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<document id="BA1370E04FCBB077148491F40F51B19A" ID-DOI="10.11646/zootaxa.4991.3.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="5042656" ID-ZooBank="F0FAE07E-4A90-4602-BCC5-6D80E158BB6C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624989159860" checkinUser="plazi" docAuthor="Suksai, Benjamart, Gustafson, Grey T., Sites, Robert W. &amp; Sangpradub, Narumon" docDate="2021" docId="03BC87C1FFE022470CC07F31FB5354CE" docLanguage="en" docName="zootaxa.4991.3.8.pdf" docOrigin="Zootaxa 4991 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Patrus garuda Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="573" masterDocId="FF85FFB9FFEA224B0C577A23FFDF537D" masterDocTitle="Review of the Patrus landaisi species group from Thailand with description of three new species (Coleoptera, Gyrinidae)" masterLastPageNumber="591" masterPageNumber="561" pageNumber="571" updateTime="1698978743576" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="39179E276E3951E6A1D32F10D45D4A8F">Review of the Patrus landaisi species group from Thailand with description of three new species (Coleoptera, Gyrinidae)</mods:title>
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<mods:namePart id="A0767D8FF7C49C44502A0B85FFED6FD7">Suksai, Benjamart</mods:namePart>
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<mods:affiliation id="3D60FD32706C8517CED2D9EB2FDB0127">Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas 66045, USA. &amp; Department of Biological Sciences, Northern Arizona University, Flagstaff, Arizona, 86011, USA.</mods:affiliation>
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<mods:namePart id="1DED9117FBA1D94603FBAF59E45E13F6">Sites, Robert W.</mods:namePart>
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<mods:affiliation id="629C8987F27B7CF06222B4D492CFF10C">Enns Entomology Museum, Division of Plant Science, University of Missouri-Columbia, Missouri 65211, USA. SitesR @ missouri. edu; https: // orcid. org / 0000 - 0002 - 3895 - 813 X</mods:affiliation>
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<mods:namePart id="1490444D3A175A9FC5EB1057B2425DEA">Sangpradub, Narumon</mods:namePart>
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<mods:affiliation id="1774A7935244752093EC6F1FF4357930">Applied Taxonomic Research Center, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand &amp; Corresponding Author: narumon @ kku. ac. th; https: // orcid. org / 0000 - 0003 - 0228 - 8696</mods:affiliation>
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(
<figureCitation id="132E2A52FFE022410CC87F16FEDD5632" box="[159,258,1333,1359]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="10" pageNumber="571">Figs. 2C</figureCitation>
, 
<figureCitation id="132E2A52FFE022410D5A7F16FEEF5632" box="[269,304,1333,1359]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="10" pageNumber="571">3C</figureCitation>
, 
<figureCitation id="132E2A52FFE022410D6B7F16FE805632" box="[316,351,1333,1359]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="10" pageNumber="571">4C</figureCitation>
, 
<figureCitation id="132E2A52FFE022410D3C7F15FEA4562D" box="[363,379,1334,1360]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="10" pageNumber="571">7</figureCitation>
, 
<figureCitation id="132E2A52FFE022410DD07F16FE665632" box="[391,441,1333,1359]" captionStart="FIGURE 13" captionStartId="25.[151,260,1683,1709]" captionTargetBox="[169,1418,181,1656]" captionTargetId="figure-18@25.[169,1418,179,1659]" captionTargetPageId="25" captionText="FIGURE 13. Habitats of P. landaisi species group: (A) stream at Siriphum Waterfall, Doi Inthanon National Park, Chiang Mai Province, habitat of P. garuda sp. nov. and P. subapicalis; (B) Benjamart Suksai in habitat of P. apicalis and P. salvazai at Nam Kuen stream, Chiang Rai Province; (C) shallow stream close to Ban Thai Samakkhi, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (D) Tham Yai Waterfall Phu Kradueng National Park, Loei Province, habitat of P. phetchabunensis sp. nov.; (E) puddle in dry stream bed close to Ban Phithak Thai, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (F) P. landaisi and P. salvazai from puddle in dry stream habitat close to Ban Thai Samakkhi, Chiang Rai Province." figureDoi="http://doi.org/10.5281/zenodo.5042690" httpUri="https://zenodo.org/record/5042690/files/figure.png" pageId="10" pageNumber="571">13A</figureCitation>
, 
<figureCitation id="132E2A52FFE022410D927F16FE295632" box="[453,502,1333,1359]" captionStart="FIGURE 14" captionStartId="26.[151,260,743,769]" captionTargetBox="[152,1436,179,719]" captionTargetId="figure-25@26.[151,1436,179,719]" captionTargetPageId="26" captionText="FIGURE 14. (A–B) Distribution records of the P. landaisi species group in Thailand." figureDoi="http://doi.org/10.5281/zenodo.5042692" httpUri="https://zenodo.org/record/5042692/files/figure.png" pageId="10" pageNumber="571">14A</figureCitation>
)
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<emphasis id="B961EAC5FFE022410CC07F5BFE9B56EF" bold="true" box="[151,324,1400,1426]" pageId="10" pageNumber="571">Type material.</emphasis>
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: male (card mounted; aedeagus in microvial), “ 
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/ Prov.; Doi Inthanon Natl. / 
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; 
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/ 
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; L-401 // 
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/ 
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a.s.l.
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/ UMC &amp; CMU teams” (
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(8 exs): 
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<collectingCountry id="F3027647FFE022410DEA7FC7FD875683" box="[445,600,1508,1534]" name="Thailand" pageId="10" pageNumber="571">THAILAND</collectingCountry>
: 
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:
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, 
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, leg. CMU team, 
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; 
<materialsCitation id="3B7D3C8AFFE0224108477C2BFA47553B" collectingDate="2002-03-09" collectionCode="UMC" country="Thailand" elevation="1460" latitude="18.533333" location="Siriphum Waterfall" longLatPrecision="1275" longitude="98.51667" municipality="Doi Inthanon Natl. Park." pageId="10" pageNumber="571" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Chiang Mai" typeStatus="paratype">
same locality except for, 
<location id="8ECA600CFFE0224109637C2BFEDD553B" LSID="urn:lsid:plazi:treatment:03BC87C1FFE022470CC07F31FB5354CE:8ECA600CFFE0224109637C2BFEDD553B" country="Thailand" latitude="18.533333" longLatPrecision="1275" longitude="98.51667" name="Siriphum Waterfall" pageId="10" pageNumber="571" stateProvince="Chiang Mai">Siriphum Waterfall</location>
, 
<geoCoordinate id="EE215010FFE022410D5B7C0FFEB2553B" box="[268,365,1580,1606]" degrees="18" direction="north" minutes="32" orientation="latitude" pageId="10" pageNumber="571" precision="925" value="18.533333">18°32’N</geoCoordinate>
<geoCoordinate id="EE215010FFE022410D237C0FFE0B553B" box="[372,468,1580,1606]" degrees="98" direction="east" minutes="31" orientation="longitude" pageId="10" pageNumber="571" precision="925" value="98.51667">98°31’E</geoCoordinate>
, 
<elevation id="0038D1E4FFE022410DB77C0FFDAE553B" box="[480,625,1580,1606]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="10" pageNumber="571" unit="m" value="1460.0">
<quantity id="4CED9B32FFE022410DB77C0FFDEB553B" box="[480,564,1580,1606]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="10" pageNumber="571" unit="m" value="1460.0">1460 m</quantity>
a.s.l.
</elevation>
, 
<date id="FFAB1017FFE022410E2A7C0FFD2F553B" box="[637,752,1580,1606]" pageId="10" pageNumber="571" value="2002-03-09">
<collectingDate id="EFEFE9FFFFE022410E2A7C0FFD2F553B" box="[637,752,1580,1606]" pageId="10" pageNumber="571" value="2002-03-09">9.III.2002</collectingDate>
</date>
, leg. UMC &amp; CMU teams, L-401, 
<specimenCount id="9D13FD5EFFE0224108217C0FFB10553B" box="[1142,1231,1580,1606]" pageId="10" pageNumber="571" type="male">3 males</specimenCount>
, 
<specimenCount id="9D13FD5EFFE02241088D7C0FFAE4553B" box="[1242,1339,1580,1606]" pageId="10" pageNumber="571" type="female">1 female</specimenCount>
(
<collectionCode id="ED04AE12FFE02241091D7C0FFA4F553B" box="[1354,1424,1580,1606]" pageId="10" pageNumber="571">UMC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B7D3C8AFFE022410CC07C73FA885517" box="[151,1367,1616,1642]" collectingDate="2002-05-09" collectionCode="THNHM" country="Thailand" elevation="1460" latitude="18.533333" location="Siriphum Waterfall" longLatPrecision="1275" longitude="98.51667" municipality="Doi Inthanon Natl. Park." pageId="10" pageNumber="571" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="paratype">
same locality except for, 
<location id="8ECA600CFFE022410DE67C73FD545517" LSID="urn:lsid:plazi:treatment:03BC87C1FFE022470CC07F31FB5354CE:8ECA600CFFE022410DE67C73FD545517" box="[433,651,1616,1642]" country="Thailand" name="Siriphum Waterfall" pageId="10" pageNumber="571" stateProvince="Chiang Mai">Siriphum Waterfall</location>
, 
<date id="FFAB1017FFE022410EC17C73FD225517" box="[662,765,1616,1642]" pageId="10" pageNumber="571" value="2002-05-09">
<collectingDate id="EFEFE9FFFFE022410EC17C73FD225517" box="[662,765,1616,1642]" pageId="10" pageNumber="571" value="2002-05-09">9.V.2002</collectingDate>
</date>
, leg. UMC &amp; CMU teams, L-401, 
<specimenCount id="9D13FD5EFFE0224108D07C73FB0C5517" box="[1159,1235,1616,1642]" pageId="10" pageNumber="571" type="male">1 male</specimenCount>
(
<collectionCode id="ED04AE12FFE0224108B57C73FA905517" box="[1250,1359,1616,1642]" pageId="10" pageNumber="571">THNHM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B7D3C8AFFE0224109347C73FC2855F3" collectingDate="2002-11-14" collectionCode="THNHM, UMC" country="Thailand" elevation="1460" latitude="18.533333" location="Siriphum Waterfall" longLatPrecision="1275" longitude="98.51667" municipality="Doi Inthanon Natl. Park." pageId="10" pageNumber="571" specimenCount="2" specimenCount-female="2" stateProvince="Chiang Mai" typeStatus="paratype">
same locality except for, 
<date id="FFAB1017FFE022410D277C57FE2D55F3" box="[368,498,1652,1678]" pageId="10" pageNumber="571" value="2002-11-14">
<collectingDate id="EFEFE9FFFFE022410D277C57FE2D55F3" box="[368,498,1652,1678]" pageId="10" pageNumber="571" value="2002-11-14">14.XI.2002</collectingDate>
</date>
, leg. CMU team, 
<specimenCount id="9D13FD5EFFE022410EEE7C57FCFA55F3" box="[697,805,1652,1678]" pageId="10" pageNumber="571" type="female">2 females</specimenCount>
(
<collectionCode id="ED04AE12FFE022410F647C57FC4155F3" box="[819,926,1652,1678]" pageId="10" pageNumber="571">THNHM</collectionCode>
, 
<collectionCode id="ED04AE12FFE022410FFD7C57FC3055F3" box="[938,1007,1652,1678]" pageId="10" pageNumber="571">UMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAA36D7FFE022410C907CBBFB4955AB" blockId="10.[151,1437,1400,2038]" pageId="10" pageNumber="571">
<emphasis id="B961EAC5FFE022410C907CBBFE6255CF" bold="true" box="[199,445,1688,1714]" pageId="10" pageNumber="571">Additional material.</emphasis>
<materialsCitation id="3B7D3C8AFFE022410D917CBBFB4D55AB" collectingDate="1998-12-28" collectionCode="PMI" collectorName="P. Mazzoldi" country="Thailand" elevation="1375" location="Siriphum Falls" pageId="10" pageNumber="571" specimenCount="1" stateProvince="Chiang Mai">
<emphasis id="B961EAC5FFE022410D917CBBFCA255CF" bold="true" box="[454,893,1688,1714]" pageId="10" pageNumber="571">
<collectingCountry id="F3027647FFE022410D917CBBFDBE55CF" box="[454,609,1688,1714]" name="Thailand" pageId="10" pageNumber="571">THAILAND</collectingCountry>
: 
<collectingRegion id="49D1F835FFE022410E277CBBFCA655CF" box="[624,889,1688,1714]" country="Thailand" name="Chiang Mai" pageId="10" pageNumber="571">Chiang Mai Province</collectingRegion>
:
</emphasis>
Doi Inthanon Natl. Park., streams downstream from 
<location id="8ECA600CFFE022410C827C9FFEA255AB" LSID="urn:lsid:plazi:treatment:03BC87C1FFE022470CC07F31FB5354CE:8ECA600CFFE022410C827C9FFEA255AB" box="[213,381,1724,1750]" country="Thailand" name="Siriphum Falls" pageId="10" pageNumber="571" stateProvince="Chiang Mai">Siriphum Falls</location>
, 
<elevation id="0038D1E4FFE022410DDF7C9FFD8055AB" box="[392,607,1724,1751]" metricMagnitude="3" metricUnit="m" metricValue="1.375" metricValueMax="1.4" metricValueMin="1.35" pageId="10" pageNumber="571" unit="m" value="1375.0" valueMax="1400.0" valueMin="1350.0">
<quantity id="4CED9B32FFE022410DDF7C9FFDFC55AB" box="[392,547,1724,1751]" metricMagnitude="3" metricUnit="m" metricValue="1.375" metricValueMax="1.4" metricValueMin="1.35" pageId="10" pageNumber="571" unit="m" value="1375.0" valueMax="1400.0" valueMin="1350.0">1350–1400 m</quantity>
a.s.l.
</elevation>
(10), 
<date id="FFAB1017FFE022410EF57C9FFCF255AB" box="[674,813,1724,1750]" pageId="10" pageNumber="571" value="1998-12-28">
<collectingDate id="EFEFE9FFFFE022410EF57C9FFCF255AB" box="[674,813,1724,1750]" pageId="10" pageNumber="571" value="1998-12-28">28.XII.1998</collectingDate>
</date>
, leg. 
<collectorName id="26E05301FFE022410F3E7C9FFC2F55AB" box="[873,1008,1724,1750]" pageId="10" pageNumber="571">P. Mazzoldi</collectorName>
, 21 exs (
<collectionCode id="ED04AE12FFE0224108047C9FFB5455AB" box="[1107,1163,1724,1750]" pageId="10" pageNumber="571">PMI</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFE022400C907CC3FA5553CC" lastPageId="11" lastPageNumber="572" pageId="10" pageNumber="571" type="diagnosis">
<paragraph id="8BAA36D7FFE022400C907CC3FA5553CC" blockId="10.[151,1437,1400,2038]" lastBlockId="11.[151,1437,151,501]" lastPageId="11" lastPageNumber="572" pageId="10" pageNumber="571">
<emphasis id="B961EAC5FFE022410C907CC3FE9D5587" bold="true" box="[199,322,1760,1786]" pageId="10" pageNumber="571">Diagnosis.</emphasis>
Body form in lateral view strongly dorsoventrally convex, appearing humped in scutellar region; elytral glabrous region of male (
<figureCitation id="132E2A52FFE022410E527D27FDB75463" box="[517,616,1796,1822]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="10" pageNumber="571">Figs. 2C</figureCitation>
, 
<figureCitation id="132E2A52FFE022410E227D27FD475463" box="[629,664,1796,1822]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="10" pageNumber="571">7A</figureCitation>
) cordiform, along suture extending ca. 7/9 length of elytra, laterally expanded in anterior 2/3 to considerably wider than pronotal glabrous region, posterior 1/3 strongly acuminate with rounded margins; female elytral glabrous region (
<figureCitation id="132E2A52FFE022410E957D6FFCFC541B" box="[706,803,1868,1894]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="10" pageNumber="571">Figs. 3C</figureCitation>
, 
<figureCitation id="132E2A52FFE022410F787D6FFC8F541B" box="[815,848,1868,1894]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="10" pageNumber="571">7E</figureCitation>
) dissimilar to male, ‘nycteriform’ (batwing-shaped); elytral apex of both sexes similar, evenly rounded without apicolateral sinuation, epipleural angle indistinct; male protibia (
<figureCitation id="132E2A52FFE022410CA97DB7FE8A54D3" box="[254,341,1940,1966]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="10" pageNumber="571">Fig. 4C</figureCitation>
) wedge-shaped, protibial apex truncate with weakly prominent distolateral angle; male protarsus (
<figureCitation id="132E2A52FFE022410CF77D9BFF2954AF" box="[160,246,1976,2002]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="10" pageNumber="571">Fig. 4C</figureCitation>
) over 1/2 length of protibia, protarsomeres 1–4 mostly similar in width; mesoventrite without medial pit; aedeagus (
<figureCitation id="132E2A52FFE022410D597DFFFE58548B" box="[270,391,2012,2038]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="10" pageNumber="571">Fig. 7B–D</figureCitation>
) with median lobe weakly tapered, apex weakly constricted in lateral view (
<figureCitation id="132E2A52FFE0224108B57DFFFAE4548B" box="[1250,1339,2012,2038]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="10" pageNumber="571">Fig. 7C</figureCitation>
); female reproductive tract (
<figureCitation id="132E2A52FFE122400D3A7AB4FE1A53CC" box="[365,453,151,177]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="11" pageNumber="572">Fig. 7G</figureCitation>
) with additional sclerite between gonocoxae (
<figureCitation id="132E2A52FFE122400F9F7AB4FBC253CC" box="[968,1053,151,177]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="11" pageNumber="572">Fig. 7F</figureCitation>
), fertilization duct as in 
<figureCitation id="132E2A52FFE12240097A7AB4FA5953CC" box="[1325,1414,151,177]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="11" pageNumber="572">Fig. 7H</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFE122470C907A99FAD156BC" lastPageId="12" lastPageNumber="573" pageId="11" pageNumber="572" type="description">
<paragraph id="8BAA36D7FFE122400C907A99FEE35384" blockId="11.[151,1437,151,501]" pageId="11" pageNumber="572">
<emphasis id="B961EAC5FFE122400C907A99FE8653A9" bold="true" box="[199,345,186,212]" pageId="11" pageNumber="572">Description.</emphasis>
<emphasis id="B961EAC5FFE122400D377A98FE6653A8" box="[352,441,187,213]" italics="true" pageId="11" pageNumber="572">Habitus</emphasis>
. Size. Male: TL = 12.5–14.0 mm, TW = 7.5–8.0 mm; female: TL = 12.0–13.0 mm, TW = 
<quantity id="4CED9B32FFE122400CF97AFDFEE75384" box="[174,312,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="7.5" metricValueMin="6.8" pageId="11" pageNumber="572" unit="mm" value="7.15" valueMax="7.5" valueMin="6.8">6.8–7.5 mm</quantity>
.
</paragraph>
<paragraph id="8BAA36D7FFE122400C907B20FC5F5218" blockId="11.[151,1437,151,501]" pageId="11" pageNumber="572">Body form elongate oval, more attenuated anteriorly than posteriorly, maximum width just posterior of humeral angle; in lateral view dorsoventrally strongly convex, maximum convexity posterior to scutellar region, more depressed posteriorly than anteriorly creating humped appearance.</paragraph>
<paragraph id="8BAA36D7FFE122400C907B4CFCDE5288" blockId="11.[151,1437,151,501]" pageId="11" pageNumber="572">
<emphasis id="B961EAC5FFE122400C907B4CFE9D52F4" box="[199,322,367,393]" italics="true" pageId="11" pageNumber="572">Coloration</emphasis>
(
<figureCitation id="132E2A52FFE122400D037B4CFE6752F4" box="[340,440,367,393]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="11" pageNumber="572">Figs. 2C</figureCitation>
, 
<figureCitation id="132E2A52FFE122400D917B4CFE3552F4" box="[454,490,367,393]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="11" pageNumber="572">3C</figureCitation>
). Dorsally head, pronotum, and elytra margined with golden pubescence; glabrous regions of head pronotum and elytra black with reticulation creating brassy sheen. Pronotal and elytral lateral margins dark. Venter dark, reddish-brown to black. Middle and hind legs from trochanter to tarsus and ultimate abdominal ventrite lighter in color, red to orangish-red.
</paragraph>
<caption id="DF6A665FFFE122400CC07C64FC6A558C" ID-DOI="http://doi.org/10.5281/zenodo.5042678" ID-Zenodo-Dep="5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="11" pageNumber="572" startId="11.[151,260,1607,1633]" targetBox="[172,1416,528,1577]" targetPageId="11">
<paragraph id="8BAA36D7FFE122400CC07C64FC6A558C" blockId="11.[151,1436,1607,1777]" pageId="11" pageNumber="572">
<emphasis id="B961EAC5FFE122400CC07C64FEFE551C" bold="true" box="[151,289,1607,1633]" pageId="11" pageNumber="572">FIGURE 7.</emphasis>
<taxonomicName id="4C154D54FFE122400D7E7C6BFE13551C" authority="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub, 2021" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" box="[297,460,1607,1633]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="572" phylum="Arthropoda" rank="species" species="garuda" status="sp. nov.">
<emphasis id="B961EAC5FFE122400D7E7C6BFE13551C" box="[297,460,1607,1633]" italics="true" pageId="11" pageNumber="572">Patrus garuda</emphasis>
</taxonomicName>
<emphasis id="B961EAC5FFE122400D837C64FDF1551C" bold="true" box="[468,558,1607,1633]" pageId="11" pageNumber="572">
<taxonomicNameLabel id="A25257BEFFE122400D837C64FDF1551C" box="[468,558,1607,1633]" pageId="11" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view.
</paragraph>
</caption>
<paragraph id="8BAA36D7FFE622470C907AB4FBA152F4" blockId="12.[151,1437,150,1971]" pageId="12" pageNumber="573">
<emphasis id="B961EAC5FFE622470C907AB4FEDB53CC" box="[199,260,151,177]" italics="true" pageId="12" pageNumber="573">Head</emphasis>
. Head capsule broad, convex. Labrum short, ca. 3.5× wider than long; setose medially with setae situated in large deep punctures; apex broadly rounded. Clypeus narrow, ca. 1/2 width of labrum; strongly reticulate and densely punctured, reticulation hatched in appearance with numerous short, well-impressed transverse lines. Frons similarly with strong hatched reticulation and dense punctation; pseudofrontal ridge present; furrow beneath pseudofrontal ridge very narrow, without setae. Vertex also with strongly impressed hatch reticulation and dense punctation between dorsal eyes, reticulation becoming effaced in narrow band on occipital region. Dorsal eye delimited by strong orbital furrow; distance between dorsal eyes ca. 2.75× diameter of an eye; interorbital area strongly setose.
</paragraph>
<paragraph id="8BAA36D7FFE622470C907BB0FF6350B4" blockId="12.[151,1437,150,1971]" pageId="12" pageNumber="573">
<emphasis id="B961EAC5FFE622470C907BB0FEC752D0" box="[199,280,403,429]" italics="true" pageId="12" pageNumber="573">Thorax</emphasis>
. Pronotum (
<figureCitation id="132E2A52FFE622470DF57BB0FDDC52D0" box="[418,515,403,429]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="12" pageNumber="573">Figs. 2C</figureCitation>
, 
<figureCitation id="132E2A52FFE622470E5A7BB0FDEE52D0" box="[525,561,403,429]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="12" pageNumber="573">7A</figureCitation>
) with narrow lateral margins; pronotal transverse impressed line absent; lateral pubescence anteriorly extending onto pronotal disc as far as 2/3 width of dorsal eye; pronotal glabrous region densely and uniformly punctate with small weakly impressed punctures, strongly reticulate with uniform wellimpressed hatched reticulation similar to frons and vertex. Scutellar shield visible. Protrochanter with setose line along ventral (anterior) face. Profemur with setigerous punctures along anterior margin of ventral face, posterior margin of ventral face with row of setae along basal 2/3 ending with patch of longer, denser setae. Protibia (
<figureCitation id="132E2A52FFE6224709387864FF6651F8" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="12" pageNumber="573">Fig. 4C</figureCitation>
) wedge-shaped in males, not constricted basally, ventral (inner) margin straight; distal apex truncate, distolateral angle slightly prominent. Male protarsus (
<figureCitation id="132E2A52FFE622470ED678ACFD0451D4" box="[641,731,655,681]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="12" pageNumber="573">Fig. 4C</figureCitation>
) ca. 2.5/4 length of protibia, protarsomeres with distolateral angle round, not overlapping on following tarsomere, lateral margins weakly rounded, fifth tarsomere 2× length of fourth, with lateral margins weakly rounded. Elytra (
<figureCitation id="132E2A52FFE622470EB478F4FC9A518C" box="[739,837,727,753]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="12" pageNumber="573">Figs. 2C</figureCitation>
, 
<figureCitation id="132E2A52FFE622470F0778F4FCAB518C" box="[848,884,727,753]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="12" pageNumber="573">7A</figureCitation>
) with narrow lateral margins becoming gradually widened apically; elytral apex evenly rounded, without apicolateral sinuation, epipleural angle indistinct, sutural angle weakly rounded; elytral glabrous region along suture extending ca. 7/9 length of elytra, considerably wider than pronotal glabrous region; cordiform in males, with anterior 2/3 laterally expanded and, posterior 1/3 strongly acuminate; lateral pubescence in anterior 2/3 with rounded concave margins, posterior 1/3 convex rounded through apical acumination; reticulation and punctation similar to pronotal glabrous region. Mesoventrite without medial pit.
</paragraph>
<paragraph id="8BAA36D7FFE622470C9079F0FCCB57DC" blockId="12.[151,1437,150,1971]" pageId="12" pageNumber="573">
<emphasis id="B961EAC5FFE622470C9079F0FE7E5090" box="[199,417,979,1005]" italics="true" pageId="12" pageNumber="573">Sexual dimorphism</emphasis>
. Females smaller in size, more narrowly elongate in dorsal view (
<figureCitation id="132E2A52FFE6224708C879F0FB275090" box="[1183,1272,979,1005]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="12" pageNumber="573">Fig. 3C</figureCitation>
), with weaker lateral expansion of the elytra post humeral region resulting in narrower habitus. Glabrous region of female elytra different (
<figureCitation id="132E2A52FFE622470D5F7E38FEB35748" box="[264,364,1051,1077]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="12" pageNumber="573">Figs. 3C</figureCitation>
, 
<figureCitation id="132E2A52FFE622470D2D7E38FE425748" box="[378,413,1051,1077]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="12" pageNumber="573">7E</figureCitation>
), ‘nycteriform’ (batwing-shaped), extending ca. 7/9 length of elytra, broad in anterior 4/5 occupying 10/11 elytral width, lateral margins weakly rounded, abruptly acuminate in posterior 1/5, posterolateral corners with jagged appearance, not produced into large points, posterior margin briefly straight before becoming convex near suture. Protibia of female shorter than male.
</paragraph>
<paragraph id="8BAA36D7FFE622470C907E88FC9E5670" blockId="12.[151,1437,150,1971]" pageId="12" pageNumber="573">
Male genitalia. Aedeagus (
<figureCitation id="132E2A52FFE622470DA67E88FD9957B8" box="[497,582,1195,1221]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="12" pageNumber="573">Fig. 7B</figureCitation>
) with median lobe ca. 5/6 length of parameres, weakly tapered, apical 1/6 of aedeagus acuminate with straight lateral margins, apex weakly constricted in lateral view (
<figureCitation id="132E2A52FFE6224708C07EECFB2F5794" box="[1175,1264,1231,1257]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="12" pageNumber="573">Fig. 7C</figureCitation>
), operculum as in (
<figureCitation id="132E2A52FFE622470CEB7ED0FEC95670" box="[188,278,1267,1293]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="12" pageNumber="573">Fig. 7D</figureCitation>
); parameres narrowly rounded apically (
<figureCitation id="132E2A52FFE622470E8B7ED0FCEA5670" box="[732,821,1267,1293]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="12" pageNumber="573">Fig. 7B</figureCitation>
).
</paragraph>
<paragraph id="8BAA36D7FFE622470C907F34FB545604" blockId="12.[151,1437,150,1971]" pageId="12" pageNumber="573">
Female genitalia. Reproductive tract with large sac-like spermatheca (
<figureCitation id="132E2A52FFE622470FB27F34FB9E564C" box="[997,1089,1303,1329]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="12" pageNumber="573">Fig. 7G</figureCitation>
); fertilization duct sclerotized, highly convoluted (
<figureCitation id="132E2A52FFE622470D237F18FE105628" box="[372,463,1339,1365]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="12" pageNumber="573">Fig. 7H</figureCitation>
); small sclerite present posterior to oviduct; additional round sclerite present between gonocoxae; gonocoxae narrow, apically rounded, lateral margins weakly sinuate (
<figureCitation id="132E2A52FFE62247087E7F7CFBA15604" box="[1065,1150,1375,1401]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="12" pageNumber="573">Fig. 7F</figureCitation>
).
</paragraph>
<paragraph id="8BAA36D7FFE622470C907FA1FAD156BC" blockId="12.[151,1437,150,1971]" pageId="12" pageNumber="573">
<emphasis id="B961EAC5FFE622470C907FA1FEE056E1" bold="true" box="[199,319,1410,1436]" pageId="12" pageNumber="573">Variation.</emphasis>
There is variation in size and width among the material examined. Large males also tend to be relatively broader in total body width than smaller males. Females vary primarily in total body length.
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFE622470C907FE9FD445478" pageId="12" pageNumber="573" type="diagnosis">
<paragraph id="8BAA36D7FFE622470C907FE9FD445478" blockId="12.[151,1437,150,1971]" pageId="12" pageNumber="573">
<emphasis id="B961EAC5FFE622470C907FE9FE105699" bold="true" box="[199,463,1482,1508]" pageId="12" pageNumber="573">Differential diagnosis.</emphasis>
<taxonomicName id="4C154D54FFE622470D817FE8FDA55698" authority="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub, 2021" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" box="[470,634,1483,1509]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="573" phylum="Arthropoda" rank="species" species="garuda" status="sp. nov.">
<emphasis id="B961EAC5FFE622470D817FE8FDA55698" box="[470,634,1483,1509]" italics="true" pageId="12" pageNumber="573">Patrus garuda</emphasis>
</taxonomicName>
<emphasis id="B961EAC5FFE622470ED57FE8FD035698" bold="true" box="[642,732,1483,1509]" pageId="12" pageNumber="573">
<taxonomicNameLabel id="A25257BEFFE622470ED57FE8FD035698" box="[642,732,1483,1509]" pageId="12" pageNumber="573" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to 
<taxonomicName id="4C154D54FFE622470FE77FE8FBF85698" baseAuthorityName="Regimbart" baseAuthorityYear="1892" box="[944,1063,1483,1509]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="573" phylum="Arthropoda" rank="species" species="landaisi">
<emphasis id="B961EAC5FFE622470FE77FE8FBF85698" box="[944,1063,1483,1509]" italics="true" pageId="12" pageNumber="573">P. landaisi</emphasis>
</taxonomicName>
, though being unique among the known 
<taxonomicName id="4C154D54FFE622470CBE7FCCFE835574" baseAuthorityName="Regimbart" baseAuthorityYear="1892" box="[233,348,1519,1545]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="573" phylum="Arthropoda" rank="species" species="landaisi">
<emphasis id="B961EAC5FFE622470CBE7FCCFE835574" box="[233,348,1519,1545]" italics="true" pageId="12" pageNumber="573">P. landaisi</emphasis>
</taxonomicName>
species group in 
<collectingCountry id="F3027647FFE622470E4F7FCCFDA35574" box="[536,636,1519,1545]" name="Thailand" pageId="12" pageNumber="573">Thailand</collectingCountry>
in having broadly rounded elytral apices in both sexes (
<figureCitation id="132E2A52FFE62247088B7FCCFAE35574" box="[1244,1340,1519,1545]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="12" pageNumber="573">Figs. 2C</figureCitation>
, 
<figureCitation id="132E2A52FFE6224709117FCCFAB65574" box="[1350,1385,1519,1545]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="12" pageNumber="573">3C</figureCitation>
, 
<figureCitation id="132E2A52FFE6224709247FCCFF655550" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="12" pageNumber="573">7A, 7E</figureCitation>
). The female of 
<taxonomicName id="4C154D54FFE622470D387C30FE045550" authority="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub, 2021" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" box="[367,475,1555,1581]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="573" phylum="Arthropoda" rank="species" species="garuda" status="sp. nov.">
<emphasis id="B961EAC5FFE622470D387C30FE045550" box="[367,475,1555,1581]" italics="true" pageId="12" pageNumber="573">P. garuda</emphasis>
</taxonomicName>
<emphasis id="B961EAC5FFE622470DB67C30FDE55550" bold="true" box="[481,570,1555,1581]" pageId="12" pageNumber="573">
<taxonomicNameLabel id="A25257BEFFE622470DB67C30FDE55550" box="[481,570,1555,1581]" pageId="12" pageNumber="573" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shares with that of 
<taxonomicName id="4C154D54FFE622470F447C30FC555550" baseAuthorityName="Ochs" baseAuthorityYear="1940" box="[787,906,1555,1581]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="573" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFE622470F447C30FC555550" box="[787,906,1555,1581]" italics="true" pageId="12" pageNumber="573">P. salvazai</emphasis>
</taxonomicName>
a rounded elytral apex and indistinct epipleural angle, however, the elytral glabrous region of the new species is very different from the cordiform glabrous region in 
<taxonomicName id="4C154D54FFE622470CE37C78FEF25508" baseAuthorityName="Ochs" baseAuthorityYear="1940" box="[180,301,1627,1653]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="573" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFE622470CE37C78FEF25508" box="[180,301,1627,1653]" italics="true" pageId="12" pageNumber="573">P. salvazai</emphasis>
</taxonomicName>
(
<figureCitation id="132E2A52FFE622470D6B7C78FE7F5508" box="[316,416,1627,1653]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="12" pageNumber="573">Figs. 3G</figureCitation>
, 
<figureCitation id="132E2A52FFE622470DFA7C78FE035508" box="[429,476,1627,1653]" captionStart="FIGURE 11" captionStartId="22.[151,260,1770,1796]" captionTargetBox="[159,1430,816,1737]" captionTargetId="figure-281@22.[151,1436,810,1745]" captionTargetPageId="22" captionText="FIGURE 11. Patrus salvazai: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042686" httpUri="https://zenodo.org/record/5042686/files/figure.png" pageId="12" pageNumber="573">11E</figureCitation>
). The male of 
<taxonomicName id="4C154D54FFE622470ED77C78FD325508" authority="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub, 2021" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" box="[640,749,1627,1653]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="573" phylum="Arthropoda" rank="species" species="garuda" status="sp. nov.">
<emphasis id="B961EAC5FFE622470ED77C78FD325508" box="[640,749,1627,1653]" italics="true" pageId="12" pageNumber="573">P. garuda</emphasis>
</taxonomicName>
<emphasis id="B961EAC5FFE622470EA37C78FC965508" bold="true" box="[756,841,1627,1653]" pageId="12" pageNumber="573">
<taxonomicNameLabel id="A25257BEFFE622470EA37C78FC965508" box="[756,841,1627,1653]" pageId="12" pageNumber="573" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
shares several characteristics with that of 
<taxonomicName id="4C154D54FFE6224709717C78FA435508" baseAuthorityName="Regimbart" baseAuthorityYear="1892" box="[1318,1436,1627,1653]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="573" phylum="Arthropoda" rank="species" species="landaisi">
<emphasis id="B961EAC5FFE6224709717C78FA435508" box="[1318,1436,1627,1653]" italics="true" pageId="12" pageNumber="573">P. landaisi</emphasis>
</taxonomicName>
(
<tableCitation id="C697036CFFE622470CC87C5CFF2A55E4" box="[159,245,1663,1689]" captionStart="TABLE 2" captionStartId="13.[151,239,155,179]" captionText="TABLE 2. Morphological comparison of the eight species of Patrus landaisi species group in Thailand." pageId="12" pageNumber="573">Table 2</tableCitation>
) but can be distinguished by the rounded elytral apices and aedeagal features. The aedeagus of 
<taxonomicName id="4C154D54FFE6224709787C5CFA4355E4" authority="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub, 2021" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" box="[1327,1436,1663,1689]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="573" phylum="Arthropoda" rank="species" species="garuda" status="sp. nov.">
<emphasis id="B961EAC5FFE6224709787C5CFA4355E4" box="[1327,1436,1663,1689]" italics="true" pageId="12" pageNumber="573">P. garuda</emphasis>
</taxonomicName>
<emphasis id="B961EAC5FFE622470CC07C80FF3255C0" bold="true" box="[151,237,1699,1725]" pageId="12" pageNumber="573">
<taxonomicNameLabel id="A25257BEFFE622470CC07C80FF3255C0" box="[151,237,1699,1725]" pageId="12" pageNumber="573" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
has the median lobe more strongly tapered towards the apex, which is more strongly acuminate (
<figureCitation id="132E2A52FFE62247096A7C80FA4C55C0" box="[1341,1427,1699,1725]" captionStart="FIGURE 7" captionStartId="11.[151,260,1607,1633]" captionTargetBox="[172,1416,528,1577]" captionTargetId="figure-150@11.[164,1424,522,1582]" captionTargetPageId="11" captionText="FIGURE 7. Patrus garuda sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae and medial sclerotized structure, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042678" httpUri="https://zenodo.org/record/5042678/files/figure.png" pageId="12" pageNumber="573">Fig. 7B</figureCitation>
) compared to that of 
<taxonomicName id="4C154D54FFE622470D227CE4FE35559C" baseAuthorityName="Regimbart" baseAuthorityYear="1892" box="[373,490,1735,1761]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="573" phylum="Arthropoda" rank="species" species="landaisi">
<emphasis id="B961EAC5FFE622470D227CE4FE35559C" box="[373,490,1735,1761]" italics="true" pageId="12" pageNumber="573">P. landaisi</emphasis>
</taxonomicName>
in which the median lobe is more parallel-sided and less tapered to towards the apex, with the apex much less constricted (
<figureCitation id="132E2A52FFE622470E627CC8FD525478" box="[565,653,1771,1797]" captionStart="FIGURE 8" captionStartId="15.[151,260,1622,1648]" captionTargetBox="[157,1428,588,1589]" captionTargetId="figure-195@15.[151,1436,579,1598]" captionTargetPageId="15" captionText="FIGURE 8. Patrus landaisi: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042680" httpUri="https://zenodo.org/record/5042680/files/figure.png" pageId="12" pageNumber="573">Fig. 8B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFE622470C907D2EFB975410" pageId="12" pageNumber="573" type="etymology">
<paragraph id="8BAA36D7FFE622470C907D2EFB975410" blockId="12.[151,1437,150,1971]" pageId="12" pageNumber="573">
<emphasis id="B961EAC5FFE622470C907D2EFE91545A" bold="true" box="[199,334,1805,1831]" pageId="12" pageNumber="573">Etymology.</emphasis>
This species is named after Garuda, the legendary bird-like creature of Hindu and Buddhist mythology, and national emblem of 
<collectingCountry id="F3027647FFE622470E6B7D13FD7C5437" box="[572,675,1840,1866]" name="Thailand" pageId="12" pageNumber="573">Thailand</collectingCountry>
. The name is in reference to the wing-like glabrous region of the female elytra and is treated as a noun in the nominative case, standing in opposition.
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFE622470C907D56FB5354CE" pageId="12" pageNumber="573" type="distribution">
<paragraph id="8BAA36D7FFE622470C907D56FB5354CE" blockId="12.[151,1437,150,1971]" pageId="12" pageNumber="573">
<emphasis id="B961EAC5FFE622470C907D56FEBE54F2" bold="true" box="[199,353,1909,1935]" pageId="12" pageNumber="573">Distribution.</emphasis>
This species is currently only known from two high-elevation waterfalls and associated streams (
<figureCitation id="132E2A52FFE622470CF77DBAFEDA54CE" box="[160,261,1945,1971]" captionStart="FIGURE 13" captionStartId="25.[151,260,1683,1709]" captionTargetBox="[169,1418,181,1656]" captionTargetId="figure-18@25.[169,1418,179,1659]" captionTargetPageId="25" captionText="FIGURE 13. Habitats of P. landaisi species group: (A) stream at Siriphum Waterfall, Doi Inthanon National Park, Chiang Mai Province, habitat of P. garuda sp. nov. and P. subapicalis; (B) Benjamart Suksai in habitat of P. apicalis and P. salvazai at Nam Kuen stream, Chiang Rai Province; (C) shallow stream close to Ban Thai Samakkhi, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (D) Tham Yai Waterfall Phu Kradueng National Park, Loei Province, habitat of P. phetchabunensis sp. nov.; (E) puddle in dry stream bed close to Ban Phithak Thai, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (F) P. landaisi and P. salvazai from puddle in dry stream habitat close to Ban Thai Samakkhi, Chiang Rai Province." figureDoi="http://doi.org/10.5281/zenodo.5042690" httpUri="https://zenodo.org/record/5042690/files/figure.png" pageId="12" pageNumber="573">Fig. 13A</figureCitation>
) within Doi Inthanon National Park, 
<collectingRegion id="49D1F835FFE622470EF07DBAFC4654CE" box="[679,921,1945,1971]" country="Thailand" name="Chiang Mai" pageId="12" pageNumber="573">Chiang Mai Province</collectingRegion>
, 
<collectingCountry id="F3027647FFE622470FF37DBAFBD754CE" box="[932,1032,1945,1971]" name="Thailand" pageId="12" pageNumber="573">Thailand</collectingCountry>
(
<figureCitation id="132E2A52FFE6224708407DBAFBA054CE" box="[1047,1151,1945,1971]" captionStart="FIGURE 14" captionStartId="26.[151,260,743,769]" captionTargetBox="[152,1436,179,719]" captionTargetId="figure-25@26.[151,1436,179,719]" captionTargetPageId="26" captionText="FIGURE 14. (A–B) Distribution records of the P. landaisi species group in Thailand." figureDoi="http://doi.org/10.5281/zenodo.5042692" httpUri="https://zenodo.org/record/5042692/files/figure.png" pageId="12" pageNumber="573">Fig. 14A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>