA7ED83" box="[1236,1297,1728,1751]" pageId="11" pageNumber="10">Tejen</collectorName>
(=
<collectorName id="26845328FB79FFABFA9DF0AFFA2BED83" box="[1340,1437,1728,1751]" pageId="11" pageNumber="10">Tedzhen</collectorName>
,
<collectorName id="26845328FB79FFABFC9FF0B1FC2CEDA1" box="[830,922,1758,1781]" pageId="11" pageNumber="10">Tedshen</collectorName>
) in
<collectingCounty id="62AF4E72FB79FFABFC70F0B1FB34EDA1" box="[977,1154,1758,1781]" pageId="11" pageNumber="10">Akhal Province</collectingCounty>
, collected by
<collectorName id="26845328FB79FFABFA88F0B1FAC8EDA1" box="[1321,1406,1758,1781]" pageId="11" pageNumber="10">K. Aris</collectorName>
in
<date id="FFCF103EFB79FFABFC9FF092FC06EC40" box="[830,944,1789,1812]" pageId="11" pageNumber="10" value="1904-06">
<collectingDate id="EF8BE9D6FB79FFABFC9FF092FC06EC40" box="[830,944,1789,1812]" pageId="11" pageNumber="10" value="1904-06">June 1904</collectingDate>
</date>
(
<specimenCode id="DBD79E85FB79FFABFC64F092FB99EC40" box="[965,1071,1789,1812]" collectionCode="ZISP" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="11" pageNumber="10">ZISP 809</specimenCode>
).
<collectorName id="26845328FB79FFABFBEBF092FB2FEC40" box="[1098,1177,1789,1812]" pageId="11" pageNumber="10">Further</collectorName>
collections from
<collectingCountry id="F366766EFB79FFABFAC2F092FC28EC67" name="Turkmenistan" pageId="11" pageNumber="10">Turkmenistan</collectingCountry>
published by
<collectorName id="26845328FB79FFABFBE0F173FBD5EC67" box="[1089,1123,1820,1843]" pageId="11" pageNumber="10">Fet</collectorName>
(1989), included a series of more than
<specimenCount id="9D77FD77FB79FFABFC10F155FBF4EC05" box="[945,1090,1850,1873]" pageId="11" pageNumber="10" type="generic">20 specimens</specimenCount>
from
<collectorName id="26845328FB79FFABFB22F155FB6DEC05" box="[1155,1243,1850,1873]" pageId="11" pageNumber="10">Repetek</collectorName>
(
<collectionCode id="ED60AE3BFB79FFABFB4BF155FA92EC05" box="[1258,1316,1850,1873]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="11" pageNumber="10">ZISP</collectionCode>
,
<collectionCode id="ED60AE3BFB79FFABFA8EF155FA27EC05" box="[1327,1425,1850,1873]" pageId="11" pageNumber="10">ZMMSU</collectionCode>
). In addition,
<specimenCount id="9D77FD77FB79FFABFC6FF136FBF7EC24" box="[974,1089,1881,1904]" pageId="11" pageNumber="10" type="male">two males</specimenCount>
from Repetek are housed in
<collectionCode id="ED60AE3BFB7EFFACFF61F2ACFEA1EF8E" box="[192,279,1219,1242]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="11" type="Museum">NHMW</collectionCode>
and one in
<collectionCode id="ED60AE3BFB7EFFACFE3EF2ACFE02EF8E" box="[415,436,1219,1242]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="11">R</collectionCode>
. Kinzelbach's collection (
<collectionCode id="ED60AE3BFB7EFFACFD6CF2ACFCA1EF8E" box="[717,791,1219,1242]" pageId="12" pageNumber="11">ZSRO</collectionCode>
). Some of these specimens are illustrated here in
<figureCitation id="134A2A7BFB7EFFACFD6BF28DFF79EE43" captionStart="Figures 4–7" captionStartId="6.[192,278,1734,1757]" captionTargetBox="[207,1403,201,1681]" captionTargetId="figure-12@6.[192,1439,188,1704]" captionTargetPageId="6" captionText="Figures 4–7: Anomalobuthus rickmersi from Turkmenistan: adult male from Repetek (4–5) and adult female from Chainury (6– 7), full-body views, dorsal (4, 6) and ventral (5, 7). Scale bar = 10 mm." pageId="12" pageNumber="11">Figs. 4– 5</figureCitation>
,
<figureCitation id="134A2A7BFB7EFFACFF42F36FFE9FEE43" box="[227,297,1280,1303]" captionStart="Figures 27–33" captionStartId="10.[192,278,1394,1417]" captionTargetBox="[192,1437,192,1364]" captionTargetId="figure-12@10.[192,1437,192,1364]" captionTargetPageId="10" captionText="Figures 27–33: Anomalobuthus rickmersi from Turkmenistan, adult male from Repetek (27–30) and adult female from Chainury (31–33): metasoma and telson, lateral (27, 31), dorsal (28, 32) and ventral (29, 33), telson, lateral (30)." pageId="12" pageNumber="11">27–30</figureCitation>
,
<figureCitation id="134A2A7BFB7EFFACFE9AF36FFE77EE43" box="[315,449,1280,1303]" captionStart-0="Figures 101–106" captionStart-1="Figures 107–119" captionStartId-0="32.[192,278,1242,1265]" captionStartId-1="33.[192,278,1734,1757]" captionTargetBox-0="[192,1425,198,1183]" captionTargetBox-1="[191,1440,216,1705]" captionTargetId-0="figure-12@32.[192,1437,168,1212]" captionTargetId-1="figure-6@33.[192,1439,217,1704]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionText-0="Figures 101–106: Anomalobuthus talebii, adult female holotype: right pedipalp trochanter and femur, dorsal (101), right pedipalp patella, dorsal (102) and external (103), right pedipalp chela, dorsal (104) and external (105), right pedipalp movable finger, dorsal (106). Black dots depict trichobothria." captionText-1="Figures 107–119: Anomalobuthus spp., from ZISP historical collection: adult male topotypes of A. pavlovskyi sp. n. (107– 112), full-body dorsal views (107, 110), labels (108, 111) and bottles (109, 112); adult male (113) and adult female (114) A. pavlovskyi sp. n. from Kunya-Urgench, Turkmenistan, full-body dorsal views (113, 114), labels (115) and bottle (116); adult male A. rickmersi from Tejen, Turkmenistan (117–119), full-body dorsal view (117), labels (118) and bottle (119). Photographs courtesy Alexander Koval." pageId="12" pageNumber="11">103–115. A</figureCitation>
few specimens from northern
<collectingCountry id="F366766EFB7EFFACFF61F370FEE2EE62" box="[192,340,1311,1334]" name="Turkmenistan" pageId="12" pageNumber="11">Turkmenistan</collectingCountry>
(Kunya-Urgench,
<specimenCode id="DBD79E85FB7EFFACFD95F370FD03EE62" box="[564,693,1311,1334]" collectionCode="ZISP" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="12" pageNumber="11">ZISP 1699</specimenCode>
; Shakhsenem,
<collectionCode id="ED60AE3BFB7EFFACFEABF351FEDAEE01" box="[266,364,1342,1365]" pageId="12" pageNumber="11">ZMMSU</collectionCode>
Tb-374), published by Fet (1989) as
<taxonomicName id="4C714D7DFB7EFFACFCAAF351FE93EE27" authorityName="Kraepelin" authorityYear="1899" class="Arachnida" family="Gylippidae" genus="Gylippus" kingdom="Animalia" order="Solifugae" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="rickmersi">
<emphasis id="B905EAECFB7EFFACFCAAF351FE93EE27" italics="true" pageId="12" pageNumber="11">A. rickmersi</emphasis>
</taxonomicName>
, actually correspond to a different species (
<taxonomicName id="4C714D7DFB7EFFACFCAAF333FE87EEC6" authority="Teruel &amp; Kovařík &amp; Fet, 2018" authorityName="Teruel &amp; Kovařík &amp; Fet" authorityYear="2018" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="pavlovskyi" status="sp. nov.">
<emphasis id="B905EAECFB7EFFACFCAAF333FE87EEC6" italics="true" pageId="12" pageNumber="11">A. pavlovskyi</emphasis>
</taxonomicName>
<emphasis id="B905EAECFB7EFFACFE96F314FEC5EEC6" bold="true" box="[311,371,1403,1426]" pageId="12" pageNumber="11">
<taxonomicNameLabel id="A2365797FB7EFFACFE96F314FEC5EEC6" box="[311,371,1403,1426]" pageId="12" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, see below)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCE36FEFB7EFFACFEE4F689FAB0EF60" pageId="12" pageNumber="11">
<table id="F971C45EFB7E005FFEA2F6BEFAF5EF60" box="[259,1347,209,1076]" gridcols="5" gridrows="20" pageId="12" pageNumber="11">
<tr id="354134BCFB7E005FFEA2F6BEFAF5EA4D" box="[259,1347,209,281]" gridrow="0" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F6BEFDE9EA4D" box="[259,607,209,281]" colspan="2" colspanRight="1" gridcol="0" gridrow="0" pageId="12" pageNumber="11">
<emphasis id="B905EAECFB7EFFACFEE4F689FD94EA54" bold="true" box="[325,546,230,256]" pageId="12" pageNumber="11">Dimensions (mm)</emphasis>
</th>
<th id="76905DC0FB7E005FFDD9F6BEFCC1EA4D" box="[632,887,209,281]" gridcol="2" gridrow="0" pageId="12" pageNumber="11">
<emphasis id="B905EAECFB7EFFACFD3AF6BEFCE2EBBF" bold="true" box="[667,852,209,235]" pageId="12" pageNumber="11">TJK, Baljuvon</emphasis>
♂
<emphasis id="B905EAECFB7EFFACFD37F690FCC1EA4D" bold="true" box="[662,887,255,281]" pageId="12" pageNumber="11">holotype (ZMUH)</emphasis>
</th>
<th id="76905DC0FB7E005FFC3AF6BEFAF5EA4D" box="[923,1347,209,281]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="12" pageNumber="11">
<emphasis id="B905EAECFB7EFFACFBACF6BEFB6DEBBF" bold="true" box="[1037,1243,209,235]" pageId="12" pageNumber="11">TKM, Chainury</emphasis>
♀
<emphasis id="B905EAECFB7EFFACFC72F690FB80EA4D" bold="true" box="[979,1078,255,281]" pageId="12" pageNumber="11">(FKCP)</emphasis>
♀
<emphasis id="B905EAECFB7EFFACFB64F690FA9EEA4D" bold="true" box="[1221,1320,255,281]" pageId="12" pageNumber="11">(FKCP)</emphasis>
</th>
</tr>
<tr id="354134BCFB7E005FFEA2F741FAF5EA11" box="[259,1347,302,325]" gridrow="1" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F741FE6EEA11" box="[259,472,302,325]" gridcol="0" gridrow="1" pageId="12" pageNumber="11">Carapace</th>
<td id="76905DC0FB7E005FFE58F741FDE9EA11" box="[505,607,302,325]" gridcol="1" gridrow="1" pageId="12" pageNumber="11">L / Wp</td>
<td id="76905DC0FB7E005FFDD9F741FCC1EA11" box="[632,887,302,325]" gridcol="2" gridrow="1" pageId="12" pageNumber="11">3.05 / 3.80</td>
<td id="76905DC0FB7E005FFC3AF741FBE7EA11" box="[923,1105,302,325]" gridcol="3" gridrow="1" pageId="12" pageNumber="11">3.67 / 4.30</td>
<td id="76905DC0FB7E005FFB2CF741FAF5EA11" box="[1165,1347,302,325]" gridcol="4" gridrow="1" pageId="12" pageNumber="11">3.55 / 4.03</td>
</tr>
<tr id="354134BCFB7E005FFEA2F737FAF5EA3B" box="[259,1347,344,367]" gridrow="2" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F737FE6EEA3B" box="[259,472,344,367]" gridcol="0" gridrow="2" pageId="12" pageNumber="11">Mesosoma</th>
<td id="76905DC0FB7E005FFE58F737FDE9EA3B" box="[505,607,344,367]" gridcol="1" gridrow="2" pageId="12" pageNumber="11">L</td>
<td id="76905DC0FB7E005FFDD9F737FCC1EA3B" box="[632,887,344,367]" gridcol="2" gridrow="2" pageId="12" pageNumber="11">7.70</td>
<td id="76905DC0FB7E005FFC3AF737FBE7EA3B" box="[923,1105,344,367]" gridcol="3" gridrow="2" pageId="12" pageNumber="11">9.40</td>
<td id="76905DC0FB7E005FFB2CF737FAF5EA3B" box="[1165,1347,344,367]" gridcol="4" gridrow="2" pageId="12" pageNumber="11">7.10</td>
</tr>
<tr id="354134BCFB7E005FFEA2F7EEFAF5EACC" box="[259,1347,385,408]" gridrow="3" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F7EEFE6EEACC" box="[259,472,385,408]" gridcol="0" gridrow="3" pageId="12" pageNumber="11">Tergite VII</th>
<td id="76905DC0FB7E005FFE58F7EEFDE9EACC" box="[505,607,385,408]" gridcol="1" gridrow="3" pageId="12" pageNumber="11">L / W</td>
<td id="76905DC0FB7E005FFDD9F7EEFCC1EACC" box="[632,887,385,408]" gridcol="2" gridrow="3" pageId="12" pageNumber="11">2.15 / 3.50</td>
<td id="76905DC0FB7E005FFC3AF7EEFBE7EACC" box="[923,1105,385,408]" gridcol="3" gridrow="3" pageId="12" pageNumber="11">2.35 / 4.10</td>
<td id="76905DC0FB7E005FFB2CF7EEFAF5EACC" box="[1165,1347,385,408]" gridcol="4" gridrow="3" pageId="12" pageNumber="11">1.79 / 4.18</td>
</tr>
<tr id="354134BCFB7E005FFEA2F7C5FAF5EA95" box="[259,1347,426,449]" gridrow="4" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F7C5FE6EEA95" box="[259,472,426,449]" gridcol="0" gridrow="4" pageId="12" pageNumber="11">Metasoma + Telson</th>
<td id="76905DC0FB7E005FFE58F7C5FDE9EA95" box="[505,607,426,449]" gridcol="1" gridrow="4" pageId="12" pageNumber="11">L</td>
<td id="76905DC0FB7E005FFDD9F7C5FCC1EA95" box="[632,887,426,449]" gridcol="2" gridrow="4" pageId="12" pageNumber="11">23.38</td>
<td id="76905DC0FB7E005FFC3AF7C5FBE7EA95" box="[923,1105,426,449]" gridcol="3" gridrow="4" pageId="12" pageNumber="11">23.41</td>
<td id="76905DC0FB7E005FFB2CF7C5FAF5EA95" box="[1165,1347,426,449]" gridcol="4" gridrow="4" pageId="12" pageNumber="11">21.48</td>
</tr>
<tr id="354134BCFB7E005FFEA2F7BBFAF5EABF" box="[259,1347,468,491]" gridrow="5" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F7BBFE6EEABF" box="[259,472,468,491]" gridcol="0" gridrow="5" pageId="12" pageNumber="11">Segment I</th>
<td id="76905DC0FB7E005FFE58F7BBFDE9EABF" box="[505,607,468,491]" gridcol="1" gridrow="5" pageId="12" pageNumber="11">L / W / D</td>
<td id="76905DC0FB7E005FFDD9F7BBFCC1EABF" box="[632,887,468,491]" gridcol="2" gridrow="5" pageId="12" pageNumber="11">2.80 / 1.61 / 1.43</td>
<td id="76905DC0FB7E005FFC3AF7BBFBE7EABF" box="[923,1105,468,491]" gridcol="3" gridrow="5" pageId="12" pageNumber="11">2.80 / 1.95 / 1.79</td>
<td id="76905DC0FB7E005FFB2CF7BBFAF5EABF" box="[1165,1347,468,491]" gridcol="4" gridrow="5" pageId="12" pageNumber="11">2.50 / 1.75 / 1.43</td>
</tr>
<tr id="354134BCFB7E005FFEA2F792FAF5E940" box="[259,1347,509,532]" gridrow="6" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F792FE6EE940" box="[259,472,509,532]" gridcol="0" gridrow="6" pageId="12" pageNumber="11">Segment II</th>
<td id="76905DC0FB7E005FFE58F792FDE9E940" box="[505,607,509,532]" gridcol="1" gridrow="6" pageId="12" pageNumber="11">L / W / D</td>
<td id="76905DC0FB7E005FFDD9F792FCC1E940" box="[632,887,509,532]" gridcol="2" gridrow="6" pageId="12" pageNumber="11">3.40 / 1.60 / 1.58</td>
<td id="76905DC0FB7E005FFC3AF792FBE7E940" box="[923,1105,509,532]" gridcol="3" gridrow="6" pageId="12" pageNumber="11">3.48 / 1.78 / 1.85</td>
<td id="76905DC0FB7E005FFB2CF792FAF5E940" box="[1165,1347,509,532]" gridcol="4" gridrow="6" pageId="12" pageNumber="11">3.00 / 1.63 / 1.58</td>
</tr>
<tr id="354134BCFB7E005FFEA2F449FAF5E969" box="[259,1347,550,573]" gridrow="7" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F449FE6EE969" box="[259,472,550,573]" gridcol="0" gridrow="7" pageId="12" pageNumber="11">Segment III</th>
<td id="76905DC0FB7E005FFE58F449FDE9E969" box="[505,607,550,573]" gridcol="1" gridrow="7" pageId="12" pageNumber="11">L / W / D</td>
<td id="76905DC0FB7E005FFDD9F449FCC1E969" box="[632,887,550,573]" gridcol="2" gridrow="7" pageId="12" pageNumber="11">3.80 / 1.73 / 1.73</td>
<td id="76905DC0FB7E005FFC3AF449FBE7E969" box="[923,1105,550,573]" gridcol="3" gridrow="7" pageId="12" pageNumber="11">3.85 / 1.85 / 1.90</td>
<td id="76905DC0FB7E005FFB2CF449FAF5E969" box="[1165,1347,550,573]" gridcol="4" gridrow="7" pageId="12" pageNumber="11">3.43 / 1.68 / 1.73</td>
</tr>
<tr id="354134BCFB7E005FFEA2F43FFAF5E933" box="[259,1347,592,615]" gridrow="8" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F43FFE6EE933" box="[259,472,592,615]" gridcol="0" gridrow="8" pageId="12" pageNumber="11">Segment IV</th>
<td id="76905DC0FB7E005FFE58F43FFDE9E933" box="[505,607,592,615]" gridcol="1" gridrow="8" pageId="12" pageNumber="11">L / W / D</td>
<td id="76905DC0FB7E005FFDD9F43FFCC1E933" box="[632,887,592,615]" gridcol="2" gridrow="8" pageId="12" pageNumber="11">4.43 / 1.73 / 1.74</td>
<td id="76905DC0FB7E005FFC3AF43FFBE7E933" box="[923,1105,592,615]" gridcol="3" gridrow="8" pageId="12" pageNumber="11">4.35 / 1.88 / 1.95</td>
<td id="76905DC0FB7E005FFB2CF43FFAF5E933" box="[1165,1347,592,615]" gridcol="4" gridrow="8" pageId="12" pageNumber="11">4.05 / 1.75 / 1.74</td>
</tr>
<tr id="354134BCFB7E005FFEA2F416FAF5E9C4" box="[259,1347,633,656]" gridrow="9" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F416FE6EE9C4" box="[259,472,633,656]" gridcol="0" gridrow="9" pageId="12" pageNumber="11">Segment V</th>
<td id="76905DC0FB7E005FFE58F416FDE9E9C4" box="[505,607,633,656]" gridcol="1" gridrow="9" pageId="12" pageNumber="11">L / W / D</td>
<td id="76905DC0FB7E005FFDD9F416FCC1E9C4" box="[632,887,633,656]" gridcol="2" gridrow="9" pageId="12" pageNumber="11">4.55 / 1.68 / 1.50</td>
<td id="76905DC0FB7E005FFC3AF416FBE7E9C4" box="[923,1105,633,656]" gridcol="3" gridrow="9" pageId="12" pageNumber="11">4.78 / 1.97 / 1.72</td>
<td id="76905DC0FB7E005FFB2CF416FAF5E9C4" box="[1165,1347,633,656]" gridcol="4" gridrow="9" pageId="12" pageNumber="11">4.45 / 1.78 / 1.50</td>
</tr>
<tr id="354134BCFB7E005FFEA2F4CDFAF5E9ED" box="[259,1347,674,697]" gridrow="10" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F4CDFE6EE9ED" box="[259,472,674,697]" gridcol="0" gridrow="10" pageId="12" pageNumber="11">Telson</th>
<td id="76905DC0FB7E005FFE58F4CDFDE9E9ED" box="[505,607,674,697]" gridcol="1" gridrow="10" pageId="12" pageNumber="11">L</td>
<td id="76905DC0FB7E005FFDD9F4CDFCC1E9ED" box="[632,887,674,697]" gridcol="2" gridrow="10" pageId="12" pageNumber="11">4.40</td>
<td id="76905DC0FB7E005FFC3AF4CDFBE7E9ED" box="[923,1105,674,697]" gridcol="3" gridrow="10" pageId="12" pageNumber="11">4.15</td>
<td id="76905DC0FB7E005FFB2CF4CDFAF5E9ED" box="[1165,1347,674,697]" gridcol="4" gridrow="10" pageId="12" pageNumber="11">4.05</td>
</tr>
<tr id="354134BCFB7E005FFEA2F4A3FAF5E9B7" box="[259,1347,716,739]" gridrow="11" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F4A3FE6EE9B7" box="[259,472,716,739]" gridcol="0" gridrow="11" pageId="12" pageNumber="11">Vesicle</th>
<td id="76905DC0FB7E005FFE58F4A3FDE9E9B7" box="[505,607,716,739]" gridcol="1" gridrow="11" pageId="12" pageNumber="11">L / W / D</td>
<td id="76905DC0FB7E005FFDD9F4A3FCC1E9B7" box="[632,887,716,739]" gridcol="2" gridrow="11" pageId="12" pageNumber="11">2.63 / 1.05 / 1.10</td>
<td id="76905DC0FB7E005FFC3AF4A3FBE7E9B7" box="[923,1105,716,739]" gridcol="3" gridrow="11" pageId="12" pageNumber="11">2.65 / 1.40 / 1.30</td>
<td id="76905DC0FB7E005FFB2CF4A3FAF5E9B7" box="[1165,1347,716,739]" gridcol="4" gridrow="11" pageId="12" pageNumber="11">2.60 / 1.30 / 1.20</td>
</tr>
<tr id="354134BCFB7E005FFEA2F49AFAF5E858" box="[259,1347,757,780]" gridrow="12" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F49AFE6EE858" box="[259,472,757,780]" gridcol="0" gridrow="12" pageId="12" pageNumber="11">Aculeus</th>
<td id="76905DC0FB7E005FFE58F49AFDE9E858" box="[505,607,757,780]" gridcol="1" gridrow="12" pageId="12" pageNumber="11">L</td>
<td id="76905DC0FB7E005FFDD9F49AFCC1E858" box="[632,887,757,780]" gridcol="2" gridrow="12" pageId="12" pageNumber="11">1.77</td>
<td id="76905DC0FB7E005FFC3AF49AFBE7E858" box="[923,1105,757,780]" gridcol="3" gridrow="12" pageId="12" pageNumber="11">1.50</td>
<td id="76905DC0FB7E005FFB2CF49AFAF5E858" box="[1165,1347,757,780]" gridcol="4" gridrow="12" pageId="12" pageNumber="11">1.45</td>
</tr>
<tr id="354134BCFB7E005FFEA2F571FAF5E861" box="[259,1347,798,821]" gridrow="13" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F571FE6EE861" box="[259,472,798,821]" gridcol="0" gridrow="13" pageId="12" pageNumber="11">Pedipalp</th>
<td id="76905DC0FB7E005FFE58F571FDE9E861" box="[505,607,798,821]" gridcol="1" gridrow="13" pageId="12" pageNumber="11">L</td>
<td id="76905DC0FB7E005FFDD9F571FCC1E861" box="[632,887,798,821]" gridcol="2" gridrow="13" pageId="12" pageNumber="11">11.08</td>
<td id="76905DC0FB7E005FFC3AF571FBE7E861" box="[923,1105,798,821]" gridcol="3" gridrow="13" pageId="12" pageNumber="11">11.15</td>
<td id="76905DC0FB7E005FFB2CF571FAF5E861" box="[1165,1347,798,821]" gridcol="4" gridrow="13" pageId="12" pageNumber="11">10.65</td>
</tr>
<tr id="354134BCFB7E005FFEA2F527FAF5E80B" box="[259,1347,840,863]" gridrow="14" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F527FE6EE80B" box="[259,472,840,863]" gridcol="0" gridrow="14" pageId="12" pageNumber="11">Femur</th>
<td id="76905DC0FB7E005FFE58F527FDE9E80B" box="[505,607,840,863]" gridcol="1" gridrow="14" pageId="12" pageNumber="11">L / W</td>
<td id="76905DC0FB7E005FFDD9F527FCC1E80B" box="[632,887,840,863]" gridcol="2" gridrow="14" pageId="12" pageNumber="11">3.03 / 0.65</td>
<td id="76905DC0FB7E005FFC3AF527FBE7E80B" box="[923,1105,840,863]" gridcol="3" gridrow="14" pageId="12" pageNumber="11">2.95 / 0.75</td>
<td id="76905DC0FB7E005FFB2CF527FAF5E80B" box="[1165,1347,840,863]" gridcol="4" gridrow="14" pageId="12" pageNumber="11">2.75 / 0.70</td>
</tr>
<tr id="354134BCFB7E005FFEA2F51EFAF5E8DC" box="[259,1347,881,904]" gridrow="15" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F51EFE6EE8DC" box="[259,472,881,904]" gridcol="0" gridrow="15" pageId="12" pageNumber="11">Patella</th>
<td id="76905DC0FB7E005FFE58F51EFDE9E8DC" box="[505,607,881,904]" gridcol="1" gridrow="15" pageId="12" pageNumber="11">L / W</td>
<td id="76905DC0FB7E005FFDD9F51EFCC1E8DC" box="[632,887,881,904]" gridcol="2" gridrow="15" pageId="12" pageNumber="11">3.40 / 0.95</td>
<td id="76905DC0FB7E005FFC3AF51EFBE7E8DC" box="[923,1105,881,904]" gridcol="3" gridrow="15" pageId="12" pageNumber="11">3.40 / 1.08</td>
<td id="76905DC0FB7E005FFB2CF51EFAF5E8DC" box="[1165,1347,881,904]" gridcol="4" gridrow="15" pageId="12" pageNumber="11">3.25 / 1.00</td>
</tr>
<tr id="354134BCFB7E005FFEA2F5F5FAF5E8E5" box="[259,1347,922,945]" gridrow="16" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F5F5FE6EE8E5" box="[259,472,922,945]" gridcol="0" gridrow="16" pageId="12" pageNumber="11">Chela</th>
<td id="76905DC0FB7E005FFE58F5F5FDE9E8E5" box="[505,607,922,945]" gridcol="1" gridrow="16" pageId="12" pageNumber="11">L</td>
<td id="76905DC0FB7E005FFDD9F5F5FCC1E8E5" box="[632,887,922,945]" gridcol="2" gridrow="16" pageId="12" pageNumber="11">4.65</td>
<td id="76905DC0FB7E005FFC3AF5F5FBE7E8E5" box="[923,1105,922,945]" gridcol="3" gridrow="16" pageId="12" pageNumber="11">4.80</td>
<td id="76905DC0FB7E005FFB2CF5F5FAF5E8E5" box="[1165,1347,922,945]" gridcol="4" gridrow="16" pageId="12" pageNumber="11">4.65</td>
</tr>
<tr id="354134BCFB7E005FFEA2F5ABFAF5E88F" box="[259,1347,964,987]" gridrow="17" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F5ABFE6EE88F" box="[259,472,964,987]" gridcol="0" gridrow="17" pageId="12" pageNumber="11">Manus</th>
<td id="76905DC0FB7E005FFE58F5ABFDE9E88F" box="[505,607,964,987]" gridcol="1" gridrow="17" pageId="12" pageNumber="11">L / W / D</td>
<td id="76905DC0FB7E005FFDD9F5ABFCC1E88F" box="[632,887,964,987]" gridcol="2" gridrow="17" pageId="12" pageNumber="11">1.35 / 0.75 / 0.70</td>
<td id="76905DC0FB7E005FFC3AF5ABFBE7E88F" box="[923,1105,964,987]" gridcol="3" gridrow="17" pageId="12" pageNumber="11">1.35 / 0.86 / 0.85</td>
<td id="76905DC0FB7E005FFB2CF5ABFAF5E88F" box="[1165,1347,964,987]" gridcol="4" gridrow="17" pageId="12" pageNumber="11">1.11 / 0.75 / 0.75</td>
</tr>
<tr id="354134BCFB7E005FFEA2F582FAF5EF50" box="[259,1347,1005,1028]" gridrow="18" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F582FE6EEF50" box="[259,472,1005,1028]" gridcol="0" gridrow="18" pageId="12" pageNumber="11">Movable finger</th>
<td id="76905DC0FB7E005FFE58F582FDE9EF50" box="[505,607,1005,1028]" gridcol="1" gridrow="18" pageId="12" pageNumber="11">L</td>
<td id="76905DC0FB7E005FFDD9F582FCC1EF50" box="[632,887,1005,1028]" gridcol="2" gridrow="18" pageId="12" pageNumber="11">3.30</td>
<td id="76905DC0FB7E005FFC3AF582FBE7EF50" box="[923,1105,1005,1028]" gridcol="3" gridrow="18" pageId="12" pageNumber="11">3.45</td>
<td id="76905DC0FB7E005FFB2CF582FAF5EF50" box="[1165,1347,1005,1028]" gridcol="4" gridrow="18" pageId="12" pageNumber="11">3.54</td>
</tr>
<tr id="354134BCFB7E005FFEA2F272FAF5EF60" box="[259,1347,1053,1076]" gridrow="19" pageId="12" pageNumber="11">
<th id="76905DC0FB7E005FFEA2F272FE6EEF60" box="[259,472,1053,1076]" gridcol="0" gridrow="19" pageId="12" pageNumber="11">Total</th>
<td id="76905DC0FB7E005FFE58F272FDE9EF60" box="[505,607,1053,1076]" gridcol="1" gridrow="19" pageId="12" pageNumber="11">L</td>
<td id="76905DC0FB7E005FFDD9F272FCC1EF60" box="[632,887,1053,1076]" gridcol="2" gridrow="19" pageId="12" pageNumber="11">34.13</td>
<td id="76905DC0FB7E005FFC3AF272FBE7EF60" box="[923,1105,1053,1076]" gridcol="3" gridrow="19" pageId="12" pageNumber="11">36.48</td>
<td id="76905DC0FB7E005FFB2CF272FAF5EF60" box="[1165,1347,1053,1076]" gridcol="4" gridrow="19" pageId="12" pageNumber="11">32.13</td>
</tr>
</table>
</paragraph>
<caption id="DF0E6676FB7EFFACFF61F231FED4EFDA" pageId="12" pageNumber="11" startId="12.[192,257,1118,1141]">
<paragraph id="8BCE36FEFB7EFFACFF61F231FED4EFDA" blockId="12.[192,1440,1118,1166]" pageId="12" pageNumber="11">
<emphasis id="B905EAECFB7EFFACFF61F231FE94EF21" bold="true" box="[192,290,1118,1141]" pageId="12" pageNumber="11">Table 1:</emphasis>
Measurements of three adults of
<taxonomicName id="4C714D7DFB7EFFACFDDAF20FFCCAEF21" authority="rickmersi." authorityName="Rickmersi." box="[635,892,1120,1141]" class="Arachnida" family="Buthidae" genus="Anomalobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAECFB7EFFACFDDAF20FFCC1EF21" box="[635,887,1120,1141]" italics="true" pageId="12" pageNumber="11">Anomalobuthus rickmersi</emphasis>
.
</taxonomicName>
Abbreviations: length (L), width (W), posterior width (Wp), depth (D).
</paragraph>
</caption>
<paragraph id="8BCE36FEFB7EFFACFF51F3F5FEC8EC29" blockId="12.[192,802,1219,1917]" pageId="12" pageNumber="11">
It is clear from A. Birula's publications that he never studied the
<typeStatus id="54CA885CFB7EFFACFEE7F3D7FE15EE9B" box="[326,419,1464,1487]" pageId="12" pageNumber="11" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C714D7DFB7EFFACFE72F3D7FDEDEE9B" authorityName="Kraepelin" authorityYear="1899" box="[467,603,1464,1487]" class="Arachnida" family="Gylippidae" genus="Gylippus" kingdom="Animalia" order="Solifugae" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="rickmersi">
<emphasis id="B905EAECFB7EFFACFE72F3D7FDEDEE9B" box="[467,603,1464,1487]" italics="true" pageId="12" pageNumber="11">A. rickmersi</emphasis>
</taxonomicName>
; however, it was seen and its trichobothrial pattern sketched by
<bibRefCitation id="EFE04B0FFB7EFFACFD6EF3B8FE81ED59" author="VACHON" pageId="12" pageNumber="11" pagination="857 - 958" refId="ref26267" refString="VACHON, M. 1974. Etudes des caracteres utilises pour classer les familles et les genres des scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, 3 e serie, 140 (Zoologie, 104): 857 - 958." type="journal article" year="1974">Vachon (1974: 908</bibRefCitation>
, fig. 37). On the other hand,
<bibRefCitation id="EFE04B0FFB7EFFACFDCFF399FC94ED59" author="LOURENCO" box="[622,802,1526,1549]" pageId="12" pageNumber="11" pagination="15 - 20" refId="ref25502" refString="LOURENCO, W. R. 2001. Un nouveau genre de Buthidae, probable vicariant geographique d'Anomalobuthus Kraepelin (Chelicerata, Scorpiones). Biogeographica, 77 (1): 15 - 20." type="journal article" year="2001">Lourenço (2001)</bibRefCitation>
claimed to have examined it; however, it is obvious that the specimen he received on loan from ZMUH was
<emphasis id="B905EAECFB7EFFACFCA1F05CFC94ED1E" box="[768,802,1587,1610]" italics="true" pageId="12" pageNumber="11">not</emphasis>
Kraepelin's original
<typeStatus id="54CA885CFB7EFFACFE07F03DFDB0ED3D" box="[422,518,1618,1641]" pageId="12" pageNumber="11" type="holotype">holotype</typeStatus>
, but a misidentified and wrongly labeled juvenile of maybe even another genus.
<bibRefCitation id="EFE04B0FFB7EFFACFF61F0E0FEC0EDF2" author="LOURENCO" box="[192,374,1679,1702]" pageId="12" pageNumber="11" pagination="15 - 20" refId="ref25502" refString="LOURENCO, W. R. 2001. Un nouveau genre de Buthidae, probable vicariant geographique d'Anomalobuthus Kraepelin (Chelicerata, Scorpiones). Biogeographica, 77 (1): 15 - 20." type="journal article" year="2001">Lourenço (2001)</bibRefCitation>
declared this specimen as an immature with a total length of
<quantity id="4C899B1BFB7EFFACFE63F0C1FD8DED91" box="[450,571,1710,1733]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="12" pageNumber="11" unit="mm" value="13.5" valueMax="14.0" valueMin="13.0">13–14 mm</quantity>
[
<emphasis id="B905EAECFB7EFFACFDF1F0C1FDD8ED91" box="[592,622,1710,1733]" italics="true" pageId="12" pageNumber="11">sic</emphasis>
], pectinal tooth count 24–26, and textually wrote: “...
<emphasis id="B905EAECFB7EFFACFDF8F0A3FDCBEC56" italics="true" pageId="12" pageNumber="11">cutting edge of the movable finger with 6 series of granules</emphasis>
[...]
<emphasis id="B905EAECFB7EFFACFD14F084FEB1EC6B" italics="true" pageId="12" pageNumber="11">the basalmost extending onto more than half the length of the finger.</emphasis>
[... Pectinal]
<emphasis id="B905EAECFB7EFFACFE03F147FCA0EC6B" box="[418,790,1832,1855]" italics="true" pageId="12" pageNumber="11">Fulcra almost vestigial and flat...</emphasis>
” [originally in French, English translation and bracketed text added by us].
</paragraph>
<paragraph id="8BCE36FEFB7EFFACFCCFF2ACFAFEED7F" blockId="12.[830,1440,1219,1579]" pageId="12" pageNumber="11">
None of these crucial characters match the
<typeStatus id="54CA885CFB7EFFACFAE2F2ACFA16EF8E" box="[1347,1440,1219,1242]" pageId="12" pageNumber="11" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C714D7DFB7EFFACFCC1F28DFC50EFAD" authorityName="Kraepelin" authorityYear="1899" box="[864,998,1250,1273]" class="Arachnida" family="Gylippidae" genus="Gylippus" kingdom="Animalia" order="Solifugae" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="rickmersi">
<emphasis id="B905EAECFB7EFFACFCC1F28DFC50EFAD" box="[864,998,1250,1273]" italics="true" pageId="12" pageNumber="11">A. rickmersi</emphasis>
</taxonomicName>
and clearly contradict the original description and illustrations published by
<bibRefCitation id="EFE04B0FFB7EFFACFB40F36FFA2AEE43" author="KRAEPELIN" box="[1249,1436,1280,1303]" pageId="12" pageNumber="11" pagination="1 - 17" refId="ref25373" refString="KRAEPELIN, K. 1900. Uber einige neue Gliederspinnen. Abhandlungen aus dem Gebiete der Naturwissenschaften. Herausgegeben vom Naturwissenschaftlichen Verein in Hamburg, 16, 1 (4): 1 - 17." type="journal article" year="1900">Kraepelin (1900)</bibRefCitation>
, which were confirmed by our own examination of the true type: it is a large adult male with a total length of
<quantity id="4C899B1BFB7EFFACFA24F351FCDCEE27" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="11" unit="mm" value="34.0">34 mm</quantity>
, pectinal tooth count 25/25, fingers with 10 rows of primary denticles of which the basal-most extends onto less than one-third of finger, and fulcra normally developed and pearl-like. Finally, a photo of the label of the specimen examined by
<bibRefCitation id="EFE04B0FFB7EFFACFBD4F3B8FA87EEBA" author="LOURENCO" box="[1141,1329,1495,1518]" pageId="12" pageNumber="11" pagination="15 - 20" refId="ref25502" refString="LOURENCO, W. R. 2001. Un nouveau genre de Buthidae, probable vicariant geographique d'Anomalobuthus Kraepelin (Chelicerata, Scorpiones). Biogeographica, 77 (1): 15 - 20." type="journal article" year="2001">Lourenço (2001)</bibRefCitation>
, provided by himself to V.F.in 2009, does not correspond to any of those accompanying the actual
<typeStatus id="54CA885CFB7EFFACFB2CF07BFB5CED7F" box="[1165,1258,1556,1579]" pageId="12" pageNumber="11" type="holotype">holotype</typeStatus>
(
<figureCitation id="134A2A7BFB7EFFACFB58F07BFA8CED7F" box="[1273,1338,1556,1579]" captionStart="Figures 1–3" captionStartId="5.[192,278,1579,1602]" captionTargetBox="[191,1440,191,1550]" captionTargetId="figure-6@5.[192,1439,192,1549]" captionTargetPageId="5" captionText="Figures 1–3: Anomalobuthus rickmersi, adult male holotype: full-body views, dorsal (1) and ventral (2), plus original labels (3). Scale bar = 10 mm (1–2). The red inset at bottom right shows the label of the specimen examined by Lourenço (2001), demonstrating it was not the actual Rickmers's type specimen." pageId="12" pageNumber="11">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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