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<document id="50C83E1F82A0F2C50346E0230262E1C2" ID-DOI="10.11646/zootaxa.4991.3.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="5042656" ID-ZooBank="F0FAE07E-4A90-4602-BCC5-6D80E158BB6C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624989159860" checkinUser="plazi" docAuthor="Suksai, Benjamart, Gustafson, Grey T., Sites, Robert W. &amp; Sangpradub, Narumon" docDate="2021" docId="03BC87C1FFE4225B0CC07D23FF2B51C6" docLanguage="en" docName="zootaxa.4991.3.8.pdf" docOrigin="Zootaxa 4991 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Patrus landaisi" docType="treatment" docVersion="3" lastPageNumber="577" masterDocId="FF85FFB9FFEA224B0C577A23FFDF537D" masterDocTitle="Review of the Patrus landaisi species group from Thailand with description of three new species (Coleoptera, Gyrinidae)" masterLastPageNumber="591" masterPageNumber="561" pageNumber="575" updateTime="1698978743576" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="4B127FB76AEA7B3240DC1D7D38EECC0D">Review of the Patrus landaisi species group from Thailand with description of three new species (Coleoptera, Gyrinidae)</mods:title>
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<mods:namePart id="36CC6AAFA007774A385547CF1E83B353">Suksai, Benjamart</mods:namePart>
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<mods:namePart id="9013F4F776E81C4CF4ABD5C6597A8177">Gustafson, Grey T.</mods:namePart>
<mods:affiliation id="FA9C8E3C14A471465432026906D116E9">Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas 66045, USA. &amp; Department of Biological Sciences, Northern Arizona University, Flagstaff, Arizona, 86011, USA.</mods:affiliation>
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<mods:namePart id="80815BCCE61B135E08B4913AEFDD38D6">Sites, Robert W.</mods:namePart>
<mods:nameIdentifier id="2DBC8C7BE22EA4908DA5BDCB4E364BEC" type="ORCID">0000-0002-3895-813X</mods:nameIdentifier>
<mods:affiliation id="982ECAC39710D4D4B043A6A5B5128E30">Enns Entomology Museum, Division of Plant Science, University of Missouri-Columbia, Missouri 65211, USA. SitesR @ missouri. edu; https: // orcid. org / 0000 - 0002 - 3895 - 813 X</mods:affiliation>
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<mods:namePart id="8DC4D69574A42C7C3E7D0ED96ADFC9EA">Sangpradub, Narumon</mods:namePart>
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<mods:affiliation id="777885EC98678A1FB77BC4DC83F41CCD">Applied Taxonomic Research Center, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand &amp; Corresponding Author: narumon @ kku. ac. th; https: // orcid. org / 0000 - 0003 - 0228 - 8696</mods:affiliation>
<mods:nameIdentifier id="3DFA3125D3C10BC05BBC589DBB0C7F7F" type="email">benjamartbenjamart@gmail.com</mods:nameIdentifier>
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<mods:date id="F0656740EB4A16E4897A383F9C4EA38B">2021</mods:date>
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<paragraph id="8BAA36D7FFE422450CC07D23FDE55464" blockId="14.[151,628,1791,1852]" box="[151,570,1791,1818]" pageId="14" pageNumber="575">
<heading id="D0E281BBFFE422450CC07D23FDE55464" bold="true" box="[151,570,1791,1818]" fontSize="11" level="1" pageId="14" pageNumber="575" reason="1">
<taxonomicName id="4C154D54FFE422450CC07D23FDE55464" authority="(Regimbart, 1892)" baseAuthorityName="Regimbart" baseAuthorityYear="1892" box="[151,570,1791,1818]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="575" phylum="Arthropoda" rank="species" species="landaisi">
<emphasis id="B961EAC5FFE422450CC07D23FDE55464" bold="true" box="[151,570,1791,1818]" pageId="14" pageNumber="575">
<emphasis id="B961EAC5FFE422450CC07D23FE915467" bold="true" box="[151,334,1791,1818]" italics="true" pageId="14" pageNumber="575">Patrus landaisi</emphasis>
(
<bibRefCitation id="EF844B26FFE422450D097CDCFDED5464" author="Regimbart, M." box="[350,562,1791,1818]" pageId="14" pageNumber="575" pagination="663 - 752" refId="ref23063" refString="Regimbart, M. (1892) Essai monographique de la famille des Gyrinidae. 2 e supplement. Annales de la Societe entomologique de France, 60 (1891), 663 - 752, pls. 618 - 619." type="journal article" year="1892">Régimbart, 1892</bibRefCitation>
)
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="C30F655CFFE422450CC07D01FDAB5441" box="[151,628,1826,1852]" pageId="14" pageNumber="575" type="description">
<paragraph id="8BAA36D7FFE422450CC07D01FDAB5441" blockId="14.[151,628,1791,1852]" box="[151,628,1826,1852]" pageId="14" pageNumber="575">
(
<figureCitation id="132E2A52FFE422450CC87D01FEDC5441" box="[159,259,1826,1852]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="14" pageNumber="575">Figs. 2D</figureCitation>
,
<figureCitation id="132E2A52FFE422450D587D01FEEC5441" box="[271,307,1826,1852]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="14" pageNumber="575">3D</figureCitation>
,
<figureCitation id="132E2A52FFE422450D687D01FEBB5441" box="[319,356,1826,1852]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="14" pageNumber="575">4D</figureCitation>
,
<figureCitation id="132E2A52FFE422450D387D01FEA15441" box="[367,382,1826,1852]" captionStart="FIGURE 8" captionStartId="15.[151,260,1622,1648]" captionTargetBox="[157,1428,588,1589]" captionTargetId="figure-195@15.[151,1436,579,1598]" captionTargetPageId="15" captionText="FIGURE 8. Patrus landaisi: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042680" httpUri="https://zenodo.org/record/5042680/files/figure.png" pageId="14" pageNumber="575">8</figureCitation>
,
<figureCitation id="132E2A52FFE422450DDC7D01FDF05441" box="[395,559,1826,1852]" captionStart="FIGURE 13" captionStartId="25.[151,260,1683,1709]" captionTargetBox="[169,1418,181,1656]" captionTargetId="figure-18@25.[169,1418,179,1659]" captionTargetPageId="25" captionText="FIGURE 13. Habitats of P. landaisi species group: (A) stream at Siriphum Waterfall, Doi Inthanon National Park, Chiang Mai Province, habitat of P. garuda sp. nov. and P. subapicalis; (B) Benjamart Suksai in habitat of P. apicalis and P. salvazai at Nam Kuen stream, Chiang Rai Province; (C) shallow stream close to Ban Thai Samakkhi, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (D) Tham Yai Waterfall Phu Kradueng National Park, Loei Province, habitat of P. phetchabunensis sp. nov.; (E) puddle in dry stream bed close to Ban Phithak Thai, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (F) P. landaisi and P. salvazai from puddle in dry stream habitat close to Ban Thai Samakkhi, Chiang Rai Province." figureDoi="http://doi.org/10.5281/zenodo.5042690" httpUri="https://zenodo.org/record/5042690/files/figure.png" pageId="14" pageNumber="575">13C, 13E, 13F</figureCitation>
,
<figureCitation id="132E2A52FFE422450E6C7D01FDB35441" box="[571,620,1826,1852]" captionStart="FIGURE 14" captionStartId="26.[151,260,743,769]" captionTargetBox="[152,1436,179,719]" captionTargetId="figure-25@26.[151,1436,179,719]" captionTargetPageId="26" captionText="FIGURE 14. (A–B) Distribution records of the P. landaisi species group in Thailand." figureDoi="http://doi.org/10.5281/zenodo.5042692" httpUri="https://zenodo.org/record/5042692/files/figure.png" pageId="14" pageNumber="575">14A</figureCitation>
)
</paragraph>
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<paragraph id="8BAA36D7FFE422450CC07D4AFCED54FC" blockId="14.[151,1040,1897,1991]" box="[151,818,1897,1921]" pageId="14" pageNumber="575">
<treatmentCitationGroup id="AB0511F9FFE422450CC07D4AFCED54FC" box="[151,818,1897,1921]" pageId="14" pageNumber="575">
<treatmentCitation id="0AB410C6FFE422450CC07D4AFD8A54FC" author="Regimbart, M." box="[151,597,1897,1921]" page="693" pageId="14" pageNumber="575" year="1892">
<taxonomicName id="4C154D54FFE422450CC07D4AFD8A54FC" authority="Regimbart, 1892: 693" authorityName="Regimbart" authorityPageNumber="693" authorityYear="1892" box="[151,597,1897,1921]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="575" phylum="Arthropoda" rank="species" species="landaisi">
<emphasis id="B961EAC5FFE422450CC07D4AFEAE54FC" box="[151,369,1897,1921]" italics="true" pageId="14" pageNumber="575">Orectochilus landaisi</emphasis>
<bibRefCitation id="EF844B26FFE422450D207D4AFD8A54FC" author="Regimbart, M." box="[375,597,1897,1921]" pageId="14" pageNumber="575" pagination="663 - 752" refId="ref23063" refString="Regimbart, M. (1892) Essai monographique de la famille des Gyrinidae. 2 e supplement. Annales de la Societe entomologique de France, 60 (1891), 663 - 752, pls. 618 - 619." type="journal article" year="1892">Régimbart, 1892: 693</bibRefCitation>
</taxonomicName>
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(original description)
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</paragraph>
<paragraph id="8BAA36D7FFE422450CC07DAFFCC254D9" blockId="14.[151,1040,1897,1991]" box="[151,797,1932,1956]" pageId="14" pageNumber="575">
<treatmentCitationGroup id="AB0511F9FFE422450CC07DAFFCC254D9" box="[151,797,1932,1956]" pageId="14" pageNumber="575">
<taxonomicName id="4C154D54FFE422450CC07DAFFD5F54D9" authority="Regimbart, 1892" authorityName="Regimbart" authorityYear="1892" box="[151,640,1932,1956]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="575" phylum="Arthropoda" rank="species" species="landaisi" subGenus="Patrus">
<emphasis id="B961EAC5FFE422450CC07DAFFEC654D9" box="[151,281,1932,1956]" italics="true" pageId="14" pageNumber="575">Orectochilus</emphasis>
(
<emphasis id="B961EAC5FFE422450D7F7DAEFEB354D9" box="[296,364,1933,1956]" italics="true" pageId="14" pageNumber="575">Patrus</emphasis>
)
<emphasis id="B961EAC5FFE422450D2D7DAFFE1354D9" box="[378,460,1932,1956]" italics="true" pageId="14" pageNumber="575">landaisi</emphasis>
<bibRefCitation id="EF844B26FFE422450D857DAFFD5F54D9" author="Regimbart, M." box="[466,640,1932,1956]" pageId="14" pageNumber="575" pagination="663 - 752" refId="ref23063" refString="Regimbart, M. (1892) Essai monographique de la famille des Gyrinidae. 2 e supplement. Annales de la Societe entomologique de France, 60 (1891), 663 - 752, pls. 618 - 619." type="journal article" year="1892">Régimbart, 1892</bibRefCitation>
</taxonomicName>
:
<treatmentCitation id="0AB410C6FFE422450EDD7DAFFCC254D9" author="Ochs" box="[650,797,1932,1956]" page="27" pageId="14" pageNumber="575" year="1930">
<bibRefCitation id="EF844B26FFE422450EDD7DAFFCC254D9" author="Ochs" box="[650,797,1932,1956]" pageId="14" pageNumber="575" pagination="1 - 39" refId="ref22521" refString="Ochs, G. (1930) Gyrinoidea. Catalogue of Indian Insects, 19, 1 - 39." type="journal article" year="1930">Ochs 1930: 27</bibRefCitation>
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</paragraph>
<paragraph id="8BAA36D7FFE422450CC07D8CFBCF54BA" blockId="14.[151,1040,1897,1991]" box="[151,1040,1967,1991]" pageId="14" pageNumber="575">
<treatmentCitationGroup id="AB0511F9FFE422450CC07D8CFBCF54BA" box="[151,1040,1967,1991]" pageId="14" pageNumber="575">
<taxonomicName id="4C154D54FFE422450CC07D8CFE3854BA" authority="Regimbart, 1892" authorityName="Regimbart" authorityYear="1892" box="[151,487,1967,1991]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="575" phylum="Arthropoda" rank="species" species="landaisi">
<emphasis id="B961EAC5FFE422450CC07D8CFEEC54BA" box="[151,307,1967,1991]" italics="true" pageId="14" pageNumber="575">Patrus landaisi</emphasis>
<bibRefCitation id="EF844B26FFE422450D6E7D8CFE3854BA" author="Regimbart, M." box="[313,487,1967,1991]" pageId="14" pageNumber="575" pagination="663 - 752" refId="ref23063" refString="Regimbart, M. (1892) Essai monographique de la famille des Gyrinidae. 2 e supplement. Annales de la Societe entomologique de France, 60 (1891), 663 - 752, pls. 618 - 619." type="journal article" year="1892">Régimbart, 1892</bibRefCitation>
</taxonomicName>
:
<treatmentCitation id="0AB410C6FFE422450DA67D8CFD0454BA" author="Hajek, J. &amp; Fery, H." box="[497,731,1967,1991]" page="29" pageId="14" pageNumber="575" year="2017">
<bibRefCitation id="EF844B26FFE422450DA67D8CFD0454BA" author="Hajek, J. &amp; Fery, H." box="[497,731,1967,1991]" pageId="14" pageNumber="575" pagination="1810" refId="ref21748" refString="Hajek, J. &amp; Fery, H. (2017) Family Gyrinidae Latreille, 1810. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Leiden, Brill, pp. 22 - 29." type="book chapter" year="2017">Hájek &amp; Fery 2017: 29</bibRefCitation>
</treatmentCitation>
(new combination, catalogue)
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</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFE522440CC07AB5FB5A5153" pageId="15" pageNumber="576" type="materials_examined">
<paragraph id="8BAA36D7FFE522440CC07AB5FD5553CD" blockId="15.[151,1437,150,558]" box="[151,650,150,179]" pageId="15" pageNumber="576">
<materialsCitation id="3B7D3C8AFFE522440CC07AB5FD5853CD" box="[151,647,150,179]" collectorName="Cao B" country="Vietnam" location="Vietnam" pageId="15" pageNumber="576" specimenCount="1" typeStatus="holotype">
<emphasis id="B961EAC5FFE522440CC07AB5FEE953CD" bold="true" box="[151,310,150,176]" pageId="15" pageNumber="576">
<typeStatus id="54AE8875FFE522440CC07AB5FF0D53CD" box="[151,210,150,176]" pageId="15" pageNumber="576">Type</typeStatus>
locality.
</emphasis>
<collectorName id="26E05301FFE522440D6B7AB5FE5D53CE" box="[316,386,150,179]" pageId="15" pageNumber="576">Cao B</collectorName>
ằng Province,
<collectingCountry id="F3027647FFE522440E737AB5FD5853CD" box="[548,647,150,176]" name="Vietnam" pageId="15" pageNumber="576">Vietnam</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAA36D7FFE522440C907A9AFAE85241" blockId="15.[151,1437,150,558]" pageId="15" pageNumber="576">
<emphasis id="B961EAC5FFE522440C907A9AFE2F53AE" bold="true" box="[199,496,185,211]" pageId="15" pageNumber="576">Type material examined.</emphasis>
<typeStatus id="54AE8875FFE522440DAF7A9AFDA953AE" box="[504,630,185,211]" pageId="15" pageNumber="576" type="syntype">SYNTYPE</typeStatus>
female ‘Tonkin int- / env. De Cao bang [beige label, handwritten black ink]’ ‘MUSEUM PARIS / COLL
<collectingCountry id="F3027647FFE522440E767AFFFD7D538B" box="[545,674,220,246]" name="Mauritius" pageId="15" pageNumber="576">MAURICE</collectingCountry>
REGIMBART / 1908 [beige label, typed black ink]’ ‘
<typeStatus id="54AE8875FFE52244094E7AFFFA43538B" box="[1305,1436,220,246]" pageId="15" pageNumber="576" type="syntype">SYNTYPE</typeStatus>
[red label, typed black ink]’ ‘
<typeStatus id="54AE8875FFE522440D8E7ADDFD835265" box="[473,604,254,280]" pageId="15" pageNumber="576" type="syntype">SYNTYPE</typeStatus>
/
<taxonomicName id="4C154D54FFE522440E257ADDFCDD5265" authorityName="Dejean" authorityYear="1833" box="[626,770,254,280]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="576" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAC5FFE522440E257ADDFCDD5265" box="[626,770,254,280]" italics="true" pageId="15" pageNumber="576">Orectochilus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C154D54FFE522440F4F7ADDFBEA5265" authority="Regimbart, 1892" authorityName="Regimbart" authorityYear="1892" box="[792,1077,254,280]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="576" phylum="Arthropoda" rank="species" species="landaisi">
<emphasis id="B961EAC5FFE522440F4F7ADDFCAD5265" box="[792,882,254,280]" italics="true" pageId="15" pageNumber="576">landaisi</emphasis>
<bibRefCitation id="EF844B26FFE522440F2E7ADDFBEA5265" author="Regimbart, M." box="[889,1077,254,280]" pageId="15" pageNumber="576" pagination="663 - 752" refId="ref23063" refString="Regimbart, M. (1892) Essai monographique de la famille des Gyrinidae. 2 e supplement. Annales de la Societe entomologique de France, 60 (1891), 663 - 752, pls. 618 - 619." type="journal article" year="1892">Régimbart, 1892</bibRefCitation>
</taxonomicName>
/ [white label, typed black ink]’ ‘MNHN / EC10156 [white label, typed black ink];
<typeStatus id="54AE8875FFE522440E837B02FC8D5246" box="[724,850,289,315]" pageId="15" pageNumber="576" type="syntype">SYNTYPE</typeStatus>
female same locality except for, EC10157.
</paragraph>
<paragraph id="8BAA36D7FFE522440C907B60FB5A5153" blockId="15.[151,1437,150,558]" pageId="15" pageNumber="576">
<emphasis id="B961EAC5FFE522440C907B60FE225220" bold="true" box="[199,509,323,349]" pageId="15" pageNumber="576">Other material examined.</emphasis>
<materialsCitation id="3B7D3C8AFFE522440E537B60FD9452DE" collectingDate="2019-05-02" collectionCode="THNHM" collectorName="B. Suksai &amp; P. Sapewisut &amp; N. Plangklang" country="Thailand" county="Stream" elevation="1089" latitude="19.750153" location="Tap Tao Subdistrict" longLatPrecision="1" longitude="100.399864" municipality="Ban Phithak Thai" pageId="15" pageNumber="576" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Rai">
<emphasis id="B961EAC5FFE522440E537B60FC6F5220" bold="true" box="[516,944,323,349]" pageId="15" pageNumber="576">
<collectingCountry id="F3027647FFE522440E537B60FD405220" box="[516,671,323,349]" name="Thailand" pageId="15" pageNumber="576">THAILAND</collectingCountry>
:
<collectingRegion id="49D1F835FFE522440EFA7B60FC745220" box="[685,939,323,349]" country="Thailand" name="Chiang Rai" pageId="15" pageNumber="576">Chiang Rai Province</collectingRegion>
:
</emphasis>
<collectingCounty id="62CB4E5BFFE522440FEF7B67FBD85223" box="[952,1031,324,350]" pageId="15" pageNumber="576">Stream</collectingCounty>
close to
<collectingMunicipality id="6BCEACADFFE5224408397B67FAE95220" box="[1134,1334,323,350]" pageId="15" pageNumber="576">Ban Phithak Thai</collectingMunicipality>
,
<location id="8ECA600CFFE5224409157B67FECC52FD" LSID="urn:lsid:plazi:treatment:03BC87C1FFE4225B0CC07D23FF2B51C6:8ECA600CFFE5224409157B67FECC52FD" country="Thailand" county="Stream" latitude="19.750153" longLatPrecision="1" longitude="100.399864" municipality="Ban Phithak Thai" name="Tap Tao Subdistrict" pageId="15" pageNumber="576" stateProvince="Chiang Rai">Tap Tao Subdistrict</location>
,
<location id="8ECA600CFFE522440D497B45FE0D52FD" LSID="urn:lsid:plazi:treatment:03BC87C1FFE4225B0CC07D23FF2B51C6:8ECA600CFFE522440D497B45FE0D52FD" box="[286,466,358,384]" country="Thailand" county="Stream" latitude="19.750153" longLatPrecision="1" longitude="100.399864" municipality="Ban Phithak Thai" name="Thoeng District" pageId="15" pageNumber="576" stateProvince="Chiang Rai">Thoeng District</location>
,
<geoCoordinate id="EE215010FFE522440D8B7B45FD5752FC" box="[476,648,358,385]" degrees="19" direction="north" minutes="45" orientation="latitude" pageId="15" pageNumber="576" precision="1" seconds="00.55" value="19.750153">19°45’00.55”N</geoCoordinate>
<geoCoordinate id="EE215010FFE522440EC77B45FC9652FC" box="[656,841,358,385]" degrees="100" direction="east" minutes="23" orientation="longitude" pageId="15" pageNumber="576" precision="1" seconds="59.53" value="100.399864">100°23’59.53”E</geoCoordinate>
,
<elevation id="0038D1E4FFE522440F027B45FC3952FD" box="[853,998,358,385]" metricMagnitude="3" metricUnit="m" metricValue="1.089" pageId="15" pageNumber="576" unit="m" value="1089.0">
<quantity id="4CED9B32FFE522440F027B45FC7552FC" box="[853,938,358,385]" metricMagnitude="3" metricUnit="m" metricValue="1.089" pageId="15" pageNumber="576" unit="m" value="1089.0">1089 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFE522440FA47B45FB8552FD" box="[1011,1114,358,384]" pageId="15" pageNumber="576" value="2019-05-02">
<collectingDate id="EFEFE9FFFFE522440FA47B45FB8552FD" box="[1011,1114,358,384]" pageId="15" pageNumber="576" value="2019-05-02">2.V.2019</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFE5224408C27B44FADC52FD" box="[1173,1283,358,385]" pageId="15" pageNumber="576">B. Suksai</collectorName>
,
<collectorName id="26E05301FFE5224409587B44FA4352FD" box="[1295,1436,358,385]" pageId="15" pageNumber="576">P. Sapewisut</collectorName>
&amp;
<collectorName id="26E05301FFE522440CE27BAAFE8652DE" box="[181,345,393,419]" pageId="15" pageNumber="576">N. Plangklang</collectorName>
,
<specimenCount id="9D13FD5EFFE522440D337BAAFE1952DE" box="[356,454,393,419]" pageId="15" pageNumber="576" type="female">1 female</specimenCount>
(
<collectionCode id="ED04AE12FFE522440D817BAAFD9C52DE" box="[470,579,393,419]" pageId="15" pageNumber="576">THNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFE522440E007BAAFE1D5294" collectingDate="2018-11-10" collectionCode="GTGC, KKU, THNHM" collectorName="B. Suksai &amp; S. Kullasoot &amp; P. Sapewisut" country="Thailand" county="Stream" elevation="1299" latitude="19.737028" location="Tap Tao Subdistrict" longLatPrecision="1" longitude="100.40061" municipality="Ban Thai Samakkhi" pageId="15" pageNumber="576" specimenCount="7" specimenCount-male="7" stateProvince="Chiang Rai">
<collectingCounty id="62CB4E5BFFE522440E007BAAFD7952DE" box="[599,678,393,419]" pageId="15" pageNumber="576">Stream</collectingCounty>
close to
<collectingMunicipality id="6BCEACADFFE522440F5A7BAAFC3052DE" box="[781,1007,393,419]" pageId="15" pageNumber="576">Ban Thai Samakkhi</collectingMunicipality>
,
<location id="8ECA600CFFE522440FAD7BAAFB0752DE" LSID="urn:lsid:plazi:treatment:03BC87C1FFE4225B0CC07D23FF2B51C6:8ECA600CFFE522440FAD7BAAFB0752DE" box="[1018,1240,393,419]" country="Thailand" county="Stream" latitude="19.737028" longLatPrecision="1" longitude="100.40061" municipality="Ban Thai Samakkhi" name="Tap Tao Subdistrict" pageId="15" pageNumber="576" stateProvince="Chiang Rai">Tap Tao Subdistrict</location>
,
<location id="8ECA600CFFE5224408B37BAAFA4752DE" LSID="urn:lsid:plazi:treatment:03BC87C1FFE4225B0CC07D23FF2B51C6:8ECA600CFFE5224408B37BAAFA4752DE" box="[1252,1432,393,419]" country="Thailand" county="Stream" latitude="19.737028" longLatPrecision="1" longitude="100.40061" municipality="Ban Thai Samakkhi" name="Thoeng District" pageId="15" pageNumber="576" stateProvince="Chiang Rai">Thoeng District</location>
,
<geoCoordinate id="EE215010FFE522440CC07B8FFEE952BB" box="[151,310,428,454]" degrees="19" direction="north" minutes="44" orientation="latitude" pageId="15" pageNumber="576" precision="1" seconds="13.3" value="19.737028">19°44’13.3”N</geoCoordinate>
<geoCoordinate id="EE215010FFE522440D177B8FFE3352BB" box="[320,492,428,454]" degrees="100" direction="east" minutes="24" orientation="longitude" pageId="15" pageNumber="576" precision="1" seconds="02.2" value="100.40061">100°24’02.2”E</geoCoordinate>
,
<elevation id="0038D1E4FFE522440DAD7B8FFD4C52B8" box="[506,659,427,454]" metricMagnitude="3" metricUnit="m" metricValue="1.299" pageId="15" pageNumber="576" unit="m" value="1299.0">
<quantity id="4CED9B32FFE522440DAD7B8FFD8C52BB" box="[506,595,428,454]" metricMagnitude="3" metricUnit="m" metricValue="1.299" pageId="15" pageNumber="576" unit="m" value="1299.0">1299 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFE522440EF47B8FFCFA52BB" box="[675,805,428,454]" pageId="15" pageNumber="576" value="2018-11-10">
<collectingDate id="EFEFE9FFFFE522440EF47B8FFCFA52BB" box="[675,805,428,454]" pageId="15" pageNumber="576" value="2018-11-10">10.XI.2018</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFE522440F307B8FFC0752B8" box="[871,984,427,454]" pageId="15" pageNumber="576">B. Suksai</collectorName>
,
<collectorName id="26E05301FFE522440FB07B8FFBAA52B8" box="[999,1141,427,454]" pageId="15" pageNumber="576">S. Kullasoot</collectorName>
&amp;
<collectorName id="26E05301FFE5224408F77B8FFAEB52B8" box="[1184,1332,427,454]" pageId="15" pageNumber="576">P. Sapewisut</collectorName>
,
<specimenCount id="9D13FD5EFFE5224409157B8FFA4352B8" box="[1346,1436,427,454]" pageId="15" pageNumber="576" type="male">7 males</specimenCount>
(
<collectionCode id="ED04AE12FFE522440CC87BEDFF2F5295" box="[159,240,462,488]" pageId="15" pageNumber="576">GTGC</collectionCode>
,
<collectionCode id="ED04AE12FFE522440D577BECFEE05294" box="[256,319,463,489]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="15" pageNumber="576">KKU</collectionCode>
,
<collectionCode id="ED04AE12FFE522440D187BECFE655294" box="[335,442,463,489]" pageId="15" pageNumber="576">THNHM</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7D3C8AFFE522440D857BECFCC45294" box="[466,795,462,489]" collectingDate="2018-11-10" collectionCode="KKU, THNHM" collectorName="B. Suksai &amp; S. Kullasoot &amp; P. Sapewisut" country="Thailand" county="Stream" elevation="1299" isTailEnumeration="true" latitude="19.737028" location="Tap Tao Subdistrict" longLatPrecision="1" longitude="100.40061" municipality="Ban Thai Samakkhi" pageId="15" pageNumber="576" specimenCount="7" specimenCount-female="7" stateProvince="Chiang Rai">
<specimenCount id="9D13FD5EFFE522440D857BECFD9C5295" box="[466,579,462,489]" pageId="15" pageNumber="576" type="female">7 females</specimenCount>
(
<collectionCode id="ED04AE12FFE522440E007BEDFD475295" box="[599,664,462,488]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="15" pageNumber="576">KKU</collectionCode>
,
<collectionCode id="ED04AE12FFE522440EFF7BECFCCB5294" box="[680,788,463,489]" pageId="15" pageNumber="576">THNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFE522440F7C7BECFC0F5176" collectingDate="2019-05-02" collectionCode="GTGC, THNHM, UMC" collectorName="B. Suksai &amp; P. Sapewisut &amp; N. Plangklang" country="Thailand" county="Stream" elevation="1299" isTailEnumeration="true" latitude="19.737028" location="Tap Tao Subdistrict" longLatPrecision="1" longitude="100.40061" municipality="Ban Thai Samakkhi" pageId="15" pageNumber="576" specimenCount="13" specimenCount-female="8" specimenCount-male="5" stateProvince="Chiang Rai">
same locality except for,
<date id="FFAB1017FFE52244080E7BEDFB1F5295" box="[1113,1216,462,488]" pageId="15" pageNumber="576" value="2019-05-02">
<collectingDate id="EFEFE9FFFFE52244080E7BEDFB1F5295" box="[1113,1216,462,488]" pageId="15" pageNumber="576" value="2019-05-02">2.V.2019</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFE5224409527BECFAA75295" box="[1285,1400,462,489]" pageId="15" pageNumber="576">B. Suksai</collectorName>
,
<collectorName id="26E05301FFE5224409DE7BECFED55176" pageId="15" pageNumber="576">P. Sapewisut</collectorName>
&amp;
<collectorName id="26E05301FFE522440D7B7BD2FE105176" box="[300,463,497,523]" pageId="15" pageNumber="576">N. Plangklang</collectorName>
,
<specimenCount id="9D13FD5EFFE522440D8E7BD2FDEE5176" box="[473,561,497,524]" pageId="15" pageNumber="576" type="male">5 males</specimenCount>
,
<specimenCount id="9D13FD5EFFE522440E6B7BD2FD785176" box="[572,679,497,523]" pageId="15" pageNumber="576" type="female">8 females</specimenCount>
(
<collectionCode id="ED04AE12FFE522440EE27BD2FCD95176" box="[693,774,497,523]" pageId="15" pageNumber="576">GTGC</collectionCode>
,
<collectionCode id="ED04AE12FFE522440F587BD2FCA65176" box="[783,889,497,523]" pageId="15" pageNumber="576">THNHM</collectionCode>
,
<collectionCode id="ED04AE12FFE522440FD47BD2FC175176" box="[899,968,497,523]" pageId="15" pageNumber="576">UMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFE522440F8D7BD2FB5E5153" collectingDate="2017-05-28" collectionCode="KKU, THNHM" collectorName="P. Sapewisut" country="Thailand" county="Stream" elevation="1263" isTailEnumeration="true" latitude="19.743181" location="Tap Tao Subdistrict" longLatPrecision="1" longitude="100.40164" municipality="Ban Thai Samakkhi" pageId="15" pageNumber="576" specimenCount="9" specimenCount-female="9" stateProvince="Chiang Rai">
same locality except for,
<geoCoordinate id="EE215010FFE5224408A77BD2FA435171" box="[1264,1436,497,524]" degrees="19" direction="north" minutes="44" orientation="latitude" pageId="15" pageNumber="576" precision="1" seconds="35.45" value="19.743181">19°44’35.45”N</geoCoordinate>
<geoCoordinate id="EE215010FFE522440CC07830FE8E5153" box="[151,337,531,558]" degrees="100" direction="east" minutes="24" orientation="longitude" pageId="15" pageNumber="576" precision="1" seconds="05.91" value="100.40164">100°24’05.91”E</geoCoordinate>
,
<elevation id="0038D1E4FFE522440D0B7830FE315150" box="[348,494,531,558]" metricMagnitude="3" metricUnit="m" metricValue="1.263" pageId="15" pageNumber="576" unit="m" value="1263.0">
<quantity id="4CED9B32FFE522440D0B7830FE6E5153" box="[348,433,531,558]" metricMagnitude="3" metricUnit="m" metricValue="1.263" pageId="15" pageNumber="576" unit="m" value="1263.0">1263 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFE522440DAD7837FDB05153" box="[506,623,532,558]" pageId="15" pageNumber="576" value="2017-05-28">
<collectingDate id="EFEFE9FFFFE522440DAD7837FDB05153" box="[506,623,532,558]" pageId="15" pageNumber="576" value="2017-05-28">28.V.2017</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFE522440EFC7837FCE45150" box="[683,827,531,558]" pageId="15" pageNumber="576">P. Sapewisut</collectorName>
,
<specimenCount id="9D13FD5EFFE522440F117837FC6D5150" box="[838,946,531,558]" pageId="15" pageNumber="576" type="female">9 females</specimenCount>
(
<collectionCode id="ED04AE12FFE522440F967837FBDD5153" box="[961,1026,532,558]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="15" pageNumber="576">KKU</collectionCode>
,
<collectionCode id="ED04AE12FFE52244085A7837FBA65153" box="[1037,1145,532,558]" pageId="15" pageNumber="576">THNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF6A665FFFE522440CC07C75FDC1547D" ID-DOI="http://doi.org/10.5281/zenodo.5042680" ID-Zenodo-Dep="5042680" httpUri="https://zenodo.org/record/5042680/files/figure.png" pageId="15" pageNumber="576" startId="15.[151,260,1622,1648]" targetBox="[157,1428,588,1589]" targetPageId="15">
<paragraph id="8BAA36D7FFE522440CC07C75FDC1547D" blockId="15.[151,1436,1622,1793]" pageId="15" pageNumber="576">
<emphasis id="B961EAC5FFE522440CC07C75FEFD550D" bold="true" box="[151,290,1622,1648]" pageId="15" pageNumber="576">FIGURE 8.</emphasis>
<taxonomicName id="4C154D54FFE522440D7D7C74FE08550D" authorityName="Regimbart" authorityYear="1892" box="[298,471,1622,1648]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="576" phylum="Arthropoda" rank="species" species="landaisi">
<emphasis id="B961EAC5FFE522440D7D7C74FE08550D" box="[298,471,1622,1648]" italics="true" pageId="15" pageNumber="576">Patrus landaisi</emphasis>
</taxonomicName>
: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view.
</paragraph>
</caption>
<subSubSection id="C30F655CFFE5225B0C907D72FE205176" lastPageId="16" lastPageNumber="577" pageId="15" pageNumber="576" type="diagnosis">
<paragraph id="8BAA36D7FFE5225B0C907D72FA8C52DE" blockId="15.[151,1437,1873,2038]" lastBlockId="16.[151,1437,151,699]" lastPageId="16" lastPageNumber="577" pageId="15" pageNumber="576">
<emphasis id="B961EAC5FFE522440C907D72FE9D5416" bold="true" box="[199,322,1873,1899]" pageId="15" pageNumber="576">Diagnosis.</emphasis>
Size: Male: 11.8–13.0 mm, female:
<quantity id="4CED9B32FFE522440E867D72FCA65411" box="[721,889,1873,1900]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.22" metricValueMin="1.08" pageId="15" pageNumber="576" unit="mm" value="11.5" valueMax="12.2" valueMin="10.8">10.8–12.2 mm</quantity>
; body form in lateral view weakly dorsoventrally convex; elytral glabrous region of male (
<figureCitation id="132E2A52FFE522440E317D57FD1354F3" box="[614,716,1908,1934]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="15" pageNumber="576">Figs. 2D</figureCitation>
,
<figureCitation id="132E2A52FFE522440E807D57FD2454F3" box="[727,763,1908,1934]" captionStart="FIGURE 8" captionStartId="15.[151,260,1622,1648]" captionTargetBox="[157,1428,588,1589]" captionTargetId="figure-195@15.[151,1436,579,1598]" captionTargetPageId="15" captionText="FIGURE 8. Patrus landaisi: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042680" httpUri="https://zenodo.org/record/5042680/files/figure.png" pageId="15" pageNumber="576">8A</figureCitation>
) cordiform, along suture extending ca. 7/9 length of elytra, laterally expanded in anterior 2/3 to considerably wider than pronotal glabrous region, posterior 1/3 strongly acuminate with rounded margins; female elytral glabrous region (
<figureCitation id="132E2A52FFE522440FF27D9AFBD154AE" box="[933,1038,1977,2003]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="15" pageNumber="576">Figs. 3D</figureCitation>
,
<figureCitation id="132E2A52FFE5224408497D9AFB9F54AE" box="[1054,1088,1977,2003]" captionStart="FIGURE 8" captionStartId="15.[151,260,1622,1648]" captionTargetBox="[157,1428,588,1589]" captionTargetId="figure-195@15.[151,1436,579,1598]" captionTargetPageId="15" captionText="FIGURE 8. Patrus landaisi: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042680" httpUri="https://zenodo.org/record/5042680/files/figure.png" pageId="15" pageNumber="576">8E</figureCitation>
) dissimilar to male, broad in anterior 2/3 occupying 10/11 elytral width, lateral margins nearly straight creating parallel-sided appearance, abruptly acuminate medially in posterior 1/3, posterolateral corners not jagged and not produced into points, posterior margin near-evenly convex toward suture; elytral apex of both sexes similar, obliquely truncate without apicolateral sinuation, epipleural angle distinct, rounded; male protibia (
<figureCitation id="132E2A52FFFA225B0FA57AFFFB90538B" box="[1010,1103,220,246]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="16" pageNumber="577">Fig. 4D</figureCitation>
) club-shaped, without basal constriction, protibial apex obliquely truncate with prominent distolateral angle; male protarsus (
<figureCitation id="132E2A52FFFA225B095A7ADCFAB65264" box="[1293,1385,255,281]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="16" pageNumber="577">Fig. 4D</figureCitation>
) ca. 1/2 length of protibia, protarsomeres 1–4 mostly similar in width, lateral margins weakly rounded; mesoventrite without medial pit; aedeagus (
<figureCitation id="132E2A52FFFA225B0DAF7B67FDAA5223" box="[504,629,324,350]" captionStart="FIGURE 8" captionStartId="15.[151,260,1622,1648]" captionTargetBox="[157,1428,588,1589]" captionTargetId="figure-195@15.[151,1436,579,1598]" captionTargetPageId="15" captionText="FIGURE 8. Patrus landaisi: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042680" httpUri="https://zenodo.org/record/5042680/files/figure.png" pageId="16" pageNumber="577">Fig. 8B–D</figureCitation>
) with median lobe weakly tapered, apical 1/6 of aedeagus acuminate with nearly straight lateral margins, apex not constricted in lateral view (
<figureCitation id="132E2A52FFFA225B0F857B44FBF452FC" box="[978,1067,359,385]" captionStart="FIGURE 8" captionStartId="15.[151,260,1622,1648]" captionTargetBox="[157,1428,588,1589]" captionTargetId="figure-195@15.[151,1436,579,1598]" captionTargetPageId="15" captionText="FIGURE 8. Patrus landaisi: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042680" httpUri="https://zenodo.org/record/5042680/files/figure.png" pageId="16" pageNumber="577">Fig. 8C</figureCitation>
); female reproductive tract (
<figureCitation id="132E2A52FFFA225B09397B44FF6452DE" captionStart="FIGURE 8" captionStartId="15.[151,260,1622,1648]" captionTargetBox="[157,1428,588,1589]" captionTargetId="figure-195@15.[151,1436,579,1598]" captionTargetPageId="15" captionText="FIGURE 8. Patrus landaisi: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042680" httpUri="https://zenodo.org/record/5042680/files/figure.png" pageId="16" pageNumber="577">Fig. 8G</figureCitation>
) without additional sclerotized structure between gonocoxae (
<figureCitation id="132E2A52FFFA225B0FD27BAAFC0352DE" box="[901,988,393,419]" captionStart="FIGURE 8" captionStartId="15.[151,260,1622,1648]" captionTargetBox="[157,1428,588,1589]" captionTargetId="figure-195@15.[151,1436,579,1598]" captionTargetPageId="15" captionText="FIGURE 8. Patrus landaisi: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042680" httpUri="https://zenodo.org/record/5042680/files/figure.png" pageId="16" pageNumber="577">Fig. 8F</figureCitation>
), fertilization duct as in
<figureCitation id="132E2A52FFFA225B08A37BAAFA9052DE" box="[1268,1359,393,419]" captionStart="FIGURE 8" captionStartId="15.[151,260,1622,1648]" captionTargetBox="[157,1428,588,1589]" captionTargetId="figure-195@15.[151,1436,579,1598]" captionTargetPageId="15" captionText="FIGURE 8. Patrus landaisi: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042680" httpUri="https://zenodo.org/record/5042680/files/figure.png" pageId="16" pageNumber="577">Fig. 8H</figureCitation>
.
</paragraph>
<paragraph id="8BAA36D7FFFA225B0C907B8FFE205176" blockId="16.[151,1437,151,699]" pageId="16" pageNumber="577">
<emphasis id="B961EAC5FFFA225B0C907B8FFE0952BB" bold="true" box="[199,470,428,454]" pageId="16" pageNumber="577">Differential diagnosis.</emphasis>
<taxonomicName id="4C154D54FFFA225B0DB77B8EFD4C52BB" authorityName="Regimbart" authorityYear="1892" box="[480,659,428,454]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="577" phylum="Arthropoda" rank="species" species="landaisi">
<emphasis id="B961EAC5FFFA225B0DB77B8EFD4C52BB" box="[480,659,428,454]" italics="true" pageId="16" pageNumber="577">Patrus landaisi</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C154D54FFFA225B0F227B8EFC3952BB" authority="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub, 2021" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" box="[885,998,428,454]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="577" phylum="Arthropoda" rank="species" species="garuda" status="sp. nov.">
<emphasis id="B961EAC5FFFA225B0F227B8EFC3952BB" box="[885,998,428,454]" italics="true" pageId="16" pageNumber="577">P. garuda</emphasis>
</taxonomicName>
<emphasis id="B961EAC5FFFA225B0FA77B8FFB9152BB" bold="true" box="[1008,1102,428,454]" pageId="16" pageNumber="577">
<taxonomicNameLabel id="A25257BEFFFA225B0FA77B8FFB9152BB" box="[1008,1102,428,454]" pageId="16" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but can be distinguished by the differential diagnoses provided above. In general, the obliquely truncate elytral apices of
<taxonomicName id="4C154D54FFFA225B08997BECFA985294" authorityName="Regimbart" authorityYear="1892" box="[1230,1351,463,489]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="577" phylum="Arthropoda" rank="species" species="landaisi">
<emphasis id="B961EAC5FFFA225B08997BECFA985294" box="[1230,1351,463,489]" italics="true" pageId="16" pageNumber="577">P. landaisi</emphasis>
</taxonomicName>
should readily distinguish this species.
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFFA225B0C907837FF2B51C6" pageId="16" pageNumber="577" type="distribution">
<paragraph id="8BAA36D7FFFA225B0C907837FF2B51C6" blockId="16.[151,1437,151,699]" pageId="16" pageNumber="577">
<emphasis id="B961EAC5FFFA225B0C907837FEBB5153" bold="true" box="[199,356,532,558]" pageId="16" pageNumber="577">Distribution.</emphasis>
This species is known from northern
<collectingCountry id="F3027647FFFA225B0F427837FCA95153" box="[789,886,532,558]" name="Vietnam" pageId="16" pageNumber="577">Vietnam</collectingCountry>
(
<bibRefCitation id="EF844B26FFFA225B0FD37837FB9C5153" author="Regimbart, M." box="[900,1091,532,558]" pageId="16" pageNumber="577" pagination="663 - 752" refId="ref23063" refString="Regimbart, M. (1892) Essai monographique de la famille des Gyrinidae. 2 e supplement. Annales de la Societe entomologique de France, 60 (1891), 663 - 752, pls. 618 - 619." type="journal article" year="1892">Régimbart 1892</bibRefCitation>
;
<bibRefCitation id="EF844B26FFFA225B08197837FB365153" author="Zaitzev, F. A." box="[1102,1257,532,558]" pageId="16" pageNumber="577" pagination="223 - 226" refId="ref23757" refString="Zaitzev, F. A. (1910) Coleopteres aquatiques nouveaux ou peu connus. Revue Russe d'Entomologie, 10, 223 - 226." type="journal article" year="1910">Zaitzev 1910</bibRefCitation>
;
<bibRefCitation id="EF844B26FFFA225B08A37837FA4F5153" author="Peschet, R." box="[1268,1424,532,558]" pageId="16" pageNumber="577" pagination="121 - 136" refId="ref22628" refString="Peschet, R. (1923) Fauna entomologique de l'Indochine Gyrinidae. Opuscules de l'Institut Scientifique de l'Indochine Saigon, 6, 121 - 136." type="journal article" year="1923">Peschet 1923</bibRefCitation>
), southeastern
<collectingCountry id="F3027647FFFA225B0D677814FEAB512C" box="[304,372,567,593]" name="China" pageId="16" pageNumber="577">China</collectingCountry>
(
<collectingRegion id="49D1F835FFFA225B0DD17814FE33512F" box="[390,492,567,594]" country="China" name="Guizhou" pageId="16" pageNumber="577">Guizhou</collectingRegion>
) (
<bibRefCitation id="EF844B26FFFA225B0E527814FD19512C" author="Regimbart, M." box="[517,710,567,593]" pageId="16" pageNumber="577" pagination="137 - 245" refId="ref23105" refString="Regimbart, M. (1907) Essai monographique de la famille des Gyrinidae. 3 e supplement. Annales de la Societe entomologique de France, 76, 137 - 245." type="journal article" year="1907">Régimbart 1907</bibRefCitation>
;
<bibRefCitation id="EF844B26FFFA225B0E847814FC9E512C" author="Wu, C. F." box="[723,833,567,593]" pageId="16" pageNumber="577" pagination="63 - 73" refId="ref23732" refString="Wu, C. F. (1931) Catalogue of Chinese Gyrinidae. Peking Natural History Bulletin, 6, 63 - 73." type="journal article" year="1931">Wu 1931</bibRefCitation>
;
<bibRefCitation id="EF844B26FFFA225B0F077814FBDF512C" author="Mazzoldi, P." box="[848,1024,566,593]" pageId="16" pageNumber="577" pagination="155 - 172" refId="ref22146" refString="Mazzoldi, P. (1995) Gyrinidae: Catalogue of Chinese Gyrinidae (Coleoptera). In: Jach, M. A. &amp; Ji, L. (Eds.), Water Beetles of China. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien, pp. 155 - 172." type="book chapter" year="1995">Mazzoldi 1995</bibRefCitation>
) and
<collectingCountry id="F3027647FFFA225B08137814FBA3512C" box="[1092,1148,567,593]" name="Laos" pageId="16" pageNumber="577">Laos</collectingCountry>
(without further details) (
<bibRefCitation id="EF844B26FFFA225B0CC8787AFEFD510E" author="Ochs" box="[159,290,601,627]" pageId="16" pageNumber="577" pagination="1 - 39" refId="ref22521" refString="Ochs, G. (1930) Gyrinoidea. Catalogue of Indian Insects, 19, 1 - 39." type="journal article" year="1930">Ochs 1930</bibRefCitation>
). The records from
<collectingRegion id="49D1F835FFFA225B0E5E787AFD24510E" box="[521,763,601,627]" country="Thailand" name="Chiang Rai" pageId="16" pageNumber="577">Chiang Rai Province</collectingRegion>
(
<figureCitation id="132E2A52FFFA225B0F5B787AFCAA510E" box="[780,885,601,627]" captionStart="FIGURE 14" captionStartId="26.[151,260,743,769]" captionTargetBox="[152,1436,179,719]" captionTargetId="figure-25@26.[151,1436,179,719]" captionTargetPageId="26" captionText="FIGURE 14. (A–B) Distribution records of the P. landaisi species group in Thailand." figureDoi="http://doi.org/10.5281/zenodo.5042692" httpUri="https://zenodo.org/record/5042692/files/figure.png" pageId="16" pageNumber="577">Fig. 14A</figureCitation>
) reported here are the first for
<collectingCountry id="F3027647FFFA225B08B6787AFA95510E" box="[1249,1354,601,627]" name="Thailand" pageId="16" pageNumber="577">Thailand</collectingCountry>
, where it was found together with
<taxonomicName id="4C154D54FFFA225B0D85785EFD9151EB" baseAuthorityName="Ochs" baseAuthorityYear="1940" box="[466,590,636,662]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="577" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFFA225B0D85785EFD9151EB" box="[466,590,636,662]" italics="true" pageId="16" pageNumber="577">P. salvazai</emphasis>
</taxonomicName>
in a shallow stream (
<figureCitation id="132E2A52FFFA225B0F1D785FFC6D51EB" box="[842,946,636,662]" captionStart="FIGURE 13" captionStartId="25.[151,260,1683,1709]" captionTargetBox="[169,1418,181,1656]" captionTargetId="figure-18@25.[169,1418,179,1659]" captionTargetPageId="25" captionText="FIGURE 13. Habitats of P. landaisi species group: (A) stream at Siriphum Waterfall, Doi Inthanon National Park, Chiang Mai Province, habitat of P. garuda sp. nov. and P. subapicalis; (B) Benjamart Suksai in habitat of P. apicalis and P. salvazai at Nam Kuen stream, Chiang Rai Province; (C) shallow stream close to Ban Thai Samakkhi, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (D) Tham Yai Waterfall Phu Kradueng National Park, Loei Province, habitat of P. phetchabunensis sp. nov.; (E) puddle in dry stream bed close to Ban Phithak Thai, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (F) P. landaisi and P. salvazai from puddle in dry stream habitat close to Ban Thai Samakkhi, Chiang Rai Province." figureDoi="http://doi.org/10.5281/zenodo.5042690" httpUri="https://zenodo.org/record/5042690/files/figure.png" pageId="16" pageNumber="577">Fig. 13C</figureCitation>
) and puddles of a drying stream bed (
<figureCitation id="132E2A52FFFA225B0939785FFF3951C6" captionStart="FIGURE 13" captionStartId="25.[151,260,1683,1709]" captionTargetBox="[169,1418,181,1656]" captionTargetId="figure-18@25.[169,1418,179,1659]" captionTargetPageId="25" captionText="FIGURE 13. Habitats of P. landaisi species group: (A) stream at Siriphum Waterfall, Doi Inthanon National Park, Chiang Mai Province, habitat of P. garuda sp. nov. and P. subapicalis; (B) Benjamart Suksai in habitat of P. apicalis and P. salvazai at Nam Kuen stream, Chiang Rai Province; (C) shallow stream close to Ban Thai Samakkhi, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (D) Tham Yai Waterfall Phu Kradueng National Park, Loei Province, habitat of P. phetchabunensis sp. nov.; (E) puddle in dry stream bed close to Ban Phithak Thai, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (F) P. landaisi and P. salvazai from puddle in dry stream habitat close to Ban Thai Samakkhi, Chiang Rai Province." figureDoi="http://doi.org/10.5281/zenodo.5042690" httpUri="https://zenodo.org/record/5042690/files/figure.png" pageId="16" pageNumber="577">Fig. 13E–F</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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