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<document id="17EE3247A2D6DA490AB12983A2D8E438" ID-CLB-Dataset="54709" ID-DOI="10.5852/ejt.2023.875.2147" ID-GBIF-Dataset="3351d966-cbeb-42a8-ac49-627b5fc261c7" ID-ISSN="2118-9773" ID-Zenodo-Dep="8083773" ID-ZooBank="32FA008C-B35D-483C-9DBE-1DCCD0868FAC" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1687788869828" checkinUser="felipe" docAuthor="Fasbender, Andrew" docDate="2023" docId="03D887BE8166363E2A67FE9F45EB0D43" docLanguage="en" docName="ejt-875_159-202.pdf" docOrigin="European Journal of Taxonomy 875" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ceratoculicoides moravicus Knoz 1987" docType="treatment" docVersion="5" lastPageNumber="179" masterDocId="FFE1FFC68175362A284DFF9741030970" masterDocTitle="Revision of the New World Ceratoculicoides Wirth &amp; Ratanaworabhan (Diptera, Ceratopogonidae, Ceratopogonini)" masterLastPageNumber="202" masterPageNumber="159" pageNumber="178" updateTime="1699530684662" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9EBAF72285A245D63916EE11C5E32C1B">Revision of the New World Ceratoculicoides Wirth &amp; Ratanaworabhan (Diptera, Ceratopogonidae, Ceratopogonini)</mods:title>
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<mods:namePart id="3FE29B9BE60041074FC57184863BB8AA">Fasbender, Andrew</mods:namePart>
<mods:affiliation id="69C110FB272A895691BE62C93C1C0EF3">Rhithron Associates Inc., 33 Fort Missoula Road, Missoula, MT 59847, USA.</mods:affiliation>
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<mods:title id="CBD66AB57990B1F6FFD8E841EC0402D3">European Journal of Taxonomy</mods:title>
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<mods:date id="5F27669AD0FF257380FB60AC8E98678C">2023</mods:date>
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<treatment id="03D887BE8166363E2A67FE9F45EB0D43" ID-DOI="http://doi.org/10.5281/zenodo.8083806" ID-GBIF-Taxon="210477700" ID-Zenodo-Dep="8083806" LSID="urn:lsid:plazi:treatment:03D887BE8166363E2A67FE9F45EB0D43" httpUri="http://treatment.plazi.org/id/03D887BE8166363E2A67FE9F45EB0D43" lastPageId="20" lastPageNumber="179" pageId="19" pageNumber="178">
<subSubSection id="C36B6523816636392A67FE9F45090853" box="[554,1034,264,291]" pageId="19" pageNumber="178" type="nomenclature">
<paragraph id="8BCE36A8816636392A67FE9F45090853" blockId="19.[554,1034,264,327]" box="[554,1034,264,291]" pageId="19" pageNumber="178">
<heading id="D08681C4816636392A67FE9F45090853" box="[554,1034,264,291]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="178" reason="2">
<taxonomicName id="4C714D2B816636392A67FE9F45090853" ID-CoL="SMSB" authority="Knoz, 1987" authorityName="Knoz" authorityYear="1987" box="[554,1034,264,291]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="178" phylum="Arthropoda" rank="species" species="moravicus">
<emphasis id="B905EABA816636392A67FE9F427A0853" bold="true" box="[554,889,264,291]" italics="true" pageId="19" pageNumber="178">Ceratoculicoidesmoravicus</emphasis>
<bibRefCitation id="EFE04B59816636392BCDFE9E45090853" author="Knoz J." box="[896,1034,265,291]" pageId="19" pageNumber="178" pagination="388 - 392" refId="ref23379" refString="Knoz J. 1987. Description of a new species of Ceratoculicoides (Diptera, Ceratopogonidae) from Czechoslovakia and a review of the genus. Acta Entomologica Bohemoslovaca 84: 388 - 392." type="journal article" year="1987">Knoz, 1987</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36B6523816636392AD8FEBA429D0837" box="[661,926,300,327]" pageId="19" pageNumber="178" type="description">
<paragraph id="8BCE36A8816636392AD8FEBA429D0837" blockId="19.[554,1034,264,327]" box="[661,926,300,327]" pageId="19" pageNumber="178">
<figureCitation id="134A2A2D816636392AD8FEBA43EC0837" box="[661,751,300,327]" captionStart="Fig" captionStartId="10.[189,232,1963,1989]" captionTargetBox="[190,1398,1040,1891]" captionTargetId="figure-160@10.[190,1398,1040,1891]" captionTargetPageId="10" captionText="Fig. 5. Male heads II. Ceratoculicoidesmoravicus group. a. C. sp. M1. b. C.moravicusKnoz, 1987. c. C.propinquus sp. nov. d. C.grogani sp. nov. e. C.borkenti sp. nov. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083789" httpUri="https://zenodo.org/record/8083789/files/figure.png" pageId="19" pageNumber="178">Figs 5b</figureCitation>
,
<figureCitation id="134A2A2D816636392AB6FEBA421B0837" box="[763,792,301,327]" captionStart="Fig" captionStartId="12.[189,230,1963,1989]" captionTargetBox="[196,1390,1017,1903]" captionTargetId="figure-171@12.[196,1390,1017,1903]" captionTargetPageId="12" captionText="Fig. 7. Female heads II.Ceratoculicoidesmoravicus group.a. C.aliciae Huerta &amp; Borkent, 2005. b. C. aliciae antenna. c. C. borkenti sp. nov. d. C.grogani sp. nov. e. C.moravicusKnoz, 1987. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083793" httpUri="https://zenodo.org/record/8083793/files/figure.png" pageId="19" pageNumber="178">7e</figureCitation>
,
<figureCitation id="134A2A2D816636392B68FEBA426F0837" box="[805,876,301,327]" captionStart="Fig" captionStartId="18.[189,231,1752,1778]" captionTargetBox="[218,1370,268,1690]" captionTargetId="figure-141@18.[218,1370,268,1690]" captionTargetPageId="18" captionText="Fig. 10. Male genitalia III. Ceratoculicoides sp. M1: a. Genitalia, aedeagus and parameres removed. b. Parameres. c. Aedeagus. C.aliciaeHuerta &amp; Borkent, 2005: d. Parameres (adapted from Huerta &amp; Borkent 2005). e. Aedeagus (adapted from Huerta &amp; Borkent 2005). C.moravicusKnoz, 1987: f. Genitalia, aedeagus and parameres removed. g. Parameres. h. Aedeagus. C. propinquus sp. nov.: i. Genitalia, aedeagus and parameres removed. j. Parameres. k. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083804" httpUri="https://zenodo.org/record/8083804/files/figure.png" pageId="19" pageNumber="178">10f–h</figureCitation>
,
<figureCitation id="134A2A2D816636392B35FEBA429D0837" box="[888,926,301,327]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[238,1398,265,1781]" captionTargetId="figure-88@24.[238,1398,265,1781]" captionTargetPageId="24" captionText="Fig. 11. Female sternite 9 I. a. Ceratoculicoides aliciae Huerta &amp; Borkent, 2005, paratype, Chamela Biological Station, Jalisco, Mexico. b. C. borkenti sp. nov., paratype, Nogales, Arizona, USA. c. C. confusus Kouchibouguac National Park, New Brunswick, Canada. d. C.confusus, paratype, Great Smoky Mountains National Park, Tennessee, USA. e. C.grogani sp. nov., paratype, Corvallis, Oregon, USA. f. C.moravicus Bornova, Izmir, Turkey. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083808" httpUri="https://zenodo.org/record/8083808/files/figure.png" pageId="19" pageNumber="178">11f</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36B65238166363928F0FEF942980B0D" pageId="19" pageNumber="178" type="reference_group">
<paragraph id="8BCE36A88166363928F0FEF9452808F8" blockId="19.[189,1067,366,393]" box="[189,1067,366,393]" pageId="19" pageNumber="178">
<emphasis id="B905EABA8166363928F0FEF9430408F8" box="[189,519,366,392]" italics="true" pageId="19" pageNumber="178">Ceratoculicoidesmoravicus</emphasis>
<bibRefCitation id="EFE04B59816636392A43FEF843D008F9" author="Knoz J." box="[526,723,367,393]" pageId="19" pageNumber="178" pagination="388 - 392" refId="ref23379" refString="Knoz J. 1987. Description of a new species of Ceratoculicoides (Diptera, Ceratopogonidae) from Czechoslovakia and a review of the genus. Acta Entomologica Bohemoslovaca 84: 388 - 392." type="journal article" year="1987">Knoz, 1987: 390</bibRefCitation>
, figs 1–19 (
<collectingCountry id="F3667638816636392B10FEF9451D08F8" box="[861,1054,366,392]" name="Czech Republic" pageId="19" pageNumber="178">Czech Republic</collectingCountry>
).
</paragraph>
<paragraph id="8BCE36A88166363928F0FE264319089E" blockId="19.[189,1398,432,637]" pageId="19" pageNumber="178">
<taxonomicName id="4C714D2B8166363928F0FE26422E08BB" ID-CoL="8Z9RV" authority="Remm, 1967: 27" authorityName="Remm" authorityPageNumber="27" authorityYear="1967" box="[189,813,432,459]" class="Insecta" family="Ceratopogonidae" genus="Ceratopogon" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="178" phylum="Arthropoda" rank="species" species="gracilipes" subGenus="Nilohelea">
<emphasis id="B905EABA8166363928F0FE26405B08BB" box="[189,344,433,459]" italics="true" pageId="19" pageNumber="178">Ceratopogon</emphasis>
(
<emphasis id="B905EABA816636392924FE2740D808BA" box="[361,475,432,458]" italics="true" pageId="19" pageNumber="178">Nilohelea</emphasis>
)
<emphasis id="B905EABA8166363929A1FE27436208BA" box="[492,609,432,458]" italics="true" pageId="19" pageNumber="178">gracilipes</emphasis>
<bibRefCitation id="EFE04B59816636392A24FE26422E08BB" author="Remm H." box="[617,813,432,459]" pageId="19" pageNumber="178" pagination="3 - 37" refId="ref23551" refString="Remm H. 1967. On the fauna of Ceratopogonidae (Diptera) of the Caucasus [in Russian]. Tartu Riikliku Ulikooli Toimetised 194: 3 - 37." type="journal article" year="1967">Remm, 1967: 27</bibRefCitation>
</taxonomicName>
(preoccupied name, subjective junior synonym), pl. 18, figs 1–7 (
<collectingCountry id="F36676388166363929E4FE43430D089E" box="[425,526,468,494]" name="Georgia" pageId="19" pageNumber="178">Georgia</collectingCountry>
).
</paragraph>
<paragraph id="8BCE36A88166363928F0FE6F45B30B46" blockId="19.[189,1398,432,637]" pageId="19" pageNumber="178">
<emphasis id="B905EABA8166363928F0FE6F40F30B62" box="[189,496,504,530]" italics="true" pageId="19" pageNumber="178">Ceratoculicoideshavelkai</emphasis>
<bibRefCitation id="EFE04B598166363929BAFE6F42020B62" author="Wirth W. W. &amp; Grogan W. L." box="[503,769,504,530]" pageId="19" pageNumber="178" pagination="1 - 160" refId="ref23900" refString="Wirth W. W. &amp; Grogan W. L. 1988. The predaceous midges of the world (Diptera: Ceratopogonidae; tribe Ceratopogonini). Flora and Fauna Handbook 4: 1 - 160." type="journal article" year="1988">Wirth &amp; Grogan, 1988</bibRefCitation>
(replacement name for
<emphasis id="B905EABA816636392C54FE6F44330B62" box="[1049,1328,504,530]" italics="true" pageId="19" pageNumber="178">Ceratopogongracilipes</emphasis>
Remm, subjective junior synonym). —
<bibRefCitation id="EFE04B59816636392AF1FD8C45330B46" author="Delecolle J. &amp; Schiegg K." box="[700,1072,539,566]" pageId="19" pageNumber="178" pagination="273 - 286" refId="ref22869" refString="Delecolle J. &amp; Schiegg K. 1998. Contribution a l'etude des ceratopogonides de Suisse. Revision et redescription des especes palearctiques du genre Ceratoculicoides Wirth &amp; Ratanaworabhan, 1971 (Diptera, Nematocera). Bulletin de la Societe entomologique de France 103: 273 - 286." type="journal article" year="1998">Delécolle &amp; Schiegg 1998: 274</bibRefCitation>
, figs 1–23.
</paragraph>
<paragraph id="8BCE36A88166363928F0FDA842980B0D" blockId="19.[189,1398,432,637]" pageId="19" pageNumber="178">
<taxonomicName id="4C714D2B8166363928F0FDA843CF0B29" ID-CoL="SMSC" authority="Gosseries, 1988: 2" authorityName="Gosseries" authorityPageNumber="2" authorityYear="1988" box="[189,716,575,601]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="178" phylum="Arthropoda" rank="species" species="remmi">
<emphasis id="B905EABA8166363928F0FDA840DF0B29" box="[189,476,575,601]" italics="true" pageId="19" pageNumber="178">Ceratoculicoides remmi</emphasis>
Gosseries, 1988: 2
</taxonomicName>
(replacement name for
<taxonomicName id="4C714D2B816636392BB5FDD7446C0B2A" ID-CoL="8Z9RV" authority="Remm" authorityName="Remm" authorityYear="1967" box="[1016,1391,575,602]" class="Insecta" family="Ceratopogonidae" genus="Ceratopogon" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="178" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B905EABA816636392BB5FDD744170B29" box="[1016,1300,575,602]" italics="true" pageId="19" pageNumber="178">Ceratopogon gracilipes</emphasis>
Remm
</taxonomicName>
, objective junior synonym of
<taxonomicName id="4C714D2B816636392A70FDF442890B0D" ID-CoL="SMS9" authority="Wirth &amp; Grogan" authorityName="Wirth &amp; Grogan" authorityYear="1988" box="[573,906,611,637]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="178" phylum="Arthropoda" rank="species" species="havelkai">
<emphasis id="B905EABA816636392A70FDF443C10B0D" box="[573,706,611,637]" italics="true" pageId="19" pageNumber="178">C.havelkai</emphasis>
Wirth &amp; Grogan
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B65238166363928F0FD3340790D3E" pageId="19" pageNumber="178" type="diagnosis">
<paragraph id="8BCE36A88166363928F0FD3340350BCE" blockId="19.[189,310,676,702]" box="[189,310,676,702]" pageId="19" pageNumber="178">
<heading id="D08681C48166363928F0FD3340350BCE" bold="true" box="[189,310,676,702]" fontSize="11" level="3" pageId="19" pageNumber="178" reason="3">
<emphasis id="B905EABA8166363928F0FD3340350BCE" bold="true" box="[189,310,676,702]" pageId="19" pageNumber="178">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36A88166363928A7FD4340290B9E" blockId="19.[189,1399,724,929]" box="[234,298,724,750]" pageId="19" pageNumber="178">
<emphasis id="B905EABA8166363928A7FD4340290B9E" bold="true" box="[234,298,724,750]" pageId="19" pageNumber="178">Male</emphasis>
</paragraph>
<paragraph id="8BCE36A88166363928F0FD6042030AD0" blockId="19.[189,1399,724,929]" pageId="19" pageNumber="178">
<taxonomicName id="4C714D2B8166363928F0FD6043070A61" authorityName="Knoz" authorityYear="1987" box="[189,516,759,785]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="178" phylum="Arthropoda" rank="species" species="moravicus">
<emphasis id="B905EABA8166363928F0FD6043070A61" box="[189,516,759,785]" italics="true" pageId="19" pageNumber="178">Ceratoculicoides moravicus</emphasis>
</taxonomicName>
can be separated from congeners by the following combination of characters: femora and tibiae brown; wing length
<quantity id="4C899B4D816636392AC7FC8C43EA0A46" box="[650,745,795,822]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="19" pageNumber="178" unit="mm" value="1.1">1.1 mm</quantity>
, apices of parameres acute, tapering distally; aedeagus lateral margins straight basally, strongly tapered at midpoint, apical and posterolateral points adjacent, 1–2 accessory spines between apical point and posterolateral point of aedeagus, posterolateral point subtriangular, apex directed anteriorly (
<figureCitation id="134A2A2D816636392AC0FC1143F10AD0" box="[653,754,902,928]" captionStart="Fig" captionStartId="18.[189,231,1752,1778]" captionTargetBox="[218,1370,268,1690]" captionTargetId="figure-141@18.[218,1370,268,1690]" captionTargetPageId="18" captionText="Fig. 10. Male genitalia III. Ceratoculicoides sp. M1: a. Genitalia, aedeagus and parameres removed. b. Parameres. c. Aedeagus. C.aliciaeHuerta &amp; Borkent, 2005: d. Parameres (adapted from Huerta &amp; Borkent 2005). e. Aedeagus (adapted from Huerta &amp; Borkent 2005). C.moravicusKnoz, 1987: f. Genitalia, aedeagus and parameres removed. g. Parameres. h. Aedeagus. C. propinquus sp. nov.: i. Genitalia, aedeagus and parameres removed. j. Parameres. k. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083804" httpUri="https://zenodo.org/record/8083804/files/figure.png" pageId="19" pageNumber="178">Fig. 10h</figureCitation>
).
</paragraph>
<paragraph id="8BCE36A88166363928A7FC5F40460A92" blockId="19.[189,1398,968,1102]" box="[234,325,968,994]" pageId="19" pageNumber="178">
<emphasis id="B905EABA8166363928A7FC5F40460A92" bold="true" box="[234,325,968,994]" pageId="19" pageNumber="178">Female</emphasis>
</paragraph>
<paragraph id="8BCE36A88166363928F0FC7B40790D3E" blockId="19.[189,1398,968,1102]" pageId="19" pageNumber="178">
Only species of
<taxonomicName id="4C714D2B816636392933FC7B43450D76" authorityName=", Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[382,582,1004,1030]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B905EABA816636392933FC7B43450D76" box="[382,582,1004,1030]" italics="true" pageId="19" pageNumber="178">Ceratoculicoides</emphasis>
</taxonomicName>
with the following combination of characters: FR 1.62–1.83; femora and tibiae brown; wing length ~
<quantity id="4C899B4D816636392A7AFB8743970D5A" box="[567,660,1040,1066]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="178" unit="mm" value="1.2">1.2 mm</quantity>
; single major spermatheca 75–77; medial margin of 9
<superScript id="7C049BE0816636392D41FB98441A0D6D" attach="left" box="[1292,1305,1039,1053]" fontSize="6" pageId="19" pageNumber="178">th</superScript>
sternite deeply concave.
</paragraph>
</subSubSection>
<subSubSection id="C36B65238166363928F0FBE242B10D92" pageId="19" pageNumber="178" type="materials_examined">
<paragraph id="8BCE36A88166363928F0FBE240A90DFF" blockId="19.[189,426,1141,1167]" box="[189,426,1141,1167]" pageId="19" pageNumber="178">
<heading id="D08681C48166363928F0FBE240A90DFF" bold="true" box="[189,426,1141,1167]" fontSize="11" level="3" pageId="19" pageNumber="178" reason="3">
<emphasis id="B905EABA8166363928F0FBE240A90DFF" bold="true" box="[189,426,1141,1167]" pageId="19" pageNumber="178">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36A88166363928F0FB3242B10D92" blockId="19.[189,1398,1188,1250]" pageId="19" pageNumber="178">
<materialsCitation id="3B193CF58166363928F0FB3240000D92" ID-GBIF-Occurrence="4135858347" collectingDate="1962-05" collectionCode="CNCI" collectorName="T. Curtin" country="Turkey" location="Bornova" pageId="19" pageNumber="178" specimenCount="1" specimenCount-female="1" stateProvince="Izmir">
<collectingCountry id="F36676388166363928F0FB3240300DCF" box="[189,307,1189,1215]" name="Turkey" pageId="19" pageNumber="178">TURKEY</collectingCountry>
•
<specimenCount id="9D77FD21816636392900FB3340790DCF" box="[333,378,1188,1215]" pageId="19" pageNumber="178" type="female">1 ♀</specimenCount>
;
<collectingRegion id="49B5F84A8166363929C4FB3340C90DCE" box="[393,458,1188,1214]" country="Turkey" name="Izmir" pageId="19" pageNumber="178">Izmir</collectingRegion>
,
<location id="8EAE607381663639299AFB3243430DCF" LSID="urn:lsid:plazi:treatment:03D887BE8166363E2A67FE9F45EB0D43:8EAE607381663639299AFB3243430DCF" box="[471,576,1189,1215]" country="Turkey" name="Bornova" pageId="19" pageNumber="178" stateProvince="Izmir">Bornova</location>
;
<date id="FFCF1068816636392A01FB3243C40DCE" box="[588,711,1188,1215]" pageId="19" pageNumber="178" value="1962-05">
<collectingDate id="EF8BE980816636392A01FB3243C40DCE" box="[588,711,1188,1215]" pageId="19" pageNumber="178" value="1962-05">May 1962</collectingDate>
</date>
;
<collectorName id="2684537E816636392A99FB32423E0DCE" box="[724,829,1188,1215]" pageId="19" pageNumber="178">T. Curtin</collectorName>
leg.; light, originally identified as
<taxonomicName id="4C714D2B816636392C9BFB32446D0DCE" authorityName="Remm" authorityYear="1967" box="[1238,1390,1188,1215]" class="Insecta" family="Ceratopogonidae" genus="Ceratopogon" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="178" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B905EABA816636392C9BFB32446D0DCE" box="[1238,1390,1188,1215]" italics="true" pageId="19" pageNumber="178">C.gracilipes</emphasis>
</taxonomicName>
;
<collectionCode id="ED60AE6D8166363928F0FB5F40000D92" box="[189,259,1224,1250]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="19" pageNumber="178">CNCI</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B193CF5816636392951FB5F42B10D92" ID-GBIF-Occurrence="4135858326" box="[284,946,1224,1250]" collectionCode="USNM" pageId="19" pageNumber="178" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D77FD21816636392951FB5F40490D92" box="[284,330,1224,1250]" pageId="19" pageNumber="178" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D77FD2181663639291AFB5F40870D92" box="[343,388,1224,1250]" pageId="19" pageNumber="178" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED60AE6D816636392B18FB5F42AD0D92" box="[853,942,1224,1250]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="178" type="Museum">USNM</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36B65238166363E28F0FA9E421308DC" lastPageId="20" lastPageNumber="179" pageId="19" pageNumber="178" type="description">
<paragraph id="8BCE36A88166363928F0FA9E404D0C53" blockId="19.[189,334,1289,1315]" box="[189,334,1289,1315]" pageId="19" pageNumber="178">
<heading id="D08681C48166363928F0FA9E404D0C53" bold="true" box="[189,334,1289,1315]" fontSize="11" level="3" pageId="19" pageNumber="178" reason="3">
<emphasis id="B905EABA8166363928F0FA9E404D0C53" bold="true" box="[189,334,1289,1315]" pageId="19" pageNumber="178">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36A88166363928A7FAAE40290C23" blockId="19.[189,1399,1337,1470]" box="[234,298,1337,1363]" pageId="19" pageNumber="178">
<emphasis id="B905EABA8166363928A7FAAE40290C23" bold="true" box="[234,298,1337,1363]" pageId="19" pageNumber="178">Male</emphasis>
</paragraph>
<paragraph id="8BCE36A88166363928F0FACA40AB0CCE" blockId="19.[189,1399,1337,1470]" pageId="19" pageNumber="178">
<smallCapsWord id="8D28A0748166363928F0FACA406F0C05" baselines="1392,1392" box="[189,364,1373,1399]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="19" pageNumber="178">MEASUREMENTS</smallCapsWord>
(n = 1). Head width 289; flagellomeres 43, 27, 30, 29, 24, 25, 24, 32, 30, 36, 95, 76, 85; AR 0.85; FR 1.92; wing length
<quantity id="4C899B4D816636392A62FA1743980CEB" box="[559,667,1408,1435]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="19" pageNumber="178" unit="mm" value="1.16">1.16 mm</quantity>
; wing width
<quantity id="4C899B4D816636392B7FFA17429D0CEB" box="[818,926,1408,1435]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="19" pageNumber="178" unit="mm" value="0.37">0.37 mm</quantity>
; costal ratio 0.52; GCR 1.65; GSR 1.15; aedeagus ratio 1.33.
</paragraph>
<paragraph id="8BCE36A88166363928F0FA7142410F53" blockId="19.[189,1399,1510,1572]" pageId="19" pageNumber="178">
<smallCapsWord id="8D28A0748166363928F0FA71401B0C8E" baselines="1529,1529" box="[189,280,1510,1536]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="19" pageNumber="178">THORAX</smallCapsWord>
. Dorsocentral punctations possibly absent, at most a few spots of thinned cuticle among dorsocentral setae. Legs with femora and tibiae brown.
</paragraph>
<paragraph id="8BCE36A88166363928F0F9DC40480E68" blockId="19.[189,1399,1611,1816]" pageId="19" pageNumber="178">
<smallCapsWord id="8D28A0748166363928F0F9DC402D0F14" baselines="1630,1631" box="[189,302,1611,1637]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Genitalia" pageId="19" pageNumber="178">GENITALIA</smallCapsWord>
(
<figureCitation id="134A2A2D81663639290FF9DC40C20F15" box="[322,449,1611,1637]" captionStart="Fig" captionStartId="18.[189,231,1752,1778]" captionTargetBox="[218,1370,268,1690]" captionTargetId="figure-141@18.[218,1370,268,1690]" captionTargetPageId="18" captionText="Fig. 10. Male genitalia III. Ceratoculicoides sp. M1: a. Genitalia, aedeagus and parameres removed. b. Parameres. c. Aedeagus. C.aliciaeHuerta &amp; Borkent, 2005: d. Parameres (adapted from Huerta &amp; Borkent 2005). e. Aedeagus (adapted from Huerta &amp; Borkent 2005). C.moravicusKnoz, 1987: f. Genitalia, aedeagus and parameres removed. g. Parameres. h. Aedeagus. C. propinquus sp. nov.: i. Genitalia, aedeagus and parameres removed. j. Parameres. k. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083804" httpUri="https://zenodo.org/record/8083804/files/figure.png" pageId="19" pageNumber="178">Fig. 10f–h</figureCitation>
). Distal portion of parameres tapering gradually to acute apex. Aedeagus lateral margins tapering, constricted noticeably at midlength; posterior margin a smooth concave arc, hyaline medial incision broad posteriorly, tapering anteriorly; base of posterolateral point directed laterally, apex rounded, directed anterolaterally; apical point subacute, directed posterolaterally, adjacent to posterolateral point, with 1–2 accessory spines between apical and posterolateral points, similar in size and shape to apical point.
</paragraph>
<paragraph id="8BCE36A88166363928A7F8D740460E2A" blockId="19.[189,1399,1856,2025]" box="[234,325,1856,1882]" pageId="19" pageNumber="178">
<emphasis id="B905EABA8166363928A7F8D740460E2A" bold="true" box="[234,325,1856,1882]" pageId="19" pageNumber="178">Female</emphasis>
</paragraph>
<paragraph id="8BCE36A88166363928F0F8F3456E0E98" blockId="19.[189,1399,1856,2025]" pageId="19" pageNumber="178">
<smallCapsWord id="8D28A0748166363928F0F8F3406D0E0C" baselines="1911,1911" box="[189,366,1892,1918]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="19" pageNumber="178">MEASUREMENTS</smallCapsWord>
(n = 2). Head width 272–314; flagellomeres 33–41, 23–27, 21–25, 26–32, 26–32, 28– 30, 31, 33–34, 46–50, 48–51, 54–55, 51–53, 62–63; AR 1.11–1.14; FR 1.62–1.83; wing length
<quantity id="4C899B4D816636392D79F81040280EB5" metricMagnitude="-3" metricUnit="m" metricValue="1.265" metricValueMax="1.28" metricValueMin="1.25" pageId="19" pageNumber="178" unit="mm" value="1.265" valueMax="1.28" valueMin="1.25">1.25– 1.28 mm</quantity>
; wing width
<quantity id="4C899B4D816636392985F83D436A0EB5" box="[456,617,1962,1989]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="19" pageNumber="178" unit="mm" value="0.49" valueMax="0.5" valueMin="0.48">0.48–0.5 mm</quantity>
; costal ratio 0.52–0.54; spermathecal length 75–77; spermathecal width 66–70; spermathecal neck (n = 1) 11; spermatheca/neck ratio (n = 1) 0.14.
</paragraph>
<paragraph id="8BCE36A88161363E28F0FE9C43C80855" blockId="20.[189,715,267,293]" box="[189,715,267,293]" pageId="20" pageNumber="179">
<smallCapsWord id="8D28A0748161363E28F0FE9C401A0853" baselines="286,286" box="[189,281,267,293]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="20" pageNumber="179">THORAX</smallCapsWord>
. Legs with femora and tibiae brown.
</paragraph>
<paragraph id="8BCE36A88161363E28F0FEDB421308DC" blockId="20.[187,1398,331,429]" pageId="20" pageNumber="179">
<smallCapsWord id="8D28A0748161363E28F0FEDB40320815" baselines="351,352" box="[189,305,332,358]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Genitalia" pageId="20" pageNumber="179">GENITALIA</smallCapsWord>
(
<figureCitation id="134A2A2D8161363E290FFEDB40A30816" box="[322,416,332,358]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[238,1398,265,1781]" captionTargetId="figure-88@24.[238,1398,265,1781]" captionTargetPageId="24" captionText="Fig. 11. Female sternite 9 I. a. Ceratoculicoides aliciae Huerta &amp; Borkent, 2005, paratype, Chamela Biological Station, Jalisco, Mexico. b. C. borkenti sp. nov., paratype, Nogales, Arizona, USA. c. C. confusus Kouchibouguac National Park, New Brunswick, Canada. d. C.confusus, paratype, Great Smoky Mountains National Park, Tennessee, USA. e. C.grogani sp. nov., paratype, Corvallis, Oregon, USA. f. C.moravicus Bornova, Izmir, Turkey. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083808" httpUri="https://zenodo.org/record/8083808/files/figure.png" pageId="20" pageNumber="179">Fig. 11f</figureCitation>
). 9
<superScript id="7C049BE08161363E298FFEDC40CC0829" attach="left" box="[450,463,331,345]" fontSize="6" pageId="20" pageNumber="179">th</superScript>
sternite anterior branch broad, evenly rounded, apices nearly touching medially; posterior branch directed straight posteromedially without curve, spiniform, weakly tapering, tip pointed or with minute hook at apex. 1 major spermatheca.
</paragraph>
</subSubSection>
<subSubSection id="C36B65238161363E28F0FE4340AF0B6C" box="[189,428,468,541]" pageId="20" pageNumber="179" type="distribution">
<paragraph id="8BCE36A88161363E28F0FE434055089E" blockId="20.[189,342,468,494]" box="[189,342,468,494]" pageId="20" pageNumber="179">
<heading id="D08681C48161363E28F0FE434055089E" bold="true" box="[189,342,468,494]" fontSize="11" level="3" pageId="20" pageNumber="179" reason="3">
<emphasis id="B905EABA8161363E28F0FE434055089E" bold="true" box="[189,342,468,494]" pageId="20" pageNumber="179">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36A88161363E28F0FD9440AF0B6C" blockId="20.[189,428,514,541]" box="[189,428,514,541]" pageId="20" pageNumber="179">Western Palaearctic.</paragraph>
</subSubSection>
<subSubSection id="C36B65238161363E28F0FDD345EB0D43" pageId="20" pageNumber="179" type="discussion">
<paragraph id="8BCE36A88161363E28F0FDD3402C0B2E" blockId="20.[189,303,580,606]" box="[189,303,580,606]" pageId="20" pageNumber="179">
<heading id="D08681C48161363E28F0FDD3402C0B2E" bold="true" box="[189,303,580,606]" fontSize="11" level="3" pageId="20" pageNumber="179" reason="3">
<emphasis id="B905EABA8161363E28F0FDD3402C0B2E" bold="true" box="[189,303,580,606]" pageId="20" pageNumber="179">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36A88161363E28F0FDE545EB0D43" blockId="20.[188,1399,626,1075]" pageId="20" pageNumber="179">
This extralimital species is diagnosed here to aid in differentiating it from other members of the
<taxonomicName id="4C714D2B8161363E28F0FD01405C0BDF" authorityName="Knoz" authorityYear="1987" box="[189,351,662,688]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="179" phylum="Arthropoda" rank="species" species="moravicus">
<emphasis id="B905EABA8161363E28F0FD01405C0BDF" box="[189,351,662,688]" italics="true" pageId="20" pageNumber="179">C. moravicus</emphasis>
</taxonomicName>
group, particularly
<emphasis id="B905EABA8161363E2A0BFD0143620BC0" box="[582,609,662,688]" italics="true" pageId="20" pageNumber="179">C.</emphasis>
sp. M1 (see above). Delécolle &amp; Schiegg synonymized
<taxonomicName id="4C714D2B8161363E2CBCFD0144750BC0" authorityName="Wirth &amp; Grogan" authorityYear="1988" box="[1265,1398,662,688]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="179" phylum="Arthropoda" rank="species" species="havelkai">
<emphasis id="B905EABA8161363E2CBCFD0144750BC0" box="[1265,1398,662,688]" italics="true" pageId="20" pageNumber="179">C.havelkai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C714D2B8161363E28BEFD2E40970BA3" authorityName="Knoz" authorityYear="1987" box="[243,404,697,723]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="179" phylum="Arthropoda" rank="species" species="moravicus">
<emphasis id="B905EABA8161363E28BEFD2E40970BA3" box="[243,404,697,723]" italics="true" pageId="20" pageNumber="179">C. moravicus</emphasis>
</taxonomicName>
without examining either
<typeStatus id="54CA880A8161363E2A91FD2E42450BA3" box="[732,838,697,723]" pageId="20" pageNumber="179" type="holotype">holotype</typeStatus>
, based solely on specimens from
<collectingCountry id="F36676388161363E2CA7FD2E44750BA3" box="[1258,1398,697,723]" name="Switzerland" pageId="20" pageNumber="179">Switzerland</collectingCountry>
and
<collectingCountry id="F36676388161363E28BCFD4B405D0B86" box="[241,350,732,758]" name="Germany" pageId="20" pageNumber="179">Germany</collectingCountry>
(
<bibRefCitation id="EFE04B598161363E293DFD4B43B70B86" author="Delecolle J. &amp; Schiegg K." box="[368,692,732,758]" pageId="20" pageNumber="179" pagination="273 - 286" refId="ref22869" refString="Delecolle J. &amp; Schiegg K. 1998. Contribution a l'etude des ceratopogonides de Suisse. Revision et redescription des especes palearctiques du genre Ceratoculicoides Wirth &amp; Ratanaworabhan, 1971 (Diptera, Nematocera). Bulletin de la Societe entomologique de France 103: 273 - 286." type="journal article" year="1998">Delécolle &amp; Schiegg 1998</bibRefCitation>
). Any future work on the Palaearctic fauna should involve examining both type specimens to confirm they are conspecific. Females of this species are the only known
<taxonomicName id="4C714D2B8161363E2955FCB540E30A4C" authorityName=", Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[280,480,802,828]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="179" phylum="Arthropoda" rank="genus">
<emphasis id="B905EABA8161363E2955FCB540E30A4C" box="[280,480,802,828]" italics="true" pageId="20" pageNumber="179">Ceratoculicoides</emphasis>
</taxonomicName>
with a single major spermatheca. The males are extremely similar to those of
<taxonomicName id="4C714D2B8161363E28ADFCD140550A10" authorityName="Huerta &amp; Borkent" authorityYear="2005" box="[224,342,838,864]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="179" phylum="Arthropoda" rank="species" species="aliciae">
<emphasis id="B905EABA8161363E28ADFCD140550A10" box="[224,342,838,864]" italics="true" pageId="20" pageNumber="179">C.aliciae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C714D2B8161363E2924FCD143170A2F" authority="Fasbender, 2023" authorityName="Fasbender" authorityYear="2023" box="[361,532,838,864]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="179" phylum="Arthropoda" rank="species" species="propinquus" status="sp. nov.">
<emphasis id="B905EABA8161363E2924FCD143170A2F" box="[361,532,838,864]" italics="true" pageId="20" pageNumber="179">C. propinquus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657C18161363E2A53FCD1437E0A10" box="[542,637,838,864]" pageId="20" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
and
<emphasis id="B905EABA8161363E2AF0FCD143D40A10" box="[701,727,838,864]" italics="true" pageId="20" pageNumber="179">C.</emphasis>
sp. M1.
<taxonomicName id="4C714D2B8161363E2B07FCD1459B0A2F" authorityName="Knoz" authorityYear="1987" box="[842,1176,838,864]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="179" phylum="Arthropoda" rank="species" species="moravicus">
<emphasis id="B905EABA8161363E2B07FCD1459B0A2F" box="[842,1176,838,864]" italics="true" pageId="20" pageNumber="179">Ceratoculicoides moravicus</emphasis>
</taxonomicName>
(
<figureCitation id="134A2A2D8161363E2CE1FCD144100A10" box="[1196,1299,838,864]" captionStart="Fig" captionStartId="18.[189,231,1752,1778]" captionTargetBox="[218,1370,268,1690]" captionTargetId="figure-141@18.[218,1370,268,1690]" captionTargetPageId="18" captionText="Fig. 10. Male genitalia III. Ceratoculicoides sp. M1: a. Genitalia, aedeagus and parameres removed. b. Parameres. c. Aedeagus. C.aliciaeHuerta &amp; Borkent, 2005: d. Parameres (adapted from Huerta &amp; Borkent 2005). e. Aedeagus (adapted from Huerta &amp; Borkent 2005). C.moravicusKnoz, 1987: f. Genitalia, aedeagus and parameres removed. g. Parameres. h. Aedeagus. C. propinquus sp. nov.: i. Genitalia, aedeagus and parameres removed. j. Parameres. k. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083804" httpUri="https://zenodo.org/record/8083804/files/figure.png" pageId="20" pageNumber="179">Fig. 10h</figureCitation>
) can be distinguished from
<taxonomicName id="4C714D2B8161363E29D6FCFE430E0AF3" authorityName="Huerta &amp; Borkent" authorityYear="2005" box="[411,525,873,899]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="179" phylum="Arthropoda" rank="species" species="aliciae">
<emphasis id="B905EABA8161363E29D6FCFE430E0AF3" box="[411,525,873,899]" italics="true" pageId="20" pageNumber="179">C.aliciae</emphasis>
</taxonomicName>
(
<figureCitation id="134A2A2D8161363E2A56FCFE437F0AF3" box="[539,636,873,899]" captionStart="Fig" captionStartId="18.[189,231,1752,1778]" captionTargetBox="[218,1370,268,1690]" captionTargetId="figure-141@18.[218,1370,268,1690]" captionTargetPageId="18" captionText="Fig. 10. Male genitalia III. Ceratoculicoides sp. M1: a. Genitalia, aedeagus and parameres removed. b. Parameres. c. Aedeagus. C.aliciaeHuerta &amp; Borkent, 2005: d. Parameres (adapted from Huerta &amp; Borkent 2005). e. Aedeagus (adapted from Huerta &amp; Borkent 2005). C.moravicusKnoz, 1987: f. Genitalia, aedeagus and parameres removed. g. Parameres. h. Aedeagus. C. propinquus sp. nov.: i. Genitalia, aedeagus and parameres removed. j. Parameres. k. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083804" httpUri="https://zenodo.org/record/8083804/files/figure.png" pageId="20" pageNumber="179">Fig. 10e</figureCitation>
) by the posterior margin of the aedeagus being evenly concave (vs with acute medial notch), from
<taxonomicName id="4C714D2B8161363E2A61FC1B43D00AD6" authorityName="Fasbender" authorityYear="2023" box="[556,723,908,934]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="179" phylum="Arthropoda" rank="species" species="propinquus">
<emphasis id="B905EABA8161363E2A61FC1B43D00AD6" box="[556,723,908,934]" italics="true" pageId="20" pageNumber="179">C.propinquus</emphasis>
</taxonomicName>
(
<figureCitation id="134A2A2D8161363E2AAFFC1B42450AD6" box="[738,838,908,934]" captionStart="Fig" captionStartId="18.[189,231,1752,1778]" captionTargetBox="[218,1370,268,1690]" captionTargetId="figure-141@18.[218,1370,268,1690]" captionTargetPageId="18" captionText="Fig. 10. Male genitalia III. Ceratoculicoides sp. M1: a. Genitalia, aedeagus and parameres removed. b. Parameres. c. Aedeagus. C.aliciaeHuerta &amp; Borkent, 2005: d. Parameres (adapted from Huerta &amp; Borkent 2005). e. Aedeagus (adapted from Huerta &amp; Borkent 2005). C.moravicusKnoz, 1987: f. Genitalia, aedeagus and parameres removed. g. Parameres. h. Aedeagus. C. propinquus sp. nov.: i. Genitalia, aedeagus and parameres removed. j. Parameres. k. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083804" httpUri="https://zenodo.org/record/8083804/files/figure.png" pageId="20" pageNumber="179">Fig. 10k</figureCitation>
) by the accessory spines between the apical and posterolateral points (spines absent in
<taxonomicName id="4C714D2B8161363E2A33FC2742260AB9" authorityName="Fasbender" authorityYear="2023" box="[638,805,944,970]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="179" phylum="Arthropoda" rank="species" species="propinquus">
<emphasis id="B905EABA8161363E2A33FC2742260AB9" box="[638,805,944,970]" italics="true" pageId="20" pageNumber="179">C.propinquus</emphasis>
</taxonomicName>
), and
<emphasis id="B905EABA8161363E2B23FC27428A0ABA" box="[878,905,944,970]" italics="true" pageId="20" pageNumber="179">C.</emphasis>
sp. M1 (
<figureCitation id="134A2A2D8161363E2BBFFC3845570AB9" box="[1010,1108,943,969]" captionStart="Fig" captionStartId="18.[189,231,1752,1778]" captionTargetBox="[218,1370,268,1690]" captionTargetId="figure-141@18.[218,1370,268,1690]" captionTargetPageId="18" captionText="Fig. 10. Male genitalia III. Ceratoculicoides sp. M1: a. Genitalia, aedeagus and parameres removed. b. Parameres. c. Aedeagus. C.aliciaeHuerta &amp; Borkent, 2005: d. Parameres (adapted from Huerta &amp; Borkent 2005). e. Aedeagus (adapted from Huerta &amp; Borkent 2005). C.moravicusKnoz, 1987: f. Genitalia, aedeagus and parameres removed. g. Parameres. h. Aedeagus. C. propinquus sp. nov.: i. Genitalia, aedeagus and parameres removed. j. Parameres. k. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083804" httpUri="https://zenodo.org/record/8083804/files/figure.png" pageId="20" pageNumber="179">Fig. 10c</figureCitation>
) by having the aedeagus tapering distinctly at midlength (vs lateral margins straight). The differences between
<taxonomicName id="4C714D2B8161363E2CEEFC4444460A9C" authorityName="Knoz" authorityYear="1987" box="[1187,1349,979,1005]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="179" phylum="Arthropoda" rank="species" species="moravicus">
<emphasis id="B905EABA8161363E2CEEFC4444460A9C" box="[1187,1349,979,1005]" italics="true" pageId="20" pageNumber="179">C. moravicus</emphasis>
</taxonomicName>
and
<emphasis id="B905EABA8161363E28F1FC6141D40D60" box="[188,215,1014,1040]" italics="true" pageId="20" pageNumber="179">C.</emphasis>
sp. M1 may be the result of the overly compressed slide preparation of that specimen. Additional material is needed to ascertain whether the latter provisional morphospecies is conspecific.
</paragraph>
</subSubSection>
</treatment>
</document>
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