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<document id="78DD0076B12A6859E1157CA4CD466137" ID-DOI="10.5852/ejt.2017.265" ID-GBIF-Dataset="ea6ba28f-024d-4115-a4b0-6aa0c06521fd" ID-ISSN="2118-9773" ID-Zenodo-Dep="3823163" ID-ZooBank="C289F7F5-268C-4935-A539-9A20F2F64F31" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1589314954388" checkinUser="carolina" docAuthor="Bresseel, Joachim &amp; Constant, Jérôme" docDate="2017" docId="03BC87FFFF86FF91FF70F98CFACBFEF8" docLanguage="en" docName="ejt-2017-265.pdf.imf" docOrigin="European Journal of Taxonomy 265 (265)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Diapheromeridae Kirby 1904, gen. nov." docType="key" docVersion="2" lastPageNumber="5" masterDocId="FF85FF87FF84FF95FFCDFFD9FFBCFFAC" masterDocTitle="Philippine mossy forest stick insects: first record of the genus Otraleus Günther, 1935 in the country, with four new species, and the new genus Capuyanus gen. nov. (Phasmida, Diapheromeridae, Necrosciinae)" masterLastPageNumber="31" masterPageNumber="1" pageNumber="3" updateTime="1733924272154" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="7A98E08887C1CE35921BB6242DCCEF74">Philippine mossy forest stick insects: first record of the genus Otraleus Günther, 1935 in the country, with four new species, and the new genus Capuyanus gen. nov. (Phasmida, Diapheromeridae, Necrosciinae)</mods:title>
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<mods:name id="D48CBCB776C27800A6F27AD1874C3DF3" type="personal">
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<mods:namePart id="6C02F50A255B50F08702324AC92660AC">Bresseel, Joachim</mods:namePart>
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<mods:affiliation id="FA7D7C8A19441E9DB442A870EFEC7DC0">Royal Belgian Institute of Natural Sciences, O. D. Phylogeny and Taxonomy, Entomology, Vautierstraat 29, B- 1000 Brussels, Belgium.</mods:affiliation>
<mods:nameIdentifier id="8B1B6F59C188C5BC195248A615570ED1" type="email">joachimbresseel@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F3F71A0AB4B7B627BDCCDF018C402122">Constant, Jérôme</mods:namePart>
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<mods:affiliation id="1FBEBBA0849DDB5BF5506B0B6A3A75DF">Royal Belgian Institute of Natural Sciences, O. D. Phylogeny and Taxonomy, Entomology, Vautierstraat 29, B- 1000 Brussels, Belgium.</mods:affiliation>
<mods:nameIdentifier id="43E06F9FE95C70105000D086DECD4372" type="email">jerome.constant@naturalsciences.be</mods:nameIdentifier>
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<mods:title id="3C3E6E56E9FCB529552322BD61835989">European Journal of Taxonomy</mods:title>
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<mods:date id="CE4501EE56C333CE4C84F304E7B20AB8">2017</mods:date>
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<treatment id="03BC87FFFF86FF91FF70F98CFACBFEF8" ID-DOI="http://doi.org/10.5281/zenodo.3844393" ID-GBIF-Taxon="164126462" ID-Zenodo-Dep="3844393" LSID="urn:lsid:plazi:treatment:03BC87FFFF86FF91FF70F98CFACBFEF8" httpUri="http://treatment.plazi.org/id/03BC87FFFF86FF91FF70F98CFACBFEF8" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" scope_family="Diapheromeridae" scope_genus="Otraleus" scope_order="Phasmida" scope_subFamily="Necrosciinae" scope_tribe="Necrosciini">
<subSubSection id="C30F6562FF86FF97FF70F98CFC53F9C3" box="[189,1007,1620,1647]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BAA36E9FF86FF97FF70F98CFC53F9C3" blockId="2.[189,1007,1620,1647]" box="[189,1007,1620,1647]" pageId="2" pageNumber="3">
<heading id="D0E28185FF86FF97FF70F98CFC53F9C3" bold="true" box="[189,1007,1620,1647]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B961EAFBFF86FF97FF70F98CFC53F9C3" bold="true" box="[189,1007,1620,1647]" pageId="2" pageNumber="3">
Key to the Philippine species of
<taxonomicName id="4C154D6AFF86FF97FD86F98DFCC6F9C3" ID-CoL="8NKK2" authority="Kirby, 1904" authorityName="Kirby" authorityYear="1904" box="[587,890,1620,1647]" class="Insecta" family="Diapheromeridae" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="family">
<emphasis id="B961EAFBFF86FF97FD86F98DFD0FF9C3" bold="true" box="[587,691,1620,1647]" italics="true" pageId="2" pageNumber="3">Otraleus</emphasis>
and
<emphasis id="B961EAFBFF86FF97FD3CF98CFCC6F9C3" bold="true" box="[753,890,1621,1647]" italics="true" pageId="2" pageNumber="3">Capuyanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2525780FF86FF97FC4CF98CFC53F9C3" box="[897,1007,1621,1647]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30F6562FF86FF96FF70F95AFACBFE7A" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="key">
<key id="FF8461D8FF86FF96FF70F95AFACBFE7A" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<keyStep id="30E12E4CFF86FF97FF70F95AFACAF8E1" pageId="2" pageNumber="3">
<paragraph id="8BAA36E9FF86FF97FF70F95AFACBF948" blockId="2.[189,1399,1667,1870]" pageId="2" pageNumber="3">
<keyLead id="30E495DCFF86FF97FF70F95AFACBF948" pageId="2" pageNumber="3">
1. Abdominal terga without postero-lateral lobes; females with subgenital plate strongly tapered, projecting over apex of tergum X; males with swollen and conical thornpads on anal segment (genus
<taxonomicName id="4C154D6AFF86FF97FF27F913FED2F948" authorityName="Bresseel &amp; Constant" authorityYear="2017" box="[234,366,1738,1764]" class="Insecta" family="Diapheromeridae" genus="Capuyanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAFBFF86FF97FF27F913FED2F948" box="[234,366,1738,1764]" italics="true" pageId="2" pageNumber="3">Capuyanus</emphasis>
</taxonomicName>
) .............................................................................
<taxonomicName id="4C154D6AFF86FF97FC08F910FB48F948" authority="Bresseel &amp; Constant, 2017" authorityName="Bresseel &amp; Constant" authorityYear="2017" box="[965,1268,1737,1764]" class="Insecta" family="Diapheromeridae" genus="Capuyanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="magwilangi" status="sp. nov.">
<emphasis id="B961EAFBFF86FF97FC08F910FB48F948" bold="true" box="[965,1268,1737,1764]" italics="true" pageId="2" pageNumber="3">Capuyanus magwilangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2525780FF86FF97FAC1F913FACBF948" box="[1292,1399,1738,1764]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAA36E9FF86FF97FF0CF934FACAF8E1" blockId="2.[189,1399,1667,1870]" pageId="2" pageNumber="3">
<keyLead id="30E495DCFF86FF97FF0CF934FACAF8E1" pageId="2" pageNumber="3">
Abdominal terga with postero-lateral lobes; females with subgenital plate not strongly tapered, not projecting over apex of tergum X; males without swollen and conical thornpads on lower posterior margin of anal segment (genus
<taxonomicName id="4C154D6AFF86FF97FB81F8EAFB0FF8E1" authorityName="Gunther" authorityYear="1935" box="[1100,1203,1843,1869]" class="Insecta" family="Diapheromeridae" genus="Otraleus" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAFBFF86FF97FB81F8EAFB0FF8E1" box="[1100,1203,1843,1869]" italics="true" pageId="2" pageNumber="3">Otraleus</emphasis>
</taxonomicName>
)....................2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E12E4CFF86FF97FF70F8ACFACAF81E" pageId="2" pageNumber="3">
<paragraph id="8BAA36E9FF86FF97FF70F8ACFACBF823" blockId="2.[189,1399,1908,1970]" box="[189,1399,1908,1935]" pageId="2" pageNumber="3">
<keyLead id="30E495DCFF86FF97FF70F8ACFACBF823" box="[189,1399,1908,1935]" pageId="2" pageNumber="3">
2. Elytra and alae present ......................................................................
<taxonomicName id="4C154D6AFF86FF97FBDCF8ADFAB0F823" authority="Bresseel &amp; Constant, 2017" authorityName="Bresseel &amp; Constant" authorityYear="2017" box="[1041,1292,1908,1935]" class="Insecta" family="Diapheromeridae" genus="Otraleus" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="christianae" status="sp. nov.">
<emphasis id="B961EAFBFF86FF97FBDCF8ADFAB0F823" bold="true" box="[1041,1292,1908,1935]" italics="true" pageId="2" pageNumber="3">Otraleus christianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2525780FF86FF97FAD5F8ACFACBF823" box="[1304,1399,1909,1935]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAA36E9FF86FF97FF0CF841FACAF81E" blockId="2.[189,1399,1908,1970]" box="[193,1398,1944,1970]" pageId="2" pageNumber="3">
<keyLead id="30E495DCFF86FF97FF0CF841FACAF81E" box="[193,1398,1944,1970]" pageId="2" pageNumber="3"> Elytra and alae absent...........................................................................................................3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E12E4CFF87FF96FF70FED2FACBFEE5" pageId="3" pageNumber="4">
<paragraph id="8BAA36E9FF87FF96FF70FED2FACBFE89" blockId="3.[189,1399,267,329]" box="[189,1399,267,294]" pageId="3" pageNumber="4">
<keyLead id="30E495DCFF87FF96FF70FED2FACBFE89" box="[189,1399,267,294]" pageId="3" pageNumber="4">3. Abdominal terga armed with spines or tubercles posteromedially..................................4</keyLead>
</paragraph>
<paragraph id="8BAA36E9FF87FF96FF70FEF6FACBFEE5" blockId="3.[189,1399,267,329]" box="[189,1399,302,329]" pageId="3" pageNumber="4">
<keyLead id="30E495DCFF87FF96FF70FEF6FACBFEE5" box="[189,1399,302,329]" pageId="3" pageNumber="4">
Abdominal terga smooth .........................................................................
<taxonomicName id="4C154D6AFF87FF96FB8EFEF7FABAFEE5" authority="Bresseel &amp; Constant, 2017" authorityName="Bresseel &amp; Constant" authorityYear="2017" box="[1091,1286,302,329]" class="Insecta" family="Diapheromeridae" genus="Otraleus" kingdom="Animalia" order="Phasmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="applai" status="sp. nov.">
<emphasis id="B961EAFBFF87FF96FB8EFEF7FABAFEE5" bold="true" box="[1091,1286,302,329]" italics="true" pageId="3" pageNumber="4">Otraleus applai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2525780FF87FF96FAD8FEF6FACBFEE5" box="[1301,1399,303,329]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E12E4CFF87FF96FF70FE8BFACBFE7A" pageId="3" pageNumber="4">
<paragraph id="8BAA36E9FF87FF96FF70FE8BFACBFE23" blockId="3.[189,1399,338,470]" pageId="3" pageNumber="4">
<keyLead id="30E495DCFF87FF96FF70FE8BFACBFE23" pageId="3" pageNumber="4">
4. Vertex elongated and slightly conical; meso- and metanotum with a definite, vertical, elongated blunt spine posteromedially ............................................
<taxonomicName id="4C154D6AFF87FF96FC08FEADFB5FFE23" authority="Bresseel &amp; Constant, 2017" authorityName="Bresseel &amp; Constant" authorityYear="2017" box="[965,1251,372,399]" class="Insecta" family="Diapheromeridae" genus="Otraleus" kingdom="Animalia" order="Phasmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bellemansae" status="sp. nov.">
<emphasis id="B961EAFBFF87FF96FC08FEADFB5FFE23" bold="true" box="[965,1251,372,399]" italics="true" pageId="3" pageNumber="4">Otraleus bellemansae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2525780FF87FF96FACEFEACFACBFE23" box="[1283,1399,373,399]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAA36E9FF87FF96FF0CFE40FACBFE7A" blockId="3.[189,1399,338,470]" pageId="3" pageNumber="4">
<keyLead id="30E495DCFF87FF96FF0CFE40FACBFE7A" pageId="3" pageNumber="4">
Vertex not conical; meso- and metanotum at best with a short conical spine posteromedially ... ......................................................................................................
<taxonomicName id="4C154D6AFF87FF96FC1BFE62FB59FE7A" authority="Bresseel &amp; Constant, 2017" authorityName="Bresseel &amp; Constant" authorityYear="2017" box="[982,1253,443,470]" class="Insecta" family="Diapheromeridae" genus="Otraleus" kingdom="Animalia" order="Phasmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="elizabethae" status="sp. nov.">
<emphasis id="B961EAFBFF87FF96FC1BFE62FB59FE7A" bold="true" box="[982,1253,443,470]" italics="true" pageId="3" pageNumber="4">Otraleus elizabethae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2525780FF87FF96FAC9FE65FACBFE7A" box="[1284,1399,444,470]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<caption id="DF6A6661FF87FF96FF70F874FACBF847" ID-DOI="http://doi.org/10.5281/zenodo.3823165" ID-Zenodo-Dep="3823165" httpUri="https://zenodo.org/record/3823165/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1965,1991]" targetBox="[253,1334,540,1919]" targetPageId="3">
<paragraph id="8BAA36E9FF87FF96FF70F874FACBF847" blockId="3.[189,1399,1965,2027]" pageId="3" pageNumber="4">
<emphasis id="B961EAFBFF87FF96FF70F874FEB6F86B" bold="true" box="[189,266,1965,1991]" pageId="3" pageNumber="4">Fig. 1.</emphasis>
<taxonomicName id="4C154D6AFF87FF96FEDDF874FD5EF864" authority="Gunther, 1935" authorityName="Gunther" authorityYear="1935" box="[272,738,1965,1992]" class="Insecta" family="Diapheromeridae" genus="Otraleus" kingdom="Animalia" order="Phasmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="hypsimelathrus">
<emphasis id="B961EAFBFF87FF96FEDDF874FD8DF86B" box="[272,561,1965,1991]" italics="true" pageId="3" pageNumber="4">Otraleus hypsimelathrus</emphasis>
<bibRefCitation id="EF844B18FF87FF96FDF5F874FD5EF864" author="Gunther K." box="[568,738,1965,1992]" pageId="3" pageNumber="4" pagination="1 - 29" refId="ref9157" refString="Gunther K. 1935. Die von Gerd Heinrich 1930 - 32 auf Celebes gesammelten Phasmoiden. Mitteilungen aus dem Zoologischen Museum, Berlin 21 (1): 1 - 29." type="journal article" year="1935">Günther, 1935</bibRefCitation>
</taxonomicName>
, ♂, holotype (ZMHB).
<emphasis id="B961EAFBFF87FF96FC38F874FBB6F86B" bold="true" box="[1013,1034,1965,1991]" pageId="3" pageNumber="4">A</emphasis>
. Habitus, lateral view.
<emphasis id="B961EAFBFF87FF96FADAF874FA97F86B" bold="true" box="[1303,1323,1965,1991]" pageId="3" pageNumber="4">B</emphasis>
. Apex of abdomen, lateral view.
<emphasis id="B961EAFBFF87FF96FE28F809FE46F846" bold="true" box="[485,506,2000,2026]" pageId="3" pageNumber="4">C</emphasis>
. Distribution map in Sulawesi. (Photographs © O. Conle and F. Hennemann.)
</paragraph>
</caption>
<subSubSection id="C30F6562FF80FF91FF70FED2FACBFEF8" box="[189,1399,267,340]" pageId="4" pageNumber="5" type="distribution">
<paragraph id="8BAA36E9FF80FF91FF70FED2FEEAFE89" blockId="4.[189,342,267,293]" box="[189,342,267,293]" pageId="4" pageNumber="5">
<heading id="D0E28185FF80FF91FF70FED2FEEAFE89" bold="true" box="[189,342,267,293]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B961EAFBFF80FF91FF70FED2FEEAFE89" bold="true" box="[189,342,267,293]" pageId="4" pageNumber="5">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAA36E9FF80FF91FF70FEE0FACBFEF8" blockId="4.[189,1399,313,340]" box="[189,1399,313,340]" pageId="4" pageNumber="5">
Currently known from
<collectingRegion id="49D1F80BFF80FF91FE07FEE0FD88FEFF" box="[458,564,313,339]" country="Indonesia" name="Sulawesi Utara" pageId="4" pageNumber="5">Sulawesi</collectingRegion>
(
<collectingCountry id="F3027679FF80FF91FD8EFEE0FD07FEF8" box="[579,699,313,340]" name="Indonesia" pageId="4" pageNumber="5">Indonesia</collectingCountry>
), Borneo (
<collectingCountry id="F3027679FF80FF91FCFEFEE0FC19FEFF" box="[819,933,313,339]" name="Malaysia" pageId="4" pageNumber="5">Malaysia</collectingCountry>
:
<collectingRegion id="49D1F80BFF80FF91FC62FEE0FC44FEF8" box="[943,1016,313,340]" country="Malaysia" name="Sabah" pageId="4" pageNumber="5">Sabah</collectingRegion>
) and Luzon Island (
<collectingCountry id="F3027679FF80FF91FB2DFEE0FAD7FEF8" box="[1248,1387,313,340]" name="Philippines" pageId="4" pageNumber="5">Philippines</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
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