607 lines
107 KiB
XML
607 lines
107 KiB
XML
<document id="6C4A65418EA23C66FAC3FCBFFFAB0129" ID-CLB-Dataset="295176" ID-DOI="10.4202/app.00922.2021" ID-GBIF-Dataset="3051108e-586a-4af3-a53b-4e30fc817abb" ID-ISSN="1732-2421" ID-Zenodo-Dep="10626754" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1707250717328" checkinUser="felipe" docAuthor="Westrop, Stephen R. & Dengler, Alyce A." docDate="2022" docId="03D887C549697F58FC89FD7B5DD574AB" docLanguage="en" docName="ActaPalaeontolPol.67.1.27-33.pdf" docOrigin="Acta Palaeontologica Polonica 67 (1)" docSource="http://dx.doi.org/10.4202/app.00922.2021" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.4:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="4" docTitle="Catillicephala cifellii Westrop & Dengler 2022, sp. nov." docType="treatment" docVersion="4" lastPageNumber="32" masterDocId="FFE1FFBD49687F5DFFA1FF925E6B701A" masterDocTitle="A new Cambrian catillicephalid trilobite from the Shallow Bay Formation of western Newfoundland, Canada" masterLastPageNumber="33" masterPageNumber="27" pageNumber="28" updateTime="1714489733347" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2BCB2F470E1EE8843502B6E51A993AFE">A new Cambrian catillicephalid trilobite from the Shallow Bay Formation of western Newfoundland, Canada</mods:title>
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<mods:namePart id="22FE4EE8DD064C038A1D7F97026E4BA3">Westrop, Stephen R.</mods:namePart>
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<mods:roleTerm id="79DD67A823A6D1982CF5BF49F91AF7B2">Author</mods:roleTerm>
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<mods:namePart id="063F53140346482AEDB3611D95A2AE7A">Dengler, Alyce A.</mods:namePart>
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<mods:typeOfResource id="6CE66A1614B946BF3478B10586E03646">text</mods:typeOfResource>
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<mods:title id="A028C8056B6660196DF318C6B24477E1">Acta Palaeontologica Polonica</mods:title>
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<mods:part id="B4D39F0CBB970A003D018C77E64F627D">
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<mods:date id="AC2F6DA18BC9CBA1D41E13FEE32A9AFE">2022</mods:date>
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<mods:number id="F9235A48E8D40A2234EC9109E0E4669E">2022-03-30</mods:number>
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<mods:url id="FED24B627C0A3BF1D50DCEA840216E6B">http://dx.doi.org/10.4202/app.00922.2021</mods:url>
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<treatment id="03D887C549697F58FC89FD7B5DD574AB" ID-DOI="http://doi.org/10.5281/zenodo.11093738" ID-Zenodo-Dep="11093738" LSID="urn:lsid:plazi:treatment:03D887C549697F58FC89FD7B5DD574AB" httpUri="http://treatment.plazi.org/id/03D887C549697F58FC89FD7B5DD574AB" lastPageId="5" lastPageNumber="32" pageId="1" pageNumber="28">
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<subSubSection id="C36B655849697F5CFC89FD7B5AC8731F" box="[808,1187,745,774]" pageId="1" pageNumber="28" type="nomenclature">
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<paragraph id="8BCE36D349697F5CFC89FD7B5AC8731F" blockId="1.[808,1187,745,774]" box="[808,1187,745,774]" pageId="1" pageNumber="28">
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<heading id="D08681BF49697F5CFC89FD7B5AC8731F" box="[808,1187,745,774]" fontSize="12" level="2" pageId="1" pageNumber="28" reason="2">
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<taxonomicName id="4C714D5049697F5CFC89FD7B5A52731C" authority="Westrop & Dengler, 2022" authorityName="Westrop & Dengler" authorityYear="2022" box="[808,1081,745,774]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="cifellii" status="sp. nov.">
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<emphasis id="B905EAC149697F5CFC89FD7B5A52731C" box="[808,1081,745,774]" italics="true" pageId="1" pageNumber="28">Catillicephala cifellii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23657BA49697F5CFBE0FD7B5AC8731F" box="[1089,1187,745,773]" pageId="1" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B655849697F5CFC89FC875D957374" pageId="1" pageNumber="28" type="description">
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<paragraph id="8BCE36D349697F5CFC89FC875DED7331" blockId="1.[808,902,789,811]" box="[808,902,789,811]" pageId="1" pageNumber="28">
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<figureCitation id="134A2A5649697F5CFC89FC875DE97331" box="[808,898,789,811]" captionStart-0="Fig 1" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="2.[151,181,1711,1733]" captionStartId-1="3.[98,131,1712,1734]" captionStartId-2="3.[98,131,1852,1874]" captionTargetBox-0="[154,1533,161,1686]" captionTargetBox-2="[98,1478,431,1687]" captionTargetId-0="figure-81@2.[154,1533,161,1686]" captionTargetId-1="figure-155@3.[98,1478,431,1687]" captionTargetId-2="figure-155@3.[98,1478,431,1687]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionText-0="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." captionText-1="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." captionText-2="Fig.3. Pygidia of catillicephalid trilobite Catillicephalacifellii sp.nov.from the Downes Point Member,Shallow BayFormation,Guzhangian,Miaolingian, → Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142520, paratype; dorsal (A1), lateral (A2), and posterior (A3) views. B. GSC 142521, paratype; posterior (B1), dorsal (B2), and lateral (B3) views. C. GSC 142522, paratype; posterior (C1) and dorsal (C2) views. D. GSC 142523, paratype; posterior (D1) and dorsal (D2) views. E. GSC 142524, paratype; dorsal view. Scale bars 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10626756" figureDoi-1="http://doi.org/10.5281/zenodo.11093736" figureDoi-2="http://doi.org/10.5281/zenodo.10626760" httpUri-0="https://zenodo.org/record/10626756/files/figure.png" httpUri-1="https://zenodo.org/record/11093736/files/figure.png" httpUri-2="https://zenodo.org/record/10626760/files/figure.png" pageId="1" pageNumber="28">Figs. 1–3</figureCitation>
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.
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</paragraph>
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<paragraph id="8BCE36D349697F5CFC89FCAE5D957374" blockId="1.[808,1481,828,878]" pageId="1" pageNumber="28">
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<emphasis id="B905EAC149697F5CFC89FCAE5DAA7348" box="[808,961,828,850]" italics="true" pageId="1" pageNumber="28">Zoobank LCID</emphasis>
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:
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<uri id="FFE03AD149697F5CFC74FCAE5D957374" pageId="1" pageNumber="28">
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urn:lsid:zoobank.org:act:
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<uuid id="FFD70C0649697F5CFB63FCAE5D957374" pageId="1" pageNumber="28">4B4919F0-DFB7-4569-B8 B1-1A2DE79BACFD</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B655849697F5CFC89FCE95B6473B7" pageId="1" pageNumber="28" type="etymology">
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<paragraph id="8BCE36D349697F5CFC89FCE95B6473B7" blockId="1.[808,1481,891,941]" pageId="1" pageNumber="28">
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<emphasis id="B905EAC149697F5CFC89FCE95DE4738A" box="[808,911,891,912]" italics="true" pageId="1" pageNumber="28">Etymology</emphasis>
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: In honour of Richard Cifelli upon his retirement and in recognition of his studies of Cretaceous mammals.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36B655849697F5CFC89FC285AEE7533" pageId="1" pageNumber="28" type="materials_examined">
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||
<paragraph id="8BCE36D349697F5CFC89FC285B6E7447" blockId="1.[808,1481,953,1117]" pageId="1" pageNumber="28">
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||
<emphasis id="B905EAC149697F5CFC89FC285DD973D4" box="[808,946,953,975]" italics="true" pageId="1" pageNumber="28">Type material</emphasis>
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||
:
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||
<typeStatus id="54CA887149697F5CFC67FC2B5A4873D5" box="[966,1059,953,975]" pageId="1" pageNumber="28">Holotype</typeStatus>
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||
: GSC 142512, well preserved cranidium (
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||
<figureCitation id="134A2A5649697F5CFC8EFC475D1073F1" box="[815,891,981,1003]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="1" pageNumber="28">Fig. 1A</figureCitation>
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).
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<materialsCitation id="3B193C8E49697F5CFC2DFC445AC9743E" collectionCode="BPS" pageId="1" pageNumber="28" specimenCode="GSC 142513-142515, GSC 142517, GSC 142519, GSC 142520- 142524, BPS 467" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54CA887149697F5CFC2DFC445D8673F1" box="[908,1005,982,1003]" pageId="1" pageNumber="28">Paratypes</typeStatus>
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||
: five cranidia (
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||
<specimenCode id="DBD79EA849697F5CFBDBFC475B2573F1" box="[1146,1358,981,1003]" pageId="1" pageNumber="28">GSC 142513–142515</specimenCode>
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,
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<figureCitation id="134A2A5649697F5CFAF6FC475BE473F0" box="[1367,1423,981,1003]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="1" pageNumber="28">Fig. 1</figureCitation>
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;
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<specimenCode id="DBD79EA849697F5CFA3BFC475D197412" pageId="1" pageNumber="28">GSC 142517</specimenCode>
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,
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<figureCitation id="134A2A5649697F5CFCDCFC605DA3741D" box="[893,968,1010,1032]" captionStart="Fig" captionStartId="3.[98,131,1712,1734]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="1" pageNumber="28">Fig. 2B</figureCitation>
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;
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<specimenCode id="DBD79EA849697F5CFC72FC605A387412" box="[979,1107,1010,1032]" pageId="1" pageNumber="28">GSC 142519</specimenCode>
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,
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<figureCitation id="134A2A5649697F5CFBFCFC605AC17412" box="[1117,1194,1010,1032]" captionStart="Fig" captionStartId="3.[98,131,1712,1734]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="1" pageNumber="28">Fig. 2D</figureCitation>
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), five pygidia (
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<specimenCode id="DBD79EA849697F5CFA9FFC605D19743E" pageId="1" pageNumber="28">GSC 142520– 142524</specimenCode>
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,
|
||
<figureCitation id="134A2A5649697F5CFCDCFB9C5DDE743E" box="[893,949,1038,1060]" captionStart="Fig" captionStartId="3.[98,131,1852,1874]" captionTargetBox="[98,1478,431,1687]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig.3. Pygidia of catillicephalid trilobite Catillicephalacifellii sp.nov.from the Downes Point Member,Shallow BayFormation,Guzhangian,Miaolingian, → Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142520, paratype; dorsal (A1), lateral (A2), and posterior (A3) views. B. GSC 142521, paratype; posterior (B1), dorsal (B2), and lateral (B3) views. C. GSC 142522, paratype; posterior (C1) and dorsal (C2) views. D. GSC 142523, paratype; posterior (D1) and dorsal (D2) views. E. GSC 142524, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626760" httpUri="https://zenodo.org/record/10626760/files/figure.png" pageId="1" pageNumber="28">Fig. 3</figureCitation>
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||
) from boulder
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||
<specimenCode id="DBD79EA849697F5CFBE8FB9C5AC9743E" box="[1097,1186,1038,1060]" collectionCode="BPS" pageId="1" pageNumber="28">BPS 467</specimenCode>
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||
</materialsCitation>
|
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;
|
||
<materialsCitation id="3B193C8E49697F5CFB0DFB9D5A3E745B" collectionCode="BPS" pageId="1" pageNumber="28" specimenCode="GSC 142516, BPS 468" specimenCount="1" typeStatus="paratype">
|
||
one cranidium (
|
||
<specimenCode id="DBD79EA849697F5CFAE5FB9C5BAE743E" box="[1348,1477,1038,1060]" pageId="1" pageNumber="28">GSC 142516</specimenCode>
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||
,
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||
<figureCitation id="134A2A5649697F5CFC89FBB95D19745B" box="[808,882,1067,1089]" captionStart="Fig" captionStartId="3.[98,131,1712,1734]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="1" pageNumber="28">Fig. 2A</figureCitation>
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||
) from boulder
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||
<specimenCode id="DBD79EA849697F5CFC5EFBB95A3E745B" box="[1023,1109,1067,1089]" collectionCode="BPS" pageId="1" pageNumber="28">BPS 468</specimenCode>
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||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B193C8E49697F5CFBFFFBB95A6F7447" collectionCode="BPS" pageId="1" pageNumber="28" specimenCode="GSC 142518, BPS 458" specimenCount="1" typeStatus="paratype">
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||
one cranidium (
|
||
<specimenCode id="DBD79EA849697F5CFB53FBB95B1A745A" box="[1266,1393,1066,1088]" pageId="1" pageNumber="28">GSC 142518</specimenCode>
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||
,
|
||
<figureCitation id="134A2A5649697F5CFADBFBB95BA9745B" box="[1402,1474,1067,1089]" captionStart="Fig" captionStartId="3.[98,131,1712,1734]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="1" pageNumber="28">Fig. 2C</figureCitation>
|
||
) from boulder
|
||
<specimenCode id="DBD79EA849697F5CFC0DFBD55A6F7447" box="[940,1028,1095,1117]" collectionCode="BPS" pageId="1" pageNumber="28">BPS 458</specimenCode>
|
||
</materialsCitation>
|
||
. All from the type locality.
|
||
</paragraph>
|
||
<paragraph id="8BCE36D349697F5CFC89FBFB5BC47465" blockId="1.[808,1455,1129,1151]" box="[808,1455,1129,1151]" pageId="1" pageNumber="28">
|
||
<materialsCitation id="3B193C8E49697F5CFC89FBFB5BC07465" box="[808,1451,1129,1151]" country="Canada" location="Broom Point South" municipality="western Newfoundland" pageId="1" pageNumber="28" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B905EAC149697F5CFC89FBFB5DC87464" box="[808,931,1129,1150]" italics="true" pageId="1" pageNumber="28">
|
||
<typeStatus id="54CA887149697F5CFC89FBFB5D3E7464" box="[808,853,1129,1150]" pageId="1" pageNumber="28">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
:
|
||
<location id="8EAE600849697F5CFC11FBFB5A007465" LSID="urn:lsid:plazi:treatment:03D887C549697F58FC89FD7B5DD574AB:8EAE600849697F5CFC11FBFB5A007465" box="[944,1131,1129,1151]" country="Canada" municipality="western Newfoundland" name="Broom Point South" pageId="1" pageNumber="28">Broom Point South</location>
|
||
,
|
||
<collectingMunicipality id="6BAAACA949697F5CFBD4FBFB5B3C7465" box="[1141,1367,1129,1151]" pageId="1" pageNumber="28">western Newfoundland</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F366764349697F5CFAC1FBFB5BC07465" box="[1376,1451,1129,1151]" name="Canada" pageId="1" pageNumber="28">Canada</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCE36D349697F5CFC89FB195A9474A7" blockId="1.[808,1481,1162,1213]" pageId="1" pageNumber="28">
|
||
<materialsCitation id="3B193C8E49697F5CFC89FB195A9074A7" collectionCode="BPS" county="Cambrian" location="Downes Point Member" municipality="Miaolingian" pageId="1" pageNumber="28" specimenCode="BPS 467" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B905EAC149697F5CFC89FB195DCF74BA" box="[808,932,1163,1184]" italics="true" pageId="1" pageNumber="28">
|
||
<typeStatus id="54CA887149697F5CFC89FB195D3E74BA" box="[808,853,1163,1184]" pageId="1" pageNumber="28">Type</typeStatus>
|
||
horizon
|
||
</emphasis>
|
||
: Boulder
|
||
<specimenCode id="DBD79EA849697F5CFBA5FB195A3174BA" box="[1028,1114,1162,1184]" collectionCode="BPS" pageId="1" pageNumber="28">BPS 467</specimenCode>
|
||
,
|
||
<location id="8EAE600849697F5CFBC5FB195B2F74BB" LSID="urn:lsid:plazi:treatment:03D887C549697F58FC89FD7B5DD574AB:8EAE600849697F5CFBC5FB195B2F74BB" box="[1124,1348,1163,1185]" county="Cambrian" municipality="Miaolingian" name="Downes Point Member" pageId="1" pageNumber="28">Downes Point Member</location>
|
||
,
|
||
<location id="8EAE600849697F5CFAEFFB195DE474A7" LSID="urn:lsid:plazi:treatment:03D887C549697F58FC89FD7B5DD574AB:8EAE600849697F5CFAEFFB195DE474A7" county="Cambrian" municipality="Miaolingian" name="Shallow Bay Formation" pageId="1" pageNumber="28">
|
||
<location id="8EAE600849697F5CFAEFFB195BA274BA" LSID="urn:lsid:plazi:treatment:03D887C549697F58FC89FD7B5DD574AB:8EAE600849697F5CFAEFFB195BA274BA" box="[1358,1481,1163,1185]" county="Cambrian" municipality="Miaolingian" name="Shallow Bay" pageId="1" pageNumber="28">Shallow Bay</location>
|
||
Formation
|
||
</location>
|
||
,
|
||
<location id="8EAE600849697F5CFC39FB355A6574A7" LSID="urn:lsid:plazi:treatment:03D887C549697F58FC89FD7B5DD574AB:8EAE600849697F5CFC39FB355A6574A7" box="[920,1038,1191,1213]" county="Cambrian" municipality="Miaolingian" name="Guzhangian" pageId="1" pageNumber="28">Guzhangian</location>
|
||
,
|
||
<collectingMunicipality id="6BAAACA949697F5CFBB9FB355AFB74A7" box="[1048,1168,1191,1213]" pageId="1" pageNumber="28">Miaolingian</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62AF4E5F49697F5CFB38FB355A9074A7" box="[1177,1275,1191,1213]" pageId="1" pageNumber="28">Cambrian</collectingCounty>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCE36D349697F5CFC89FB5D5AEE7533" blockId="1.[808,1481,1231,1322]" pageId="1" pageNumber="28">
|
||
<emphasis id="B905EAC149697F5CFC89FB5D5DE174F3" box="[808,906,1231,1257]" italics="true" pageId="1" pageNumber="28">Material</emphasis>
|
||
.—
|
||
<materialsCitation id="3B193C8E49697F5CFC0AFB5D5AE97533" collectionCode="BPS" pageId="1" pageNumber="28" specimenCode="BPS 467, BPS 458" specimenCount="1">
|
||
The type material and two pygidia from boulder
|
||
<specimenCode id="DBD79EA849697F5CFC89FB7D5DE57513" box="[808,910,1263,1289]" collectionCode="BPS" pageId="1" pageNumber="28">BPS 467</specimenCode>
|
||
; two cranidia and two pygidia from boulder
|
||
<specimenCode id="DBD79EA849697F5CFA39FB7D5D3F7530" collectionCode="BPS" pageId="1" pageNumber="28">BPS 458</specimenCode>
|
||
. All from the type locality
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B655849697F5CFC89FAA95B0F758F" pageId="1" pageNumber="28" type="diagnosis">
|
||
<paragraph id="8BCE36D349697F5CFC89FAA95B0F758F" blockId="1.[808,1481,1339,1430]" pageId="1" pageNumber="28">
|
||
<emphasis id="B905EAC149697F5CFC89FAA95DF0754E" box="[808,923,1339,1364]" italics="true" pageId="1" pageNumber="28">Diagnosis</emphasis>
|
||
.—
|
||
<taxonomicName id="4C714D5049697F5CFC61FAA95A08754F" authorityName="Raymond" authorityYear="1938" box="[960,1123,1339,1365]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B905EAC149697F5CFC61FAA95A08754F" box="[960,1123,1339,1365]" italics="true" pageId="1" pageNumber="28">Catillicephala</emphasis>
|
||
</taxonomicName>
|
||
with ovoid glabella, occipital ring subtriangular, drawn out posteriorly into stout spine, and short, narrow anterior border tapered laterally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B655849697F58FC89FA355B1772DA" lastPageId="5" lastPageNumber="32" pageId="1" pageNumber="28" type="description">
|
||
<paragraph id="8BCE36D349697F58FC89FA355FEE7121" blockId="1.[808,1482,1447,1960]" lastBlockId="5.[98,771,158,935]" lastPageId="5" lastPageNumber="32" pageId="1" pageNumber="28">
|
||
<emphasis id="B905EAC149697F5CFC89FA355DDB75DA" box="[808,944,1447,1472]" italics="true" pageId="1" pageNumber="28">Description</emphasis>
|
||
.—Strongly convex cranidium subtrapezoidal in outline (excluding LO), with curved anterior margin; preoccipital glabella length 54% (51–58%) maximum cranidial width across posterolateral projections. Frontal area represented only by very narrow, rim-like anterior border. Anterior border furrow finely etched, merges with preglabellar furrow; axial furrows shallow but clearly defined grooves, bowed gently outwards. Glabella barrel-shaped in outline, with outwardly curved flanks and gently rounded anteriorly; inflated, standing well above adjacent fixigenae in anterior view (e.g.,
|
||
<figureCitation id="134A2A5649697F5CFB86F9795AE3771F" box="[1063,1160,1771,1797]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="1" pageNumber="28">Figs. 1A</figureCitation>
|
||
<figureCitation id="134A2A5649697F5CFB29F96A5AF97713" box="[1160,1170,1784,1801]" captionStart="Fig" captionStartId="3.[98,131,1852,1874]" captionTargetBox="[98,1478,431,1687]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig.3. Pygidia of catillicephalid trilobite Catillicephalacifellii sp.nov.from the Downes Point Member,Shallow BayFormation,Guzhangian,Miaolingian, → Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142520, paratype; dorsal (A1), lateral (A2), and posterior (A3) views. B. GSC 142521, paratype; posterior (B1), dorsal (B2), and lateral (B3) views. C. GSC 142522, paratype; posterior (C1) and dorsal (C2) views. D. GSC 142523, paratype; posterior (D1) and dorsal (D2) views. E. GSC 142524, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626760" httpUri="https://zenodo.org/record/10626760/files/figure.png" pageId="1" pageNumber="28">
|
||
<subScript id="17F5349649697F5CFB29F96A5AF97713" attach="left" box="[1160,1170,1784,1801]" fontSize="7" pageId="1" pageNumber="28">3</subScript>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="134A2A5649697F5CFB03F9795AAB771F" box="[1186,1216,1771,1797]" captionStart="Fig" captionStartId="3.[98,131,1712,1734]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="1" pageNumber="28">2B</figureCitation>
|
||
<figureCitation id="134A2A5649697F5CFB60F96A5AA17713" box="[1217,1226,1784,1801]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="1" pageNumber="28">
|
||
<subScript id="17F5349649697F5CFB60F96A5AA17713" attach="right" box="[1217,1226,1784,1801]" fontSize="7" pageId="1" pageNumber="28">1</subScript>
|
||
</figureCitation>
|
||
); width at anterior tip of S1 equal to 89% (81–94%; lower values in smaller specimens) preoccipital length. LO extended into stout, upwardly curved, triangular occipital spine in larger individuals (e.g.,
|
||
<figureCitation id="134A2A5649697F5CFC89F8FF5DF97791" box="[808,914,1901,1931]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="1" pageNumber="28">
|
||
Figs. 1A
|
||
<subScript id="17F5349649697F5CFC28F8E85DF97791" attach="left" box="[905,914,1914,1931]" fontSize="7" pageId="1" pageNumber="28">1</subScript>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="134A2A5649697F5CFC02F8FF5DA3779D" box="[931,968,1901,1927]" captionStart="Fig" captionStartId="3.[98,131,1712,1734]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="1" pageNumber="28">2D</figureCitation>
|
||
). SO curved gently backwards, shallow, defined in part by break in slope between LO and preoccipital glabella, particularly on larger specimens (e.g.,
|
||
<figureCitation id="134A2A56496A7F5FFD71F8BE5D40775C" box="[720,811,1836,1862]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="2" pageNumber="29">Fig. 1A</figureCitation>
|
||
). Lateral glabellar furrows barely perceptible on all but the smallest specimens (e.g.,
|
||
<figureCitation id="134A2A56496A7F5FFE61F8FE5C407790" box="[448,555,1900,1930]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="2" pageNumber="29">
|
||
Figs. 1A
|
||
<subScript id="17F53496496A7F5FFD83F8EB5C407790" attach="left" box="[546,555,1913,1930]" fontSize="7" pageId="2" pageNumber="29">1</subScript>
|
||
</figureCitation>
|
||
, B,
|
||
<figureCitation id="134A2A56496A7F5FFDC3F8FE5CE9779C" box="[610,642,1900,1926]" captionStart="Fig" captionStartId="3.[98,131,1712,1734]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="2" pageNumber="29">2B</figureCitation>
|
||
<figureCitation id="134A2A56496A7F5FFD23F8EB5CE77790" box="[642,652,1913,1930]" captionStart="Fig" captionStartId="3.[98,131,1852,1874]" captionTargetBox="[98,1478,431,1687]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig.3. Pygidia of catillicephalid trilobite Catillicephalacifellii sp.nov.from the Downes Point Member,Shallow BayFormation,Guzhangian,Miaolingian, → Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142520, paratype; dorsal (A1), lateral (A2), and posterior (A3) views. B. GSC 142521, paratype; posterior (B1), dorsal (B2), and lateral (B3) views. C. GSC 142522, paratype; posterior (C1) and dorsal (C2) views. D. GSC 142523, paratype; posterior (D1) and dorsal (D2) views. E. GSC 142524, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626760" httpUri="https://zenodo.org/record/10626760/files/figure.png" pageId="2" pageNumber="29">
|
||
<subScript id="17F53496496A7F5FFD23F8EB5CE77790" attach="left" box="[642,652,1913,1930]" fontSize="7" pageId="2" pageNumber="29">3</subScript>
|
||
</figureCitation>
|
||
, D). S1 geniculate, termination short of LO; S2 curved gently forward; S3 straight, directed obliquely forward from axial furrow. L1–L3 without independent convexity, maximum lengths (exsag.) roughly equal. Palpebral area of fixigenae narrow, equal to 25% (21–32%; higher values in smaller specimens) of glabellar width at anterior tip of S1, curved downwards in anterior view (e.g.,
|
||
<figureCitation id="134A2A56496B7F5EFEF6FF2C5FDE70C2" box="[343,437,190,216]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="3" pageNumber="30">Figs. 1A</figureCitation>
|
||
<figureCitation id="134A2A56496B7F5EFE14FF595FD470C6" box="[437,447,203,220]" captionStart="Fig" captionStartId="3.[98,131,1852,1874]" captionTargetBox="[98,1478,431,1687]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig.3. Pygidia of catillicephalid trilobite Catillicephalacifellii sp.nov.from the Downes Point Member,Shallow BayFormation,Guzhangian,Miaolingian, → Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142520, paratype; dorsal (A1), lateral (A2), and posterior (A3) views. B. GSC 142521, paratype; posterior (B1), dorsal (B2), and lateral (B3) views. C. GSC 142522, paratype; posterior (C1) and dorsal (C2) views. D. GSC 142523, paratype; posterior (D1) and dorsal (D2) views. E. GSC 142524, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626760" httpUri="https://zenodo.org/record/10626760/files/figure.png" pageId="3" pageNumber="30">
|
||
<subScript id="17F53496496B7F5EFE14FF595FD470C6" attach="left" box="[437,447,203,220]" fontSize="7" pageId="3" pageNumber="30">3</subScript>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="134A2A56496B7F5EFE6CFF2C5F8070C2" box="[461,491,190,216]" captionStart="Fig" captionStartId="3.[98,131,1712,1734]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="3" pageNumber="30">2B</figureCitation>
|
||
<figureCitation id="134A2A56496B7F5EFE4AFF595F9F70C6" box="[491,500,203,220]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="3" pageNumber="30">
|
||
<subScript id="17F53496496B7F5EFE4AFF595F9F70C6" attach="left" box="[491,500,203,220]" fontSize="7" pageId="3" pageNumber="30">1</subScript>
|
||
</figureCitation>
|
||
). Palpebral lobe narrow tr.), gently curved, length equal to 20% (16–24%) of preoccipital glabella length and centred opposite S2; palpebral furrow shallow, defined in part by break in slope between lobe and adjacent palpebral area. Palpebral ridge weakly convex, extends obliquely forward to intersect axial furrow near anterior end of glabella. Anterior branches of facial suture short, convergent; posterior branches divergent before curving backward at posterior border furrow to become subparallel. Postocular fixigenae broadly triangular in outline, cranidial width at palpebral lobe 77% (74–80%) maximum width of cranidium; posterolateral projection flexed sharply downward in anterior view (e.g.,
|
||
<figureCitation id="134A2A56496B7F5EFB50FEAC5B2F7142" box="[1265,1348,318,344]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="3" pageNumber="30">Fig. 1A</figureCitation>
|
||
<figureCitation id="134A2A56496B7F5EFAE4FED95B257146" box="[1349,1358,331,348]" captionStart="Fig" captionStartId="3.[98,131,1852,1874]" captionTargetBox="[98,1478,431,1687]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig.3. Pygidia of catillicephalid trilobite Catillicephalacifellii sp.nov.from the Downes Point Member,Shallow BayFormation,Guzhangian,Miaolingian, → Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142520, paratype; dorsal (A1), lateral (A2), and posterior (A3) views. B. GSC 142521, paratype; posterior (B1), dorsal (B2), and lateral (B3) views. C. GSC 142522, paratype; posterior (C1) and dorsal (C2) views. D. GSC 142523, paratype; posterior (D1) and dorsal (D2) views. E. GSC 142524, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626760" httpUri="https://zenodo.org/record/10626760/files/figure.png" pageId="3" pageNumber="30">
|
||
<subScript id="17F53496496B7F5EFAE4FED95B257146" attach="right" box="[1349,1358,331,348]" fontSize="7" pageId="3" pageNumber="30">3</subScript>
|
||
</figureCitation>
|
||
). Posterior border furrow narrow, oblique, deep groove adaxially, shallowing abruptly abaxially and curving forward. Posterior weakly convex, expanding abaxially, with minimum length near axial furrow 28% (22–33%) of length at point where border furrow shallows. External surface other than furrows finely pitted, augmented with coarse granules on smaller specimens (e.g.,
|
||
<figureCitation id="134A2A56496D7F58FEBAFEB25F077120" box="[283,364,288,314]" captionStart="Fig" captionStartId="3.[98,131,1712,1734]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="5" pageNumber="32">Fig. 2C</figureCitation>
|
||
<figureCitation id="134A2A56496D7F58FECDFEBF5F1E7124" box="[364,373,301,318]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="5" pageNumber="32">
|
||
<subScript id="17F53496496D7F58FECDFEBF5F1E7124" attach="left" box="[364,373,301,318]" fontSize="7" pageId="5" pageNumber="32">1</subScript>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF0E665B496A7F5FFF36F93D5C697703" ID-DOI="http://doi.org/10.5281/zenodo.10626756" ID-Zenodo-Dep="10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="2" pageNumber="29" startId="2.[151,181,1711,1733]" targetBox="[154,1533,161,1686]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BCE36D3496A7F5FFF36F93D5C697703" blockId="2.[151,1535,1711,1817]" pageId="2" pageNumber="29">
|
||
Fig 1. Cranidia of catillicephalid trilobite
|
||
<taxonomicName id="4C714D50496A7F5FFD8FF93D5C9876DE" authority="Westrop & Dengler, 2022" authorityName="Westrop & Dengler" authorityYear="2022" box="[558,755,1711,1732]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="cifellii" status="sp. nov.">
|
||
<emphasis id="B905EAC1496A7F5FFD8FF93D5C9876DE" box="[558,755,1711,1732]" italics="true" pageId="2" pageNumber="29">Catillicephala cifellii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23657BA496A7F5FFD5FF9225D2276DF" box="[766,841,1712,1733]" pageId="2" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada.
|
||
<emphasis id="B905EAC1496A7F5FFBA4F9595A7E76FA" bold="true" box="[1029,1045,1739,1760]" pageId="2" pageNumber="29">A</emphasis>
|
||
. GSC 142512, holotype; dorsal (A
|
||
<subScript id="17F53496496A7F5FFAF3F9445B3176FE" attach="right" box="[1362,1370,1750,1764]" fontSize="6" pageId="2" pageNumber="29">1</subScript>
|
||
), lateral (A
|
||
<subScript id="17F53496496A7F5FFA62F9445BA076FE" attach="right" box="[1475,1483,1750,1764]" fontSize="6" pageId="2" pageNumber="29">2</subScript>
|
||
), and anterior (A
|
||
<subScript id="17F53496496A7F5FFF5AF9605F68771A" attach="left" box="[251,259,1778,1792]" fontSize="6" pageId="2" pageNumber="29">3</subScript>
|
||
) views.
|
||
<emphasis id="B905EAC1496A7F5FFEF1F97A5F3476E7" bold="true" box="[336,351,1768,1789]" pageId="2" pageNumber="29">B</emphasis>
|
||
. GSC 142513, paratype; dorsal view.
|
||
<emphasis id="B905EAC1496A7F5FFD16F9755CAC76E7" bold="true" box="[695,711,1767,1789]" pageId="2" pageNumber="29">C</emphasis>
|
||
. GSC 142514, paratype; dorsal (C
|
||
<subScript id="17F53496496A7F5FFBA1F9605A63771A" attach="left" box="[1024,1032,1778,1792]" fontSize="6" pageId="2" pageNumber="29">1</subScript>
|
||
), lateral (C
|
||
<subScript id="17F53496496A7F5FFBD1F9605A13771A" attach="left" box="[1136,1144,1778,1792]" fontSize="6" pageId="2" pageNumber="29">2</subScript>
|
||
), and anterior (C
|
||
<subScript id="17F53496496A7F5FFAB2F9605B70771A" attach="left" box="[1299,1307,1778,1792]" fontSize="6" pageId="2" pageNumber="29">3</subScript>
|
||
) views.
|
||
<emphasis id="B905EAC1496A7F5FFAC9F97A5B1376E7" bold="true" box="[1384,1400,1768,1789]" pageId="2" pageNumber="29">D</emphasis>
|
||
. GSC 142515, paratype; dorsal view. Scale bars 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0E665B496B7F5EFFC3F9225AEB7700" ID-DOI="http://doi.org/10.5281/zenodo.11093736" ID-Zenodo-Dep="11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="3" pageNumber="30" startId="3.[98,131,1712,1734]" targetBox="[98,1478,431,1687]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BCE36D3496B7F5EFFC3F9225AEB7700" blockId="3.[98,1482,1712,1820]" pageId="3" pageNumber="30">
|
||
Fig. 2. Cranidia of catillicephalid trilobite
|
||
<taxonomicName id="4C714D50496B7F5EFE5DF9225CAA76DF" authority="Westrop & Dengler, 2022" authorityName="Westrop & Dengler" authorityYear="2022" box="[508,705,1712,1733]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="cifellii" status="sp. nov.">
|
||
<emphasis id="B905EAC1496B7F5EFE5DF9225CAA76DF" box="[508,705,1712,1733]" italics="true" pageId="3" pageNumber="30">Catillicephala cifellii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23657BA496B7F5EFD6AF9235D7D76DC" box="[715,790,1713,1734]" pageId="3" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada.
|
||
<emphasis id="B905EAC1496B7F5EFA91F95E5B2B76FB" bold="true" box="[1328,1344,1740,1761]" pageId="3" pageNumber="30">A</emphasis>
|
||
. GSC 142516, paratype; dorsal (A
|
||
<subScript id="17F53496496B7F5EFEB0F9605F72771A" attach="right" box="[273,281,1778,1792]" fontSize="6" pageId="3" pageNumber="30">1</subScript>
|
||
), lateral (A
|
||
<subScript id="17F53496496B7F5EFE23F9605FE1771A" attach="left" box="[386,394,1778,1792]" fontSize="6" pageId="3" pageNumber="30">2</subScript>
|
||
), and anterior (A
|
||
<subScript id="17F53496496B7F5EFD89F9605C5B771A" attach="right" box="[552,560,1778,1792]" fontSize="6" pageId="3" pageNumber="30">3</subScript>
|
||
) views.
|
||
<emphasis id="B905EAC1496B7F5EFDDCF97B5CE776E4" bold="true" box="[637,652,1769,1790]" pageId="3" pageNumber="30">B</emphasis>
|
||
. GSC 142517, paratype; anterior (B
|
||
<subScript id="17F53496496B7F5EFC77F9605DB5771A" attach="left" box="[982,990,1778,1792]" fontSize="6" pageId="3" pageNumber="30">1</subScript>
|
||
), lateral (B
|
||
<subScript id="17F53496496B7F5EFBE7F9605A25771A" attach="left" box="[1094,1102,1778,1792]" fontSize="6" pageId="3" pageNumber="30">2</subScript>
|
||
), and dorsal (B
|
||
<subScript id="17F53496496B7F5EFB7DF9605A8F771A" attach="left" box="[1244,1252,1778,1792]" fontSize="6" pageId="3" pageNumber="30">3</subScript>
|
||
) views.
|
||
<emphasis id="B905EAC1496B7F5EFA90F97A5B2A76E4" bold="true" box="[1329,1345,1768,1790]" pageId="3" pageNumber="30">C</emphasis>
|
||
. GSC 142518, paratype; dorsal (C
|
||
<subScript id="17F53496496B7F5EFEAEF89C5F7C7706" attach="left" box="[271,279,1806,1820]" fontSize="6" pageId="3" pageNumber="30">1</subScript>
|
||
), anterior (C
|
||
<subScript id="17F53496496B7F5EFE2CF89C5FFE7706" attach="left" box="[397,405,1806,1820]" fontSize="6" pageId="3" pageNumber="30">2</subScript>
|
||
), and lateral (C
|
||
<subScript id="17F53496496B7F5EFD82F89C5C407706" attach="left" box="[547,555,1806,1820]" fontSize="6" pageId="3" pageNumber="30">3</subScript>
|
||
) views.
|
||
<emphasis id="B905EAC1496B7F5EFDD9F8975CE37700" bold="true" box="[632,648,1797,1818]" pageId="3" pageNumber="30">D</emphasis>
|
||
. GSC 142519, paratype; dorsal view. Scale bars 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0E665B496B7F5EFFC3F8AE5AD977BC" ID-DOI="http://doi.org/10.5281/zenodo.10626760" ID-Zenodo-Dep="10626760" httpUri="https://zenodo.org/record/10626760/files/figure.png" pageId="3" pageNumber="30" startId="3.[98,131,1852,1874]" targetBox="[98,1478,431,1687]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BCE36D3496B7F5EFFC3F8AE5BA17790" blockId="3.[98,1516,1846,1960]" pageId="3" pageNumber="30">
|
||
Fig.3. Pygidia of catillicephalid trilobite
|
||
<taxonomicName id="4C714D50496B7F5EFE6FF8AE5CE6774B" authority="Westrop & Dengler, 2022" authorityName="Westrop & Dengler" authorityYear="2022" box="[462,653,1852,1873]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="cifellii" status="sp.nov.">
|
||
<emphasis id="B905EAC1496B7F5EFE6FF8AE5CE6774B" box="[462,653,1852,1873]" italics="true" pageId="3" pageNumber="30">Catillicephala cifellii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23657BA496B7F5EFD30F8AF5CBD7748" box="[657,726,1853,1874]" pageId="3" pageNumber="30" rank="species">sp.nov.</taxonomicNameLabel>
|
||
from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian,
|
||
<emphasis id="B905EAC1496B7F5EFA71F8A45B877748" bold="true" box="[1488,1516,1846,1874]" pageId="3" pageNumber="30">→</emphasis>
|
||
Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada.
|
||
<emphasis id="B905EAC1496B7F5EFCF0F8CA5D0A7777" bold="true" box="[849,865,1880,1901]" pageId="3" pageNumber="30">A</emphasis>
|
||
. GSC 142520, paratype; dorsal (A
|
||
<subScript id="17F53496496B7F5EFB3AF8F05AC8776A" attach="right" box="[1179,1187,1890,1904]" fontSize="6" pageId="3" pageNumber="30">1</subScript>
|
||
), lateral (A
|
||
<subScript id="17F53496496B7F5EFAAAF8F05B78776A" attach="left" box="[1291,1299,1890,1904]" fontSize="6" pageId="3" pageNumber="30">2</subScript>
|
||
), and posterior (A
|
||
<subScript id="17F53496496B7F5EFA18F8F05BAA776A" attach="left" box="[1465,1473,1890,1904]" fontSize="6" pageId="3" pageNumber="30">3</subScript>
|
||
) views.
|
||
<emphasis id="B905EAC1496B7F5EFF05F8E65ED87793" bold="true" box="[164,179,1908,1929]" pageId="3" pageNumber="30">B</emphasis>
|
||
. GSC 142521, paratype; posterior (B
|
||
<subScript id="17F53496496B7F5EFDAEF8EC5C7C7796" attach="left" box="[527,535,1918,1932]" fontSize="6" pageId="3" pageNumber="30">1</subScript>
|
||
), dorsal (B
|
||
<subScript id="17F53496496B7F5EFD20F8EC5CE27796" attach="left" box="[641,649,1918,1932]" fontSize="6" pageId="3" pageNumber="30">2</subScript>
|
||
), and lateral (B
|
||
<subScript id="17F53496496B7F5EFCBCF8EC5D4E7796" attach="left" box="[797,805,1918,1932]" fontSize="6" pageId="3" pageNumber="30">3</subScript>
|
||
) views.
|
||
<emphasis id="B905EAC1496B7F5EFCD7F8E65DED7790" bold="true" box="[886,902,1908,1930]" pageId="3" pageNumber="30">C</emphasis>
|
||
. GSC 142522, paratype; posterior (C
|
||
<subScript id="17F53496496B7F5EFB42F8EC5A807796" attach="left" box="[1251,1259,1918,1932]" fontSize="6" pageId="3" pageNumber="30">1</subScript>
|
||
) and dorsal (C
|
||
<subScript id="17F53496496B7F5EFAD6F8EC5B147796" attach="left" box="[1399,1407,1918,1932]" fontSize="6" pageId="3" pageNumber="30">2</subScript>
|
||
) views.
|
||
</paragraph>
|
||
<paragraph id="8BCE36D3496B7F5EFFC3F8025AD977BC" blockId="3.[98,1516,1846,1960]" box="[98,1202,1936,1960]" pageId="3" pageNumber="30">
|
||
<emphasis id="B905EAC1496B7F5EFFC3F8025E1977BF" bold="true" box="[98,114,1936,1957]" pageId="3" pageNumber="30">D</emphasis>
|
||
. GSC 142523, paratype; posterior (D
|
||
<subScript id="17F53496496B7F5EFE67F8085FA577B2" attach="left" box="[454,462,1946,1960]" fontSize="6" pageId="3" pageNumber="30">1</subScript>
|
||
) and dorsal (D
|
||
<subScript id="17F53496496B7F5EFDF6F8085C3477B2" attach="left" box="[599,607,1946,1960]" fontSize="6" pageId="3" pageNumber="30">2</subScript>
|
||
) views.
|
||
<emphasis id="B905EAC1496B7F5EFD0DF8025CD077BF" bold="true" box="[684,699,1936,1957]" pageId="3" pageNumber="30">E</emphasis>
|
||
. GSC 142524, paratype; dorsal view. Scale bars 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCE36D3496D7F58FF26FED35F3873BE" blockId="5.[98,771,158,935]" pageId="5" pageNumber="32">
|
||
Pygidium strongly arched, semicircular in outline, length equal to 70% (65–78%) of maximum width, with anterior margin angled back, and conspicuous articulating facet at anterior corner. Axis inflated, rising well above pleural field; gently tapered and well-rounded posteriorly; long, overhanging posterior margin slightly on largest specimens (e.g.,
|
||
<figureCitation id="134A2A56496D7F58FF4CFD915F297204" box="[237,322,515,542]" captionStart="Fig" captionStartId="3.[98,131,1852,1874]" captionTargetBox="[98,1478,431,1687]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig.3. Pygidia of catillicephalid trilobite Catillicephalacifellii sp.nov.from the Downes Point Member,Shallow BayFormation,Guzhangian,Miaolingian, → Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142520, paratype; dorsal (A1), lateral (A2), and posterior (A3) views. B. GSC 142521, paratype; posterior (B1), dorsal (B2), and lateral (B3) views. C. GSC 142522, paratype; posterior (C1) and dorsal (C2) views. D. GSC 142523, paratype; posterior (D1) and dorsal (D2) views. E. GSC 142524, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626760" httpUri="https://zenodo.org/record/10626760/files/figure.png" pageId="5" pageNumber="32">Fig. 3A</figureCitation>
|
||
<figureCitation id="134A2A56496D7F58FEE3FD825F20723B" box="[322,331,528,545]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="5" pageNumber="32">
|
||
<subScript id="17F53496496D7F58FEE3FD825F20723B" attach="left" box="[322,331,528,545]" fontSize="7" pageId="5" pageNumber="32">1</subScript>
|
||
</figureCitation>
|
||
, B
|
||
<figureCitation id="134A2A56496D7F58FECCFD825F1D723B" box="[365,374,528,545]" captionStart="Fig" captionStartId="3.[98,131,1712,1734]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="5" pageNumber="32">
|
||
<subScript id="17F53496496D7F58FECCFD825F1D723B" attach="left" box="[365,374,528,545]" fontSize="7" pageId="5" pageNumber="32">2</subScript>
|
||
</figureCitation>
|
||
), with width at anteriormost axial ring equal to 37% (34–40%) of maximum pygidial width. Articulating half-ring conspicuous, roughly semielliptical in outline; articulating furrow broad, roughly equal in length to half-ring. Broad, transverse ring furrows generally faint. Up to four rings and a terminal piece comprising at least two segments evident (e.g.,
|
||
<figureCitation id="134A2A56496D7F58FDABFD545C0A72FA" box="[522,609,710,736]" captionStart="Fig" captionStartId="3.[98,131,1852,1874]" captionTargetBox="[98,1478,431,1687]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig.3. Pygidia of catillicephalid trilobite Catillicephalacifellii sp.nov.from the Downes Point Member,Shallow BayFormation,Guzhangian,Miaolingian, → Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142520, paratype; dorsal (A1), lateral (A2), and posterior (A3) views. B. GSC 142521, paratype; posterior (B1), dorsal (B2), and lateral (B3) views. C. GSC 142522, paratype; posterior (C1) and dorsal (C2) views. D. GSC 142523, paratype; posterior (D1) and dorsal (D2) views. E. GSC 142524, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626760" httpUri="https://zenodo.org/record/10626760/files/figure.png" pageId="5" pageNumber="32">Fig. 3A</figureCitation>
|
||
<figureCitation id="134A2A56496D7F58FDC1FD415C0272FE" box="[608,617,723,740]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="5" pageNumber="32">
|
||
<subScript id="17F53496496D7F58FDC1FD415C0272FE" attach="left" box="[608,617,723,740]" fontSize="7" pageId="5" pageNumber="32">1</subScript>
|
||
</figureCitation>
|
||
, B
|
||
<figureCitation id="134A2A56496D7F58FD2FFD415CFC72FE" box="[654,663,723,740]" captionStart="Fig" captionStartId="3.[98,131,1712,1734]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="5" pageNumber="32">
|
||
<subScript id="17F53496496D7F58FD2FFD415CFC72FE" attach="left" box="[654,663,723,740]" fontSize="7" pageId="5" pageNumber="32">2</subScript>
|
||
</figureCitation>
|
||
). Pleural field flexed downward; one well-defined pleural furrow expressed on external surface, remainder faint, although better defined on internal moulds. Border furrow faint but clearly defined; border downsloping. External surface save for furrows finely pitted; posterior edge of border with terrace ridges (e.g.,
|
||
<figureCitation id="134A2A56496D7F58FF4BFC1B5F5173B9" box="[234,314,905,931]" captionStart="Fig" captionStartId="3.[98,131,1852,1874]" captionTargetBox="[98,1478,431,1687]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig.3. Pygidia of catillicephalid trilobite Catillicephalacifellii sp.nov.from the Downes Point Member,Shallow BayFormation,Guzhangian,Miaolingian, → Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142520, paratype; dorsal (A1), lateral (A2), and posterior (A3) views. B. GSC 142521, paratype; posterior (B1), dorsal (B2), and lateral (B3) views. C. GSC 142522, paratype; posterior (C1) and dorsal (C2) views. D. GSC 142523, paratype; posterior (D1) and dorsal (D2) views. E. GSC 142524, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626760" httpUri="https://zenodo.org/record/10626760/files/figure.png" pageId="5" pageNumber="32">Fig. 3B</figureCitation>
|
||
<figureCitation id="134A2A56496D7F58FE9BFC045F2873BD" box="[314,323,918,935]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="5" pageNumber="32">
|
||
<subScript id="17F53496496D7F58FE9BFC045F2873BD" attach="left" box="[314,323,918,935]" fontSize="7" pageId="5" pageNumber="32">1</subScript>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCE36D3496D7F58FFC3FC275A307186" blockId="5.[98,771,948,1530]" lastBlockId="5.[808,1481,158,1125]" pageId="5" pageNumber="32">
|
||
<emphasis id="B905EAC1496D7F58FFC3FC275EA673D5" box="[98,205,949,975]" italics="true" pageId="5" pageNumber="32">Ontogeny</emphasis>
|
||
.—A series of differently sized cranidia show that the occipital spine develops during holaspid ontogeny. The smallest cranidium (
|
||
<figureCitation id="134A2A56496D7F58FEEBFC675FCB7415" box="[330,416,1013,1039]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="5" pageNumber="32">Fig. 1D</figureCitation>
|
||
) has a short, aspinous LO that occupies only 21% of glabellar length, although there is an ill-defined node present. LO becomes longer (occupying 24% of glabellar length) and more triangular in outline in a somewhat larger specimen (
|
||
<figureCitation id="134A2A56496D7F58FE10FBE55C60748F" box="[433,523,1143,1173]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="5" pageNumber="32">
|
||
Fig. 1C
|
||
<subScript id="17F53496496D7F58FDA3FB165C60748F" attach="left" box="[514,523,1156,1173]" fontSize="7" pageId="5" pageNumber="32">1</subScript>
|
||
</figureCitation>
|
||
) but, although a node may be present, it is not yet spinose. By the time a distinct spine appears, LO accounts for 32% of glabellar length
|
||
<figureCitation id="134A2A56496D7F58FFCBFB4B5EAB74E9" box="[106,192,1241,1267]" captionStart="Fig" captionStartId="3.[98,131,1712,1734]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="5" pageNumber="32">Fig. 2D</figureCitation>
|
||
), and the incompletely preserved spine of the larger
|
||
<typeStatus id="54CA8871496D7F58FFC3FB6B5EAF7509" box="[98,196,1273,1299]" pageId="5" pageNumber="32">holotype</typeStatus>
|
||
is also part of a very long LO (
|
||
<figureCitation id="134A2A56496D7F58FD84FB6B5CE9750D" box="[549,642,1273,1303]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="5" pageNumber="32">
|
||
Fig. 1A
|
||
<subScript id="17F53496496D7F58FDD8FA945CE9750D" attach="left" box="[633,642,1286,1303]" fontSize="7" pageId="5" pageNumber="32">1</subScript>
|
||
</figureCitation>
|
||
). The axial furrows become increasingly bowed outwards, producing a distinctly barrel-shaped glabellar outline (compare
|
||
<figureCitation id="134A2A56496D7F58FD01FAA85EF07562" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="5" pageNumber="32">
|
||
Fig. 1A
|
||
<subScript id="17F53496496D7F58FD53FAD55C907542" attach="left" box="[754,763,1351,1368]" fontSize="7" pageId="5" pageNumber="32">1</subScript>
|
||
, B, C
|
||
<subScript id="17F53496496D7F58FF33FAF55EF07562" attach="left" box="[146,155,1383,1400]" fontSize="7" pageId="5" pageNumber="32">1</subScript>
|
||
</figureCitation>
|
||
, D). Smaller individuals have better defined lateral glabellar furrows (
|
||
<figureCitation id="134A2A56496D7F58FEAFFAE95F0C7583" box="[270,359,1403,1433]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="5" pageNumber="32">
|
||
Fig. 1C
|
||
<subScript id="17F53496496D7F58FEFFFA1A5F0C7583" attach="left" box="[350,359,1416,1433]" fontSize="7" pageId="5" pageNumber="32">1</subScript>
|
||
</figureCitation>
|
||
, D). The pygidial axis is very long in all specimens. It terminates just short of the pygidial margin in smaller specimens (e.g.,
|
||
<figureCitation id="134A2A56496D7F58FE2EFA2E5FB475CC" box="[399,479,1468,1494]" captionStart="Fig" captionStartId="3.[98,131,1852,1874]" captionTargetBox="[98,1478,431,1687]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig.3. Pygidia of catillicephalid trilobite Catillicephalacifellii sp.nov.from the Downes Point Member,Shallow BayFormation,Guzhangian,Miaolingian, → Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142520, paratype; dorsal (A1), lateral (A2), and posterior (A3) views. B. GSC 142521, paratype; posterior (B1), dorsal (B2), and lateral (B3) views. C. GSC 142522, paratype; posterior (C1) and dorsal (C2) views. D. GSC 142523, paratype; posterior (D1) and dorsal (D2) views. E. GSC 142524, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626760" httpUri="https://zenodo.org/record/10626760/files/figure.png" pageId="5" pageNumber="32">Fig. 3C</figureCitation>
|
||
<figureCitation id="134A2A56496D7F58FE7EFA5B5F8375C0" box="[479,488,1481,1498]" captionStart="Fig" captionStartId="3.[98,131,1712,1734]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="5" pageNumber="32">
|
||
<subScript id="17F53496496D7F58FE7EFA5B5F8375C0" attach="left" box="[479,488,1481,1498]" fontSize="7" pageId="5" pageNumber="32">2</subScript>
|
||
</figureCitation>
|
||
) but extends to overhang the margin slightly in larger individuals (
|
||
<figureCitation id="134A2A56496D7F58FD8AFA4E5C1675ED" box="[555,637,1500,1527]" captionStart="Fig" captionStartId="3.[98,131,1852,1874]" captionTargetBox="[98,1478,431,1687]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig.3. Pygidia of catillicephalid trilobite Catillicephalacifellii sp.nov.from the Downes Point Member,Shallow BayFormation,Guzhangian,Miaolingian, → Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142520, paratype; dorsal (A1), lateral (A2), and posterior (A3) views. B. GSC 142521, paratype; posterior (B1), dorsal (B2), and lateral (B3) views. C. GSC 142522, paratype; posterior (C1) and dorsal (C2) views. D. GSC 142523, paratype; posterior (D1) and dorsal (D2) views. E. GSC 142524, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626760" httpUri="https://zenodo.org/record/10626760/files/figure.png" pageId="5" pageNumber="32">Fig. 3A</figureCitation>
|
||
<figureCitation id="134A2A56496D7F58FDDFFA7B5CEC75E0" box="[638,647,1513,1530]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="5" pageNumber="32">
|
||
<subScript id="17F53496496D7F58FDDFFA7B5CEC75E0" attach="right" box="[638,647,1513,1530]" fontSize="7" pageId="5" pageNumber="32">1</subScript>
|
||
</figureCitation>
|
||
, B
|
||
<figureCitation id="134A2A56496D7F58FD06FA7B5CDB75E0" box="[679,688,1513,1530]" captionStart="Fig" captionStartId="3.[98,131,1712,1734]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="5" pageNumber="32">
|
||
<subScript id="17F53496496D7F58FD06FA7B5CDB75E0" attach="left" box="[679,688,1513,1530]" fontSize="7" pageId="5" pageNumber="32">2</subScript>
|
||
</figureCitation>
|
||
).
|
||
<emphasis id="B905EAC1496D7F58FFC3F9955EAF763B" box="[98,196,1543,1569]" italics="true" pageId="5" pageNumber="32">Remarks</emphasis>
|
||
.—The conspicuous glabella that overhangs a very short anterior border and pygidium with a strongly convex, gently tapered axis ally
|
||
<taxonomicName id="4C714D50496D7F58FED6F9DA5C037678" authority="Westrop & Dengler, 2022" authorityName="Westrop & Dengler" authorityYear="2022" box="[375,616,1608,1634]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="cifellii" status="sp. nov.">
|
||
<emphasis id="B905EAC1496D7F58FED6F9DA5C037678" box="[375,616,1608,1634]" italics="true" pageId="5" pageNumber="32">Catillicephala cifellii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23657BA496D7F58FDD1F9DB5CA37679" box="[624,712,1609,1635]" pageId="5" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
with the other members of the genus that were revised by
|
||
<bibRefCitation id="EFE04B22496D7F58FD05F9F85F2076BE" author="Westrop, S. R. & Dengler, A. A." pageId="5" pageNumber="32" pagination="89 - 116" refId="ref4600" refString="Westrop, S. R. and Dengler, A. A. 2014 a. The mid-Cambrian (Guzhangian; Marjuman) trilobite genus Catillicephala Raymond 1938 (Catillicephalidae) from the Cow Head Group and correlatives in eastern Canada. Memoirs of the Association of Australasian Palaeontologists 45: 89 - 116." type="journal article" year="2014">Westrop and Dengler (2014a)</bibRefCitation>
|
||
. The spinous, triangular occipital ring of larger individuals (
|
||
<figureCitation id="134A2A56496D7F58FEF3F9385FD876DE" box="[338,435,1706,1732]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="5" pageNumber="32">Figs. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="134A2A56496D7F58FE1EF9385F8A76DE" box="[447,481,1706,1732]" captionStart="Fig" captionStartId="3.[98,131,1712,1734]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="5" pageNumber="32">2D</figureCitation>
|
||
) is unique.
|
||
<taxonomicName id="4C714D50496D7F58FDC0F9385EC176FE" authority="Westrop & Dengler, 2022" authorityName="Westrop & Dengler" authorityYear="2022" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="cifellii" status="sp. nov.">
|
||
<emphasis id="B905EAC1496D7F58FDC0F9385EC176FE" italics="true" pageId="5" pageNumber="32">Catillicephala cifellii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23657BA496D7F58FF16F9595F7E76FF" box="[183,277,1739,1765]" pageId="5" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is most like
|
||
<taxonomicName id="4C714D50496D7F58FE60F9595D6976FF" authority="(Rasetti, 1946)" baseAuthorityName="Rasetti" baseAuthorityYear="1946" box="[449,770,1738,1765]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="rotunda">
|
||
<emphasis id="B905EAC1496D7F58FE60F9595FB876FF" box="[449,467,1739,1765]" italics="true" pageId="5" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="B905EAC1496D7F58FE49F9585C2876FE" box="[488,579,1738,1764]" italics="true" pageId="5" pageNumber="32">rotunda</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE04B22496D7F58FDF8F9585C9176FF" author="Rasetti, F." box="[601,762,1738,1765]" pageId="5" pageNumber="32" pagination="442 - 446" refId="ref4335" refString="Rasetti, F. 1946. Early upper Cambrian trilobites from western Gaspe. Journal of Paleontology 20: 442 - 446." type="journal article" year="1946">Rasetti, 1946</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from the Grosses-Roches Formation, Métis-sur-Mer area,
|
||
<collectingRegion id="49B5F831496D7F58FFC3F8995ED7773F" box="[98,188,1803,1829]" country="Canada" name="Quebec" pageId="5" pageNumber="32">Quebec</collectingRegion>
|
||
. The latter species has a tapered occipital ring with a rounded to bluntly pointed terminus (e.g.,
|
||
<bibRefCitation id="EFE04B22496D7F58FDD1F8BE5F63777D" author="Westrop, S. R. & Dengler, A. A." pageId="5" pageNumber="32" pagination="89 - 116" refId="ref4600" refString="Westrop, S. R. and Dengler, A. A. 2014 a. The mid-Cambrian (Guzhangian; Marjuman) trilobite genus Catillicephala Raymond 1938 (Catillicephalidae) from the Cow Head Group and correlatives in eastern Canada. Memoirs of the Association of Australasian Palaeontologists 45: 89 - 116." type="journal article" year="2014">Westrop and Dengler 2014a</bibRefCitation>
|
||
: figs. 3D, 4B, G) that resembles the condition in small specimens of
|
||
<taxonomicName id="4C714D50496D7F58FEC0F8FF5FA2779D" authorityName="Westrop & Dengler" authorityYear="2022" box="[353,457,1901,1927]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="cifellii">
|
||
<emphasis id="B905EAC1496D7F58FEC0F8FF5F18779D" box="[353,371,1901,1927]" italics="true" pageId="5" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="B905EAC1496D7F58FE23F8FF5FA2779D" box="[386,457,1901,1927]" italics="true" pageId="5" pageNumber="32">cifellii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="134A2A56496D7F58FE79F8FF5C44779D" box="[472,559,1901,1927]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="5" pageNumber="32">Fig. 1D</figureCitation>
|
||
), albeit without an occipital node. Unlike
|
||
<taxonomicName id="4C714D50496D7F58FEC2F81C5FA077BD" authorityName="Westrop & Dengler" authorityYear="2022" box="[355,459,1933,1960]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="cifellii">
|
||
<emphasis id="B905EAC1496D7F58FEC2F81C5FA077BD" box="[355,459,1933,1960]" italics="true" pageId="5" pageNumber="32">C. cifellii</emphasis>
|
||
</taxonomicName>
|
||
, the occipital ring of
|
||
<taxonomicName id="4C714D50496D7F58FD1EF81C5D0270A2" baseAuthorityName="Rasetti" baseAuthorityYear="1946" box="[703,873,158,1960]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="rotunda">
|
||
<emphasis id="B905EAC1496D7F58FD1EF81C5CBA77B2" box="[703,721,1934,1960]" italics="true" pageId="5" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="B905EAC1496D7F58FD7EF81C5D0270A2" box="[735,873,158,1959]" italics="true" pageId="5" pageNumber="32">rotunda</emphasis>
|
||
</taxonomicName>
|
||
does not develop a spine, even in the largest individuals. In addition,
|
||
<taxonomicName id="4C714D50496D7F58FC7FFF2D5A3370C3" baseAuthorityName="Rasetti" baseAuthorityYear="1946" box="[990,1112,191,217]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="rotunda">
|
||
<emphasis id="B905EAC1496D7F58FC7FFF2D5D9B70C3" box="[990,1008,191,217]" italics="true" pageId="5" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="B905EAC1496D7F58FC5FFF2D5A3370C3" box="[1022,1112,191,217]" italics="true" pageId="5" pageNumber="32">rotunda</emphasis>
|
||
</taxonomicName>
|
||
has a more bulbous glabella that is well-rounded anteriorly, and has narrower (tr.) posterior area of the fixigena. In the largest pygidia of
|
||
<taxonomicName id="4C714D50496D7F58FABEFE925BF27100" baseAuthorityName="Rasetti" baseAuthorityYear="1946" box="[1311,1433,256,282]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="rotunda">
|
||
<emphasis id="B905EAC1496D7F58FABEFE925B5A7100" box="[1311,1329,256,282]" italics="true" pageId="5" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="B905EAC1496D7F58FA9EFE925BF27100" box="[1343,1433,256,282]" italics="true" pageId="5" pageNumber="32">rotunda</emphasis>
|
||
</taxonomicName>
|
||
, the axis terminates short of the posterior margin (e.g.,
|
||
<bibRefCitation id="EFE04B22496D7F58FACAFEB35A6B7141" author="Westrop, S. R. & Dengler, A. A." pageId="5" pageNumber="32" pagination="89 - 116" refId="ref4600" refString="Westrop, S. R. and Dengler, A. A. 2014 a. The mid-Cambrian (Guzhangian; Marjuman) trilobite genus Catillicephala Raymond 1938 (Catillicephalidae) from the Cow Head Group and correlatives in eastern Canada. Memoirs of the Association of Australasian Palaeontologists 45: 89 - 116." type="journal article" year="2014">Westrop and Dengler 2014a</bibRefCitation>
|
||
: fig. 5A–C), whereas similarly sized pygidia of
|
||
<taxonomicName id="4C714D50496D7F58FC24FEF35D867161" authorityName="Westrop & Dengler" authorityYear="2022" box="[901,1005,353,380]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="cifellii">
|
||
<emphasis id="B905EAC1496D7F58FC24FEF35DFC7161" box="[901,919,353,379]" italics="true" pageId="5" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="B905EAC1496D7F58FC07FEF35D867161" box="[934,1005,353,379]" italics="true" pageId="5" pageNumber="32">cifellii</emphasis>
|
||
</taxonomicName>
|
||
have longer axes that overhang the pygidial margin (e.g.,
|
||
<figureCitation id="134A2A56496D7F58FC41FE135A387186" box="[992,1107,385,412]" captionStart="Fig" captionStartId="3.[98,131,1852,1874]" captionTargetBox="[98,1478,431,1687]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig.3. Pygidia of catillicephalid trilobite Catillicephalacifellii sp.nov.from the Downes Point Member,Shallow BayFormation,Guzhangian,Miaolingian, → Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142520, paratype; dorsal (A1), lateral (A2), and posterior (A3) views. B. GSC 142521, paratype; posterior (B1), dorsal (B2), and lateral (B3) views. C. GSC 142522, paratype; posterior (C1) and dorsal (C2) views. D. GSC 142523, paratype; posterior (D1) and dorsal (D2) views. E. GSC 142524, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626760" httpUri="https://zenodo.org/record/10626760/files/figure.png" pageId="5" pageNumber="32">Fig. 3A, B</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BCE36D3496D7F58FCEFFE305B1772DA" blockId="5.[808,1481,158,1125]" pageId="5" pageNumber="32">
|
||
Compared to
|
||
<taxonomicName id="4C714D50496D7F58FC4BFE305A3871A6" authorityName="Westrop & Dengler" authorityYear="2022" box="[1002,1107,418,444]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="cifellii">
|
||
<emphasis id="B905EAC1496D7F58FC4BFE305D9771A6" box="[1002,1020,418,444]" italics="true" pageId="5" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="B905EAC1496D7F58FBADFE305A3871A6" box="[1036,1107,418,444]" italics="true" pageId="5" pageNumber="32">cifellii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C714D50496D7F58FBC3FE305BF071A7" authority="(Rasetti, 1946)" baseAuthorityName="Rasetti" baseAuthorityYear="1946" box="[1122,1435,418,445]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="impressa">
|
||
<emphasis id="B905EAC1496D7F58FBC3FE305A1F71A6" box="[1122,1140,418,444]" italics="true" pageId="5" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="B905EAC1496D7F58FB22FE315A8271A6" box="[1155,1257,419,444]" italics="true" pageId="5" pageNumber="32">impressa</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE04B22496D7F58FB58FE305BF871A7" author="Rasetti, F." box="[1273,1427,418,445]" pageId="5" pageNumber="32" pagination="442 - 446" refId="ref4335" refString="Rasetti, F. 1946. Early upper Cambrian trilobites from western Gaspe. Journal of Paleontology 20: 442 - 446." type="journal article" year="1946">Rasetti, 1946</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
has a far more rounded preoccipital region of the glabella that is subcircular in outline and usually has a medial backward expansion that constricts the occipital furrow (e.g.,
|
||
<bibRefCitation id="EFE04B22496D7F58FACAFD965A6B7224" author="Westrop, S. R. & Dengler, A. A." pageId="5" pageNumber="32" pagination="89 - 116" refId="ref4600" refString="Westrop, S. R. and Dengler, A. A. 2014 a. The mid-Cambrian (Guzhangian; Marjuman) trilobite genus Catillicephala Raymond 1938 (Catillicephalidae) from the Cow Head Group and correlatives in eastern Canada. Memoirs of the Association of Australasian Palaeontologists 45: 89 - 116." type="journal article" year="2014">Westrop and Dengler 2014a</bibRefCitation>
|
||
: figs. 7A, I, 8E, 9B). The anterior border furrow of
|
||
<taxonomicName id="4C714D50496D7F58FC07FDD75A5A7244" baseAuthorityName="Rasetti" baseAuthorityYear="1946" box="[934,1073,581,607]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="impressa">
|
||
<emphasis id="B905EAC1496D7F58FC07FDD75DD37245" box="[934,952,581,607]" italics="true" pageId="5" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="B905EAC1496D7F58FC6AFDD75A5A7244" box="[971,1073,581,606]" italics="true" pageId="5" pageNumber="32">impressa</emphasis>
|
||
</taxonomicName>
|
||
is effaced medially across almost the entire width of the glabella (e.g.,
|
||
<bibRefCitation id="EFE04B22496D7F58FB74FDF75D067285" author="Westrop, S. R. & Dengler, A. A." pageId="5" pageNumber="32" pagination="89 - 116" refId="ref4600" refString="Westrop, S. R. and Dengler, A. A. 2014 a. The mid-Cambrian (Guzhangian; Marjuman) trilobite genus Catillicephala Raymond 1938 (Catillicephalidae) from the Cow Head Group and correlatives in eastern Canada. Memoirs of the Association of Australasian Palaeontologists 45: 89 - 116." type="journal article" year="2014">Westrop and Dengler 2014a</bibRefCitation>
|
||
: figs. 7D, 8F, I, 9C, F, I), whereas
|
||
<taxonomicName id="4C714D50496D7F58FB46FD175B257285" authorityName="Westrop & Dengler" authorityYear="2022" box="[1255,1358,645,672]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="cifellii">
|
||
<emphasis id="B905EAC1496D7F58FB46FD175A927285" box="[1255,1273,645,671]" italics="true" pageId="5" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="B905EAC1496D7F58FAA6FD175B257285" box="[1287,1358,645,671]" italics="true" pageId="5" pageNumber="32">cifellii</emphasis>
|
||
</taxonomicName>
|
||
has a shallow but complete border furrow (
|
||
<figureCitation id="134A2A56496D7F58FB3FFD375A9472DA" box="[1182,1279,677,704]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="5" pageNumber="32">Figs. 1A</figureCitation>
|
||
<figureCitation id="134A2A56496D7F58FB5EFD205B5872D9" box="[1279,1331,677,707]" captionStart="Fig" captionStartId="3.[98,131,1852,1874]" captionTargetBox="[98,1478,431,1687]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig.3. Pygidia of catillicephalid trilobite Catillicephalacifellii sp.nov.from the Downes Point Member,Shallow BayFormation,Guzhangian,Miaolingian, → Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142520, paratype; dorsal (A1), lateral (A2), and posterior (A3) views. B. GSC 142521, paratype; posterior (B1), dorsal (B2), and lateral (B3) views. C. GSC 142522, paratype; posterior (C1) and dorsal (C2) views. D. GSC 142523, paratype; posterior (D1) and dorsal (D2) views. E. GSC 142524, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626760" httpUri="https://zenodo.org/record/10626760/files/figure.png" pageId="5" pageNumber="32">
|
||
<subScript id="17F53496496D7F58FB5EFD205B6272D9" attach="left" box="[1279,1289,690,707]" fontSize="7" pageId="5" pageNumber="32">3</subScript>
|
||
, C
|
||
<subScript id="17F53496496D7F58FA88FD205B5872D9" attach="left" box="[1321,1331,690,707]" fontSize="7" pageId="5" pageNumber="32">3</subScript>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="134A2A56496D7F58FAE0FD375B0872A5" box="[1345,1379,677,703]" captionStart="Fig" captionStartId="3.[98,131,1712,1734]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig. 2. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulders BPS 468 (A), BPS 467 (B, D), BPS 458 (C), Broom Point South, western Newfoundland, Canada. A. GSC 142516, paratype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142517, paratype; anterior (B1), lateral (B2), and dorsal (B3) views. C. GSC 142518, paratype; dorsal (C1), anterior (C2), and lateral (C3) views. D. GSC 142519, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11093736" httpUri="https://zenodo.org/record/11093736/files/figure.png" pageId="5" pageNumber="32">2A</figureCitation>
|
||
<figureCitation id="134A2A56496D7F58FAC2FD205B0772D9" box="[1379,1388,690,707]" captionStart="Fig" captionStartId="3.[98,131,1852,1874]" captionTargetBox="[98,1478,431,1687]" captionTargetId="figure-155@3.[98,1478,431,1687]" captionTargetPageId="3" captionText="Fig.3. Pygidia of catillicephalid trilobite Catillicephalacifellii sp.nov.from the Downes Point Member,Shallow BayFormation,Guzhangian,Miaolingian, → Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142520, paratype; dorsal (A1), lateral (A2), and posterior (A3) views. B. GSC 142521, paratype; posterior (B1), dorsal (B2), and lateral (B3) views. C. GSC 142522, paratype; posterior (C1) and dorsal (C2) views. D. GSC 142523, paratype; posterior (D1) and dorsal (D2) views. E. GSC 142524, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626760" httpUri="https://zenodo.org/record/10626760/files/figure.png" pageId="5" pageNumber="32">
|
||
<subScript id="17F53496496D7F58FAC2FD205B0772D9" attach="left" box="[1379,1388,690,707]" fontSize="7" pageId="5" pageNumber="32">3</subScript>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B6558496D7F58FCEFFD545A5B747F" pageId="5" pageNumber="32" type="discussion">
|
||
<paragraph id="8BCE36D3496D7F58FCEFFD545A5B747F" blockId="5.[808,1481,158,1125]" pageId="5" pageNumber="32">
|
||
<taxonomicName id="4C714D50496D7F58FCEFFD545BF972FA" authority="Westrop and Dengler, 2014 a" authorityName="Westrop and Dengler" authorityYear="2014" box="[846,1426,710,736]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="shawi">
|
||
<emphasis id="B905EAC1496D7F58FCEFFD545A5072FA" box="[846,1083,710,736]" italics="true" pageId="5" pageNumber="32">Catillicephala shawi</emphasis>
|
||
<bibRefCitation id="EFE04B22496D7F58FBE7FD545BF972FA" author="Westrop, S. R. & Dengler, A. A." box="[1094,1426,710,736]" pageId="5" pageNumber="32" pagination="89 - 116" refId="ref4600" refString="Westrop, S. R. and Dengler, A. A. 2014 a. The mid-Cambrian (Guzhangian; Marjuman) trilobite genus Catillicephala Raymond 1938 (Catillicephalidae) from the Cow Head Group and correlatives in eastern Canada. Memoirs of the Association of Australasian Palaeontologists 45: 89 - 116." type="journal article" year="2014">Westrop and Dengler, 2014a</bibRefCitation>
|
||
</taxonomicName>
|
||
, differs from
|
||
<taxonomicName id="4C714D50496D7F58FC3EFD755A61731A" authorityName="Westrop & Dengler" authorityYear="2022" box="[927,1034,742,769]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="cifellii">
|
||
<emphasis id="B905EAC1496D7F58FC3EFD755DDA731B" box="[927,945,743,769]" italics="true" pageId="5" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="B905EAC1496D7F58FC62FD745A61731A" box="[963,1034,742,768]" italics="true" pageId="5" pageNumber="32">cifellii</emphasis>
|
||
</taxonomicName>
|
||
in having a subquadrate glabella that expands gently forward, an occipital furrow that shallows medially, and, as in
|
||
<taxonomicName id="4C714D50496D7F58FBBAFCBA5ACD735B" baseAuthorityName="Rasetti" baseAuthorityYear="1946" box="[1051,1190,808,834]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="impressa">
|
||
<emphasis id="B905EAC1496D7F58FBBAFCBA5A467358" box="[1051,1069,808,834]" italics="true" pageId="5" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="B905EAC1496D7F58FBE1FCBA5ACD735B" box="[1088,1190,808,833]" italics="true" pageId="5" pageNumber="32">impressa</emphasis>
|
||
</taxonomicName>
|
||
, the anterior border furrow is effaced in front of the glabella (e.g.,
|
||
<bibRefCitation id="EFE04B22496D7F58FA97FCDA5DA47398" author="Westrop, S. R. & Dengler, A. A." pageId="5" pageNumber="32" pagination="89 - 116" refId="ref4600" refString="Westrop, S. R. and Dengler, A. A. 2014 a. The mid-Cambrian (Guzhangian; Marjuman) trilobite genus Catillicephala Raymond 1938 (Catillicephalidae) from the Cow Head Group and correlatives in eastern Canada. Memoirs of the Association of Australasian Palaeontologists 45: 89 - 116." type="journal article" year="2014">Westrop and Dengler 2014a</bibRefCitation>
|
||
: figs. 14, 15). Like
|
||
<taxonomicName id="4C714D50496D7F58FB02FCFA5B617398" authorityName="Westrop & Dengler" authorityYear="2022" box="[1187,1290,872,899]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="cifellii">
|
||
<emphasis id="B905EAC1496D7F58FB02FCFA5ADE7398" box="[1187,1205,872,898]" italics="true" pageId="5" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="B905EAC1496D7F58FB62FCFA5B617398" box="[1219,1290,872,898]" italics="true" pageId="5" pageNumber="32">cifellii</emphasis>
|
||
</taxonomicName>
|
||
, the pygidium of
|
||
<taxonomicName id="4C714D50496D7F58FC89FC1B5AA173B9" authority="Westrop and Dengler, 2014 a" authorityName="Westrop and Dengler" authorityYear="2014" box="[808,1226,904,931]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="calva">
|
||
<emphasis id="B905EAC1496D7F58FC89FC1B5D5173B9" box="[808,826,905,931]" italics="true" pageId="5" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="B905EAC1496D7F58FCE9FC1A5DEE73B8" box="[840,901,904,930]" italics="true" pageId="5" pageNumber="32">calva</emphasis>
|
||
<bibRefCitation id="EFE04B22496D7F58FC2AFC1B5AA173B9" author="Westrop, S. R. & Dengler, A. A." box="[907,1226,904,931]" pageId="5" pageNumber="32" pagination="89 - 116" refId="ref4600" refString="Westrop, S. R. and Dengler, A. A. 2014 a. The mid-Cambrian (Guzhangian; Marjuman) trilobite genus Catillicephala Raymond 1938 (Catillicephalidae) from the Cow Head Group and correlatives in eastern Canada. Memoirs of the Association of Australasian Palaeontologists 45: 89 - 116." type="journal article" year="2014">Westrop and Dengler, 2014a</bibRefCitation>
|
||
</taxonomicName>
|
||
, has a convex axis that overhangs the posterior margin. The cranidia of these two species are differentiated readily. In addition to lacking an occipital spine,
|
||
<taxonomicName id="4C714D50496D7F58FC7CFC785A57741E" authorityName="Westrop and Dengler" authorityYear="2014" box="[989,1084,1002,1028]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="calva">
|
||
<emphasis id="B905EAC1496D7F58FC7CFC785D84741E" box="[989,1007,1002,1028]" italics="true" pageId="5" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="B905EAC1496D7F58FC5EFC785A57741E" box="[1023,1084,1002,1028]" italics="true" pageId="5" pageNumber="32">calva</emphasis>
|
||
</taxonomicName>
|
||
has a relatively narrow, forwardly expanding glabella that contrasts with the barrel-shaped glabella of
|
||
<taxonomicName id="4C714D50496D7F58FC07FBB95A65745F" authorityName="Westrop & Dengler" authorityYear="2022" box="[934,1038,1067,1093]" class="Trilobita" family="Catillicephalidae" genus="Catillicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="cifellii">
|
||
<emphasis id="B905EAC1496D7F58FC07FBB95DD3745F" box="[934,952,1067,1093]" italics="true" pageId="5" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="B905EAC1496D7F58FC66FBB95A65745F" box="[967,1038,1067,1093]" italics="true" pageId="5" pageNumber="32">cifellii</emphasis>
|
||
</taxonomicName>
|
||
(compare
|
||
<figureCitation id="134A2A56496D7F58FB24FBB95A91745F" box="[1157,1274,1067,1093]" captionStart="Fig 1" captionStartId="2.[151,181,1711,1733]" captionTargetBox="[154,1533,161,1686]" captionTargetId="figure-81@2.[154,1533,161,1686]" captionTargetPageId="2" captionText="Fig 1. Cranidia of catillicephalid trilobite Catillicephala cifellii sp. nov. from the Downes Point Member, Shallow Bay Formation, Guzhangian, Miaolingian, Cambrian; boulder BPS 467, Broom Point South, western Newfoundland, Canada. A. GSC 142512, holotype; dorsal (A1), lateral (A2), and anterior (A3) views. B. GSC 142513, paratype; dorsal view. C. GSC 142514, paratype; dorsal (C1), lateral (C2), and anterior (C3) views. D. GSC 142515, paratype; dorsal view. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10626756" httpUri="https://zenodo.org/record/10626756/files/figure.png" pageId="5" pageNumber="32">Fig. 1A, B</figureCitation>
|
||
with
|
||
<bibRefCitation id="EFE04B22496D7F58FA9BFBB95DA4747F" author="Westrop, S. R. & Dengler, A. A." pageId="5" pageNumber="32" pagination="89 - 116" refId="ref4600" refString="Westrop, S. R. and Dengler, A. A. 2014 a. The mid-Cambrian (Guzhangian; Marjuman) trilobite genus Catillicephala Raymond 1938 (Catillicephalidae) from the Cow Head Group and correlatives in eastern Canada. Memoirs of the Association of Australasian Palaeontologists 45: 89 - 116." type="journal article" year="2014">Westrop and Dengler 2014a</bibRefCitation>
|
||
: fig. 19).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B6558496D7F58FC89FBE45DD574AB" pageId="5" pageNumber="32" type="biology_ecology">
|
||
<paragraph id="8BCE36D3496D7F58FC89FBE45DD574AB" blockId="5.[808,1481,1142,1201]" pageId="5" pageNumber="32">
|
||
<emphasis id="B905EAC1496D7F58FC89FBE45ABE748A" box="[808,1237,1142,1168]" italics="true" pageId="5" pageNumber="32">Stratigraphic and geographic range</emphasis>
|
||
.—
|
||
<typeStatus id="54CA8871496D7F58FB57FBE55B5B748B" box="[1270,1328,1143,1169]" pageId="5" pageNumber="32">Type</typeStatus>
|
||
locality and horizon only.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |