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<mods:title id="0AAE79D0DC7D13D42BF5F6D236327563">The genus Polistes Latreille, 1802 (Hymenoptera: Vespidae) of Brazil: morphological diagnoses, identification key, and geographical distribution</mods:title>
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<mods:namePart id="FEE4D52CC68DC8DA514273E654E82BEE">Brito, Samanta</mods:namePart>
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<mods:namePart id="3B4212295AFEF5CC2A6C9460FBB2D719">Oliveira, Marcio L.</mods:namePart>
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<mods:affiliation id="6D8D0B2DE4FD357577587C881595DF8F">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade, Manaus, AM, Brazil</mods:affiliation>
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<mods:namePart id="D2392114096B9D91417648A70412C64F">Carpenter, James M.</mods:namePart>
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<mods:affiliation id="72171728B8EEC84390E2F20136DBF230">American Museum of Natural History, Division of Invertebrate Zoology, New York, USA</mods:affiliation>
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<mods:namePart id="1AC26D5791860810A255CF906C66C610">Somavilla, Alexandre</mods:namePart>
<mods:affiliation id="3E1FAD3C8E41E039FE334F55C1C627D8">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade, Manaus, AM, Brazil</mods:affiliation>
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<mods:date id="34360226E1C63B4AB6D27F7972A96E9D">2024</mods:date>
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<paragraph id="8BAA36A6FFBBFFF7CCEEFD734993FD1D" blockId="14.[151,739,654,717]" box="[151,739,654,681]" pageId="14" pageNumber="523">
<heading id="D0E281CAFFBBFFF7CCEEFD734993FD1D" bold="true" box="[151,739,654,681]" fontSize="11" level="1" pageId="14" pageNumber="523" reason="1">
<taxonomicName id="4C154D25FFBBFFF7CCEEFD734993FD1D" ID-CoL="4KY85" authority="Richards, 1951" authorityName="Richards" authorityYear="1951" box="[151,739,654,681]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="523" phylum="Arthropoda" rank="species" species="angulinus" subGenus="Epicnemius">
<emphasis id="B961EAB4FFBBFFF7CCEEFD734993FD1D" bold="true" box="[151,739,654,681]" pageId="14" pageNumber="523">
<emphasis id="B961EAB4FFBBFFF7CCEEFD734B84FD1C" bold="true" box="[151,244,654,681]" italics="true" pageId="14" pageNumber="523">Polistes</emphasis>
(
<emphasis id="B961EAB4FFBBFFF7CD7CFD724AE4FD1C" bold="true" box="[261,404,655,681]" italics="true" pageId="14" pageNumber="523">Epicnemius</emphasis>
)
<emphasis id="B961EAB4FFBBFFF7CDDFFD734950FD1C" bold="true" box="[422,544,654,681]" italics="true" pageId="14" pageNumber="523">angulinus</emphasis>
<bibRefCitation id="EF844B57FFBBFFF7CE5EFD734993FD1D" author="Richards, O. W. &amp; Richards, M. J." box="[551,739,654,680]" pageId="14" pageNumber="523" pagination="1 - 170" refId="ref27784" refString="Richards, O. W. &amp; Richards, M. J. (1951) Observations on the social wasps of South America (Hymenoptera, Vespidae). Transactions of the Royal Entomological Society of London, 102, 1 - 170. https: // doi. org / 10.1111 / j. 1365 - 2311.1951. tb 01241. x" type="journal article" year="1951">Richards, 1951</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8BAA36A6FFBBFFF7CCEEFD4E4A57FD78" blockId="14.[151,739,654,717]" box="[151,295,691,717]" pageId="14" pageNumber="523">
(
<figureCitation id="132E2A23FFBBFFF7CCE6FD4E4A50FD78" box="[159,288,691,717]" captionStart="FIGURES 3744" captionStartId="15.[152,270,1896,1920]" captionTargetBox="[158,1428,189,1865]" captionTargetId="figure-17@15.[151,1436,181,1872]" captionText="FIGURES 3744. Polistes spp. 37, 38. P. (Epicnemius) actaeon Haliday. 39, 40. P. (E.) angulinus Richards. 41, 42. P. (E.) atrox Richards. 43, 44. P. (E.) bequaertianus Willink. 37, 39, 41, 43. Dorsal view. 38, 40, 42, 44. Lateral view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13736639" httpUri="https://zenodo.org/record/13736639/files/figure.png" pageId="14" pageNumber="523">Figs 39, 40</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F652DFFBBFFF7CCEEFD0A489FFCBA" box="[151,1007,759,783]" pageId="14" pageNumber="523" type="reference_group">
<paragraph id="8BAA36A6FFBBFFF7CCEEFD0A489FFCBA" blockId="14.[151,1007,759,783]" box="[151,1007,759,783]" pageId="14" pageNumber="523">
<treatmentCitationGroup id="AB051188FFBBFFF7CCEEFD0A489FFCBA" box="[151,1007,759,783]" pageId="14" pageNumber="523">
<taxonomicName id="4C154D25FFBBFFF7CCEEFD0A491EFCBA" ID-CoL="4KY85" authority="Richards" authorityName="Richards" box="[151,622,759,783]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="523" phylum="Arthropoda" rank="variety" species="erythrogaster" variety="angulinus">
<emphasis id="B961EAB4FFBBFFF7CCEEFD0A4A07FCBA" box="[151,375,759,783]" italics="true" pageId="14" pageNumber="523">Polistes erythrogaster</emphasis>
var.
<emphasis id="B961EAB4FFBBFFF7CDD3FD0A497EFCBA" box="[426,526,759,783]" italics="true" pageId="14" pageNumber="523">angulinus</emphasis>
Richards
</taxonomicName>
in
<treatmentCitation id="0AB410B7FFBBFFF7CEF6FD0A48BCFCBA" author="Richards, O. W. &amp; Richards, M. J." box="[655,972,759,783]" page="92" pageId="14" pageNumber="523" year="1951">
<bibRefCitation id="EF844B57FFBBFFF7CEF6FD0A48BCFCBA" author="Richards, O. W. &amp; Richards, M. J." box="[655,972,759,783]" pageId="14" pageNumber="523" pagination="1 - 170" refId="ref27784" refString="Richards, O. W. &amp; Richards, M. J. (1951) Observations on the social wasps of South America (Hymenoptera, Vespidae). Transactions of the Royal Entomological Society of London, 102, 1 - 170. https: // doi. org / 10.1111 / j. 1365 - 2311.1951. tb 01241. x" type="journal article" year="1951">Richards &amp; Richards, 1951: 92</bibRefCitation>
</treatmentCitation>
,
<specimenCount id="9D13FD2FFFBBFFF7CFAFFD05489AFCBA" box="[982,1002,760,783]" count="1" pageId="14" pageNumber="523" type="female"></specimenCount>
.
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</paragraph>
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<subSubSection id="C30F652DFFBBFFF7CCEEFCC04923FBC2" pageId="14" pageNumber="523" type="diagnosis">
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<emphasis id="B961EAB4FFBBFFF7CCEEFCC04A03FCE2" bold="true" box="[151,371,829,855]" pageId="14" pageNumber="523">Diagnosis.Female.</emphasis>
Body dark brown to black, with the mesepisternum lighter than the body color.Wing pigmentation hyaline and matte. Relatively small size, maximum
<quantity id="4CED9B43FFBBFFF7CE9FFC9C4847FCC9" box="[742,823,865,892]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="523" unit="mm" value="15.0">15 mm</quantity>
in total length. Clypeus longer than wide, covered by short bristles at its base becoming long toward the apex, with medium-sized and deep punctures, present only in the apical region; apex rounded, clypeus-eye touching for an extent equal to or greater than width of antennal socket, gena width at midlength equal to or slightly wider than eye; occipital carina strong, extending to middle of the eye; humeri slightly projecting; mesepisternum with short bristles, punctation small and shallow; propodeal concavity and striae absent; tergum I longer than wide, bristles short and dense, punctures strong, medium sized, and deep; tergum II as wide as long or wider than long, with short bristles, weakly punctate.
<emphasis id="B961EAB4FFBBFFF7C857FBC44F02FBE6" bold="true" box="[1070,1138,1081,1107]" pageId="14" pageNumber="523">Male.</emphasis>
Similar to females, except for the size, with
<quantity id="4CED9B43FFBBFFF7CD23FBA04AD9FBCD" box="[346,425,1117,1144]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="523" unit="mm" value="17.0">17 mm</quantity>
in total length.
</paragraph>
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<subSubSection id="C30F652DFFBBFFF7CCBEFB7C4881FB2E" box="[199,1009,1153,1180]" pageId="14" pageNumber="523" type="materials_examined">
<paragraph id="8BAA36A6FFBBFFF7CCBEFB7C4881FB2E" blockId="14.[151,1436,829,1216]" box="[199,1009,1153,1180]" pageId="14" pageNumber="523">
<emphasis id="B961EAB4FFBBFFF7CCBEFB7C4AD9FB2E" bold="true" box="[199,425,1153,1179]" pageId="14" pageNumber="523">Material examined</emphasis>
.
<materialsCitation id="3B7D3CFBFFBBFFF7CDCEFB7C4842FB2E" box="[439,818,1153,1179]" country="Brazil" location="Rondonia" pageId="14" pageNumber="523" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Rondonia">
<specimenCount id="9D13FD2FFFBBFFF7CDCEFB7C4AAAFB2E" box="[439,474,1153,1179]" count="1" pageId="14" pageNumber="523" type="female">1♀</specimenCount>
,
<specimenCount id="9D13FD2FFFBBFFF7CD91FB7C497BFB2E" box="[488,523,1153,1179]" count="1" pageId="14" pageNumber="523" type="male">1♂</specimenCount>
in total.
<collectingCountry id="F3027636FFBBFFF7CE16FB7C49C8FB2E" box="[623,696,1153,1179]" name="Brazil" pageId="14" pageNumber="523">Brazil</collectingCountry>
:
<collectingRegion id="49D1F844FFBBFFF7CEBBFB7C4842FB2E" box="[706,818,1153,1179]" country="Brazil" name="Rondonia" pageId="14" pageNumber="523">Rondônia</collectingRegion>
</materialsCitation>
,
<materialsCitation id="3B7D3CFBFFBBFFF7CF44FB7C489DFB2E" box="[829,1005,1153,1179]" collectionCode="INPA" country="Brazil" pageId="14" pageNumber="523" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Rondonia">
<specimenCount id="9D13FD2FFFBBFFF7CF44FB7C4810FB2E" box="[829,864,1153,1179]" count="1" pageId="14" pageNumber="523" type="female">1♀</specimenCount>
,
<specimenCount id="9D13FD2FFFBBFFF7CF17FB7C48E1FB2E" box="[878,913,1153,1179]" count="1" pageId="14" pageNumber="523" type="male">1♂</specimenCount>
(
<collectionCode id="ED04AE63FFBBFFF7CFD9FB7C4896FB2E" box="[928,998,1153,1179]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="14" pageNumber="523">INPA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F652DFFBBFFF7CCBEFB584868FB0A" box="[199,792,1189,1216]" pageId="14" pageNumber="523" type="distribution">
<paragraph id="8BAA36A6FFBBFFF7CCBEFB584868FB0A" blockId="14.[151,1436,829,1216]" box="[199,792,1189,1216]" pageId="14" pageNumber="523">
<emphasis id="B961EAB4FFBBFFF7CCBEFB584A11FB0A" bold="true" box="[199,353,1189,1215]" pageId="14" pageNumber="523">Distribution.</emphasis>
<collectingCountry id="F3027636FFBBFFF7CD1EFB584AC0FB0A" box="[359,432,1189,1215]" name="Brazil" pageId="14" pageNumber="523">Brazil</collectingCountry>
: Amazonas,
<collectingRegion id="49D1F844FFBBFFF7CE44FB5849DEFB0A" box="[573,686,1189,1215]" country="Brazil" name="Rondonia" pageId="14" pageNumber="523">Rondônia</collectingRegion>
;
<collectingCountry id="F3027636FFBBFFF7CEC0FB584864FB0A" box="[697,788,1189,1215]" name="Guyana" pageId="14" pageNumber="523">Guyana</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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