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<document id="E695B4172F236C5521C3924A8D10AF27" ID-CLB-Dataset="3079" ID-DOI="10.1649/072.068.0mo4.1" ID-GBIF-Dataset="62322993-ca80-4b6c-ac95-861cbb236abd" ID-ISSN="1938-4394" ID-Zenodo-Dep="4907735" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623093043626" checkinUser="carolina" docAuthor="Park, J. - S. &amp; Carlton, C. E." docDate="2014" docId="03D82A3EBD4C11693D48FA0EFB45FA9E" docLanguage="en" docName="TheColeopBull.68.mo13.1-156.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 13) 68" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Sagola pulchra Broun 1880" docType="treatment" docVersion="23" lastPageNumber="38" masterDocId="FFE15246BD69114C3C45FFF3FFD6FF9A" masterDocTitle="A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae)" masterLastPageNumber="156" masterPageNumber="1" pageNumber="38" updateTime="1699001444256" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="683C343A0DAB17001E8EA6120C5DE2B3">A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae)</mods:title>
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<mods:namePart id="DE00E923A23CFA1FDC4CBE5D8476DCC3">Park, J. - S.</mods:namePart>
<mods:affiliation id="978BB129C16DEA638C465957A4698FDD">Louisiana State Arthropod Museum, Department of Entomology, LSB 404 Louisiana State University Agric</mods:affiliation>
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<mods:namePart id="48C498CCE9A10C3064C47E54E516E5EE">Carlton, C. E.</mods:namePart>
<mods:affiliation id="D24225052F3A4644B15FA14A072446F9">Louisiana State Arthropod Museum, Department of Entomology, LSB 404 Louisiana State University Agric</mods:affiliation>
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<mods:title id="DFAB4B6EF44D71A5D6D0034A09DD4256">The Coleopterists Bulletin</mods:title>
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<mods:date id="0234CCF305C4165911890596302548BB">2014</mods:date>
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<mods:title id="8446F930F7056D36438AA873B34229D6">mo 13</mods:title>
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<treatment id="03D82A3EBD4C11693D48FA0EFB45FA9E" ID-DOI="http://doi.org/10.5281/zenodo.5765087" ID-GBIF-Taxon="191245050" ID-Zenodo-Dep="5765087" LSID="urn:lsid:plazi:treatment:03D82A3EBD4C11693D48FA0EFB45FA9E" httpUri="http://treatment.plazi.org/id/03D82A3EBD4C11693D48FA0EFB45FA9E" lastPageNumber="38" pageId="37" pageNumber="38">
<subSubSection id="C36BC8A3BD4C11693D48FA0EFDDFF988" box="[269,521,1533,1554]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="8BCE9B28BD4C11693D48FA0EFDDFF988" blockId="37.[269,521,1533,1581]" box="[269,521,1533,1554]" pageId="37" pageNumber="38">
<heading id="D0862C44BD4C11693D48FA0EFDDFF988" box="[269,521,1533,1554]" centered="true" fontSize="10" level="2" pageId="37" pageNumber="38" reason="2">
<emphasis id="B905473ABD4C11693D48FA0EFDDFF988" bold="true" box="[269,521,1533,1554]" pageId="37" pageNumber="38">
5.
<taxonomicName id="4C71E0ABBD4C11693D6DFA0EFEA2F988" ID-CoL="4TZ2J" authorityName="Broun" authorityYear="1880" box="[296,372,1533,1554]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B905473ABD4C11693D6DFA0EFEA2F988" bold="true" box="[296,372,1533,1554]" italics="true" pageId="37" pageNumber="38">pulchra</emphasis>
</taxonomicName>
species-group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD4C11693D13F9EBFE68F9B7" box="[342,446,1560,1581]" pageId="37" pageNumber="38" type="description">
<paragraph id="8BCE9B28BD4C11693D13F9EBFE68F9B7" blockId="37.[269,521,1533,1581]" box="[342,446,1560,1581]" pageId="37" pageNumber="38">(4 species)</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD4C11693CEAF9B1FC9CFD8C" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="8BCE9B28BD4C11693CEAF9B1FC9CFD8C" blockId="37.[151,638,1602,1729]" lastBlockId="37.[685,1172,194,534]" pageId="37" pageNumber="38">
<emphasis id="B905473ABD4C11693CEAF9B1FECEF9CD" bold="true" box="[175,280,1602,1623]" pageId="37" pageNumber="38">Diagnosis.</emphasis>
The members of the
<taxonomicName id="4C71E0ABBD4C11693E6DF9B1FDA8F9CD" authorityName="Broun" authorityYear="1880" box="[552,638,1602,1623]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B905473ABD4C11693E6DF9B1FDA8F9CD" box="[552,638,1602,1623]" italics="true" pageId="37" pageNumber="38">pulchra</emphasis>
</taxonomicName>
species-group can be distinguished from other
<taxonomicName id="4C71E0ABBD4C11693CD2F984FF0DF916" box="[151,219,1655,1676]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B905473ABD4C11693CD2F984FF0DF916" box="[151,219,1655,1676]" italics="true" pageId="37" pageNumber="38">Sagola</emphasis>
</taxonomicName>
species-groups by the following combination of characters: large body length
<quantity id="4C8936CDBD4C11693E43F961FDACF93D" box="[518,634,1681,1703]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="4.2" metricValueMin="3.2" pageId="37" pageNumber="38" unit="mm" value="3.7" valueMax="4.2" valueMin="3.2">3.24.2 mm</quantity>
; antennomere 1 approximately 2.5 times longer than wide with dull surface; gular region of male head with transverse depression (
<figureCitation id="134A87ADBD4C11693FFAFF2EFBC6FF68" box="[959,1040,221,242]" captionStart="Fig" captionStartId="38.[197,229,1021,1040]" captionTargetBox="[183,1193,187,998]" captionTargetId="figure-353@38.[173,1193,183,1008]" captionTargetPageId="38" captionText="Fig. 19. Sagola species in the pulchra species-group. Habitus, dorsal views: a) S. pulchra, b) S. keejeongi, c) S. ramsayi, d) S. poortmani. Aedeagi, dorsal views: e) S. pulchra, f) S. keejeongi, g) S. ramsayi, h) S. poortmani. S. keejeongi, male: i) Dorsal surface of head, j) Ventral surface of head (arrow indicates lateral process), k) Mid-leg, lateral view. S. pulchra, female: l) Ventrite VIII, ventral view, m) Ventrite IX, ventral view. Scale bars ad = 1 mm, em = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907795" httpUri="https://zenodo.org/record/4907795/files/figure.png" pageId="37" pageNumber="38">Fig. 19j</figureCitation>
); ventral surface of male head with a pair of processes laterally (
<figureCitation id="134A87ADBD4C11693EF0FEE1FCDCFEBD" box="[693,778,274,295]" captionStart="Fig" captionStartId="38.[197,229,1021,1040]" captionTargetBox="[183,1193,187,998]" captionTargetId="figure-353@38.[173,1193,183,1008]" captionTargetPageId="38" captionText="Fig. 19. Sagola species in the pulchra species-group. Habitus, dorsal views: a) S. pulchra, b) S. keejeongi, c) S. ramsayi, d) S. poortmani. Aedeagi, dorsal views: e) S. pulchra, f) S. keejeongi, g) S. ramsayi, h) S. poortmani. S. keejeongi, male: i) Dorsal surface of head, j) Ventral surface of head (arrow indicates lateral process), k) Mid-leg, lateral view. S. pulchra, female: l) Ventrite VIII, ventral view, m) Ventrite IX, ventral view. Scale bars ad = 1 mm, em = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907795" httpUri="https://zenodo.org/record/4907795/files/figure.png" pageId="37" pageNumber="38">Fig. 19j</figureCitation>
: arrow); hind wings reduced to small pads; fore and mid-femur with semicircular depression (
<figureCitation id="134A87ADBD4C11693EA0FEB4FCEDFEC6" box="[741,827,327,348]" captionStart="Fig" captionStartId="38.[197,229,1021,1040]" captionTargetBox="[183,1193,187,998]" captionTargetId="figure-353@38.[173,1193,183,1008]" captionTargetPageId="38" captionText="Fig. 19. Sagola species in the pulchra species-group. Habitus, dorsal views: a) S. pulchra, b) S. keejeongi, c) S. ramsayi, d) S. poortmani. Aedeagi, dorsal views: e) S. pulchra, f) S. keejeongi, g) S. ramsayi, h) S. poortmani. S. keejeongi, male: i) Dorsal surface of head, j) Ventral surface of head (arrow indicates lateral process), k) Mid-leg, lateral view. S. pulchra, female: l) Ventrite VIII, ventral view, m) Ventrite IX, ventral view. Scale bars ad = 1 mm, em = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907795" httpUri="https://zenodo.org/record/4907795/files/figure.png" pageId="37" pageNumber="38">Fig. 19k</figureCitation>
); male mid-femur at least twice as broad as tibia (
<figureCitation id="134A87ADBD4C11693F06FE92FC4EFEEC" box="[835,920,353,374]" captionStart="Fig" captionStartId="38.[197,229,1021,1040]" captionTargetBox="[183,1193,187,998]" captionTargetId="figure-353@38.[173,1193,183,1008]" captionTargetPageId="38" captionText="Fig. 19. Sagola species in the pulchra species-group. Habitus, dorsal views: a) S. pulchra, b) S. keejeongi, c) S. ramsayi, d) S. poortmani. Aedeagi, dorsal views: e) S. pulchra, f) S. keejeongi, g) S. ramsayi, h) S. poortmani. S. keejeongi, male: i) Dorsal surface of head, j) Ventral surface of head (arrow indicates lateral process), k) Mid-leg, lateral view. S. pulchra, female: l) Ventrite VIII, ventral view, m) Ventrite IX, ventral view. Scale bars ad = 1 mm, em = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907795" httpUri="https://zenodo.org/record/4907795/files/figure.png" pageId="37" pageNumber="38">Fig. 19k</figureCitation>
); Male tergite IV without microtrichial patch; abdominal tergites IVVI with discal carinae; female abdominal ventrite VIII triangularly produced posteriorly (
<figureCitation id="134A87ADBD4C11693FA2FE42FBE1FE5C" box="[999,1079,433,454]" captionStart="Fig" captionStartId="38.[197,229,1021,1040]" captionTargetBox="[183,1193,187,998]" captionTargetId="figure-353@38.[173,1193,183,1008]" captionTargetPageId="38" captionText="Fig. 19. Sagola species in the pulchra species-group. Habitus, dorsal views: a) S. pulchra, b) S. keejeongi, c) S. ramsayi, d) S. poortmani. Aedeagi, dorsal views: e) S. pulchra, f) S. keejeongi, g) S. ramsayi, h) S. poortmani. S. keejeongi, male: i) Dorsal surface of head, j) Ventral surface of head (arrow indicates lateral process), k) Mid-leg, lateral view. S. pulchra, female: l) Ventrite VIII, ventral view, m) Ventrite IX, ventral view. Scale bars ad = 1 mm, em = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907795" httpUri="https://zenodo.org/record/4907795/files/figure.png" pageId="37" pageNumber="38">Fig. 19l</figureCitation>
), ventrite IX deeply emarginate posteriorly (
<figureCitation id="134A87ADBD4C11693867FE3FFB5EFE7B" box="[1058,1160,460,481]" captionStart="Fig" captionStartId="38.[197,229,1021,1040]" captionTargetBox="[183,1193,187,998]" captionTargetId="figure-353@38.[173,1193,183,1008]" captionTargetPageId="38" captionText="Fig. 19. Sagola species in the pulchra species-group. Habitus, dorsal views: a) S. pulchra, b) S. keejeongi, c) S. ramsayi, d) S. poortmani. Aedeagi, dorsal views: e) S. pulchra, f) S. keejeongi, g) S. ramsayi, h) S. poortmani. S. keejeongi, male: i) Dorsal surface of head, j) Ventral surface of head (arrow indicates lateral process), k) Mid-leg, lateral view. S. pulchra, female: l) Ventrite VIII, ventral view, m) Ventrite IX, ventral view. Scale bars ad = 1 mm, em = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907795" httpUri="https://zenodo.org/record/4907795/files/figure.png" pageId="37" pageNumber="38">
Fig.
<quantity id="4C8936CDBD4C1169381DFE3FFB5EFE7B" box="[1112,1160,460,481]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="37" pageNumber="38" unit="m" value="19.0">19m</quantity>
</figureCitation>
); present on North Island, not known from South Island (
<figureCitation id="134A87ADBD4C11693EB3FDF2FC96FD8C" box="[758,832,513,534]" captionStart="Fig" captionStartId="39.[175,207,1662,1681]" captionTargetBox="[154,636,943,1636]" captionTargetId="figure-658@39.[151,637,938,1639]" captionTargetPageId="39" captionText="Fig. 20. Known collection localities of New Zealand Sagola species. S. pulchra: black circles; S. keejeongi: triangles; S. ramsayi: black square; S. poortmani: star." figureDoi="http://doi.org/10.5281/zenodo.4907799" httpUri="https://zenodo.org/record/4907799/files/figure.png" pageId="37" pageNumber="38">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD4C11693EB4FDB3FB45FA9E" pageId="37" pageNumber="38" type="key">
<paragraph id="8BCE9B28BD4C11693EB4FDB3FC25FDE9" blockId="37.[753,1103,576,628]" pageId="37" pageNumber="38">
<emphasis id="B905473ABD4C11693EB4FDB3FC25FDE9" bold="true" pageId="37" pageNumber="38">
KEY TO SPECIES OF THE
<emphasis id="B905473ABD4C11693FBDFDB6FB99FDCF" bold="true" box="[1016,1103,581,597]" italics="true" pageId="37" pageNumber="38">PULCHRA</emphasis>
SPECIES- GROUP
</emphasis>
</paragraph>
<paragraph id="8BCE9B28BD4C11693E80FD79FCF1FD4E" blockId="37.[685,1172,650,724]" pageId="37" pageNumber="38">The key is based on genitalia because most specimens are indistinguishable based on external morphology.</paragraph>
<key id="FFE0CC19BD4C11693EE8FD1AFB45FA9E" pageId="37" pageNumber="38">
<keyStep id="3085838DBD4C11693EE8FD1AFB42FC11" pageId="37" pageNumber="38">
<paragraph id="8BCE9B28BD4C11693EE8FD1AFB42FCA9" blockId="37.[685,1172,745,1284]" pageId="37" pageNumber="38">
<keyLead id="3080381DBD4C11693EE8FD1AFB42FCA9" pageId="37" pageNumber="38">
1. Median lobe of genitalia without minor lobe; broader paramere without branch from base (
<figureCitation id="134A87ADBD4C11693F2EFCEDFC6FFCA9" box="[875,953,798,819]" captionStart="Fig" captionStartId="38.[197,229,1021,1040]" captionTargetBox="[183,1193,187,998]" captionTargetId="figure-353@38.[173,1193,183,1008]" captionTargetPageId="38" captionText="Fig. 19. Sagola species in the pulchra species-group. Habitus, dorsal views: a) S. pulchra, b) S. keejeongi, c) S. ramsayi, d) S. poortmani. Aedeagi, dorsal views: e) S. pulchra, f) S. keejeongi, g) S. ramsayi, h) S. poortmani. S. keejeongi, male: i) Dorsal surface of head, j) Ventral surface of head (arrow indicates lateral process), k) Mid-leg, lateral view. S. pulchra, female: l) Ventrite VIII, ventral view, m) Ventrite IX, ventral view. Scale bars ad = 1 mm, em = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907795" httpUri="https://zenodo.org/record/4907795/files/figure.png" pageId="37" pageNumber="38">Fig. 19e</figureCitation>
)......
<taxonomicName id="4C71E0ABBD4C11693FA2FCEDFB42FCA9" ID-CoL="4TZ2J" authority="Broun" authorityName="Broun" authorityYear="1880" box="[999,1172,798,819]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B905473ABD4C11693FA2FCEDFB42FCA9" bold="true" box="[999,1172,798,819]" pageId="37" pageNumber="38">
<emphasis id="B905473ABD4C11693FA2FCEDFB9BFCA9" bold="true" box="[999,1101,798,819]" italics="true" pageId="37" pageNumber="38">S. pulchra</emphasis>
Broun
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCE9B28BD4C11693EE8FCB2FB42FC11" blockId="37.[685,1172,745,1284]" pageId="37" pageNumber="38">
<keyLead id="3080381DBD4C11693EE8FCB2FB42FC11" pageId="37" pageNumber="38">1. Median lobe of genitalia with minor lobe; broader paramere with 1 or 2 branches from base .................................................2</keyLead>
</paragraph>
</keyStep>
<keyStep id="3085838DBD4C11693EE8FC6AFB42FBBA" pageId="37" pageNumber="38">
<paragraph id="8BCE9B28BD4C11693EE8FC6AFB42FC52" blockId="37.[685,1172,745,1284]" pageId="37" pageNumber="38">
<keyLead id="3080381DBD4C11693EE8FC6AFB42FC52" pageId="37" pageNumber="38">2(1). Branches of apical lobe of genitalia approximately equal............................................... 3</keyLead>
</paragraph>
<paragraph id="8BCE9B28BD4C11693EE8FC25FB42FBBA" blockId="37.[685,1172,745,1284]" pageId="37" pageNumber="38">
<keyLead id="3080381DBD4C11693EE8FC25FB42FBBA" pageId="37" pageNumber="38">
2. Apical lobe of genitalia with longer major branch and shorter minor branch (
<figureCitation id="134A87ADBD4C11693878FC03FB5BFB9F" box="[1085,1165,1008,1029]" captionStart="Fig" captionStartId="38.[197,229,1021,1040]" captionTargetBox="[183,1193,187,998]" captionTargetId="figure-353@38.[173,1193,183,1008]" captionTargetPageId="38" captionText="Fig. 19. Sagola species in the pulchra species-group. Habitus, dorsal views: a) S. pulchra, b) S. keejeongi, c) S. ramsayi, d) S. poortmani. Aedeagi, dorsal views: e) S. pulchra, f) S. keejeongi, g) S. ramsayi, h) S. poortmani. S. keejeongi, male: i) Dorsal surface of head, j) Ventral surface of head (arrow indicates lateral process), k) Mid-leg, lateral view. S. pulchra, female: l) Ventrite VIII, ventral view, m) Ventrite IX, ventral view. Scale bars ad = 1 mm, em = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907795" httpUri="https://zenodo.org/record/4907795/files/figure.png" pageId="37" pageNumber="38">Fig. 19h</figureCitation>
) ..........................
<emphasis id="B905473ABD4C11693FDDFBF8FB42FBBA" bold="true" box="[920,1172,1035,1056]" pageId="37" pageNumber="38">
<taxonomicName id="4C71E0ABBD4C11693FDDFBF8FBCEFBBA" ID-CoL="4TZ2D" box="[920,1048,1035,1056]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="poortmani" status="sp. nov.">
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3(2). Apical lobe of genitalia with U-shaped emargination; right paramere with 2 branches from base (
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) .................. .........................
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<taxonomicName id="4C71E0ABBD4C11693FDEFB8EFBC3FB08" ID-CoL="4TYYR" box="[923,1045,1149,1170]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="keejeongi" status="sp. nov.">
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3. Apical lobe of genitalia with V-shaped emargination; left paramere with 1 branch from base (
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Fig.
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) ................................. ............................
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<taxonomicName id="4C71E0ABBD4C11693FEEFB1CFBC0FA9E" ID-CoL="4TZ2P" box="[939,1046,1263,1284]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="ramsayi" status="sp. nov.">
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