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<document id="4D5DDAE31E407FE1A8CAF4926F17AA2A" ID-DOI="10.5281/zenodo.276689" ID-GBIF-Dataset="b6781430-d288-4cb4-b178-db6192a61dde" ID-ISSN="1175-5326" ID-Zenodo-Dep="276689" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460230557710" checkinUser="plazi" docAuthor="Enghoff, Henrik" docDate="2011" docId="03BC87EAFFBA3178FF22FF6704DEF8B5" docLanguage="en" docName="zt02753p041.pdf" docOrigin="Zootaxa 2753" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Parabolus dimorphus Carl 1909, n.comb." docType="treatment" docVersion="9" lastPageNumber="25" masterDocId="FF85FF92FFAD3160FFB5FFF00761FFF9" masterDocTitle="East African giant millipedes of the tribe Pachybolini (Diplopoda, Spirobolida, Pachybolidae)" masterLastPageNumber="41" masterPageNumber="1" pageNumber="24" updateTime="1698238933756" updateUser="plazi">
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<mods:title id="60749788BED751042D9FF06FF1FC12FD">East African giant millipedes of the tribe Pachybolini (Diplopoda, Spirobolida, Pachybolidae)</mods:title>
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<mods:namePart id="4667AB0E0E91F6E5F636434573A917F7">Enghoff, Henrik</mods:namePart>
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<mods:typeOfResource id="933B117CE331218EB15F50EA2471CDBA">text</mods:typeOfResource>
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<mods:title id="A1D0EF9A484715F3FF356C8FB8587A7C">Zootaxa</mods:title>
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<mods:date id="416BBB8F2EF4A19443D5B34F8B545411">2011</mods:date>
<mods:detail id="59524AB318588B8057AED451CC449A64" type="volume">
<mods:number id="9F1B593C0C3DC699276083B673AC070E">2753</mods:number>
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<mods:identifier id="8E1ABC558FA54879CB2279242AB225CD" type="DOI">10.5281/zenodo.276689</mods:identifier>
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<treatment id="03BC87EAFFBA3178FF22FF6704DEF8B5" ID-DOI="http://doi.org/10.5281/zenodo.5612662" ID-GBIF-Taxon="119387227" ID-Zenodo-Dep="5612662" LSID="urn:lsid:plazi:treatment:03BC87EAFFBA3178FF22FF6704DEF8B5" httpUri="http://treatment.plazi.org/id/03BC87EAFFBA3178FF22FF6704DEF8B5" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<subSubSection id="C30F6577FFBA3177FF22FF670630FF2B" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8BAA36FCFFBA3177FF22FF6705CAFF48" blockId="23.[151,683,151,210]" box="[151,683,151,177]" pageId="23" pageNumber="24">
<heading id="D0E28190FFBA3177FF22FF6705CAFF48" bold="true" box="[151,683,151,177]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<emphasis id="B961EAEEFFBA3177FF22FF6705CAFF48" bold="true" box="[151,683,151,177]" pageId="23" pageNumber="24">
<taxonomicName id="4C154D7FFFBA3177FF22FF67055BFF48" ID-CoL="4CMT6" authority="Carl, 1909" authorityName="Carl" authorityYear="1909" box="[151,570,151,177]" class="Diplopoda" family="Pachybolidae" genus="Parabolus" kingdom="Animalia" order="Spirobolida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="dimorphus" status="comb. nov.">
<emphasis id="B961EAEEFFBA3177FF22FF6706FDFF48" bold="true" box="[151,412,151,177]" italics="true" pageId="23" pageNumber="24">Parabolus dimorphus</emphasis>
(
<bibRefCitation id="EF844B0DFFBA3177FE19FF670553FF48" author="Carl" box="[428,562,151,177]" pageId="23" pageNumber="40" refString="Carl, J. (1909) Reise von Dr. J. Carl im nordlichen central-afrikanischen Seengebit. Diplopoden. Revue Suisse de Zoologie, 17, 281 - 365, pl. 6 - 8." type="journal article" year="1909">Carl, 1909</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A2525795FFBA3177FDF2FF6705CAFF48" box="[583,683,151,177]" pageId="23" pageNumber="24">n.comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAA36FCFFBA3177FF22FF4A0630FF2B" blockId="23.[151,683,151,210]" box="[151,337,186,210]" pageId="23" pageNumber="24">
(
<figureCitation id="132E2A79FFBA3177FF2AFF4A0641FF2B" box="[159,288,186,210]" captionStart-0="FIGURES 44 – 46" captionStart-1="FIGURES 47 – 50" captionStart-2="FIGURES 51 – 53" captionStartId-0="23.[151,264,1730,1753]" captionStartId-1="25.[151,264,1891,1914]" captionStartId-2="26.[151,264,1672,1695]" captionTargetBox-0="[193,1402,1066,1696]" captionTargetBox-1="[225,1359,198,1865]" captionTargetBox-2="[189,1398,183,1651]" captionTargetId-0="figure@23.[175,1411,1058,1708]" captionTargetId-1="figure@25.[223,1361,193,1868]" captionTargetId-2="figure@26.[187,1400,181,1653]" captionTargetPageId-0="23" captionTargetPageId-1="25" captionTargetPageId-2="26" captionText-0="FIGURES 44 – 46. Parabolus dimorphus, original drawings from Carl (1909), original labelling removed. 44: right posterior gonopod, anterior view, 45: left anterior gonopod + anterior gonopod sternum, anterior view, 46: anterior gonopods, posterior view. tr. a.: tracheal apodeme." captionText-1="FIGURES 47 – 50. Parabolus dimorphus. 47: male from the vicinity of Dar-es-Salaam, Brinkmann leg., gonopods (without left anterior gonopod coxa and telopodite), anterior view, 48: do., posterior view, 49: male from ” Deutsch Ost-Afrika ”, Eichelbaum leg., right posterior gonopod, anterior view, 50: do., posterior view. Scales 2 cm. acp: apical clublike process, bs: basal sclerite, cx: coxa, is: intermedite segment, k: keel, sl: serrate lobe, st: sternum, tlp: telopodite." captionText-2="FIGURES 51 – 53. Parabolus dimorphus, female from the vicinity of Dar-es-Salaam, Brinkmann leg., right vulva. 51: posteriolateral view, 52: basis of vulva with operculum; 53, apical appendix. Scales: 1 mm (51), 0.1 mm (52 – 53). ap: apical appendix, op: operculum. SEM." httpUri-0="https://zenodo.org/record/276706/files/figure.png" httpUri-1="https://zenodo.org/record/276707/files/figure.png" httpUri-2="https://zenodo.org/record/276708/files/figure.png" pageId="23" pageNumber="24">Figs 44–53</figureCitation>
,
<figureCitation id="132E2A79FFBA3177FE9EFF4A0629FF2B" box="[299,328,186,210]" captionStart="FIGURE 84" captionStartId="36.[151,249,1479,1502]" captionTargetBox="[456,1120,410,1444]" captionTargetId="figure@36.[456,1122,407,1445]" captionTargetPageId="36" captionText="FIGURE 84. Known distribution of East African Pachybolini, except Hadrobolus crassicollis and Epibolus pulchripes. dots (●): Crurifarcimen vagans, filled diamonds (♦): Hyperbolus apicomplexus, open diamonds (♢): Hyperbolus morogoroensis, filled triangle (˔): Parabolus dimorphus, inverted open triangle (): Parabolus calceus, inverted filled triangle (ˑ): Pachybolus tectus,, filled square (․): Pachybolini indet. Based on Yahoo maps." httpUri="https://zenodo.org/record/276715/files/figure.png" pageId="23" pageNumber="24">84</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F6577FFBA3177FF22FF0A0550FED6" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="8BAA36FCFFBA3177FF22FF0A068EFEE9" blockId="23.[151,561,250,303]" box="[151,495,250,272]" pageId="23" pageNumber="24">
<taxonomicName id="4C154D7FFFBA3177FF22FF0A068EFEE9" authority="Carl, 1909" authorityName="Carl" authorityYear="1909" box="[151,495,250,272]" class="Diplopoda" family="Trigoniulidae" genus="Trigoniulus" kingdom="Animalia" order="Spirobolida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="dimorphus">
<emphasis id="B961EAEEFFBA3177FF22FF0A061FFEE9" box="[151,382,250,272]" italics="true" pageId="23" pageNumber="24">Trigoniulus dimorphus</emphasis>
<bibRefCitation id="EF844B0DFFBA3177FE31FF0A068EFEE9" author="Carl" box="[388,495,250,272]" pageId="23" pageNumber="40" refString="Carl, J. (1909) Reise von Dr. J. Carl im nordlichen central-afrikanischen Seengebit. Diplopoden. Revue Suisse de Zoologie, 17, 281 - 365, pl. 6 - 8." type="journal article" year="1909">Carl, 1909</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BAA36FCFFBA3177FF22FEE90550FED6" blockId="23.[151,561,250,303]" box="[151,561,281,303]" pageId="23" pageNumber="24">
<taxonomicName id="4C154D7FFFBA3177FF22FEE90550FED6" authority="Hoffman (1965)" authorityName="Hoffman" authorityYear="1965" box="[151,561,281,303]" class="Diplopoda" family="Pachybolidae" genus="Pachybolus" kingdom="Animalia" order="Spirobolida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="dimorphus">
<emphasis id="B961EAEEFFBA3177FF22FEE9061EFED6" box="[151,383,281,303]" italics="true" pageId="23" pageNumber="24">Pachybolus dimorphus</emphasis>
: Hoffman (1965)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30F6577FFBA3177FF22FEA6030BFE20" pageId="23" pageNumber="24" type="description">
<paragraph id="8BAA36FCFFBA3177FF22FEA603D3FE97" blockId="23.[151,1436,342,1005]" box="[151,1202,342,367]" pageId="23" pageNumber="24">
<typeStatus id="54AE885EFFBA3177FF22FEA60648FE97" box="[151,297,342,366]" pageId="23" pageNumber="24" type="syntype">SYNTYPES</typeStatus>
: 2 3, 1 Ƥ
<collectingCountry id="F302766CFFBA3177FE04FEA70522FE96" box="[433,579,343,367]" name="Tanzania" pageId="23" pageNumber="24">TANZANIA</collectingCountry>
, Dar-es-Salaam,
<geoCoordinate id="EE21503BFFBA3177FCB4FEA6042EFE97" box="[769,847,342,366]" direction="south" orientation="latitude" pageId="23" pageNumber="24" precision="925" value="-6.8333335">6°50’S</geoCoordinate>
<geoCoordinate id="EE21503BFFBA3177FCE3FEA604D4FE97" box="[854,949,342,366]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="925" value="39.333332">39°20’E</geoCoordinate>
, J. Carl leg. (MHNG).
</paragraph>
<paragraph id="8BAA36FCFFBA3177FF70FE8A030BFE20" blockId="23.[151,1436,342,1005]" pageId="23" pageNumber="24">
ADDITIONAL MATERIAL: 1 Ƥ,
<specimenCount id="9D13FD75FFBA3177FDE1FE8A05ABFE6B" box="[596,714,378,402]" pageId="23" pageNumber="24" type="subadult">1 subadult</specimenCount>
Ƥ
<collectingCountry id="F302766CFFBA3177FD45FE8A04E3FE6B" box="[752,898,378,402]" name="Tanzania" pageId="23" pageNumber="24">TANZANIA</collectingCountry>
, Dar-es-Salaam (ZMUH). – 1 3 “Deutsch Ost- Afrika” 1903, F. Eichelbaum leg., R.L. Hoffman det., 1964 (ZMUH). – 1 3 “Ost-Afrika”, plantations and roads in vicinity of Dar-es-Salaam, 1929, H. Brinkmann leg, R.L. Hoffman det., 1964 (ZMUH).
</paragraph>
</subSubSection>
<subSubSection id="C30F6577FFBA3177FF72FE150609FCC2" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="8BAA36FCFFBA3177FF72FE150609FCC2" blockId="23.[151,1436,342,1005]" pageId="23" pageNumber="24">
<emphasis id="B961EAEEFFBA3177FF72FE1506A8FE04" bold="true" box="[199,457,484,509]" pageId="23" pageNumber="24">Notes to the material.</emphasis>
The
<typeStatus id="54AE885EFFBA3177FDB6FE150552FE04" box="[515,563,485,509]" pageId="23" pageNumber="24">type</typeStatus>
series (
<collectionCode id="ED04AE39FFBA3177FD3FFE140582FE05" box="[650,739,484,508]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="23" pageNumber="24">MHNG</collectionCode>
) contains two 3 and one Ƥ. Since they were all in the same container, and the gonopods of both 3 had been dissected, it was not possible to associate the two sets of gonopods (one of which lacks one of the posterior gonopods) with the two male bodies. The latter could be reconstructed based on Carl’s information about body ring number (“57 Segmente”). Each of the three individuals has now been placed in a separate container, and the two sets of gonopods in two further, separate containers. The two 3 are obviously conspecific, and given the uncertainty about which gonopods belong to which body, I have not selected a
<typeStatus id="54AE885EFFBA3177FEBBFD490619FD28" box="[270,376,697,721]" pageId="23" pageNumber="24" type="lectotype">lectotype</typeStatus>
. The female and subadult female from Dar-es-Salaam (
<collectionCode id="ED04AE39FFBA3177FC5BFD490325FD28" box="[1006,1092,697,721]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="23" pageNumber="24">ZMUH</collectionCode>
) are labelled as
<typeStatus id="54AE885EFFBA3177FAB5FD4A020AFD2B" box="[1280,1387,698,722]" pageId="23" pageNumber="24" type="paratype">paratypes</typeStatus>
and listed as “Paratypoid” by
<bibRefCitation id="EF844B0DFFBA3177FE0FFD2C050CFD0D" author="Weidner" box="[442,621,732,756]" pageId="23" pageNumber="41" refString="Weidner, H. (1960) Die entomologischen Sammlungen des zoologischen Staatsinstituts und zoologischen Museums Hamburg III. Teil Chilopoda und Progoneata. Mitteilungen des hamburgischen zoologischen Museums und Instituts, 58, 57 - 104." type="journal article" year="1960">Weidner (1960)</bibRefCitation>
, but it is uncertain if they belong to the material studied by
<bibRefCitation id="EF844B0DFFBA3177FAACFD2C02FDFD0D" author="Carl" box="[1305,1436,732,756]" pageId="23" pageNumber="40" refString="Carl, J. (1909) Reise von Dr. J. Carl im nordlichen central-afrikanischen Seengebit. Diplopoden. Revue Suisse de Zoologie, 17, 281 - 365, pl. 6 - 8." type="journal article" year="1909">Carl (1909)</bibRefCitation>
who did not indicate how many specimens he had studied; the hand-writing on the label is different from that on the type-vial label.
</paragraph>
</subSubSection>
<subSubSection id="C30F6577FFBA3177FF72FCB60697FC14" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="8BAA36FCFFBA3177FF72FCB60654FC5F" blockId="23.[151,1436,342,1005]" pageId="23" pageNumber="24">
<emphasis id="B961EAEEFFBA3177FF72FCB60620FCA6" bold="true" box="[199,321,838,863]" pageId="23" pageNumber="24">Diagnosis.</emphasis>
A species of
<taxonomicName id="4C154D7FFFBA3177FE62FCB7052AFCA6" box="[471,587,839,863]" class="Diplopoda" family="Pachybolidae" genus="Parabolus" kingdom="Animalia" order="Spirobolida" pageId="23" pageNumber="23" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B961EAEEFFBA3177FE62FCB7052AFCA6" box="[471,587,839,863]" italics="true" pageId="23" pageNumber="24">Parabolus</emphasis>
</taxonomicName>
in which the anterior gonopod coxa has no distinct distolateral shoulder and the terminal segment of the posterior gonopod has a coarsely serrated mesobasal lobe and a slender, curved, blunt apical process
</paragraph>
<paragraph id="8BAA36FCFFBA3177FF72FC410697FC14" blockId="23.[151,1436,342,1005]" pageId="23" pageNumber="24">
<emphasis id="B961EAEEFFBA3177FF72FC4106FFFC33" bold="true" box="[199,414,945,970]" pageId="23" pageNumber="24">Descriptive notes.</emphasis>
Carl’s original description is quite good. In particular, his illustrations are fine and are reproduced here as
<figureCitation id="132E2A79FFBA3177FEC4FC250693FC14" box="[369,498,981,1005]" captionStart="FIGURES 44 – 46" captionStartId="23.[151,264,1730,1753]" captionTargetBox="[193,1402,1066,1696]" captionTargetId="figure@23.[175,1411,1058,1708]" captionTargetPageId="23" captionText="FIGURES 44 – 46. Parabolus dimorphus, original drawings from Carl (1909), original labelling removed. 44: right posterior gonopod, anterior view, 45: left anterior gonopod + anterior gonopod sternum, anterior view, 46: anterior gonopods, posterior view. tr. a.: tracheal apodeme." httpUri="https://zenodo.org/record/276706/files/figure.png" pageId="23" pageNumber="24">Figs 44–46</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="DF6A6674FFBA3177FF22F93206A1F8EC" httpUri="https://zenodo.org/record/276706/files/figure.png" pageId="23" pageNumber="24" targetBox="[193,1402,1066,1696]" targetPageId="23">
<paragraph id="8BAA36FCFFBA3177FF22F93206A1F8EC" blockId="23.[151,1436,1730,1813]" pageId="23" pageNumber="24">
<emphasis id="B961EAEEFFBA3177FF22F9320637F921" bold="true" box="[151,342,1730,1753]" pageId="23" pageNumber="24">FIGURES 44–46.</emphasis>
<taxonomicName id="4C154D7FFFBA3177FEE8F932055BF921" box="[349,570,1730,1752]" class="Diplopoda" family="Pachybolidae" genus="Parabolus" kingdom="Animalia" order="Spirobolida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="dimorphus">
<emphasis id="B961EAEEFFBA3177FEE8F932055BF921" box="[349,570,1730,1752]" italics="true" pageId="23" pageNumber="24">Parabolus dimorphus</emphasis>
</taxonomicName>
, original drawings from Carl (1909), original labelling removed. 44: right posterior gonopod, anterior view, 45: left anterior gonopod + anterior gonopod sternum, anterior view, 46: anterior gonopods, posterior view. tr.a.: tracheal apodeme.
</paragraph>
</caption>
<subSubSection id="C30F6577FFBA3178FF70F8B303B5F8D1" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="description">
<paragraph id="8BAA36FCFFBA3177FF70F8B3055BF8A3" blockId="23.[197,570,1858,1883]" box="[197,570,1858,1883]" pageId="23" pageNumber="24">
As
<taxonomicName id="4C154D7FFFBA3177FF5EF8B20682F8A3" box="[235,483,1858,1882]" class="Diplopoda" family="Pachybolidae" genus="Crurifarcimen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="vagans">
<emphasis id="B961EAEEFFBA3177FF5EF8B20682F8A3" box="[235,483,1858,1882]" italics="true" pageId="23" pageNumber="24">Crurifarcimen vagans</emphasis>
</taxonomicName>
except:
</paragraph>
<paragraph id="8BAA36FCFFBA3177FF22F87B05BAF83F" blockId="23.[151,1436,1930,2026]" pageId="23" pageNumber="24">
- Adults with 56 podous rings, no apodous rings. Length ca.
<quantity id="4CED9B19FFBA3177FCEFF87A04C2F85B" box="[858,931,1930,1954]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="24" unit="cm" value="12.0">12 cm</quantity>
. Width: 11.9–12.4 mm (3) / 11.9–12.2 mm (Ƥ). Ten rows of ocelli in 5–6 horizontal series.
</paragraph>
<paragraph id="8BAA36FCFFBA3177FF22F82302EEF813" blockId="23.[151,1436,1930,2026]" box="[151,1423,2002,2026]" pageId="23" pageNumber="24">- 2+2 labral teeth, one 3 with ca. 10+10 labral setae in bight above labral teeth, other specimens without setae.</paragraph>
<paragraph id="8BAA36FCFFB53178FF22FF6804ECFF0E" blockId="24.[151,1436,151,536]" pageId="24" pageNumber="25">- Mandibles with 4 pectinate lamellae plus a variable number of teeth in a jumbled group basally; ventral margin of gnathal lobe pectinate area not concave; a sclerotized smooth-edged ridge between pectinate lamellae and mola, ridge dorsally with a row of small spines; 5 molar ridges.</paragraph>
<paragraph id="8BAA36FCFFB53178FF22FEF5027DFEE5" blockId="24.[151,1436,151,536]" box="[151,1308,260,284]" pageId="24" pageNumber="25">- Gnathohilarium with 3+3 distal stipital and 4–5+4–5 distal lingual lamellar setae in transverse row.</paragraph>
<paragraph id="8BAA36FCFFB53178FF22FED803C1FEC6" blockId="24.[151,1436,151,536]" box="[151,1184,295,319]" pageId="24" pageNumber="25">- Collum smooth, no striae, with more or less marked marginal furrow at anterior margin.</paragraph>
<paragraph id="8BAA36FCFFB53178FF22FEBD0482FE9D" blockId="24.[151,1436,151,536]" box="[151,995,332,356]" pageId="24" pageNumber="25">- Body rings: tergo-pleural suture visible on pro-, meso- and metazonite.</paragraph>
<paragraph id="8BAA36FCFFB53178FF22FE800662FE55" blockId="24.[151,1436,151,536]" pageId="24" pageNumber="25">- Terga smooth, except for fine oblique striae ventrally on mesozona and fine, dense longitudinal striae on metazona.</paragraph>
<paragraph id="8BAA36FCFFB53178FF22FE48051AFE36" blockId="24.[151,1436,151,536]" box="[151,635,439,463]" pageId="24" pageNumber="25">- Ozopores 1 diameter in front of suture.</paragraph>
<paragraph id="8BAA36FCFFB53178FF22FE2D0456FE0D" blockId="24.[151,1436,151,536]" box="[151,823,476,500]" pageId="24" pageNumber="25">
- Telson as in
<taxonomicName id="4C154D7FFFB53178FEE6FE2C0556FE0D" box="[339,567,476,500]" class="Diplopoda" family="Pachybolidae" genus="Crurifarcimen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="variety" variety="but">
<emphasis id="B961EAEEFFB53178FEE6FE2C056CFE0D" box="[339,525,476,500]" italics="true" pageId="24" pageNumber="25">Crurifarcimen v.</emphasis>
but
</taxonomicName>
dorsal profile straight.
</paragraph>
<paragraph id="8BAA36FCFFB53178FF22FDF00520FDEE" blockId="24.[151,1436,151,536]" box="[151,577,511,536]" pageId="24" pageNumber="25">- Legs. Length 63% of body width.</paragraph>
<paragraph id="8BAA36FCFFB53178FF72FDB8063DFD5E" blockId="24.[151,1437,583,1220]" pageId="24" pageNumber="25">
<emphasis id="B961EAEEFFB53178FF72FDB80683FD99" bold="true" box="[199,482,584,608]" pageId="24" pageNumber="25">Male sexual characters.</emphasis>
Leg modifications as in
<taxonomicName id="4C154D7FFFB53178FD4FFDB8040BFD99" box="[762,874,584,608]" class="Diplopoda" family="Pachybolidae" genus="Crurifarcimen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="vagans">
<emphasis id="B961EAEEFFB53178FD4FFDB8040BFD99" box="[762,874,584,608]" italics="true" pageId="24" pageNumber="25">C. vagans</emphasis>
</taxonomicName>
. The coxal lobes on legs 6–7 were not mentioned by
<bibRefCitation id="EF844B0DFFB53178FF0CFD9C065CFD7D" author="Carl" box="[185,317,620,644]" pageId="24" pageNumber="40" refString="Carl, J. (1909) Reise von Dr. J. Carl im nordlichen central-afrikanischen Seengebit. Diplopoden. Revue Suisse de Zoologie, 17, 281 - 365, pl. 6 - 8." type="journal article" year="1909">Carl (1909)</bibRefCitation>
. In the two males of the
<typeStatus id="54AE885EFFB53178FDFBFD9D051FFD7C" box="[590,638,621,645]" pageId="24" pageNumber="25">type</typeStatus>
series, these legs are badly damaged, but in the males from ZMUH, the lobes are evident.
</paragraph>
<paragraph id="8BAA36FCFFB53178FF72FD44020EFBCD" blockId="24.[151,1437,583,1220]" pageId="24" pageNumber="25">
Anterior gonopods (
<figureCitation id="132E2A79FFB53178FE16FD440545FD35" box="[419,548,692,716]" captionStart-0="FIGURES 44 – 46" captionStart-1="FIGURES 47 – 50" captionStartId-0="23.[151,264,1730,1753]" captionStartId-1="25.[151,264,1891,1914]" captionTargetBox-0="[193,1402,1066,1696]" captionTargetBox-1="[225,1359,198,1865]" captionTargetId-0="figure@23.[175,1411,1058,1708]" captionTargetId-1="figure@25.[223,1361,193,1868]" captionTargetPageId-0="23" captionTargetPageId-1="25" captionText-0="FIGURES 44 – 46. Parabolus dimorphus, original drawings from Carl (1909), original labelling removed. 44: right posterior gonopod, anterior view, 45: left anterior gonopod + anterior gonopod sternum, anterior view, 46: anterior gonopods, posterior view. tr. a.: tracheal apodeme." captionText-1="FIGURES 47 – 50. Parabolus dimorphus. 47: male from the vicinity of Dar-es-Salaam, Brinkmann leg., gonopods (without left anterior gonopod coxa and telopodite), anterior view, 48: do., posterior view, 49: male from ” Deutsch Ost-Afrika ”, Eichelbaum leg., right posterior gonopod, anterior view, 50: do., posterior view. Scales 2 cm. acp: apical clublike process, bs: basal sclerite, cx: coxa, is: intermedite segment, k: keel, sl: serrate lobe, st: sternum, tlp: telopodite." httpUri-0="https://zenodo.org/record/276706/files/figure.png" httpUri-1="https://zenodo.org/record/276707/files/figure.png" pageId="24" pageNumber="25">Figs 45–48</figureCitation>
): sternum concave on oral face, with high, broad, triangular mesal process reaching ca. 2/3 the height of the coxae; two halves of process meeting under blunt angle in midline, forming a midline ridge; tip of process with small median indentation; sternum laterally continuing as narrow band around side of gonopod, reaching well along base of coxa on aboral side, articulating with a narrow sclerite at base of telopodite (“ bridetrachéenne” of
<bibRefCitation id="EF844B0DFFB53178FEFEFCB5069CFCA5" author="Demange" box="[331,509,836,861]" pageId="24" pageNumber="40" refString="Demange, J. M. (1967) Recherches sur la segmentation du tronc des Chilopodes et des Diplopodes Chilognathes (Myriapodes). Memoires du Museum national d'Histoire naturelle Paris, nouvelle serie, serie A. Zoologie, 44, 1 - 188." type="journal article" year="1967">Demange 1967</bibRefCitation>
). Coxa in oral view kidney- to sausage-shaped, lateral margins evenly convex, mesally with very big slender process directed distad parallel to mesal margin of main part of coxa; internal surface of coxa (hidden from view in undissected specimens) with ridge at base of mesal process. Coxa extending to aboral side of gonopod where it appears as a narrow lateral band. Telopodite (
<figureCitation id="132E2A79FFB53178FC05FC5F049AFC3E" box="[944,1019,943,967]" captionStart="FIGURES 47 – 50" captionStartId="25.[151,264,1891,1914]" captionTargetBox="[225,1359,198,1865]" captionTargetId="figure@25.[223,1361,193,1868]" captionTargetPageId="25" captionText="FIGURES 47 – 50. Parabolus dimorphus. 47: male from the vicinity of Dar-es-Salaam, Brinkmann leg., gonopods (without left anterior gonopod coxa and telopodite), anterior view, 48: do., posterior view, 49: male from ” Deutsch Ost-Afrika ”, Eichelbaum leg., right posterior gonopod, anterior view, 50: do., posterior view. Scales 2 cm. acp: apical clublike process, bs: basal sclerite, cx: coxa, is: intermedite segment, k: keel, sl: serrate lobe, st: sternum, tlp: telopodite." httpUri="https://zenodo.org/record/276707/files/figure.png" pageId="24" pageNumber="25">Fig 48</figureCitation>
) extending slightly beyond coxal tip, with a distolateral incision and a shallow impression running from incision to base of low mesal process which very slightly overlaps that of opposite telopodite; tip of telopodite distally broadly rounded; oral surface of telopodite (hidden from view in undissected specimens) with strong, oblique ridge delimiting a narrow furrow.
</paragraph>
<paragraph id="8BAA36FCFFB53178FF72FBCF026BFB3D" blockId="24.[151,1437,583,1220]" pageId="24" pageNumber="25">
Posterior gonopods (
<figureCitation id="132E2A79FFB53178FE12FBCF069AFBAE" box="[423,507,1087,1111]" captionStart="FIGURES 44 – 46" captionStartId="23.[151,264,1730,1753]" captionTargetBox="[193,1402,1066,1696]" captionTargetId="figure@23.[175,1411,1058,1708]" captionTargetPageId="23" captionText="FIGURES 44 – 46. Parabolus dimorphus, original drawings from Carl (1909), original labelling removed. 44: right posterior gonopod, anterior view, 45: left anterior gonopod + anterior gonopod sternum, anterior view, 46: anterior gonopods, posterior view. tr. a.: tracheal apodeme." httpUri="https://zenodo.org/record/276706/files/figure.png" pageId="24" pageNumber="25">Figs 44</figureCitation>
,
<figureCitation id="132E2A79FFB53178FDB0FBCF052DFBAE" box="[517,588,1087,1111]" captionStart="FIGURES 47 – 50" captionStartId="25.[151,264,1891,1914]" captionTargetBox="[225,1359,198,1865]" captionTargetId="figure@25.[223,1361,193,1868]" captionTargetPageId="25" captionText="FIGURES 47 – 50. Parabolus dimorphus. 47: male from the vicinity of Dar-es-Salaam, Brinkmann leg., gonopods (without left anterior gonopod coxa and telopodite), anterior view, 48: do., posterior view, 49: male from ” Deutsch Ost-Afrika ”, Eichelbaum leg., right posterior gonopod, anterior view, 50: do., posterior view. Scales 2 cm. acp: apical clublike process, bs: basal sclerite, cx: coxa, is: intermedite segment, k: keel, sl: serrate lobe, st: sternum, tlp: telopodite." httpUri="https://zenodo.org/record/276707/files/figure.png" pageId="24" pageNumber="25">49–50</figureCitation>
)
<emphasis id="B961EAEEFFB53178FDECFBB105FAFBA1" box="[601,667,1089,1112]" italics="true" pageId="24" pageNumber="25">in situ</emphasis>
supported by ridge at base of mesal coxal process and oblique ridge on oral surface of telopodite of anterior gonopod; visible in oral and aboral view as narrow bands along mesal margin of coxa and telopodite, respectively, connected by median sternal remnant consisting of mesal membranous region flanked by a pair of rounded-triangular sclerites. Each posterior gonopod clearly divisible into four parts:
</paragraph>
<paragraph id="8BAA36FCFFB53178FF22FB040374FAF5" blockId="24.[151,1437,1268,1868]" box="[151,1045,1268,1292]" pageId="24" pageNumber="25">
1. a long, slender, almost straight
<emphasis id="B961EAEEFFB53178FD91FB04058FFAF5" box="[548,750,1268,1292]" italics="true" pageId="24" pageNumber="25">tracheal apodeme</emphasis>
, flattened towards its end,
</paragraph>
<paragraph id="8BAA36FCFFB53178FF22FAE80366FAAD" blockId="24.[151,1437,1268,1868]" pageId="24" pageNumber="25">
2. a
<emphasis id="B961EAEEFFB53178FF6FFAE8060EFAC9" box="[218,367,1304,1328]" italics="true" pageId="24" pageNumber="25">basal sclerite</emphasis>
which articulates with the tracheal apodeme, is roughly pentagonal in outline and has a distinct notch on its distal margin, next to articulation with median sternal sclerite,
</paragraph>
<paragraph id="8BAA36FCFFB53178FF22FA900443FA65" blockId="24.[151,1437,1268,1868]" pageId="24" pageNumber="25">
3. an
<emphasis id="B961EAEEFFB53178FF5CFA9006BBFA81" box="[233,474,1376,1400]" italics="true" pageId="24" pageNumber="25">intermediate segment</emphasis>
originating from lateral half of basal sclerite, parallel-sided, with extensive membranous areas, especially on mesal part of aboral surface,
</paragraph>
<paragraph id="8BAA36FCFFB53178FF22FA5806F1F960" blockId="24.[151,1437,1268,1868]" pageId="24" pageNumber="25">
4. a
<emphasis id="B961EAEEFFB53178FF69FA5806FEFA39" box="[220,415,1448,1472]" italics="true" pageId="24" pageNumber="25">terminal segment</emphasis>
separated from intermediate segment by distinct impression across lateral and aboral surface. Overall shape of terminal segment triangular (or like the profile of a hadrosaur head); with a small laterobasal, stout spine at articulation with intermediate segment, a coarsely serrate mesobasal lobe resting (
<emphasis id="B961EAEEFFB53178FAF8FA0102F3F9F1" box="[1357,1426,1521,1544]" italics="true" pageId="24" pageNumber="25">in situ</emphasis>
) on convex ridge at base of mesal coxal process and a slender, curved, apically clublike rounded process; aboral surface smooth, oral surface with a longitudinal keel running from base to tip. Efferent groove running along mesal edge of intermediate segment, then making an abrupt turn and terminating behind longitudinal keel of terminal segment.
</paragraph>
<paragraph id="8BAA36FCFFB53178FF72F95402FDF945" blockId="24.[151,1437,1268,1868]" box="[199,1436,1700,1724]" pageId="24" pageNumber="25">
<emphasis id="B961EAEEFFB53178FF72F9540562F945" bold="true" box="[199,515,1700,1724]" pageId="24" pageNumber="25">Female sexual characters.</emphasis>
Distal margin of lateral coxosternal extensions projecting as right-angled lobe.
</paragraph>
<paragraph id="8BAA36FCFFB53178FF22F93802FCF919" blockId="24.[151,1437,1268,1868]" box="[151,1437,1736,1760]" pageId="24" pageNumber="25">
Vulva (
<figureCitation id="132E2A79FFB53178FF5DF9380606F919" box="[232,359,1736,1760]" captionStart="FIGURES 51 – 53" captionStartId="26.[151,264,1672,1695]" captionTargetBox="[189,1398,183,1651]" captionTargetId="figure@26.[187,1400,181,1653]" captionTargetPageId="26" captionText="FIGURES 51 – 53. Parabolus dimorphus, female from the vicinity of Dar-es-Salaam, Brinkmann leg., right vulva. 51: posteriolateral view, 52: basis of vulva with operculum; 53, apical appendix. Scales: 1 mm (51), 0.1 mm (52 – 53). ap: apical appendix, op: operculum. SEM." httpUri="https://zenodo.org/record/276708/files/figure.png" pageId="24" pageNumber="25">Figs 51–53</figureCitation>
) less elongate than in
<taxonomicName id="4C154D7FFFB53178FDE8F9380435F919" box="[605,852,1736,1760]" class="Diplopoda" family="Pachybolidae" genus="Crurifarcimen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="vagans">
<emphasis id="B961EAEEFFB53178FDE8F9380435F919" box="[605,852,1736,1760]" italics="true" pageId="24" pageNumber="25">Crurifarcimen vagans</emphasis>
</taxonomicName>
, valves quite regularly sub-ovoid in shape, with free
</paragraph>
<paragraph id="8BAA36FCFFB53178FF22F91C02FDF8FD" blockId="24.[151,1437,1268,1868]" box="[151,1436,1772,1797]" pageId="24" pageNumber="25">margins only very shallowly sinuous, oral valve much longer than aboral valve, with large, rounded-subpyramidal</paragraph>
<paragraph id="8BAA36FCFFB53178FF22F8E003B5F8D1" blockId="24.[151,1437,1268,1868]" box="[151,1236,1808,1832]" pageId="24" pageNumber="25">
apical appendix (
<figureCitation id="132E2A79FFB53178FEE2F8E006C9F8D1" box="[343,424,1808,1832]" captionStart="FIGURES 51 – 53" captionStartId="26.[151,264,1672,1695]" captionTargetBox="[189,1398,183,1651]" captionTargetId="figure@26.[187,1400,181,1653]" captionTargetPageId="26" captionText="FIGURES 51 – 53. Parabolus dimorphus, female from the vicinity of Dar-es-Salaam, Brinkmann leg., right vulva. 51: posteriolateral view, 52: basis of vulva with operculum; 53, apical appendix. Scales: 1 mm (51), 0.1 mm (52 – 53). ap: apical appendix, op: operculum. SEM." httpUri="https://zenodo.org/record/276708/files/figure.png" pageId="24" pageNumber="25">Fig. 53</figureCitation>
)
<emphasis id="B961EAEEFFB53178FE02F8E1069DF8D1" box="[439,508,1809,1832]" italics="true" pageId="24" pageNumber="25">in situ</emphasis>
lying close to lateral coxosternal extension. Crest not protruding.
</paragraph>
</subSubSection>
<subSubSection id="C30F6577FFB53178FF72F8C304DEF8B5" box="[199,959,1843,1868]" pageId="24" pageNumber="25" type="distribution">
<paragraph id="8BAA36FCFFB53178FF72F8C304DEF8B5" blockId="24.[151,1437,1268,1868]" box="[199,959,1843,1868]" pageId="24" pageNumber="25">
<emphasis id="B961EAEEFFB53178FF72F8C30601F8B5" bold="true" box="[199,352,1843,1868]" pageId="24" pageNumber="25">Distribution.</emphasis>
Only known from Dar-es-Salaam,
<collectingCountry id="F302766CFFB53178FD5FF8C4042FF8B5" box="[746,846,1844,1868]" name="Tanzania" pageId="24" pageNumber="25">Tanzania</collectingCountry>
(
<figureCitation id="132E2A79FFB53178FCEBF8C404D2F8B5" box="[862,947,1844,1868]" captionStart="FIGURE 84" captionStartId="36.[151,249,1479,1502]" captionTargetBox="[456,1120,410,1444]" captionTargetId="figure@36.[456,1122,407,1445]" captionTargetPageId="36" captionText="FIGURE 84. Known distribution of East African Pachybolini, except Hadrobolus crassicollis and Epibolus pulchripes. dots (●): Crurifarcimen vagans, filled diamonds (♦): Hyperbolus apicomplexus, open diamonds (♢): Hyperbolus morogoroensis, filled triangle (˔): Parabolus dimorphus, inverted open triangle (): Parabolus calceus, inverted filled triangle (ˑ): Pachybolus tectus,, filled square (․): Pachybolini indet. Based on Yahoo maps." httpUri="https://zenodo.org/record/276715/files/figure.png" pageId="24" pageNumber="25">Fig. 84</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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