190 lines
19 KiB
XML
190 lines
19 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4759018" ID-GBIF-Dataset="e094e473-f8d3-49f6-8d86-49ad57e85df0" ID-GBIF-Taxon="182067989" ID-ISSN="1854-0392" ID-Zenodo-Dep="4759018" checkinTime="1620933308184" checkinUser="felipe" docAuthor="Stark, Bill P. & Sivec, Ignac" docDate="2005" docId="03D88787FFDAFFFFFC8074F1B02A30A0" docLanguage="en" docName="Illiesia.1.1.1-7.pdf" docOrigin="Illiesia 1 (1)" docStyle="DocumentStyle:1DEF1196F206C1C297CF0540A5C6768F.1:Illesia.2005-2008.journal_article" docStyleId="1DEF1196F206C1C297CF0540A5C6768F" docStyleName="Illesia.2005-2008.journal_article" docStyleVersion="1" docTitle="Tyloperla courtneyi Stark & Sivec 2005, sp. n." docType="treatment" docVersion="4" lastPageId="2" lastPageNumber="3" masterDocId="FFE1FFFFFFDAFFFDFFE07731B54D320E" masterDocTitle="New Species Of Tyloperla (Plecoptera: Perlidae) From Vietnam And Thailand" masterLastPageNumber="7" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1643588373908" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New Species Of Tyloperla (Plecoptera: Perlidae) From Vietnam And Thailand</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Stark, Bill P.</mods:namePart>
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<mods:affiliation>Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sivec, Ignac</mods:namePart>
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<mods:affiliation>Slovenian Museum of Natural History, Prešernova 20, P. O. Box 290, SLO- 1001 Ljubljana, Slovenia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Illiesia</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2005</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2005-01-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>7</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4758557</mods:identifier>
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<mods:identifier type="GBIF-Dataset">e094e473-f8d3-49f6-8d86-49ad57e85df0</mods:identifier>
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<mods:identifier type="ISSN">1854-0392</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4758557</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4759018" ID-GBIF-Taxon="182067989" ID-Zenodo-Dep="4759018" LSID="urn:lsid:plazi:treatment:03D88787FFDAFFFFFC8074F1B02A30A0" httpUri="http://treatment.plazi.org/id/03D88787FFDAFFFFFC8074F1B02A30A0" lastPageId="2" lastPageNumber="3" pageId="0" pageNumber="1">
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<subSubSection box="[864,1352,960,985]" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph blockId="0.[812,1405,960,1953]" box="[864,1352,960,985]" pageId="0" pageNumber="1">
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<heading box="[864,1352,960,985]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="1" reason="3">
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<emphasis bold="true" box="[864,1166,960,985]" pageId="0" pageNumber="1">
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<taxonomicName authority="Stark & Sivec, 2005" authorityName="Stark & Sivec" authorityYear="2005" box="[864,1091,960,984]" class="Insecta" family="Perlidae" genus="Tyloperla" kingdom="Animalia" order="Plecoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="courtneyi" status="sp. n.">
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<emphasis bold="true" box="[864,1091,960,984]" italics="true" pageId="0" pageNumber="1">Tyloperla courtneyi</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel box="[1104,1166,961,985]" pageId="0" pageNumber="1" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation box="[1181,1251,960,985]" captionStart="Figs" captionStartId="1.[184,234,1870,1894]" captionTargetBox="[164,1264,263,1800]" captionTargetId="figure-75@1.[158,930,915,1264]" captionTargetPageId="1" captionText="Figs. 1-6. Tyloperla courtneyi. 1. Head and pronotum. 2. Male terminalia. 3. Hemitergal process, dorsolateral. 4. Aedeagus, lateral. 5. Female terminalia. 6. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758559" httpUri="https://zenodo.org/record/4758559/files/figure.png" pageId="0" pageNumber="1">Figs. 1</figureCitation>
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-6,21-23)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph blockId="0.[812,1405,960,1953]" pageId="0" pageNumber="1">
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<emphasis bold="true" box="[812,912,995,1019]" pageId="0" pageNumber="1">Material</emphasis>
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.
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<materialsCitation ID-GBIF-Occurrence="3111217301" collectingDate="2002-05-08" collectingDateMax="2002-05-09" collectingDateMin="2002-05-08" collectionCode="PMSL" country="Thailand" county="Doi Inthanon National Park" location="University Team" municipality="Siriphum" pageId="0" pageNumber="1" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="holotype">
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<typeStatus box="[936,1044,995,1019]" pageId="0" pageNumber="1">Holotype</typeStatus>
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<specimenCount box="[1062,1081,995,1019]" pageId="0" pageNumber="1" type="male">♂</specimenCount>
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and
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<specimenCount box="[1161,1210,995,1020]" pageId="0" pageNumber="1" type="female">1 ♀</specimenCount>
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<typeStatus box="[1228,1330,996,1020]" pageId="0" pageNumber="1">paratype</typeStatus>
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from
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<collectingCountry box="[812,917,1030,1054]" name="Thailand" pageId="0" pageNumber="1">Thailand</collectingCountry>
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,
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<collectingRegion box="[928,1066,1030,1054]" country="Thailand" name="Chiang Mai" pageId="0" pageNumber="1">Chiang Mai</collectingRegion>
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,
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<collectingCounty box="[1077,1399,1030,1054]" pageId="0" pageNumber="1">Doi Inthanon National Park</collectingCounty>
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,
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<collectingMunicipality box="[812,920,1065,1089]" pageId="0" pageNumber="1">Siriphum</collectingMunicipality>
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waterfall,
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<date box="[1063,1237,1065,1089]" pageId="0" pageNumber="1" value="2002-05-08" valueMax="2002-05-09" valueMin="2002-05-08">
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<collectingDate box="[1063,1237,1065,1089]" pageId="0" pageNumber="1" value="2002-05-08" valueMax="2002-05-09" valueMin="2002-05-08">8-9 May 2002</collectingDate>
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</date>
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,
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<collectingRegion box="[1258,1404,1065,1089]" country="Thailand" name="Chiang Mai" pageId="0" pageNumber="1">Chiang Mai</collectingRegion>
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<location LSID="urn:lsid:plazi:treatment:03D88787FFDAFFFFFC8074F1B02A30A0:8EAE604AFFDAFFFDFCCC737AB6A1366D" box="[812,1004,1099,1123]" country="Thailand" county="Doi Inthanon National Park" municipality="Siriphum" name="University Team" pageId="0" pageNumber="1" stateProvince="Chiang Mai">University Team</location>
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(
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<collectionCode box="[1018,1093,1099,1123]" collectionName="Slovenia, Ljubljana, Slovenian Museum of Natural History" country="Slovenia" httpUri="http://grbio.org/cool/4mab-159w" name="Slovenian Museum of Natural History (Prirodosloveni Muzej Slovenije)" pageId="0" pageNumber="1">PMSL</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="3" pageId="0" pageNumber="1" type="description">
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<paragraph blockId="0.[812,1405,960,1953]" pageId="0" pageNumber="1">
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<emphasis bold="true" box="[812,979,1134,1158]" pageId="0" pageNumber="1">Adult habitus.</emphasis>
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Triocellate. Head yellowish with dark brown spot between ocelli expanded along anterior margin near callosities; anterior region of frons dusky brown forward of M-line; pronotum brown with darker rugosities (
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<figureCitation box="[1024,1088,1272,1296]" captionStart="Figs" captionStartId="1.[184,234,1870,1894]" captionTargetBox="[164,1264,263,1800]" captionTargetId="figure-75@1.[158,930,915,1264]" captionTargetPageId="1" captionText="Figs. 1-6. Tyloperla courtneyi. 1. Head and pronotum. 2. Male terminalia. 3. Hemitergal process, dorsolateral. 4. Aedeagus, lateral. 5. Female terminalia. 6. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758559" httpUri="https://zenodo.org/record/4758559/files/figure.png" pageId="0" pageNumber="1">Fig. 1</figureCitation>
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). Palpi, antennae and cerci brown. Legs brown with distal end of femora and proximal end of tibiae slightly darker. Wing membrane and veins brown.
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</paragraph>
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<paragraph blockId="0.[812,1405,960,1953]" pageId="0" pageNumber="1">
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<emphasis bold="true" box="[812,879,1411,1435]" pageId="0" pageNumber="1">Male.</emphasis>
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Forewing length
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<quantity box="[1076,1157,1411,1435]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="0" pageNumber="1" unit="mm" value="17.0">17 mm</quantity>
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. Hairbrushes reduced on abdominal sternum 5 but well developed on sterna 6-7 and on thoracic metasternum. Tergum 8 and 9 without distinct lobes but each with a mesal low mound covered with sensilla basiconica (
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<figureCitation box="[1327,1391,1549,1573]" captionStart="Figs" captionStartId="1.[184,234,1870,1894]" captionTargetBox="[164,1264,263,1800]" captionTargetId="figure-75@1.[158,930,915,1264]" captionTargetPageId="1" captionText="Figs. 1-6. Tyloperla courtneyi. 1. Head and pronotum. 2. Male terminalia. 3. Hemitergal process, dorsolateral. 4. Aedeagus, lateral. 5. Female terminalia. 6. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758559" httpUri="https://zenodo.org/record/4758559/files/figure.png" pageId="0" pageNumber="1">Fig. 2</figureCitation>
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). Basal cushion of hemitergal process about 1/3 of inner length of process; process rounded dorsally, grooved along inner surface and pointed dorsoapically (
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<figureCitation box="[978,1039,1687,1711]" captionStart="Figs" captionStartId="1.[184,234,1870,1894]" captionTargetBox="[164,1264,263,1800]" captionTargetId="figure-75@1.[158,930,915,1264]" captionTargetPageId="1" captionText="Figs. 1-6. Tyloperla courtneyi. 1. Head and pronotum. 2. Male terminalia. 3. Hemitergal process, dorsolateral. 4. Aedeagus, lateral. 5. Female terminalia. 6. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758559" httpUri="https://zenodo.org/record/4758559/files/figure.png" pageId="0" pageNumber="1">Fig. 3</figureCitation>
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). Aedeagal sac apically recurved and bearing a pair of small basolateral, spiny lobes (
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<figureCitation box="[820,894,1756,1780]" captionStart="Figs" captionStartId="1.[184,234,1870,1894]" captionTargetBox="[164,1264,263,1800]" captionTargetId="figure-75@1.[158,930,915,1264]" captionTargetPageId="1" captionText="Figs. 1-6. Tyloperla courtneyi. 1. Head and pronotum. 2. Male terminalia. 3. Hemitergal process, dorsolateral. 4. Aedeagus, lateral. 5. Female terminalia. 6. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758559" httpUri="https://zenodo.org/record/4758559/files/figure.png" pageId="0" pageNumber="1">Fig. 4</figureCitation>
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); sac apex offset by a distinct subapical constriction; armature well developed as a broad patch of small spines over apex and extending beyond subapical constriction over lateral and ventral surfaces to near midlength of sac. Cercal segments with a single enlarged ventral bristle.
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</paragraph>
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<paragraph blockId="1.[1282,1307,1639,1684]" box="[1282,1307,1639,1684]" pageId="1" pageNumber="2">
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<emphasis bold="true" box="[1282,1307,1639,1684]" pageId="1" pageNumber="2">5</emphasis>
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.4758559" ID-Zenodo-Dep="4758559" httpUri="https://zenodo.org/record/4758559/files/figure.png" pageId="1" pageNumber="2" startId="1.[184,234,1870,1894]" targetBox="[164,1264,263,1800]" targetPageId="1">
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<paragraph blockId="1.[184,1404,1870,1928]" pageId="1" pageNumber="2">
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Figs. 1-6.
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<taxonomicName authorityName="Stark & Sivec" authorityYear="2005" box="[288,492,1870,1894]" class="Insecta" family="Perlidae" genus="Tyloperla" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="courtneyi">
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<emphasis box="[288,492,1870,1894]" italics="true" pageId="1" pageNumber="2">Tyloperla courtneyi</emphasis>
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</taxonomicName>
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. 1. Head and pronotum. 2. Male terminalia. 3. Hemitergal process, dorsolateral. 4. Aedeagus, lateral. 5. Female terminalia. 6. Vagina and spermatheca.
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</paragraph>
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</caption>
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<paragraph blockId="2.[184,776,316,824]" pageId="2" pageNumber="3">
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<emphasis bold="true" box="[184,276,316,340]" pageId="2" pageNumber="3">Female.</emphasis>
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Forewing length
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<quantity box="[488,573,316,340]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="3" unit="mm" value="24.0">24 mm</quantity>
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. Subgenital plate arcuate with a median U-shaped notch extending deeply into plate; plate produced over ca. ¼ or less of sternum 9 (
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<figureCitation box="[310,374,419,443]" captionStart="Figs" captionStartId="1.[184,234,1870,1894]" captionTargetBox="[164,1264,263,1800]" captionTargetId="figure-75@1.[158,930,915,1264]" captionTargetPageId="1" captionText="Figs. 1-6. Tyloperla courtneyi. 1. Head and pronotum. 2. Male terminalia. 3. Hemitergal process, dorsolateral. 4. Aedeagus, lateral. 5. Female terminalia. 6. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758559" httpUri="https://zenodo.org/record/4758559/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
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). Vagina about as wide as long with major anterior chamber weakly lined along lateral wrinkles with brown setae. Spermatheca hook shaped (
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<figureCitation box="[281,346,523,547]" captionStart="Figs" captionStartId="1.[184,234,1870,1894]" captionTargetBox="[164,1264,263,1800]" captionTargetId="figure-75@1.[158,930,915,1264]" captionTargetPageId="1" captionText="Figs. 1-6. Tyloperla courtneyi. 1. Head and pronotum. 2. Male terminalia. 3. Hemitergal process, dorsolateral. 4. Aedeagus, lateral. 5. Female terminalia. 6. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758559" httpUri="https://zenodo.org/record/4758559/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
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).
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</paragraph>
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<paragraph blockId="2.[184,776,316,824]" pageId="2" pageNumber="3">
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<emphasis bold="true" box="[184,235,558,582]" pageId="2" pageNumber="3">Egg.</emphasis>
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Length ca.
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<quantity box="[368,469,558,582]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="3" unit="mm" value="0.42">0.42 mm</quantity>
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, width ca.
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<quantity box="[593,694,558,582]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="3" unit="mm" value="0.38">0.38 mm</quantity>
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. Collar short, ca.
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<quantity box="[290,386,593,617]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="3" unit="mm" value="0.03">0.03 mm</quantity>
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in length, and wide, ca.
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<quantity box="[653,749,593,617]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="3" unit="mm" value="0.2">0.20 mm</quantity>
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in diameter (
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<figureCitation box="[305,389,627,651]" captionStart="Figs" captionStartId="5.[812,862,1835,1859]" captionTargetBox="[810,1405,339,1772]" captionTargetId="figure-85@5.[808,1405,824,1287]" captionTargetPageId="5" captionText="Figs. 21-23. Tyloperla courtneyi egg structures. 21. Entire egg, lateral. 22. Egg collar, polar aspect. 23. Opercular ring and lid." figureDoi="http://doi.org/10.5281/zenodo.4758927" httpUri="https://zenodo.org/record/4758927/files/figure.png" pageId="2" pageNumber="3">Fig. 22</figureCitation>
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); collar rim smooth and slightly flanged, sides with a series of obscure vertical ridges (
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<figureCitation box="[192,274,696,720]" captionStart="Figs" captionStartId="5.[812,862,1835,1859]" captionTargetBox="[810,1405,339,1772]" captionTargetId="figure-85@5.[808,1405,824,1287]" captionTargetPageId="5" captionText="Figs. 21-23. Tyloperla courtneyi egg structures. 21. Entire egg, lateral. 22. Egg collar, polar aspect. 23. Opercular ring and lid." figureDoi="http://doi.org/10.5281/zenodo.4758927" httpUri="https://zenodo.org/record/4758927/files/figure.png" pageId="2" pageNumber="3">Fig. 21</figureCitation>
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). Chorion covered with irregularly shaped large pits with ornately punctate floors. Opercular ring distinct as a narrow groove separating lid from egg body (
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<figureCitation box="[305,383,800,824]" captionStart="Figs" captionStartId="5.[812,862,1835,1859]" captionTargetBox="[810,1405,339,1772]" captionTargetId="figure-85@5.[808,1405,824,1287]" captionTargetPageId="5" captionText="Figs. 21-23. Tyloperla courtneyi egg structures. 21. Entire egg, lateral. 22. Egg collar, polar aspect. 23. Opercular ring and lid." figureDoi="http://doi.org/10.5281/zenodo.4758927" httpUri="https://zenodo.org/record/4758927/files/figure.png" pageId="2" pageNumber="3">Fig. 23</figureCitation>
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). Micropyles not observed.
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</paragraph>
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<paragraph blockId="2.[812,1404,316,686]" box="[812,1014,316,340]" pageId="2" pageNumber="3">
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<emphasis bold="true" box="[812,885,316,340]" pageId="2" pageNumber="3">Larva.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="diagnosis">
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<paragraph blockId="2.[812,1404,316,686]" pageId="2" pageNumber="3">
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<emphasis bold="true" box="[812,931,350,374]" pageId="2" pageNumber="3">Diagnosis</emphasis>
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. Males of this species are similar to those of
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<taxonomicName authority="(Okamoto)" baseAuthorityName="Okamoto" baseAuthorityYear="1912" box="[844,1118,385,409]" class="Insecta" family="Perlidae" genus="Tyloperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="formosana">
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<emphasis box="[844,984,385,409]" italics="true" pageId="2" pageNumber="3">T. formosana</emphasis>
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(Okamoto)
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</taxonomicName>
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and
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<taxonomicName authorityName="Stark & Sivec" authorityYear="2005" box="[1180,1274,385,409]" class="Insecta" family="Perlidae" genus="Tyloperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="khang">
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<emphasis box="[1180,1274,385,409]" italics="true" pageId="2" pageNumber="3">T. khang</emphasis>
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</taxonomicName>
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(described below) in external genitalic features. It differs from the former in having shorter, more rounded hemitergal calluses and in lacking small subapical lobes on the aedeagal sac.
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<taxonomicName authorityName="Stark & Sivec" authorityYear="2005" box="[1102,1267,523,547]" class="Insecta" family="Perlidae" genus="Tyloperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="khang">
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<emphasis box="[1102,1267,523,547]" italics="true" pageId="2" pageNumber="3">Tyloperla khang</emphasis>
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</taxonomicName>
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lacks lateral lobes on the aedeagal sac but a pair of these are present basolaterally in
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<taxonomicName authorityName="Stark & Sivec" authorityYear="2005" box="[1085,1214,593,616]" class="Insecta" family="Perlidae" genus="Tyloperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="courtneyi">
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<emphasis box="[1085,1214,593,616]" italics="true" pageId="2" pageNumber="3">T. courtneyi</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="etymology">
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<paragraph blockId="2.[812,1404,316,686]" pageId="2" pageNumber="3">
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<emphasis bold="true" box="[812,939,627,651]" pageId="2" pageNumber="3">Etymology</emphasis>
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. The patronym honors G. Courtney for his leadership in the study of Thai aquatic insects.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |