treatments-xml/data/03/CD/62/03CD6212D56AFFD1FD78FCA2E9C2F975.xml
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<document id="4ECE2660FC7E465C85DF951298F02495" ID-DOI="10.1080/0022293021000055441" ID-ISSN="1464-5262" ID-Zenodo-Dep="4653568" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617217246595" checkinUser="carolina" docAuthor="Peart, R. A." docDate="2004" docId="03CD6212D56AFFD1FD78FCA2E9C2F975" docLanguage="en" docName="JNATHIST.38.301-336.pdf" docOrigin="Journal of Natural History 38 (3)" docSource="http://www.tandfonline.com/doi/abs/10.1080/0022293021000055441" docStyle="DocumentStyle:443144A73DAE3E092C51A378E4CD0138.2:JNatHist.2003-2004.journal_article.1cover" docStyleId="443144A73DAE3E092C51A378E4CD0138" docStyleName="JNatHist.2003-2004.journal_article.1cover" docStyleVersion="2" docTitle="Cymadusa ledoyeri Peart 2004, n. sp." docType="treatment" docVersion="4" lastPageNumber="313" masterDocId="FFF41A6AD560FFDCFFF0FFFBEA3AFFBD" masterDocTitle="A revision of the Cymadusa filosa complex (Crustacea: Amphipoda: Corophioidea: Ampithoidae)" masterLastPageNumber="336" masterPageNumber="301" pageNumber="310" updateTime="1698898288008" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="BDE217C4A38DC9CC87EFDC597D0409E7">A revision of the Cymadusa filosa complex (Crustacea: Amphipoda: Corophioidea: Ampithoidae)</mods:title>
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<mods:namePart id="8BCD33998A88423718D3CD02A17A0101">Peart, R. A.</mods:namePart>
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<mods:title id="D12C8FC843923248CD3BF1C288A1F099">Journal of Natural History</mods:title>
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<mods:date id="83F60CF0E5B9A2FC0F6F0F6040BB18C7">2004</mods:date>
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<mods:number id="3815A2D6A8604265C400F2657B009868">2004-02-10</mods:number>
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<treatment id="03CD6212D56AFFD1FD78FCA2E9C2F975" ID-DOI="http://doi.org/10.5281/zenodo.4653842" ID-Zenodo-Dep="4653842" LSID="urn:lsid:plazi:treatment:03CD6212D56AFFD1FD78FCA2E9C2F975" httpUri="http://treatment.plazi.org/id/03CD6212D56AFFD1FD78FCA2E9C2F975" lastPageId="13" lastPageNumber="313" pageId="10" pageNumber="310">
<subSubSection id="C37E808FD56AFFD6FD78FCA2E95AFC2C" pageId="10" pageNumber="310" type="nomenclature">
<paragraph id="8BDBD304D56AFFD6FD78FCA2E99FFCCC" blockId="10.[313,996,857,945]" box="[648,933,857,881]" pageId="10" pageNumber="310">
<heading id="D0936468D56AFFD6FD78FCA2E99FFCCC" box="[648,933,857,881]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="310" reason="3">
<taxonomicName id="4C64A887D56AFFD6FD78FCA2E967FCCC" ID-CoL="6BV4Z" authority="Peart, 2004" authorityName="Peart" authorityYear="2004" box="[648,861,857,881]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="310" phylum="Arthropoda" rank="species" species="ledoyeri" status="sp. nov.">
<emphasis id="B9100F16D56AFFD6FD78FCA2E967FCCC" bold="true" box="[648,861,857,881]" italics="true" pageId="10" pageNumber="310">Cymadusa ledoyeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223B26DD56AFFD6FC96FCA1E99FFCCC" box="[870,933,858,881]" pageId="10" pageNumber="310" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BDBD304D56AFFD6FD3EFC82E95AFC2C" blockId="10.[313,996,857,945]" box="[718,864,888,913]" pageId="10" pageNumber="310">(figures 57)</paragraph>
</subSubSection>
<subSubSection id="C37E808FD56AFFD6FEC9FC60E9DEFC0D" box="[313,996,922,945]" pageId="10" pageNumber="310" type="reference_group">
<paragraph id="8BDBD304D56AFFD6FEC9FC60E9DEFC0D" blockId="10.[313,996,857,945]" box="[313,996,922,945]" pageId="10" pageNumber="310">
<treatmentCitationGroup id="AB74F42AD56AFFD6FEC9FC60E9DEFC0D" box="[313,996,922,945]" pageId="10" pageNumber="310">
<treatmentCitation id="0AC5F515D56AFFD6FEC9FC60E8DCFC0C" author="LEDOYER, M." box="[313,742,922,945]" page="135" pageId="10" pageNumber="310" year="1982">
<taxonomicName id="4C64A887D56AFFD6FEC9FC60E8DCFC0C" ID-CoL="335PS" authority=": Ledoyer, 1982: 130 - 135" authorityName=": Ledoyer" authorityPageNumber="130 - 135" authorityYear="1982" box="[313,742,922,945]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="310" phylum="Arthropoda" rank="species" species="filosa">
<emphasis id="B9100F16D56AFFD6FEC9FC60EBE4FC0D" box="[313,478,923,944]" italics="true" pageId="10" pageNumber="310">Cymadusa filosa</emphasis>
:
<bibRefCitation id="EFF5AEF5D56AFFD6FE1DFC60E8DCFC0C" author="LEDOYER, M." box="[493,742,922,945]" pageId="10" pageNumber="310" pagination="1 - 598" refId="ref11601" refString="LEDOYER, M., 1982, Crustaces amphipodes gammariens familles des Acanthonotozomatidae a Gammaridae, Faune de Madagascar, 59 (1), 1 - 598." type="journal article" year="1982">Ledoyer, 1982: 130135</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figures 4446 (form B).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37E808FD56AFFD1FE91FC36E9C2F975" lastPageId="13" lastPageNumber="313" pageId="10" pageNumber="310" type="materials_examined">
<paragraph id="8BDBD304D56AFFD6FE91FC36E810FBDE" blockId="10.[313,1269,972,1315]" pageId="10" pageNumber="310">
<emphasis id="B9100F16D56AFFD6FE91FC36E83EFC59" box="[353,516,972,996]" italics="true" pageId="10" pageNumber="310">Type material.</emphasis>
<materialsCitation id="3B0CD959D56AFFD6FDFAFC37EE2FFBB9" collectorName="Caye-Thomassin." country="Madagascar" location="Madagascar" pageId="10" pageNumber="310" specimenCount="1" typeStatus="holotype">
<typeStatus id="54DF6DA6D56AFFD6FDFAFC37E845FC59" box="[522,639,972,996]" pageId="10" pageNumber="310" type="holotype">HOLOTYPE</typeStatus>
: male,
<quantity id="4C9C7EE1D56AFFD6FD21FC37E926FC5E" box="[721,796,972,996]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="310" unit="mm" value="13.0">13 mm</quantity>
length, MNHN-AM3297 (Paris), Tuléar,
<collectingCountry id="F3739394D56AFFD6FEC9FC17EBFDFBB9" box="[313,455,1004,1028]" name="Madagascar" pageId="10" pageNumber="310">Madagascar</collectingCountry>
, 23°12∞S, 43°24.6∞E, collected by
<collectorName id="2691B6D2D56AFFD6FCBFFC17EE2FFBB9" box="[847,1045,1004,1028]" pageId="10" pageNumber="310">Caye-Thomassin.</collectorName>
</materialsCitation>
<typeStatus id="54DF6DA6D56AFFD6FBEFFC17EEA2FBB9" box="[1055,1176,1004,1028]" pageId="10" pageNumber="310" type="paratype">PARATYPES</typeStatus>
: female (one specimen), unknown length, MNHN-AM3300 (Paris), collection data as for holotype;
<materialsCitation id="3B0CD959D56AFFD6FE5CFBD0E81CFBDE" collectorName="Caye-Thomassin." country="Madagascar" location="Madagascar" pageId="10" pageNumber="310" specimenCount="1" typeStatus="paratype">males (two specimens), MNHN-AM3298 and AM3299 (Paris), collection data as for holotype</materialsCitation>
.
</paragraph>
<paragraph id="8BDBD304D56AFFD6FE91FB97E8CBFB3E" blockId="10.[313,1269,972,1315]" box="[353,753,1131,1155]" pageId="10" pageNumber="310">
<materialsCitation id="3B0CD959D56AFFD6FE91FB97E8D7FB3E" box="[353,749,1131,1155]" country="Madagascar" location="Tulear" pageId="10" pageNumber="310" specimenCount="1" typeStatus="holotype">
<emphasis id="B9100F16D56AFFD6FE91FB97EBC1FB3E" box="[353,507,1131,1155]" italics="true" pageId="10" pageNumber="310">
<typeStatus id="54DF6DA6D56AFFD6FE91FB97EBA3FB3E" box="[353,409,1132,1155]" pageId="10" pageNumber="310">Type</typeStatus>
locality.
</emphasis>
<location id="8EBB85DFD56AFFD6FDF3FB90E86AFB3E" LSID="urn:lsid:plazi:treatment:03CD6212D56AFFD1FD78FCA2E9C2F975:8EBB85DFD56AFFD6FDF3FB90E86AFB3E" box="[515,592,1131,1155]" country="Madagascar" name="Tulear" pageId="10" pageNumber="310">Tulear</location>
,
<collectingCountry id="F3739394D56AFFD6FDAEFB90E8D7FB3E" box="[606,749,1131,1155]" name="Madagascar" pageId="10" pageNumber="310">Madagascar</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDBD304D56AFFD6FE91FB77EE95FA9E" blockId="10.[313,1269,972,1315]" pageId="10" pageNumber="310">
<emphasis id="B9100F16D56AFFD6FE91FB77EBEFFB1E" box="[353,469,1164,1187]" italics="true" pageId="10" pageNumber="310">Diagnosis.</emphasis>
Antenna 1 with a two-articulate accessory flagellum. Antenna 2 not documented. Gnathopod 1 with poorly setose margins; merus slightly produced to form a small, subacute anteroventral lobe; carpus subtriangular. Adult male gnathopod 2 with poorly setose margins; propodal palm defined by large, rounded posterodistal tooth, mid-palmar tooth absent; dactylus longer than the palm.
</paragraph>
<paragraph id="8BDBD304D56AFFD6FEC9FAA9EB82FAD4" blockId="10.[313,1268,1362,1736]" box="[313,440,1362,1385]" pageId="10" pageNumber="310">
<emphasis id="B9100F16D56AFFD6FEC9FAA9EB82FAD4" box="[313,440,1362,1385]" italics="true" pageId="10" pageNumber="310">Description</emphasis>
</paragraph>
<paragraph id="8BDBD304D56AFFD6FE91FA8AE960FA34" blockId="10.[313,1268,1362,1736]" box="[353,858,1393,1417]" pageId="10" pageNumber="310">
Based on the
<typeStatus id="54DF6DA6D56AFFD6FE0FFA8AE85FFA34" box="[511,613,1393,1417]" pageId="10" pageNumber="310" type="holotype">holotype</typeStatus>
, male,
<quantity id="4C9C7EE1D56AFFD6FD49FA8AE93EFA35" box="[697,772,1393,1417]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="310" unit="mm" value="13.0">13 mm</quantity>
length.
</paragraph>
<paragraph id="8BDBD304D56AFFD6FE91FA6AEE5FF9DA" blockId="10.[313,1268,1362,1736]" pageId="10" pageNumber="310">
<emphasis id="B9100F16D56AFFD6FE91FA6AE825FA14" box="[353,543,1425,1449]" italics="true" pageId="10" pageNumber="310">Head. Antenna 1</emphasis>
peduncular article 1 subequal to article 2, article 2 longer than article 3, article 3 shorter than article 1; primary flagellum with 50 articles; accessory flagellum with two articles. Antenna 2 not documented.
<emphasis id="B9100F16D56AFFD6FC4CFA2AEE1CFA55" box="[956,1062,1488,1512]" italics="true" pageId="10" pageNumber="310">Lower lip</emphasis>
outer plate, outer lobe longer than inner lobe.
<emphasis id="B9100F16D56AFFD6FD7CFA0BE8CFF9B5" box="[652,757,1520,1544]" italics="true" pageId="10" pageNumber="310">Mandible</emphasis>
molar setal row with 10 robust setae; palp article 1 shorter than article 2, article 2 shorter than article 3, article 3 longer than article 1, setae on posterior margin and apex.
<emphasis id="B9100F16D56AFFD6FCCFF9CBE996F9F5" box="[831,940,1584,1608]" italics="true" pageId="10" pageNumber="310">Maxilla 1</emphasis>
inner plate with four slender setae.
<emphasis id="B9100F16D56AFFD6FE71F9ABEBCAF9D5" box="[385,496,1616,1640]" italics="true" pageId="10" pageNumber="310">Maxilla 2</emphasis>
outer lobe broader than inner lobe.
<emphasis id="B9100F16D56AFFD6FC68F9ABEE2BF9D5" box="[920,1041,1616,1640]" italics="true" pageId="10" pageNumber="310">Maxilliped</emphasis>
setose.
</paragraph>
<paragraph id="8BDBD304D56AFFD7FE91F98AE9F1FAB4" blockId="10.[313,1268,1362,1736]" lastBlockId="11.[313,1269,978,1736]" lastPageId="11" lastPageNumber="311" pageId="10" pageNumber="310">
<emphasis id="B9100F16D56AFFD6FE91F98AE86AF935" box="[353,592,1648,1672]" italics="true" pageId="10" pageNumber="310">Pereon. Gnathopod 1</emphasis>
smaller than gnathopod 2; poorly setose; coxa produced forward, anterior margin straight, ventral margin with a tuft of a few simple setae on posterior corner; basis subequal in length to the coxa, without long plumose setal fringe, distal lobe present; merus produced to form a small, subacute anteroventral lobe; carpus longer than merus, longer than propodus, margins divergent at distal end, carpal lobe truncated with rounded corners; propodus ovoid; palm entire, straight, defining posterodistal tooth absent, mid-palmar tooth absent; dactylus subequal to palm length.
<emphasis id="B9100F16D56BFFD7FD93FBA9E8C9FBD7" box="[611,755,1106,1130]" italics="true" pageId="11" pageNumber="311">Gnathopod 2</emphasis>
with sparse setae; coxa ventral margin with a tuft of simple setae on posterior corner; basis longer than coxa, without setae on anterior margin, distal lobe present; merus produced to form a small, subacute anteroventral lobe; carpus subequal to merus, shorter than propodus; propodus ovoid; palm entire, straight, posterodistal tooth rounded and large, mid-palmar tooth absent; dactylus longer than palm, tapering evenly.
</paragraph>
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<paragraph id="8BDBD304D56BFFD7FEC9FCA6EEA8FC1A" blockId="11.[313,1268,860,935]" pageId="11" pageNumber="311">
MAP 1. Global distribution of
<taxonomicName id="4C64A887D56BFFD7FD7CFCA7E9F4FCCF" authority="Savigny, 1816" authorityName="Savigny" authorityYear="1816" box="[652,974,860,882]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="311" phylum="Arthropoda" rank="species" species="filosa">
<emphasis id="B9100F16D56BFFD7FD7CFCA7E909FCCC" box="[652,819,860,881]" italics="true" pageId="11" pageNumber="311">Cymadusa filosa</emphasis>
<bibRefCitation id="EFF5AEF5D56BFFD7FCCDFCA7E9F4FCCF" author="SAVIGNY, J. - C." box="[829,974,860,882]" pageId="11" pageNumber="311" pagination="39 - 117" refId="ref12134" refString="SAVIGNY, J. - C., 1816, Observations generales sur la bouche des arachnides, des crustaces et des entomostraces, Memories sur les Animaux sans Vertebres, Second Memorie, 1, 39 - 117." type="journal article" year="1816">Savigny, 1816</bibRefCitation>
</taxonomicName>
(
<emphasis id="B9100F16D56BFFD7FC10FCA6E9C2FCCF" bold="true" box="[992,1016,861,882]" pageId="11" pageNumber="311">*</emphasis>
),
<taxonomicName id="4C64A887D56BFFD7FBE5FCA7EEECFCCC" authority="Peart, 2004" authorityName="Peart" authorityYear="2004" box="[1045,1238,860,881]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="311" phylum="Arthropoda" rank="species" species="ledoyeri" status="sp. nov.">
<emphasis id="B9100F16D56BFFD7FBE5FCA7EEECFCCC" box="[1045,1238,860,881]" italics="true" pageId="11" pageNumber="311">Cymadusa ledoyeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223B26DD56BFFD7FB10FCA6EBACFC31" pageId="11" pageNumber="311" rank="species">n. sp.</taxonomicNameLabel>
(
<emphasis id="B9100F16D56BFFD7FE55FC83EB87FC30" bold="true" box="[421,445,888,909]" pageId="11" pageNumber="311"></emphasis>
),
<taxonomicName id="4C64A887D56BFFD7FE23FC8CE898FC31" authority="Peart, 2004" authorityName="Peart" authorityYear="2004" box="[467,674,887,908]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="311" phylum="Arthropoda" rank="species" species="tattersalli" status="sp. nov.">
<emphasis id="B9100F16D56BFFD7FE23FC8CE898FC31" box="[467,674,887,908]" italics="true" pageId="11" pageNumber="311">Cymadusa tattersalli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223B26DD56BFFD7FD59FC8CE8DBFC31" box="[681,737,887,908]" pageId="11" pageNumber="311" rank="species">n. sp.</taxonomicNameLabel>
(
<emphasis id="B9100F16D56BFFD7FD00FC83E932FC30" bold="true" box="[752,776,888,909]" pageId="11" pageNumber="311">*</emphasis>
),
<taxonomicName id="4C64A887D56BFFD7FCEEFC8CEEFAFC31" authority="Rabindranath, 1972" authorityName="Rabindranath" authorityYear="1972" box="[798,1216,887,909]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="311" phylum="Arthropoda" rank="species" species="imbroglio">
<emphasis id="B9100F16D56BFFD7FCEEFC8CE9D6FC31" box="[798,1004,887,908]" italics="true" pageId="11" pageNumber="311">Cymadusa imbroglio</emphasis>
<bibRefCitation id="EFF5AEF5D56BFFD7FC03FC8CEEFAFC31" author="RABINDRANATH, P." box="[1011,1216,887,909]" pageId="11" pageNumber="311" pagination="161 - 178" refId="ref11935" refString="RABINDRANATH, P., 1972, Marine Gammaridea (Crustacea: Amphipoda) from the Indian region. Family Ampithoidae, Marine Biology, 14, 161 - 178." type="journal article" year="1972">Rabindranath, 1972</bibRefCitation>
</taxonomicName>
(
<emphasis id="B9100F16D56BFFD7FB3FFC83EEDFFC30" bold="true" box="[1231,1253,888,909]" pageId="11" pageNumber="311">Δ</emphasis>
),
<taxonomicName id="4C64A887D56BFFD7FE89FC6AE8EBFC1A" authority="(Haswell, 1879)" baseAuthorityName="Haswell" baseAuthorityYear="1879" box="[377,721,913,935]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="311" phylum="Arthropoda" rank="species" species="setosa">
<emphasis id="B9100F16D56BFFD7FE89FC6AE81CFC1A" box="[377,550,913,935]" italics="true" pageId="11" pageNumber="311">Cymadusa setosa</emphasis>
(
<bibRefCitation id="EFF5AEF5D56BFFD7FDC8FC6AE8F1FC1A" author="HASWELL, W. A." box="[568,715,913,935]" pageId="11" pageNumber="311" pagination="245 - 279" refId="ref11436" refString="HASWELL, W. A., 1879, On Australian Amphipoda, Proceedings of the Linnean Society of New South Wales, 4, 245 - 279." type="journal article" year="1879">Haswell, 1879</bibRefCitation>
)
</taxonomicName>
(
<emphasis id="B9100F16D56BFFD7FD11FC69E8C3FC1A" bold="true" box="[737,761,914,935]" pageId="11" pageNumber="311">v</emphasis>
),
<taxonomicName id="4C64A887D56BFFD7FCE3FC6AEE61FC1A" authority="Imbach, 1967" authorityName="Imbach" authorityYear="1967" box="[787,1115,913,935]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="311" phylum="Arthropoda" rank="species" species="vadosa">
<emphasis id="B9100F16D56BFFD7FCE3FC6AE9FDFC1B" box="[787,967,913,934]" italics="true" pageId="11" pageNumber="311">Cymadusa vadosa</emphasis>
<bibRefCitation id="EFF5AEF5D56BFFD7FC3FFC6AEE61FC1A" author="IMBACH, M. C." box="[975,1115,913,935]" pageId="11" pageNumber="311" pagination="39 - 167" refId="ref11465" refString="IMBACH, M. C., 1967, Gammaridean Amphipoda from the South China Sea, Naga Report, 4 (1), 39 - 167." type="journal article" year="1967">Imbach, 1967</bibRefCitation>
</taxonomicName>
(
<emphasis id="B9100F16D56BFFD7FB9BFC69EEB9FC1A" bold="true" box="[1131,1155,914,935]" pageId="11" pageNumber="311">&amp;</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BDBD304D56BFFD7FE91FAEAEE04FA15" blockId="11.[313,1269,978,1736]" pageId="11" pageNumber="311">
<emphasis id="B9100F16D56BFFD7FE91FAEAEBD2FA94" box="[353,488,1297,1321]" italics="true" pageId="11" pageNumber="311">Pereopods 3</emphasis>
and
<emphasis id="B9100F16D56BFFD7FDD5FAEAE808FA94" box="[549,562,1297,1321]" italics="true" pageId="11" pageNumber="311">4</emphasis>
similar in shape; coxae with a row of small setules along the ventral margin and a tuft of longer setae on the posterior corner.
<emphasis id="B9100F16D56BFFD7FBDFFACAEE91FAF5" box="[1071,1195,1329,1353]" italics="true" pageId="11" pageNumber="311">Pereopod 5</emphasis>
coxae with a row of small setules along the ventral margin, but absent from the medial surface.
<emphasis id="B9100F16D56BFFD7FE68FA8AE828FA34" box="[408,530,1393,1417]" italics="true" pageId="11" pageNumber="311">Pereopod 6</emphasis>
basis with sparse medial setae.
<emphasis id="B9100F16D56BFFD7FC85FA8AE9CAFA35" box="[885,1008,1393,1417]" italics="true" pageId="11" pageNumber="311">Pereopod 7</emphasis>
basis ovoid.
<emphasis id="B9100F16D56BFFD7FB73FA8AEB7CFA14" italics="true" pageId="11" pageNumber="311">Pereopods 6</emphasis>
and
<emphasis id="B9100F16D56BFFD7FE73FA6AEBAAFA15" box="[387,400,1425,1448]" italics="true" pageId="11" pageNumber="311">7</emphasis>
much longer than pereopods 35; distal articles with setae.
</paragraph>
<paragraph id="8BDBD304D56BFFD7FE91FA4BE9EBF9DA" blockId="11.[313,1269,978,1736]" pageId="11" pageNumber="311">
<emphasis id="B9100F16D56BFFD7FE91FA4BE815FA74" box="[353,559,1456,1481]" italics="true" pageId="11" pageNumber="311">Pleon. Epimeron 3</emphasis>
subquadrate.
<emphasis id="B9100F16D56BFFD7FD22FA4BE901FA74" box="[722,827,1456,1481]" italics="true" pageId="11" pageNumber="311">Uropod 1</emphasis>
peduncle with nine robust setae; outer ramus longer than inner ramus.
<emphasis id="B9100F16D56BFFD7FD4CFA2BE913FA54" box="[700,809,1488,1513]" italics="true" pageId="11" pageNumber="311">Uropod 2</emphasis>
outer ramus longer than inner ramus.
<emphasis id="B9100F16D56BFFD7FEC9FA0BEB99F9B5" box="[313,419,1520,1544]" italics="true" pageId="11" pageNumber="311">Uropod 3</emphasis>
with eight distal peduncular robust setae; outer ramus shorter than inner ramus, both hooks strongly recurved, setal fringe present on dorso-lateral margin; inner ramus with both apical robust and slender setae.
<taxonomicName id="4C64A887D56BFFD7FC42F9CBE9C0F9F5" authorityName="Pearse" authorityYear="1952" box="[946,1018,1584,1608]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="11" pageNumber="311" phylum="Arthropoda" rank="genus">
<emphasis id="B9100F16D56BFFD7FC42F9CBE9C0F9F5" box="[946,1018,1584,1608]" italics="true" pageId="11" pageNumber="311">Telson</emphasis>
</taxonomicName>
truncated, apical and lateral slender setae in oblique rows, apical cusps present.
</paragraph>
<paragraph id="8BDBD304D56BFFD7FE91F98BEEF7F935" blockId="11.[313,1269,978,1736]" box="[353,1229,1648,1672]" pageId="11" pageNumber="311">
<emphasis id="B9100F16D56BFFD7FE91F98BEB82F935" box="[353,440,1648,1672]" italics="true" pageId="11" pageNumber="311">Female.</emphasis>
Based on
<typeStatus id="54DF6DA6D56BFFD7FDC3F98BE8A0F935" box="[563,666,1648,1672]" pageId="11" pageNumber="311" type="paratype">paratype</typeStatus>
, unknown length, sexually dimorphic characters.
</paragraph>
<paragraph id="8BDBD304D56BFFD0FE91F96AEEDBFA42" blockId="11.[313,1269,978,1736]" lastBlockId="12.[313,1269,1447,1535]" lastPageId="12" lastPageNumber="312" pageId="11" pageNumber="311">
<emphasis id="B9100F16D56BFFD7FE91F96AE871F915" box="[353,587,1680,1704]" italics="true" pageId="11" pageNumber="311">Pereon. Gnathopod 1</emphasis>
coxa anterior margin concave; basis shorter than the coxa; carpus subequal to propodus, margins divergent at distal end, carpal lobe rounded; propodal palm entire, convex; dactylus subequal to or slightly longer than palm.
<emphasis id="B9100F16D56CFFD0FEC9FA3CEBFFFA62" box="[313,453,1479,1503]" italics="true" pageId="12" pageNumber="312">Gnathopod 2</emphasis>
weakly setose; basis shorter than coxa; propodus subquadrate; propodal palm defining posterodistal tooth subacute and small; dactylus subequal to palm.
</paragraph>
<caption id="DF1B838CD56CFFD0FEC9FAE4E9E1FAF2" ID-DOI="http://doi.org/10.5281/zenodo.4653580" ID-Zenodo-Dep="4653580" httpUri="https://zenodo.org/record/4653580/files/figure.png" pageId="12" pageNumber="312" startId="12.[313,329,1311,1332]" targetBox="[316,1266,185,1281]" targetPageId="12">
<paragraph id="8BDBD304D56CFFD0FEC9FAE4E9E1FAF2" blockId="12.[313,1268,1310,1360]" pageId="12" pageNumber="312">
FIG. 5.
<taxonomicName id="4C64A887D56CFFD0FE65FAE4E86CFA89" authority="Peart, 2004" authorityName="Peart" authorityYear="2004" box="[405,598,1311,1332]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="312" phylum="Arthropoda" rank="species" species="ledoyeri" status="sp. nov.">
<emphasis id="B9100F16D56CFFD0FE65FAE4E86CFA89" box="[405,598,1311,1332]" italics="true" pageId="12" pageNumber="312">Cymadusa ledoyeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223B26DD56CFFD0FDAFFAE4E8AAFA89" box="[607,656,1311,1332]" pageId="12" pageNumber="312" rank="species">n.sp.</taxonomicNameLabel>
Holotype male, 13 mm, MNHN-AM3297 (Paris). Tuléar, Madagascar. Scales represent 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BDBD304D56CFFD0FEC9F9CBEBA6F9F5" blockId="12.[313,1269,1584,1736]" box="[313,412,1584,1608]" pageId="12" pageNumber="312">
<emphasis id="B9100F16D56CFFD0FEC9F9CBEBA6F9F5" box="[313,412,1584,1608]" italics="true" pageId="12" pageNumber="312">Remarks</emphasis>
</paragraph>
<paragraph id="8BDBD304D56CFFD1FE91F9ABEBF0F9F5" blockId="12.[313,1269,1584,1736]" lastBlockId="13.[313,1268,1488,1736]" lastPageId="13" lastPageNumber="313" pageId="12" pageNumber="312">
<bibRefCitation id="EFF5AEF5D56CFFD0FE91F9ABE82EF9D5" author="LEDOYER, M." box="[353,532,1616,1640]" pageId="12" pageNumber="312" pagination="1 - 598" refId="ref11601" refString="LEDOYER, M., 1982, Crustaces amphipodes gammariens familles des Acanthonotozomatidae a Gammaridae, Faune de Madagascar, 59 (1), 1 - 598." type="journal article" year="1982">Ledoyer (1982)</bibRefCitation>
notes that this form is very close to the species
<taxonomicName id="4C64A887D56CFFD0FBA5F9ABE859F935" authority="Chevreux, 1901" authorityName="Chevreux" authorityYear="1901" class="Malacostraca" family="Ampithoidae" genus="Grubia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="312" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="B9100F16D56CFFD0FBA5F9ABEB9AF935" italics="true" pageId="12" pageNumber="312">Grubia microphthalma</emphasis>
<bibRefCitation id="EFF5AEF5D56CFFD0FE5BF98BE859F935" author="CHEVREUX, E." box="[427,611,1648,1672]" pageId="12" pageNumber="312" pagination="388 - 438" refId="ref10982" refString="CHEVREUX, E., 1901, Crustaces amphipodes mission scientifique de M. Ch. Alluad aux Iles Sechelles (Mars, Avril, Mai, 1892), Memoirs de la Societe de France, 14, 388 - 438." type="journal article" year="1901">Chevreux, 1901</bibRefCitation>
</taxonomicName>
. However,
<taxonomicName id="4C64A887D56CFFD0FD1AF98BE98FF935" authorityName="Chevreux" authorityYear="1901" box="[746,949,1648,1672]" class="Malacostraca" family="Ampithoidae" genus="Grubia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="312" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="B9100F16D56CFFD0FD1AF98BE98FF935" box="[746,949,1648,1672]" italics="true" pageId="12" pageNumber="312">C. microphthalma</emphasis>
</taxonomicName>
has only been recorded as females (illustrations and specimens), making it difficult to identify.
<taxonomicName id="4C64A887D56CFFD0FB70F96BEBABF975" authority="Peart, 2004" authorityName="Peart" authorityYear="2004" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="312" phylum="Arthropoda" rank="species" species="ledoyeri" status="sp. nov.">
<emphasis id="B9100F16D56CFFD0FB70F96BEBABF975" italics="true" pageId="12" pageNumber="312">Cymadusa ledoyeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223B26DD56CFFD0FE6EF94BEBDBF97A" box="[414,481,1712,1735]" pageId="12" pageNumber="312" rank="species">n. sp.</taxonomicNameLabel>
differs from
<taxonomicName id="4C64A887D56CFFD0FD75F94BE97BF975" authorityName="Savigny" authorityYear="1816" box="[645,833,1712,1736]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="312" phylum="Arthropoda" rank="species" species="filosa">
<emphasis id="B9100F16D56CFFD0FD75F94BE97BF975" box="[645,833,1712,1736]" italics="true" pageId="12" pageNumber="312">Cymadusa filosa</emphasis>
</taxonomicName>
by having poorly setose antenna 2, gnathopods 1 and 2 (
<taxonomicName id="4C64A887D56DFFD1FDC5FA2AE8AFFA55" authorityName="Savigny" authorityYear="1816" box="[565,661,1488,1512]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="313" phylum="Arthropoda" rank="species" species="filosa">
<emphasis id="B9100F16D56DFFD1FDC5FA2AE8AFFA55" box="[565,661,1488,1512]" italics="true" pageId="13" pageNumber="313">C. filosa</emphasis>
</taxonomicName>
has densely setae antenna 2, gnathopods 1 and 2); gnathopod 2 palm is defined by a large rounded posterodistal tooth (gnathopod 2 palm defined by a small, subacute posterodistal tooth) and uropod 3 has fewer robust setae.
</paragraph>
<caption id="DF1B838CD56DFFD1FEC9FAACEEB7FA3A" ID-DOI="http://doi.org/10.5281/zenodo.4653584" ID-Zenodo-Dep="4653584" httpUri="https://zenodo.org/record/4653584/files/figure.png" pageId="13" pageNumber="313" startId="13.[313,329,1367,1388]" targetBox="[317,1266,182,1335]" targetPageId="13">
<paragraph id="8BDBD304D56DFFD1FEC9FAACEEB7FA3A" blockId="13.[313,1268,1367,1416]" pageId="13" pageNumber="313">
FIG. 6.
<taxonomicName id="4C64A887D56DFFD1FE65FAACE869FAD1" authority="Peart, 2004" authorityName="Peart" authorityYear="2004" box="[405,595,1367,1388]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="313" phylum="Arthropoda" rank="species" species="ledoyeri" status="sp. nov.">
<emphasis id="B9100F16D56DFFD1FE65FAACE869FAD1" box="[405,595,1367,1388]" italics="true" pageId="13" pageNumber="313">Cymadusa ledoyeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223B26DD56DFFD1FDA9FAACE8ABFAD1" box="[601,657,1367,1388]" pageId="13" pageNumber="313" rank="species">n. sp.</taxonomicNameLabel>
Holotype male, 13 mm, MNHN-AM3297 (Paris). Paratype female, MNHN-AM3300, Tuléar, Madagascar. Scales represent 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BDBD304D56DFFD1FE91F9ABE91BF935" blockId="13.[313,1268,1488,1736]" pageId="13" pageNumber="313">
<emphasis id="B9100F16D56DFFD1FE91F9ABEBDBF9D5" box="[353,481,1616,1640]" italics="true" pageId="13" pageNumber="313">Etymology.</emphasis>
This species is named for Michel Ledoyer who contributed so much to the systematics of this group of species.
</paragraph>
<paragraph id="8BDBD304D56DFFD1FE91F96BE9CAF915" blockId="13.[313,1268,1488,1736]" box="[353,1008,1680,1704]" pageId="13" pageNumber="313">
<emphasis id="B9100F16D56DFFD1FE91F96BEB85F915" box="[353,447,1680,1704]" italics="true" pageId="13" pageNumber="313">Habitat.</emphasis>
These specimens were collected from seagrasses.
</paragraph>
<paragraph id="8BDBD304D56DFFD1FE91F94BE9C2F975" blockId="13.[313,1268,1488,1736]" box="[353,1016,1712,1736]" pageId="13" pageNumber="313">
<emphasis id="B9100F16D56DFFD1FE91F94BEBD4F975" box="[353,494,1712,1736]" italics="true" pageId="13" pageNumber="313">Distribution.</emphasis>
Tuléar,
<collectingCountry id="F3739394D56DFFD1FDA0F94BE8E4F975" box="[592,734,1712,1736]" name="Madagascar" pageId="13" pageNumber="313">Madagascar</collectingCountry>
, Indian Ocean (
<figureCitation id="135FCF81D56DFFD1FC68F94BE9DFF975" box="[920,997,1712,1736]" captionStart="MAP" captionStartId="11.[313,337,861,882]" captionTargetBox="[345,1236,179,831]" captionTargetId="figure-8@11.[343,1239,177,834]" captionTargetPageId="11" captionText="MAP 1. Global distribution of Cymadusa filosa Savigny, 1816 (*), Cymadusa ledoyeri n. sp. (•), Cymadusa tattersalli n. sp. (*), Cymadusa imbroglio Rabindranath, 1972 (Δ), Cymadusa setosa (Haswell, 1879) (v), Cymadusa vadosa Imbach, 1967 (&amp;)." figureDoi="http://doi.org/10.5281/zenodo.4653578" httpUri="https://zenodo.org/record/4653578/files/figure.png" pageId="13" pageNumber="313">Map 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>