287 lines
36 KiB
XML
287 lines
36 KiB
XML
<document id="764468C79E9C7905253E340C47E952FE" ID-CLB-Dataset="22824" ID-DOI="10.11646/zootaxa.4764.1.1" ID-GBIF-Dataset="55f6b110-0a37-45fe-be1c-34e9a035fe0e" ID-ISSN="1175-5334" ID-Zenodo-Dep="3764923" ID-ZooBank="urn:lsid:zoobank.org:pub:53C5289A-156C-4AF1-B84A-73099A332C05" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587733789776" checkinUser="carolina" docAuthor="Van Ofwegen, Leen P." docDate="2020" docId="03BCB717231EFF95FF35825CFE12FDD2" docLanguage="en" docName="zootaxa.4764.1.1.pdf.imf" docOrigin="Zootaxa 4764 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Litophyton snakeensis Van 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="70" masterDocId="FF85CF6F2321FFD0FFA2856EFFC1FFB1" masterDocTitle="The genus Litophyton Forskål, 1775 (Octocorallia: Alcyonacea: Nephtheidae) from Australia" masterLastPageNumber="131" masterPageNumber="1" pageNumber="64" updateTime="1698837010817" updateUser="ExternalLinkService">
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<mods:mods id="6FBBEB2192C1606967076B0099A85FFB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B221253581D5D20DE07A2A4769FD59B8">
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<mods:title id="F1DCDBBABFD1B2E4BB2B32120A552C61">The genus Litophyton Forskål, 1775 (Octocorallia: Alcyonacea: Nephtheidae) from Australia</mods:title>
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</mods:titleInfo>
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<mods:name id="533A35FF674D7760CA44B94355D5B767" type="personal">
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<mods:role id="13231FBA4737371AF77767C28202F5F7">
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<mods:roleTerm id="B9778BECF564746E83AD8209EB7E2418">Author</mods:roleTerm>
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<mods:namePart id="6A383DE90B89C402CF3D34C9B7F8AB09">Van Ofwegen, Leen P.</mods:namePart>
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<mods:typeOfResource id="CE4E6966A40EAE6596234C860F666F13">text</mods:typeOfResource>
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<mods:titleInfo id="C062E7D765846699B05A13D86DE9F63F">
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<mods:title id="04BA25D9EBA5CBAA72FC92111535AFFC">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3300793D3F0F19461923C47E57385999">
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<mods:date id="B344D6899C3A64634B0109354F7682DE">2020</mods:date>
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<mods:detail id="93F93382E240C5ABD9E51C82AEFE4A5D" type="pubDate">
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<mods:number id="68C50434E708DA402EC58814227231C7">2020-04-16</mods:number>
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</mods:detail>
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<mods:detail id="D4CEDE964BA2C093ADA336B044B3A6C8" type="volume">
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<mods:number id="647D7FC923697FCA9976C6B5B15B72F6">4764</mods:number>
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</mods:detail>
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<mods:detail id="042CFE4EABDC723E019889CC29A45382" type="issue">
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<mods:number id="F2464CC56662FE07A6F4BC86C5758FBD">1</mods:number>
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<mods:start id="7AF7BB97D8D3155A5B2272759402BA3A">1</mods:start>
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<mods:end id="DE5F7C010004274E80E131244BB8609C">131</mods:end>
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<mods:classification id="A58C26483C9B9446B35CD7982E75933D">journal article</mods:classification>
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<mods:identifier id="667EDEA5C2623D21C15503EF3EC2421B" type="CLB-Dataset">22824</mods:identifier>
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<mods:identifier id="6634686645B3C5162EF3414F91CB3153" type="DOI">10.11646/zootaxa.4764.1.1</mods:identifier>
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<mods:identifier id="3D03C556C3EEAA1D7885475258D3E87B" type="GBIF-Dataset">55f6b110-0a37-45fe-be1c-34e9a035fe0e</mods:identifier>
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<mods:identifier id="18236B39AC3AD368BBC47FED8A2139EE" type="ISSN">1175-5334</mods:identifier>
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<mods:identifier id="5A6A75356ECA8D1B8D2484BAEC7B6923" type="Zenodo-Dep">3764923</mods:identifier>
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<mods:identifier id="FF415F341C78EA9A33E4B0E7DF75DDB8" type="ZooBank">urn:lsid:zoobank.org:pub:53C5289A-156C-4AF1-B84A-73099A332C05</mods:identifier>
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<treatment id="03BCB717231EFF95FF35825CFE12FDD2" ID-DOI="http://doi.org/10.5281/zenodo.3803546" ID-GBIF-Taxon="163737713" ID-Zenodo-Dep="3803546" LSID="urn:lsid:plazi:treatment:03BCB717231EFF95FF35825CFE12FDD2" httpUri="http://treatment.plazi.org/id/03BCB717231EFF95FF35825CFE12FDD2" lastPageId="69" lastPageNumber="70" pageId="63" pageNumber="64">
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<subSubSection id="C30F558A231EFFEFFF35825CFDC1F8FD" box="[151,512,1842,1868]" pageId="63" pageNumber="64" type="nomenclature">
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<paragraph id="8BAA0601231EFFEFFF35825CFDC1F8FD" blockId="63.[151,512,1842,1904]" box="[151,512,1842,1868]" pageId="63" pageNumber="64">
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<taxonomicName id="4C157D82231EFFEFFF35825CFE5AF8FD" ID-CoL="866NC" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[151,411,1842,1868]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="63" pageNumber="64" phylum="Cnidaria" rank="species" species="snakeensis" status="sp. nov.">
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<emphasis id="B961DA13231EFFEFFF35825CFE5AF8FD" box="[151,411,1842,1868]" italics="true" pageId="63" pageNumber="64">Litophyton snakeensis</emphasis>
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</taxonomicName>
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<emphasis id="B961DA13231EFFEFFE00825CFDC1F8FD" bold="true" box="[418,512,1842,1868]" pageId="63" pageNumber="64">
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<taxonomicNameLabel id="A2526768231EFFEFFE00825CFDC1F8FD" box="[418,512,1842,1868]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F558A231EFFEFFF358238FE51F8C1" box="[151,400,1877,1904]" pageId="63" pageNumber="64" type="description">
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<paragraph id="8BAA0601231EFFEFFF358238FE51F8C1" blockId="63.[151,512,1842,1904]" box="[151,400,1877,1904]" pageId="63" pageNumber="64">
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(
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<figureCitation id="132E1A84231EFFEFFF3D8238FEC6F8C1" box="[159,263,1878,1904]" captionStart="FIGURE 49" captionStartId="64.[151,250,1967,1992]" captionTargetBox="[203,1385,273,1941]" captionTargetId="figure@64.[203,1385,273,1941]" captionTargetPageId="64" captionText="FIGURE 49. Litophyton type specimens: a, L. simplex n. sp., holotype NTM C3949; b, L. snakeensis n. sp., holotype NTM C10439; c, L. squamatum n. sp., holotype NTM C 4136. Scale at c 1 cm, a-b, natural size." figureDoi="http://doi.org/10.5281/zenodo.3765027" httpUri="https://zenodo.org/record/3765027/files/figure.png" pageId="63" pageNumber="64">Figs. 49b</figureCitation>
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,
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<figureCitation id="132E1A84231EFFEFFEB18238FEFDF8C1" box="[275,316,1878,1904]" captionStart="FIGURE 42" captionStartId="55.[151,250,1910,1935]" captionTargetBox="[187,1390,185,1880]" captionTargetId="figure@55.[178,1409,181,1884]" captionTargetPageId="55" captionText="FIGURE 42. Lateral views polyp armature of Litophyton species: a, L. brewerensis n. sp., holotype NTM C4050; b, L. legiopolypum (Verseveldt & Alderslade, 1982) holotype AMG 14806; c, L. oxleyensis n. sp., holotype NTM C4669; d, L. simplex n. sp., holotype NTM C3949; e, L. snakeensis n. sp., holotype NTM C10439; f, L. squamatum n. sp., holotype NTM C 4136; g-h, L. compactum (Verseveldt, 1966) holotype RMNH 2236 (after Verseveldt; magnification was not indicated by him)." figureDoi="http://doi.org/10.5281/zenodo.3765013" httpUri="https://zenodo.org/record/3765013/files/figure.png" pageId="63" pageNumber="64">42e</figureCitation>
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,
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<figureCitation id="132E1A84231EFFEFFEE5823BFE49F8C1" box="[327,392,1877,1904]" captionStart-0="FIGURE 52" captionStart-1="FIGURE 53" captionStartId-0="67.[151,248,1945,1970]" captionStartId-1="68.[151,248,1760,1785]" captionTargetBox-0="[178,1409,181,1919]" captionTargetBox-1="[151,1435,181,1734]" captionTargetId-0="figure@67.[178,1409,181,1919]" captionTargetId-1="figure@68.[151,1435,181,1734]" captionTargetPageId-0="67" captionTargetPageId-1="68" captionText-0="FIGURE 52. Litophyton snakeensis n. sp., holotype NTM C10439; a, abaxial and lateral point spindles; b-c, adaxial, tentacle, and polyp stalk sclerites; d, supporting bundle spindles; e, sclerites of surface layer top of stalk. Scale at a also applies to d-e." captionText-1="FIGURE 53. Litophyton snakeensis n. sp., holotype NTM C10439; a-b, sclerites of surface layer base of stalk; c-d, spindles of interior base of stalk; e, tuberculation of interior base stalk spindle. Scale at b also applies to d." figureDoi-0="http://doi.org/10.5281/zenodo.3765033" figureDoi-1="http://doi.org/10.5281/zenodo.3765035" httpUri-0="https://zenodo.org/record/3765033/files/figure.png" httpUri-1="https://zenodo.org/record/3765035/files/figure.png" pageId="63" pageNumber="64">52-53</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F558A231EFF90FF3582F3FF3AFF4A" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64" type="materials_examined">
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<paragraph id="8BAA0601231EFF90FF3582F3FF3AFF4A" blockId="63.[151,1437,1949,2013]" lastBlockId="64.[151,1437,152,251]" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64">
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<emphasis id="B961DA13231EFFEFFF3582F3FEB4F809" bold="true" box="[151,373,1949,1976]" pageId="63" pageNumber="64">Material examined</emphasis>
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.
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<materialsCitation id="3B7D0C5C231EFFEFFE2782F0FE71F86C" ID-GBIF-Occurrence="2608549231" collectingDate="1990-12-14" collectionCode="NTM" collectorName="J. Jan" elevation="11" location="Snake" municipality="Great Barrier Reef" pageId="63" pageNumber="64" specimenCode="NTM C10439" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
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<typeStatus id="54AEB8A3231EFFEFFE2782F0FE2AF809" box="[389,491,1950,1976]" pageId="63" pageNumber="64" type="holotype">Holotype</typeStatus>
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<specimenCode id="DBB3AE7A231EFFEFFE5082F1FD54F809" box="[498,661,1950,1977]" pageId="63" pageNumber="64">NTM C10439</specimenCode>
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,
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<collectingRegion id="49D1C8E3231EFFEFFD0082F0FCE7F809" box="[674,806,1950,1976]" country="Australia" name="Queensland" pageId="63" pageNumber="64">Queensland</collectingRegion>
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,
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<collectingMunicipality id="6BCE9C7B231EFFEFFC9082F0FBC3F809" box="[818,1026,1950,1976]" pageId="63" pageNumber="64">Great Barrier Reef</collectingMunicipality>
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,
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<location id="8ECA50DA231EFFEFFBAD82F0FB92F809" LSID="urn:lsid:plazi:treatment:03BCB717231EFF95FF35825CFE12FDD2:8ECA50DA231EFFEFFBAD82F0FB92F809" box="[1039,1107,1950,1976]" municipality="Great Barrier Reef" name="Snake" pageId="63" pageNumber="64" stateProvince="Queensland">Snake</location>
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reef, depth
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<quantity id="4CEDABE4231EFFEFFB7C82F0FA81F808" box="[1246,1344,1950,1977]" metricMagnitude="1" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="63" pageNumber="64" unit="m" value="11.0" valueMax="12.0" valueMin="10.0">
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<elevation id="0038E132231EFFEFFB7C82F0FA81F808" box="[1246,1344,1950,1977]" metricMagnitude="1" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="63" pageNumber="64" unit="m" value="11.0" valueMax="12.0" valueMin="10.0">10-12 m</elevation>
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</quantity>
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,
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<date id="FFAB20C1231EFFEFFAEC82F0FEEDF86D" pageId="63" pageNumber="64" value="1990-12-14">
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<collectingDate id="EFEFD929231EFFEFFAEC82F0FEEDF86D" pageId="63" pageNumber="64" value="1990-12-14">14 December 1990</collectingDate>
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</date>
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, coll.
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<collectorName id="26E063D7231EFFEFFECD82ADFE71F86C" box="[367,432,1987,2013]" pageId="63" pageNumber="64">J. Jan</collectorName>
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</materialsCitation>
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;
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<typeStatus id="54AEB8A3231EFFEFFE1E82ADFDEDF86C" box="[444,556,1987,2013]" pageId="63" pageNumber="64" type="paratype">paratypes</typeStatus>
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: NTM C3988,
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<collectingRegion id="49D1C8E3231EFFEFFD7282ACFC95F86D" box="[720,852,1986,2012]" country="Australia" name="Queensland" pageId="63" pageNumber="64">Queensland</collectingRegion>
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, Great Barrier Reef, Rib reef, depth
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<quantity id="4CEDABE4231EFFEFFB7C82AFFAFCF86C" box="[1246,1341,1985,2013]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="63" pageNumber="64" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">10-15 m</quantity>
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,
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<date id="FFAB20C1231EFF90FAEA82ACFF11FF02" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64" value="1981-05-26">26 May 1981</date>
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, coll. Z. Dinesen; NTM C4011,
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<collectingRegion id="49D1C8E32361FF90FD8D85F7FD72FF02" box="[559,691,153,179]" country="Australia" name="Queensland" pageId="64" pageNumber="65">Queensland</collectingRegion>
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, Great Barrier Reef, Davies reef, depth
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<quantity id="4CEDABE42361FF90FBC285F6FB7FFF02" box="[1120,1214,152,179]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="64" pageNumber="65" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">10-15 m</quantity>
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,
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<date id="FFAB20C12361FF90FB6B85F6FAA3FF02" box="[1225,1378,152,179]" pageId="64" pageNumber="65" value="1981-04-15">15 April 1981</date>
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, coll. Z. Dinesen; NTM C4340,
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<collectingRegion id="49D1C8E32361FF90FE6385D3FD89FF66" box="[449,584,189,215]" country="Australia" name="Queensland" pageId="64" pageNumber="65">Queensland</collectingRegion>
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, Great Barrier Reef, Myrmidon reef, depth
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<quantity id="4CEDABE42361FF90FB9B85D2FB41FF66" box="[1081,1152,188,215]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="64" pageNumber="65" unit="m" value="2.5" valueMax="5.0" valueMin="0.0">0-5 m</quantity>
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;
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<date id="FFAB20C12361FF90FB2F85D3FAFCFF66" box="[1165,1341,189,215]" pageId="64" pageNumber="65" value="1981-01-07">7 January 1981</date>
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, coll. Z. Dinesen.
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</paragraph>
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</subSubSection>
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<caption id="DF6A56892361FF90FF3582C1FBEAF85D" ID-DOI="http://doi.org/10.5281/zenodo.3765027" ID-Zenodo-Dep="3765027" httpUri="https://zenodo.org/record/3765027/files/figure.png" pageId="64" pageNumber="65" startId="64.[151,250,1967,1992]" targetBox="[203,1385,273,1941]" targetPageId="64">
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<paragraph id="8BAA06012361FF90FF3582C1FBEAF85D" blockId="64.[151,1436,1967,2028]" pageId="64" pageNumber="65">
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<emphasis id="B961DA132361FF90FF3582C1FEDAF879" bold="true" box="[151,283,1967,1992]" pageId="64" pageNumber="65">FIGURE 49</emphasis>
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.
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<taxonomicName id="4C157D822361FF90FE8B82DEFE56F879" authorityName="Forskal" authorityYear="1775" box="[297,407,1968,1992]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="64" pageNumber="65" phylum="Cnidaria" rank="genus">
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<emphasis id="B961DA132361FF90FE8B82DEFE56F879" box="[297,407,1968,1992]" italics="true" pageId="64" pageNumber="65">Litophyton</emphasis>
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</taxonomicName>
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type specimens: a,
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<taxonomicName id="4C157D822361FF90FDC682DEFD0DF879" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[612,716,1968,1992]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="64" pageNumber="65" phylum="Cnidaria" rank="species" species="simplex" status="sp. nov.">
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<emphasis id="B961DA132361FF90FDC682DEFDB3F876" box="[612,626,1968,1991]" italics="true" pageId="64" pageNumber="65">L</emphasis>
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.
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<emphasis id="B961DA132361FF90FD2282DEFD0DF879" box="[640,716,1968,1992]" italics="true" pageId="64" pageNumber="65">simplex</emphasis>
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</taxonomicName>
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<emphasis id="B961DA132361FF90FD7182DEFCD2F879" bold="true" box="[723,787,1968,1992]" pageId="64" pageNumber="65">
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<taxonomicNameLabel id="A25267682361FF90FD7182DEFCCFF879" box="[723,782,1968,1992]" pageId="64" pageNumber="65" rank="species">n. sp.</taxonomicNameLabel>
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,
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</emphasis>
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holotype NTM C3949; b,
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<taxonomicName id="4C157D822361FF90FB8882DEFB73F879" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[1066,1202,1968,1992]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="64" pageNumber="65" phylum="Cnidaria" rank="species" species="snakeensis" status="sp. nov.">
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<emphasis id="B961DA132361FF90FB8882DEFB73F879" box="[1066,1202,1968,1992]" italics="true" pageId="64" pageNumber="65">L. snakeensis</emphasis>
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</taxonomicName>
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<emphasis id="B961DA132361FF90FB1882DEFB3AF879" bold="true" box="[1210,1275,1968,1992]" pageId="64" pageNumber="65">
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<taxonomicNameLabel id="A25267682361FF90FB1882DEFB37F879" box="[1210,1270,1968,1992]" pageId="64" pageNumber="65" rank="species">n. sp.</taxonomicNameLabel>
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,
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</emphasis>
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holotype NTM C10439; c,
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<taxonomicName id="4C157D822361FF90FEAF82BAFE5DF85A" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[269,412,2004,2027]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="64" pageNumber="65" phylum="Cnidaria" rank="species" species="squamatum" status="sp. nov.">
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<emphasis id="B961DA132361FF90FEAF82BAFE5DF85A" box="[269,412,2004,2027]" italics="true" pageId="64" pageNumber="65">L. squamatum</emphasis>
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</taxonomicName>
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<emphasis id="B961DA132361FF90FE0182BAFE23F85D" bold="true" box="[419,482,2004,2028]" pageId="64" pageNumber="65">
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<taxonomicNameLabel id="A25267682361FF90FE0182BAFE1CF85D" box="[419,477,2004,2028]" pageId="64" pageNumber="65" rank="species">n. sp.</taxonomicNameLabel>
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,
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</emphasis>
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holotype NTM C 4136. Scale at c 1 cm, a-b, natural size.
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</paragraph>
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</caption>
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<caption id="DF6A56892360FF91FF358218FD60F84A" ID-DOI="http://doi.org/10.5281/zenodo.3765029" ID-Zenodo-Dep="3765029" httpUri="https://zenodo.org/record/3765029/files/figure.png" pageId="65" pageNumber="66" startId="65.[151,250,1910,1935]" targetBox="[190,1397,181,1884]" targetPageId="65">
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<paragraph id="8BAA06012360FF91FF358218FD60F84A" blockId="65.[151,1437,1910,2043]" pageId="65" pageNumber="66">
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<emphasis id="B961DA132360FF91FF358218FED8F83E" bold="true" box="[151,281,1910,1935]" pageId="65" pageNumber="66">FIGURE 50</emphasis>
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|
.
|
|
<taxonomicName id="4C157D822360FF91FE858219FE26F83E" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[295,487,1911,1935]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="65" pageNumber="66" phylum="Cnidaria" rank="species" species="simplex" status="sp. nov.">
|
|
<emphasis id="B961DA132360FF91FE858219FE26F83E" box="[295,487,1911,1935]" italics="true" pageId="65" pageNumber="66">Litophyton simplex</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="B961DA132360FF91FE4F8219FDEDF83E" bold="true" box="[493,556,1911,1935]" pageId="65" pageNumber="66">
|
|
<taxonomicNameLabel id="A25267682360FF91FE4F8219FDE6F83E" box="[493,551,1911,1935]" pageId="65" pageNumber="66" rank="species">n. sp.</taxonomicNameLabel>
|
|
,
|
|
</emphasis>
|
|
holotype NTM C3949; a, abaxial point spindles; b-c, lateral, adaxial point, tentacle, and polyp stalk sclerites; d, supporting bundle spindle, partly; e-f, sclerites of surface layer top of stalk; g-h, sclerites of surface layer base of stalk; i-j, spindles of interior base of stalk; k, tuberculation of interior base stalk spindle. Scale at a also applies to d, e, h, j; scale at i also applies to f; scale at b also applies to g.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="DF6A56892363FF92FF358218FEFAF84B" ID-DOI="http://doi.org/10.5281/zenodo.3765031" ID-Zenodo-Dep="3765031" httpUri="https://zenodo.org/record/3765031/files/figure.png" pageId="66" pageNumber="67" startId="66.[151,248,1910,1935]" targetBox="[194,1393,184,1879]" targetPageId="66">
|
|
<paragraph id="8BAA06012363FF92FF358218FEFAF84B" blockId="66.[151,1437,1910,2043]" pageId="66" pageNumber="67">
|
|
<emphasis id="B961DA132363FF92FF358218FED7F83E" bold="true" box="[151,278,1910,1935]" pageId="66" pageNumber="67">FIGURE 51</emphasis>
|
|
.
|
|
<taxonomicName id="4C157D822363FF92FE838219FD4EF83E" authority="(Roxas, 1933)" baseAuthorityName="Roxas" baseAuthorityYear="1933" box="[289,655,1911,1935]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="66" pageNumber="67" phylum="Cnidaria" rank="species" species="nigrescens">
|
|
<emphasis id="B961DA132363FF92FE838219FE3FF83E" box="[289,510,1911,1935]" italics="true" pageId="66" pageNumber="67">Litophyton nigrescens</emphasis>
|
|
(
|
|
<bibRefCitation id="EF847BF02363FF92FDA88219FD49F83E" author="Roxas" box="[522,648,1911,1935]" firstAuthor="Roxas" pageId="66" pageNumber="67" pagination="345 - 470" refId="ref42531" refString="Roxas, H. A. (1933) Philippine Alcyonaria, II. The families Alcyoniidae and Nephthyidae. Philippine Journal of Science, 50 (4), 345 - 470." type="journal article" year="1933">Roxas, 1933</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
, holotype SMF 61; a, lateral views polyp armature; b-c, abaxial point spindles; d, tentacle scales; e, lateral, adaxial, and polyp stalk scales; f, supporting bundle spindles, partly; g, sclerites of surface layer top of stalk; h, sclerites of surface layer base of stalk; i-j, spindles of interior base of stalk. Scale at h also applies to b-c, f-g, j; scale at d also applies to e.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="DF6A56892362FF93FF3582F7FAF8F867" ID-DOI="http://doi.org/10.5281/zenodo.3765033" ID-Zenodo-Dep="3765033" httpUri="https://zenodo.org/record/3765033/files/figure.png" pageId="67" pageNumber="68" startId="67.[151,248,1945,1970]" targetBox="[178,1409,181,1919]" targetPageId="67">
|
|
<paragraph id="8BAA06012362FF93FF3582F7FAF8F867" blockId="67.[151,1436,1945,2006]" pageId="67" pageNumber="68">
|
|
<emphasis id="B961DA132362FF93FF3582F7FED7F803" bold="true" box="[151,278,1945,1970]" pageId="67" pageNumber="68">FIGURE 52</emphasis>
|
|
.
|
|
<taxonomicName id="4C157D822362FF93FE8382F4FDC1F803" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[289,512,1946,1970]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="67" pageNumber="68" phylum="Cnidaria" rank="species" species="snakeensis" status="sp. nov.">
|
|
<emphasis id="B961DA132362FF93FE8382F4FDC1F803" box="[289,512,1946,1970]" italics="true" pageId="67" pageNumber="68">Litophyton snakeensis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="B961DA132362FF93FDA482F4FD85F803" bold="true" box="[518,580,1946,1970]" pageId="67" pageNumber="68">
|
|
<taxonomicNameLabel id="A25267682362FF93FDA482F4FDFEF803" box="[518,575,1946,1970]" pageId="67" pageNumber="68" rank="species">n. sp.</taxonomicNameLabel>
|
|
,
|
|
</emphasis>
|
|
holotype NTM C10439; a, abaxial and lateral point spindles; b-c, adaxial, tentacle, and polyp stalk sclerites; d, supporting bundle spindles; e, sclerites of surface layer top of stalk. Scale at a also applies to d-e.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="DF6A56892365FF94FF35838EFBF8F8AC" ID-DOI="http://doi.org/10.5281/zenodo.3765035" ID-Zenodo-Dep="3765035" httpUri="https://zenodo.org/record/3765035/files/figure.png" pageId="68" pageNumber="69" startId="68.[151,248,1760,1785]" targetBox="[151,1435,181,1734]" targetPageId="68">
|
|
<paragraph id="8BAA06012365FF94FF35838EFBF8F8AC" blockId="68.[151,1437,1760,1821]" pageId="68" pageNumber="69">
|
|
<emphasis id="B961DA132365FF94FF35838EFED9F948" bold="true" box="[151,280,1760,1785]" pageId="68" pageNumber="69">FIGURE 53</emphasis>
|
|
.
|
|
<taxonomicName id="4C157D822365FF94FE86838FFDC7F948" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[292,518,1761,1785]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="68" pageNumber="69" phylum="Cnidaria" rank="species" species="snakeensis" status="sp. nov.">
|
|
<emphasis id="B961DA132365FF94FE86838FFDC7F948" box="[292,518,1761,1785]" italics="true" pageId="68" pageNumber="69">Litophyton snakeensis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="B961DA132365FF94FDAF838FFD8CF948" bold="true" box="[525,589,1761,1785]" pageId="68" pageNumber="69">
|
|
<taxonomicNameLabel id="A25267682365FF94FDAF838FFD89F948" box="[525,584,1761,1785]" pageId="68" pageNumber="69" rank="species">n. sp.</taxonomicNameLabel>
|
|
,
|
|
</emphasis>
|
|
holotype NTM C10439; a-b, sclerites of surface layer base of stalk; c-d, spindles of interior base of stalk; e, tuberculation of interior base stalk spindle. Scale at b also applies to d.
|
|
</paragraph>
|
|
</caption>
|
|
<subSubSection id="C30F558A2365FF95FF658225FEE1FE62" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69" type="description">
|
|
<paragraph id="8BAA06012365FF94FF658225FBA5F838" blockId="68.[151,1437,1867,2038]" pageId="68" pageNumber="69">
|
|
<emphasis id="B961DA132365FF94FF658225FE98F8D7" bold="true" box="[199,345,1867,1894]" pageId="68" pageNumber="69">Description.</emphasis>
|
|
The
|
|
<typeStatus id="54AEB8A32365FF94FE318225FE34F8D4" box="[403,501,1867,1893]" pageId="68" pageNumber="69" type="holotype">holotype</typeStatus>
|
|
NTM C10439 is
|
|
<quantity id="4CEDABE42365FF94FD1E8222FCC0F8D7" box="[700,769,1868,1894]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="68" pageNumber="69" unit="cm" value="12.0">12 cm</quantity>
|
|
high and
|
|
<quantity id="4CEDABE42365FF94FCD38225FC0DF8D7" box="[881,972,1867,1894]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" pageId="68" pageNumber="69" unit="cm" value="12.5">12.5 cm</quantity>
|
|
wide, the stalk is
|
|
<quantity id="4CEDABE42365FF94FB3A8225FB25F8D7" box="[1176,1252,1867,1894]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="68" pageNumber="69" unit="cm" value="3.5">3.5 cm</quantity>
|
|
high (
|
|
<figureCitation id="132E1A842365FF94FA8F8225FA51F8D7" box="[1325,1424,1867,1894]" captionStart="FIGURE 49" captionStartId="64.[151,250,1967,1992]" captionTargetBox="[203,1385,273,1941]" captionTargetId="figure@64.[203,1385,273,1941]" captionTargetPageId="64" captionText="FIGURE 49. Litophyton type specimens: a, L. simplex n. sp., holotype NTM C3949; b, L. snakeensis n. sp., holotype NTM C10439; c, L. squamatum n. sp., holotype NTM C 4136. Scale at c 1 cm, a-b, natural size." figureDoi="http://doi.org/10.5281/zenodo.3765027" httpUri="https://zenodo.org/record/3765027/files/figure.png" pageId="68" pageNumber="69">Fig. 49b</figureCitation>
|
|
). The polyps are up to
|
|
<quantity id="4CEDABE42365FF94FE26821EFE29F83B" box="[388,488,1904,1930]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="68" pageNumber="69" unit="mm" value="0.8">0.80 mm</quantity>
|
|
high and
|
|
<quantity id="4CEDABE42365FF94FDF58201FD7AF83B" box="[599,699,1903,1930]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="68" pageNumber="69" unit="mm" value="0.5">0.50 mm</quantity>
|
|
wide, and grouped in conical catkins.
|
|
</paragraph>
|
|
<paragraph id="8BAA06012365FF95FF6582FAFE15FF66" blockId="68.[151,1437,1867,2038]" lastBlockId="69.[151,1437,152,611]" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69">
|
|
<emphasis id="B961DA132365FF94FF6582FAFED5F81F" bold="true" box="[199,276,1940,1966]" pageId="68" pageNumber="69">Polyps</emphasis>
|
|
. Without clear arrangement of sclerites (
|
|
<figureCitation id="132E1A842365FF94FD4682FDFC84F81F" box="[740,837,1939,1966]" captionStart="FIGURE 42" captionStartId="55.[151,250,1910,1935]" captionTargetBox="[187,1390,185,1880]" captionTargetId="figure@55.[178,1409,181,1884]" captionTargetPageId="55" captionText="FIGURE 42. Lateral views polyp armature of Litophyton species: a, L. brewerensis n. sp., holotype NTM C4050; b, L. legiopolypum (Verseveldt & Alderslade, 1982) holotype AMG 14806; c, L. oxleyensis n. sp., holotype NTM C4669; d, L. simplex n. sp., holotype NTM C3949; e, L. snakeensis n. sp., holotype NTM C10439; f, L. squamatum n. sp., holotype NTM C 4136; g-h, L. compactum (Verseveldt, 1966) holotype RMNH 2236 (after Verseveldt; magnification was not indicated by him)." figureDoi="http://doi.org/10.5281/zenodo.3765013" httpUri="https://zenodo.org/record/3765013/files/figure.png" pageId="68" pageNumber="69">Fig. 42e</figureCitation>
|
|
). Abaxial and lateral point sclerites are spindles with simple tubercles, up to
|
|
<quantity id="4CEDABE42365FF94FE3782D6FE36F863" box="[405,503,1976,2002]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="68" pageNumber="69" unit="mm" value="0.4">0.40 mm</quantity>
|
|
long (
|
|
<figureCitation id="132E1A842365FF94FD9F82D9FD5AF863" box="[573,667,1975,2002]" captionStart="FIGURE 52" captionStartId="67.[151,248,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@67.[178,1409,181,1919]" captionTargetPageId="67" captionText="FIGURE 52. Litophyton snakeensis n. sp., holotype NTM C10439; a, abaxial and lateral point spindles; b-c, adaxial, tentacle, and polyp stalk sclerites; d, supporting bundle spindles; e, sclerites of surface layer top of stalk. Scale at a also applies to d-e." figureDoi="http://doi.org/10.5281/zenodo.3765033" httpUri="https://zenodo.org/record/3765033/files/figure.png" pageId="68" pageNumber="69">Fig. 52a</figureCitation>
|
|
), adaxial scales and rods are present, which are also present in the tentacles (
|
|
<figureCitation id="132E1A842365FF94FF4782B5FEF4F847" box="[229,309,2011,2038]" captionStart="FIGURE 52" captionStartId="67.[151,248,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@67.[178,1409,181,1919]" captionTargetPageId="67" captionText="FIGURE 52. Litophyton snakeensis n. sp., holotype NTM C10439; a, abaxial and lateral point spindles; b-c, adaxial, tentacle, and polyp stalk sclerites; d, supporting bundle spindles; e, sclerites of surface layer top of stalk. Scale at a also applies to d-e." figureDoi="http://doi.org/10.5281/zenodo.3765033" httpUri="https://zenodo.org/record/3765033/files/figure.png" pageId="68" pageNumber="69">Fig. 52</figureCitation>
|
|
b-c). Several of these scales and rods have a crystalline texture. Supporting bundle with spindles, up to
|
|
<quantity id="4CEDABE42364FF95FF3585F7FF39FF02" box="[151,248,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="69" pageNumber="70" unit="mm" value="1.4">1.40 mm</quantity>
|
|
long, with simple tubercles (
|
|
<figureCitation id="132E1A842364FF95FD9285F7FD4EFF02" box="[560,655,152,179]" captionStart="FIGURE 52" captionStartId="67.[151,248,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@67.[178,1409,181,1919]" captionTargetPageId="67" captionText="FIGURE 52. Litophyton snakeensis n. sp., holotype NTM C10439; a, abaxial and lateral point spindles; b-c, adaxial, tentacle, and polyp stalk sclerites; d, supporting bundle spindles; e, sclerites of surface layer top of stalk. Scale at a also applies to d-e." figureDoi="http://doi.org/10.5281/zenodo.3765033" httpUri="https://zenodo.org/record/3765033/files/figure.png" pageId="69" pageNumber="70">Fig. 52d</figureCitation>
|
|
), and not projecting beyond the polyp head. Several polyp sclerites have an optically black inner core.
|
|
</paragraph>
|
|
<paragraph id="8BAA06012364FF95FF65858EFEB3FEAE" blockId="69.[151,1437,152,611]" pageId="69" pageNumber="70">
|
|
<emphasis id="B961DA132364FF95FF65858EFE0DFF4A" bold="true" box="[199,460,224,251]" pageId="69" pageNumber="70">Surface layer top stalk</emphasis>
|
|
. Spindles and unilaterally spinose spindles, up to
|
|
<quantity id="4CEDABE42364FF95FC44858FFB86FF4A" box="[998,1095,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="69" pageNumber="70" unit="mm" value="0.8">0.80 mm</quantity>
|
|
long, with simple and complex tubercles (
|
|
<figureCitation id="132E1A842364FF95FEAA846BFEA7FEAE" box="[264,358,260,287]" captionStart="FIGURE 52" captionStartId="67.[151,248,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@67.[178,1409,181,1919]" captionTargetPageId="67" captionText="FIGURE 52. Litophyton snakeensis n. sp., holotype NTM C10439; a, abaxial and lateral point spindles; b-c, adaxial, tentacle, and polyp stalk sclerites; d, supporting bundle spindles; e, sclerites of surface layer top of stalk. Scale at a also applies to d-e." figureDoi="http://doi.org/10.5281/zenodo.3765033" httpUri="https://zenodo.org/record/3765033/files/figure.png" pageId="69" pageNumber="70">Fig. 52e</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8BAA06012364FF95FF658446FC6AFE3A" blockId="69.[151,1437,152,611]" pageId="69" pageNumber="70">
|
|
<emphasis id="B961DA132364FF95FF658446FE18FEF2" bold="true" box="[199,473,296,323]" pageId="69" pageNumber="70">Surface layer base stalk</emphasis>
|
|
. Capstans, derivatives of capstans, spindles and unilaterally spinose spindles, the smaller sclerites with simple tubercles, the larger ones also with complex tubercles (
|
|
<figureCitation id="132E1A842364FF95FC7B8423FBE9FED6" box="[985,1064,332,359]" captionStart="FIGURE 53" captionStartId="68.[151,248,1760,1785]" captionTargetBox="[151,1435,181,1734]" captionTargetId="figure@68.[151,1435,181,1734]" captionTargetPageId="68" captionText="FIGURE 53. Litophyton snakeensis n. sp., holotype NTM C10439; a-b, sclerites of surface layer base of stalk; c-d, spindles of interior base of stalk; e, tuberculation of interior base stalk spindle. Scale at b also applies to d." figureDoi="http://doi.org/10.5281/zenodo.3765035" httpUri="https://zenodo.org/record/3765035/files/figure.png" pageId="69" pageNumber="70">Fig. 53</figureCitation>
|
|
a-b). The capstans and derivatives of capstans are
|
|
<quantity id="4CEDABE42364FF95FE9D841EFE18FE3A" box="[319,473,368,395]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="69" pageNumber="70" unit="mm" value="0.1" valueMax="0.15" valueMin="0.05">0.05-0.15 mm</quantity>
|
|
long, the spindles are up to
|
|
<quantity id="4CEDABE42364FF95FCA9841FFCADFE3A" box="[779,876,369,395]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="69" pageNumber="70" unit="mm" value="0.6">0.60 mm</quantity>
|
|
long.
|
|
</paragraph>
|
|
<paragraph id="8BAA06012364FF95FF6584FAFEE1FE62" blockId="69.[151,1437,152,611]" pageId="69" pageNumber="70">
|
|
<emphasis id="B961DA132364FF95FF6584FAFEA3FE1E" bold="true" box="[199,354,404,431]" pageId="69" pageNumber="70">Interior stalk</emphasis>
|
|
. Spindles, up to
|
|
<quantity id="4CEDABE42364FF95FDB884FAFDAFFE1E" box="[538,622,404,431]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="69" pageNumber="70" unit="mm" value="1.5">1.5 mm</quantity>
|
|
long, with simple tubercles (
|
|
<figureCitation id="132E1A842364FF95FC0984FBFBC9FE1E" box="[939,1032,404,431]" captionStart="FIGURE 53" captionStartId="68.[151,248,1760,1785]" captionTargetBox="[151,1435,181,1734]" captionTargetId="figure@68.[151,1435,181,1734]" captionTargetPageId="68" captionText="FIGURE 53. Litophyton snakeensis n. sp., holotype NTM C10439; a-b, sclerites of surface layer base of stalk; c-d, spindles of interior base of stalk; e, tuberculation of interior base stalk spindle. Scale at b also applies to d." figureDoi="http://doi.org/10.5281/zenodo.3765035" httpUri="https://zenodo.org/record/3765035/files/figure.png" pageId="69" pageNumber="70">Fig. 53e</figureCitation>
|
|
), and pointed ends; several branched (
|
|
<figureCitation id="132E1A842364FF95FF3D84D7FF2FFE62" box="[159,238,440,467]" captionStart="FIGURE 53" captionStartId="68.[151,248,1760,1785]" captionTargetBox="[151,1435,181,1734]" captionTargetId="figure@68.[151,1435,181,1734]" captionTargetPageId="68" captionText="FIGURE 53. Litophyton snakeensis n. sp., holotype NTM C10439; a-b, sclerites of surface layer base of stalk; c-d, spindles of interior base of stalk; e, tuberculation of interior base stalk spindle. Scale at b also applies to d." figureDoi="http://doi.org/10.5281/zenodo.3765035" httpUri="https://zenodo.org/record/3765035/files/figure.png" pageId="69" pageNumber="70">Fig. 53</figureCitation>
|
|
c-d).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C30F558A2364FF95FF6584B3FD54FE46" box="[199,661,477,503]" pageId="69" pageNumber="70" type="etymology">
|
|
<paragraph id="8BAA06012364FF95FF6584B3FD54FE46" blockId="69.[151,1437,152,611]" box="[199,661,477,503]" pageId="69" pageNumber="70">
|
|
<emphasis id="B961DA132364FF95FF6584B3FE84FE46" bold="true" box="[199,325,477,503]" pageId="69" pageNumber="70">Etymology</emphasis>
|
|
. Named after the
|
|
<typeStatus id="54AEB8A32364FF95FDAB84B3FDF9FE46" box="[521,568,477,503]" pageId="69" pageNumber="70">type</typeStatus>
|
|
locality.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C30F558A2364FF95FF65876EFB98FDAA" box="[199,1113,512,539]" pageId="69" pageNumber="70" type="distribution">
|
|
<paragraph id="8BAA06012364FF95FF65876EFB98FDAA" blockId="69.[151,1437,152,611]" box="[199,1113,512,539]" pageId="69" pageNumber="70">
|
|
<emphasis id="B961DA132364FF95FF65876EFE94FDAA" bold="true" box="[199,341,512,539]" pageId="69" pageNumber="70">Distribution</emphasis>
|
|
. Great Barrier Reef: Snake reef, Rib reef, Davies reef, Myrmidon reef.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C30F558A2364FF95FF65874BFE12FDD2" pageId="69" pageNumber="70" type="discussion">
|
|
<paragraph id="8BAA06012364FF95FF65874BFE12FDD2" blockId="69.[151,1437,152,611]" pageId="69" pageNumber="70">
|
|
<emphasis id="B961DA132364FF95FF65874BFEF8FD8E" bold="true" box="[199,313,549,575]" pageId="69" pageNumber="70">Remarks.</emphasis>
|
|
The big interior spindles and many unilateral spinose spindles in the surface of the base of the stalk are characteristic for the species.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |