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<document id="B6265492ED5DED8804900B8BCE0E21AD" ID-CLB-Dataset="27769" ID-DOI="10.11646/zootaxa.4531.4.3" ID-GBIF-Dataset="c953a470-a1b6-47b0-a403-9a6a828951e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="2614887" ID-ZooBank="05141281-0CC7-451B-B459-0498EBCEDA30" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553787445763" checkinUser="plazi" docAuthor="Selis, Marco" docDate="2018" docId="03BC87A3FFD6C409FF50FA5211BA8E1B" docLanguage="en" docName="zootaxa.4531.4.3.pdf" docOrigin="Zootaxa 4531 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Polistes (Gyrostoma) nigrosericans Selis 2018, stat. nov." docType="treatment" docVersion="7" lastPageNumber="510" masterDocId="FF85FFDBFFD7C40AFFC7FFEA1226881D" masterDocTitle="Revision of the genus Polistes Latreille (Hymenoptera: Vespidae: Polistinae) in the Philippine Islands" masterLastPageNumber="531" masterPageNumber="507" pageNumber="508" updateTime="1698714915165" updateUser="ExternalLinkService">
<mods:mods id="2504AB5A7DD8B3D1D3DFCF9D67436381" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EE73BFC790480E53F44E89C89DAFD965">
<mods:title id="DF70F03DE0AAC2F69287D84DCEAF3DEB">Revision of the genus Polistes Latreille (Hymenoptera: Vespidae: Polistinae) in the Philippine Islands</mods:title>
</mods:titleInfo>
<mods:name id="6FEC2B33F1EC187DCEAEBD74795BB145" type="personal">
<mods:role id="6FDABBC11F48866CBC50B9F2C587A2B5">
<mods:roleTerm id="1680CD9B5CEDBEE3823F01BEFAF00C90">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1B15F61541F63DD985D7111FE8313372">Selis, Marco</mods:namePart>
</mods:name>
<mods:typeOfResource id="B0C662127AE9C55DB47C1DEBD248D975">text</mods:typeOfResource>
<mods:relatedItem id="9B89A30BB132B2E5CD529F4206D45E79" type="host">
<mods:titleInfo id="1678777C495CE5C6048D6C40946B96D5">
<mods:title id="EF8A557116FE035C78C5C073FD8939F8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0FE3EFFB4A934183BA3BCB08F413AADB">
<mods:date id="A31F6ACD6AC0845D12F34BFC4C6A8449">2018</mods:date>
<mods:detail id="334271401FB78893561428CE6080C353" type="pubDate">
<mods:number id="5A3AF081F4B90788BBF9D0760C76A289">2018-12-14</mods:number>
</mods:detail>
<mods:detail id="1D19C6B8F8EC3F21B7C8BCA747A838BF" type="volume">
<mods:number id="A06E534D56070B31C2A65C901E42B677">4531</mods:number>
</mods:detail>
<mods:detail id="A87A76AC8880CA97CFB43BB078AEAD3E" type="issue">
<mods:number id="430C3A38E2479E9C7B824577AF822492">4</mods:number>
</mods:detail>
<mods:extent id="0C480B3466342F3810A69CAA63BB6025" unit="page">
<mods:start id="BABB2DE0D7BF03302A8C9BE8746A237F">507</mods:start>
<mods:end id="0260D0190F94A74016D23AB63424EF6D">531</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6988ED28432753C014A640E131D92DB1">journal article</mods:classification>
<mods:identifier id="5DC8069BF2647C833901683D01E17FDC" type="CLB-Dataset">27769</mods:identifier>
<mods:identifier id="B261904F9BFFBAC90DBFCAF475626E56" type="DOI">10.11646/zootaxa.4531.4.3</mods:identifier>
<mods:identifier id="B4BFD0122E24CD93F8E8C2F7976184EF" type="GBIF-Dataset">c953a470-a1b6-47b0-a403-9a6a828951e4</mods:identifier>
<mods:identifier id="2F08DB69667800B918A7AD4598228286" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="03D2507FF9A5276D87C1DA1688A8A195" type="Zenodo-Dep">2614887</mods:identifier>
<mods:identifier id="897E613A67F77B3EED9C5125697127BC" type="ZooBank">05141281-0CC7-451B-B459-0498EBCEDA30</mods:identifier>
</mods:mods>
<treatment id="03BC87A3FFD6C409FF50FA5211BA8E1B" ID-DOI="http://doi.org/10.5281/zenodo.5958539" ID-GBIF-Taxon="154521009" ID-Zenodo-Dep="5958539" LSID="urn:lsid:plazi:treatment:03BC87A3FFD6C409FF50FA5211BA8E1B" httpUri="http://treatment.plazi.org/id/03BC87A3FFD6C409FF50FA5211BA8E1B" lastPageId="3" lastPageNumber="510" pageId="1" pageNumber="508">
<subSubSection id="C30F653EFFD6C40BFF50FA52115F8DCE" box="[151,889,1464,1491]" pageId="1" pageNumber="508" type="nomenclature">
<paragraph id="8BAA36B5FFD6C40BFF50FA52115F8DCE" blockId="1.[151,889,1464,1525]" box="[151,889,1464,1491]" pageId="1" pageNumber="508">
<heading id="D0E281D9FFD6C40BFF50FA52115F8DCE" bold="true" box="[151,889,1464,1491]" fontSize="11" level="1" pageId="1" pageNumber="508" reason="1">
<emphasis id="B961EAA7FFD6C40BFF50FA52115F8DCE" bold="true" box="[151,889,1464,1491]" pageId="1" pageNumber="508">
<taxonomicName id="4C154D36FFD6C40BFF50FA5211258DCE" authority="Bequaert, 1940" authorityName="Selis" authorityYear="2018" baseAuthorityName="Bequaert" baseAuthorityYear="1940" box="[151,771,1464,1491]" class="Insecta" family="Vespidae" genus="Polistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="species" species="nigrosericans" status="stat. nov." subGenus="Gyrostoma">
<emphasis id="B961EAA7FFD6C40BFF50FA5212D28DCF" bold="true" box="[151,244,1464,1490]" italics="true" pageId="1" pageNumber="508">Polistes</emphasis>
(
<emphasis id="B961EAA7FFD6C40BFEC3FA5213AE8DCF" bold="true" box="[260,392,1464,1490]" italics="true" pageId="1" pageNumber="508">Gyrostoma</emphasis>
)
<emphasis id="B961EAA7FFD6C40BFE5DFA5210188DCF" bold="true" box="[410,574,1464,1490]" italics="true" pageId="1" pageNumber="508">nigrosericans</emphasis>
<bibRefCitation id="EF844B44FFD6C40BFD82FA5311258DCE" author="Bequaert, J." box="[581,771,1465,1491]" pageId="1" pageNumber="508" pagination="265 - 272" refId="ref11389" refString="Bequaert, J. (1940) Notes on Oriental Polistes wasps (Hymenoptera: Vespidae). Transactions of the American Entomological Society, 66, 265 - 272." type="journal article" year="1940">Bequaert, 1940</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="A25257DCFFD6C40BFCCDFA53115F8DCE" box="[778,889,1465,1491]" pageId="1" pageNumber="508">stat. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30F653EFFD6C40BFF50FA3613348DE8" box="[151,274,1500,1525]" pageId="1" pageNumber="508" type="description">
<paragraph id="8BAA36B5FFD6C40BFF50FA3613348DE8" blockId="1.[151,889,1464,1525]" box="[151,274,1500,1525]" pageId="1" pageNumber="508">
(
<figureCitation id="132E2A30FFD6C40BFF58FA36132F8DE8" box="[159,265,1500,1525]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–6" captionStartId-0="2.[151,264,1982,2005]" captionStartId-1="3.[151,264,699,722]" captionTargetBox-0="[197,1377,206,1965]" captionTargetBox-1="[151,1436,193,678]" captionTargetId-0="figure@2.[197,1377,193,1965]" captionTargetId-1="figure@3.[151,1436,193,678]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–4. Polistes (Gyrostoma) nigrosericans Bequaert, 1940. 1, 3–4, ♂. 2, ♀. 1–2, Habitus. 3, Head in frontal view. 4, Head in lateral view." captionText-1="FIGURES 5–6. Polistes (Gyrostoma) nigrosericans Bequaert, 1940. 5, ♂. 6, ♀. 5, Apex of antenna in lateral view. 6, Propodeum in postero-dorsal view." httpUri-0="https://zenodo.org/record/2614889/files/figure.png" httpUri-1="https://zenodo.org/record/2614891/files/figure.png" pageId="1" pageNumber="508">Figs. 1–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F653EFFD6C40BFF50F9CE13E88E45" pageId="1" pageNumber="508" type="reference_group">
<paragraph id="8BAA36B5FFD6C40BFF50F9CE13E88E45" blockId="1.[151,1435,1572,1624]" pageId="1" pageNumber="508">
<treatmentCitationGroup id="AB05119BFFD6C40BFF50F9CE13E88E45" pageId="1" pageNumber="508">
<taxonomicName id="4C154D36FFD6C40BFF50F9CE13578E27" authorityName="Lepeletier" authorityYear="1836" box="[151,369,1572,1594]" class="Insecta" family="Vespidae" genus="Polistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="species" species="tenebricosus">
<emphasis id="B961EAA7FFD6C40BFF50F9CE13578E27" box="[151,369,1572,1594]" italics="true" pageId="1" pageNumber="508">Polistes tenebricosus</emphasis>
</taxonomicName>
var. (or subsp.)
<taxonomicName id="4C154D36FFD6C40BFDE5F9CF11A28E27" authority="Bequaert 1940: 265" authorityName="Bequaert" authorityPageNumber="265" authorityYear="1940" box="[546,900,1572,1594]" class="Insecta" family="Vespidae" genus="Polistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="508" phylum="Arthropoda" rank="species" species="nigrosericans">
<emphasis id="B961EAA7FFD6C40BFDE5F9CF108A8E27" box="[546,684,1573,1594]" italics="true" pageId="1" pageNumber="508">nigrosericans</emphasis>
<treatmentCitation id="0AB410A4FFD6C40BFD71F9CF11A28E27" author="Bequaert, J." box="[694,900,1572,1594]" page="265" pageId="1" pageNumber="508" year="1940">
<bibRefCitation id="EF844B44FFD6C40BFD71F9CF11778E27" author="Bequaert, J." box="[694,849,1572,1594]" pageId="1" pageNumber="508" pagination="265 - 272" refId="ref11389" refString="Bequaert, J. (1940) Notes on Oriental Polistes wasps (Hymenoptera: Vespidae). Transactions of the American Entomological Society, 66, 265 - 272." type="journal article" year="1940">Bequaert 1940</bibRefCitation>
: 265
</treatmentCitation>
</taxonomicName>
(key), 266,
<specimenCount id="9D13FD3CFFD6C40BFBCEF9CF163A8E27" box="[1033,1052,1573,1594]" pageId="1" pageNumber="508" type="female">♀</specimenCount>
,
<specimenCount id="9D13FD3CFFD6C40BFBEBF9CE161B8E27" box="[1068,1085,1572,1594]" pageId="1" pageNumber="508" type="male">♂</specimenCount>
—“Los Baños, Luzon,
<collectingCountry id="F3027625FFD6C40BFAF4F9CE13368E45" name="Philippines" pageId="1" pageNumber="508">Philippine Islands</collectingCountry>
” (
<typeStatus id="54AE8817FFD6C40BFEE0F9A813A78E45" box="[295,385,1602,1624]" pageId="1" pageNumber="508" type="holotype">holotype</typeStatus>
MCZ).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30F653EFFD6C40BFF50F96F10118F17" pageId="1" pageNumber="508" type="diagnosis">
<paragraph id="8BAA36B5FFD6C40BFF50F96F10118F17" blockId="1.[151,1436,1669,1946]" pageId="1" pageNumber="508">
<emphasis id="B961EAA7FFD6C40BFF50F96F13378E83" bold="true" box="[151,273,1669,1694]" pageId="1" pageNumber="508">Diagnosis.</emphasis>
Differentiated from other species by combination of the following characters: clypeus flattened and with rounded apical margin, gena about as broad as eye in lateral view, male apical flagellomere curved and tapering at apex, pronotal carina not lamellate, posterior face of propodeum broadly and shallowly depressed, punctation coarse, and wings orange.
</paragraph>
</subSubSection>
<subSubSection id="C30F653EFFD6C40BFF00F8FF17388F6B" pageId="1" pageNumber="508" type="materials_examined">
<paragraph id="8BAA36B5FFD6C40BFF00F8FF17388F6B" blockId="1.[151,1436,1669,1946]" pageId="1" pageNumber="508">
<emphasis id="B961EAA7FFD6C40BFF00F8FF10FA8F31" bold="true" box="[199,732,1813,1838]" pageId="1" pageNumber="508">
Material examined.
<collectingCountry id="F3027625FFD6C40BFE07F8FF105E8F33" box="[448,632,1813,1838]" name="Philippines" pageId="1" pageNumber="508">PHILIPPINES</collectingCountry>
: LUZON
</emphasis>
:
<materialsCitation id="3B7D3CE8FFD6C40BFD28F8FC16EC8F33" ID-GBIF-Occurrence="2117783225" box="[751,1226,1814,1838]" collectingDate="2014-08-02" collectionCode="MSVI" country="Philippines" location="Conner" pageId="1" pageNumber="508" specimenCount="2" specimenCount-male="2" stateProvince="Apayao">
<collectingRegion id="49D1F857FFD6C40BFD28F8FC116F8F33" box="[751,841,1814,1838]" country="Philippines" name="Apayao" pageId="1" pageNumber="508">Apayao</collectingRegion>
,
<location id="8ECA606EFFD6C40BFC9EF8FC118B8F33" LSID="urn:lsid:plazi:treatment:03BC87A3FFD6C409FF50FA5211BA8E1B:8ECA606EFFD6C40BFC9EF8FC118B8F33" box="[857,941,1814,1838]" country="Philippines" name="Conner" pageId="1" pageNumber="508" stateProvince="Apayao">Conner</location>
,
<collectingDate id="EFEFE99DFFD6C40BFC7AF8FC16688F33" box="[957,1102,1814,1838]" pageId="1" pageNumber="508" value="2014-08-02">
<date id="FFAB1075FFD6C40BFC7AF8FC16688F33" box="[957,1102,1814,1838]" pageId="1" pageNumber="508" value="2014-08-02">
VIII.2014,
<specimenCount id="9D13FD3CFFD6C40BFBF8F8FC16688F33" box="[1087,1102,1814,1838]" pageId="1" pageNumber="508" type="male">2</specimenCount>
</date>
</collectingDate>
♂ (
<collectionCode id="ED04AE70FFD6C40BFBB1F8FC16E48F33" box="[1142,1218,1814,1838]" pageId="1" pageNumber="508">MSVI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3CE8FFD6C40BFB1DF8FC13B68F4F" ID-GBIF-Occurrence="2117783208" collectingDate="2014-09-03" collectionCode="MSVI" country="Philippines" location="Conner" pageId="1" pageNumber="508" specimenCount="3" specimenCount-male="3" stateProvince="Apayao">
<collectingRegion id="49D1F857FFD6C40BFB1DF8FC17128F33" box="[1242,1332,1814,1838]" country="Philippines" name="Apayao" pageId="1" pageNumber="508">Apayao</collectingRegion>
,
<location id="8ECA606EFFD6C40BFA83F8FC17BE8F33" LSID="urn:lsid:plazi:treatment:03BC87A3FFD6C409FF50FA5211BA8E1B:8ECA606EFFD6C40BFA83F8FC17BE8F33" box="[1348,1432,1814,1838]" country="Philippines" name="Conner" pageId="1" pageNumber="508" stateProvince="Apayao">Conner</location>
,
<collectingDate id="EFEFE99DFFD6C40BFF50F8D013348F4F" box="[151,274,1850,1874]" pageId="1" pageNumber="508" value="2014-09-03">
<date id="FFAB1075FFD6C40BFF50F8D013348F4F" box="[151,274,1850,1874]" pageId="1" pageNumber="508" value="2014-09-03">
IX.2014,
<specimenCount id="9D13FD3CFFD6C40BFEC2F8D013348F4F" box="[261,274,1850,1874]" pageId="1" pageNumber="508" type="male">3</specimenCount>
</date>
</collectingDate>
♂ (
<collectionCode id="ED04AE70FFD6C40BFEFBF8D013AE8F4F" box="[316,392,1850,1874]" pageId="1" pageNumber="508">MSVI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3CE8FFD6C40BFE58F8D0114B8F4F" ID-GBIF-Occurrence="2117783246" box="[415,877,1850,1874]" collectingDate="2014-10-01" collectionCode="MSVI" country="Philippines" location="St. Ana" pageId="1" pageNumber="508" specimenCount="1" specimenCount-female="1" stateProvince="Cagayan">
<collectingRegion id="49D1F857FFD6C40BFE58F8D010228F4F" box="[415,516,1850,1874]" country="Philippines" name="Cagayan" pageId="1" pageNumber="508">Cagayan</collectingRegion>
,
<location id="8ECA606EFFD6C40BFDD3F8D010488F4F" LSID="urn:lsid:plazi:treatment:03BC87A3FFD6C409FF50FA5211BA8E1B:8ECA606EFFD6C40BFDD3F8D010488F4F" box="[532,622,1850,1874]" country="Philippines" name="St. Ana" pageId="1" pageNumber="508" stateProvince="Cagayan">St. Ana</location>
,
<collectingDate id="EFEFE99DFFD6C40BFDB9F8D010D78F4F" box="[638,753,1850,1874]" pageId="1" pageNumber="508" value="2014-10-01">
<date id="FFAB1075FFD6C40BFDB9F8D010D78F4F" box="[638,753,1850,1874]" pageId="1" pageNumber="508" value="2014-10-01">
X.2014,
<specimenCount id="9D13FD3CFFD6C40BFD24F8D010D78F4F" box="[739,753,1850,1874]" pageId="1" pageNumber="508" type="female">1</specimenCount>
</date>
</collectingDate>
♀ (
<collectionCode id="ED04AE70FFD6C40BFCDEF8D011438F4F" box="[793,869,1850,1874]" pageId="1" pageNumber="508">MSVI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3CE8FFD6C40BFCBAF8D317108F4F" ID-GBIF-Occurrence="2117783218" box="[893,1334,1849,1874]" collectingDate="2014-03-01" collectionCode="MSVI" country="Philippines" location="Banaue" pageId="1" pageNumber="508" specimenCount="1" specimenCount-female="1" stateProvince="Ifugao">
<collectingRegion id="49D1F857FFD6C40BFCBAF8D311EF8F4F" box="[893,969,1849,1874]" country="Philippines" name="Ifugao" pageId="1" pageNumber="508">Ifugao</collectingRegion>
,
<location id="8ECA606EFFD6C40BFC1FF8D016098F4C" LSID="urn:lsid:plazi:treatment:03BC87A3FFD6C409FF50FA5211BA8E1B:8ECA606EFFD6C40BFC1FF8D016098F4C" box="[984,1071,1850,1873]" country="Philippines" name="Banaue" pageId="1" pageNumber="508" stateProvince="Ifugao">Banaue</location>
,
<collectingDate id="EFEFE99DFFD6C40BFBF8F8D0169C8F4F" box="[1087,1210,1850,1874]" pageId="1" pageNumber="508" value="2014-03-01">
<date id="FFAB1075FFD6C40BFBF8F8D0169C8F4F" box="[1087,1210,1850,1874]" pageId="1" pageNumber="508" value="2014-03-01">
III.2014,
<specimenCount id="9D13FD3CFFD6C40BFB6BF8D0169C8F4F" box="[1196,1210,1850,1874]" pageId="1" pageNumber="508" type="female">1</specimenCount>
</date>
</collectingDate>
♀ (
<collectionCode id="ED04AE70FFD6C40BFB25F8D017088F4F" box="[1250,1326,1850,1874]" pageId="1" pageNumber="508">MSVI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3CE8FFD6C40BFA81F8D313D08F6B" ID-GBIF-Occurrence="2117779679" collectingDate="2014-08-01" collectionCode="MSVI" country="Philippines" location="Dindin" pageId="1" pageNumber="508" specimenCount="1" specimenCount-female="1" stateProvince="Isabela">
<collectingRegion id="49D1F857FFD6C40BFA81F8D317BE8F4F" box="[1350,1432,1849,1874]" country="Philippines" name="Isabela" pageId="1" pageNumber="508">Isabela</collectingRegion>
,
<location id="8ECA606EFFD6C40BFF50F8B712CE8F6B" LSID="urn:lsid:plazi:treatment:03BC87A3FFD6C409FF50FA5211BA8E1B:8ECA606EFFD6C40BFF50F8B712CE8F6B" box="[151,232,1885,1910]" country="Philippines" name="Dindin" pageId="1" pageNumber="508" stateProvince="Isabela">Dindin</location>
,
<collectingDate id="EFEFE99DFFD6C40BFF34F8B413588F6B" box="[243,382,1886,1910]" pageId="1" pageNumber="508" value="2014-08-01">
<date id="FFAB1075FFD6C40BFF34F8B413588F6B" box="[243,382,1886,1910]" pageId="1" pageNumber="508" value="2014-08-01">
VIII.2014,
<specimenCount id="9D13FD3CFFD6C40BFEB7F8B413588F6B" box="[368,382,1886,1910]" pageId="1" pageNumber="508" type="female">1</specimenCount>
</date>
</collectingDate>
♀ (
<collectionCode id="ED04AE70FFD6C40BFE65F8B413C88F6B" box="[418,494,1886,1910]" pageId="1" pageNumber="508">MSVI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3CE8FFD6C40BFDC6F8B710708F6B" ID-GBIF-Occurrence="2117783241" box="[513,598,1885,1910]" country="Philippines" location="Pandan" pageId="1" pageNumber="508" specimenCount="1">
<location id="8ECA606EFFD6C40BFDC6F8B710708F6B" LSID="urn:lsid:plazi:treatment:03BC87A3FFD6C409FF50FA5211BA8E1B:8ECA606EFFD6C40BFDC6F8B710708F6B" box="[513,598,1885,1910]" country="Philippines" name="Pandan" pageId="1" pageNumber="508">Pandan</location>
</materialsCitation>
,
<materialsCitation id="3B7D3CE8FFD6C40BFDA5F8B7173C8F6B" ID-GBIF-Occurrence="2117783232" box="[610,1306,1885,1910]" collectingDate="2016-08" collectionCode="LDMLC" collectorName="Ismael Lumawig" country="Philippines" location="Catanduanes" pageId="1" pageNumber="508" specimenCount="1" specimenCount-female="1" stateProvince="Catanduanes">
<collectingRegion id="49D1F857FFD6C40BFDA5F8B710D28F6B" box="[610,756,1885,1910]" country="Philippines" name="Catanduanes" pageId="1" pageNumber="508">Catanduanes</collectingRegion>
,
<collectingDate id="EFEFE99DFFD6C40BFD38F8B711568F6B" box="[767,880,1885,1910]" pageId="1" pageNumber="508" value="2016-08">
<date id="FFAB1075FFD6C40BFD38F8B711568F6B" box="[767,880,1885,1910]" pageId="1" pageNumber="508" value="2016-08">VIII.2016</date>
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, leg.
<collectorName id="26E05363FFD6C40BFC6BF8B7164A8F6B" box="[940,1132,1885,1910]" pageId="1" pageNumber="508">Ismael Lumawig</collectorName>
,
<specimenCount id="9D13FD3CFFD6C40BFBB0F8B416BC8F6B" box="[1143,1178,1886,1910]" pageId="1" pageNumber="508" type="female">1♀</specimenCount>
(
<collectionCode id="ED04AE70FFD6C40BFB6EF8B417348F6B" box="[1193,1298,1886,1910]" pageId="1" pageNumber="508">LDMLC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F653EFFD6C40BFF00F86B16388F87" box="[199,1054,1921,1946]" pageId="1" pageNumber="508" type="distribution">
<paragraph id="8BAA36B5FFD6C40BFF00F86B16388F87" blockId="1.[151,1436,1669,1946]" box="[199,1054,1921,1946]" pageId="1" pageNumber="508">
<emphasis id="B961EAA7FFD6C40BFF00F86B106E8F87" bold="true" box="[199,584,1921,1946]" pageId="1" pageNumber="508">
Distribution.
<collectingCountry id="F3027625FFD6C40BFEA0F86B10658F87" box="[359,579,1921,1946]" name="Philippines" pageId="1" pageNumber="508">Philippine Islands</collectingCountry>
:
</emphasis>
Luzon (
<bibRefCitation id="EF844B44FFD6C40BFD62F86811688F87" author="Bequaert, J." box="[677,846,1922,1946]" pageId="1" pageNumber="508" pagination="265 - 272" refId="ref11389" refString="Bequaert, J. (1940) Notes on Oriental Polistes wasps (Hymenoptera: Vespidae). Transactions of the American Entomological Society, 66, 265 - 272." type="journal article" year="1940">Bequaert 1940</bibRefCitation>
;
<bibRefCitation id="EF844B44FFD6C40BFC9DF86816308F87" author="Carpenter, J. M." box="[858,1046,1921,1946]" pageId="1" pageNumber="508" pagination="1 - 39" refId="ref11471" refString="Carpenter, J. M. (1996 b) Distributional Checklist of Species of the Genus Polistes (Hymenoptera: Vespidae; Polistinae, Polistini). American Museum Novitates, 3188, 1 - 39." type="journal article" year="1996">Carpenter 1996b</bibRefCitation>
)
</paragraph>
</subSubSection>
<caption id="DF6A663DFFD5C408FF50F854134F8FE9" httpUri="https://zenodo.org/record/2614889/files/figure.png" pageId="2" pageNumber="509" startId="2.[151,264,1982,2005]" targetBox="[197,1377,206,1965]" targetPageId="2">
<paragraph id="8BAA36B5FFD5C408FF50F854134F8FE9" blockId="2.[151,1435,1982,2036]" pageId="2" pageNumber="509">
<emphasis id="B961EAA7FFD5C408FF50F854131C8FC9" bold="true" box="[151,314,1982,2005]" pageId="2" pageNumber="509">FIGURES 1–4.</emphasis>
<taxonomicName id="4C154D36FFD5C408FE85F85511768FC8" authority="Bequaert, 1940" authorityName="Bequaert" authorityYear="1940" box="[322,848,1983,2005]" class="Insecta" family="Vespidae" genus="Polistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="509" phylum="Arthropoda" rank="species" species="nigrosericans" subGenus="Gyrostoma">
<emphasis id="B961EAA7FFD5C408FE85F85513B48FC8" box="[322,402,1983,2005]" italics="true" pageId="2" pageNumber="509">Polistes</emphasis>
(
<emphasis id="B961EAA7FFD5C408FE66F82A10378FC8" box="[417,529,1984,2005]" italics="true" pageId="2" pageNumber="509">Gyrostoma</emphasis>
)
<emphasis id="B961EAA7FFD5C408FDE7F82A108C8FC8" box="[544,682,1984,2005]" italics="true" pageId="2" pageNumber="509">nigrosericans</emphasis>
Bequaert, 1940
</taxonomicName>
. 1, 3–4, ♂. 2, ♀. 1–2, Habitus. 3, Head in frontal view. 4, Head in lateral view.
</paragraph>
</caption>
<caption id="DF6A663DFFD4C409FF50FD5113DC8AED" httpUri="https://zenodo.org/record/2614891/files/figure.png" pageId="3" pageNumber="510" startId="3.[151,264,699,722]" targetBox="[151,1436,193,678]" targetPageId="3">
<paragraph id="8BAA36B5FFD4C409FF50FD5113DC8AED" blockId="3.[151,1436,699,752]" pageId="3" pageNumber="510">
<emphasis id="B961EAA7FFD4C409FF50FD5113668ACC" bold="true" box="[151,320,699,722]" pageId="3" pageNumber="510">FIGURES 5–6.</emphasis>
<taxonomicName id="4C154D36FFD4C409FE8CFD56114A8ACF" authority="Bequaert, 1940" authorityName="Bequaert" authorityYear="1940" box="[331,876,700,722]" class="Insecta" family="Vespidae" genus="Polistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="510" phylum="Arthropoda" rank="species" species="nigrosericans" subGenus="Gyrostoma">
<emphasis id="B961EAA7FFD4C409FE8CFD5613BD8ACF" box="[331,411,700,722]" italics="true" pageId="3" pageNumber="510">Polistes</emphasis>
(
<emphasis id="B961EAA7FFD4C409FE68FD5710388ACF" box="[431,542,701,722]" italics="true" pageId="3" pageNumber="510">Gyrostoma</emphasis>
)
<emphasis id="B961EAA7FFD4C409FDF4FD57109B8ACF" box="[563,701,701,722]" italics="true" pageId="3" pageNumber="510">nigrosericans</emphasis>
Bequaert, 1940
</taxonomicName>
. 5, ♂. 6, ♀. 5, Apex of antenna in lateral view. 6, Propodeum in postero-dorsal view.
</paragraph>
</caption>
<subSubSection id="C30F653EFFD4C409FF00FCF711BA8E1B" pageId="3" pageNumber="510" type="discussion">
<paragraph id="8BAA36B5FFD4C409FF00FCF711BA8E1B" blockId="3.[151,1436,797,1542]" pageId="3" pageNumber="510">
<emphasis id="B961EAA7FFD4C409FF00FCF7131D8B2B" bold="true" box="[199,315,797,822]" pageId="3" pageNumber="510">Remarks.</emphasis>
<bibRefCitation id="EF844B44FFD4C409FE84FCF413DD8B2B" author="Bequaert, J." box="[323,507,798,822]" pageId="3" pageNumber="510" pagination="265 - 272" refId="ref11389" refString="Bequaert, J. (1940) Notes on Oriental Polistes wasps (Hymenoptera: Vespidae). Transactions of the American Entomological Society, 66, 265 - 272." type="journal article" year="1940">Bequaert (1940)</bibRefCitation>
originally described this taxon as a variety or subspecies of
<taxonomicName id="4C154D36FFD4C409FB76FCF4177D8B28" authorityName="Lepeletier" authorityYear="1836" box="[1201,1371,797,821]" class="Insecta" family="Vespidae" genus="Polistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="510" phylum="Arthropoda" rank="species" species="tenebricosus">
<emphasis id="B961EAA7FFD4C409FB76FCF4177D8B28" box="[1201,1371,797,821]" italics="true" pageId="3" pageNumber="510">P. tenebricosus</emphasis>
</taxonomicName>
, then
<bibRefCitation id="EF844B44FFD4C409FF50FCAB13478B47" author="Saito, F. &amp; Kojima, J. &amp; Ubaidillah, R. &amp; Hartini, S." box="[151,353,833,858]" pageId="3" pageNumber="510" pagination="102 - 114" refId="ref12166" refString="Saito, F., Kojima, J., Ubaidillah, R. &amp; Hartini, S. (2005) Paper wasps of the genus Polistes in Eastern Lesser Sunda Islands (Hymenoptera: Vespidae). Journal of Hymenoptera Research, 14, 102 - 114." type="journal article" year="2005">
Saito
<emphasis id="B961EAA7FFD4C409FF10FCA813368B44" box="[215,272,833,857]" italics="true" pageId="3" pageNumber="510">et al.</emphasis>
(2005)
</bibRefCitation>
synonymized it along all other subspecies stating that “the subspecies category has no place in a phylogenetic system” and considering morphological characters of the subspecies
<emphasis id="B961EAA7FFD4C409FBA5FC8F169B8B60" box="[1122,1213,869,893]" italics="true" pageId="3" pageNumber="510">sulcatus</emphasis>
<bibRefCitation id="EF844B44FFD4C409FB0DFC8F177F8B63" author="Smith, F." box="[1226,1369,869,894]" pageId="3" pageNumber="510" pagination="44 - 50" refId="ref12348" refString="Smith, F. (1852) Descriptions of some hymenopterous insects captured in India, with notes on their economy, by Ezra T. Downes, Esq., who presents them to the Honourable the East India Company. Annals and Magazine of Natural History, 2 (2), 44 - 50." type="journal article" year="1852">Smith, 1852</bibRefCitation>
only. Comparing males of
<taxonomicName id="4C154D36FFD4C409FE4EFC6010F08BBC" authorityName="Bequaert" authorityYear="1940" box="[393,726,905,929]" class="Insecta" family="Vespidae" genus="Polistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="510" phylum="Arthropoda" rank="subSpecies" species="tenebricosus" subSpecies="nigrosericans">
<emphasis id="B961EAA7FFD4C409FE4EFC6010F08BBC" box="[393,726,905,929]" italics="true" pageId="3" pageNumber="510">P. tenebricosus nigrosericans</emphasis>
</taxonomicName>
from Luzon with males of the typical form from
<collectingCountry id="F3027625FFD4C409FAD2FC6317A38BBF" box="[1301,1413,905,930]" name="Indonesia" pageId="3" pageNumber="510">Indonesia</collectingCountry>
, I concluded that there are differences that allow consideration of
<taxonomicName id="4C154D36FFD4C409FCAEFC4416938BD8" authorityName="Bequaert" authorityYear="1940" box="[873,1205,941,965]" class="Insecta" family="Vespidae" genus="Polistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="510" phylum="Arthropoda" rank="subSpecies" species="tenebricosus" subSpecies="nigrosericans">
<emphasis id="B961EAA7FFD4C409FCAEFC4416938BD8" box="[873,1205,941,965]" italics="true" pageId="3" pageNumber="510">P. tenebricosus nigrosericans</emphasis>
</taxonomicName>
as a distinct species [characters of nominotypical male
<taxonomicName id="4C154D36FFD4C409FDDAFC3810E08BF4" authorityName="Lepeletier" authorityYear="1836" box="[541,710,977,1001]" class="Insecta" family="Vespidae" genus="Polistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="510" phylum="Arthropoda" rank="species" species="tenebricosus">
<emphasis id="B961EAA7FFD4C409FDDAFC3810E08BF4" box="[541,710,977,1001]" italics="true" pageId="3" pageNumber="510">P. tenebricosus</emphasis>
</taxonomicName>
in brackets]: apical margin of clypeus weakly rounded to almost truncate [with a sharp median tooth], clypeus flattened or with a weak median depression (
<figureCitation id="132E2A30FFD4C409FB18FC1F17018C13" box="[1247,1319,1013,1038]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1982,2005]" captionTargetBox="[197,1377,206,1965]" captionTargetId="figure@2.[197,1377,193,1965]" captionTargetPageId="2" captionText="FIGURES 1–4. Polistes (Gyrostoma) nigrosericans Bequaert, 1940. 1, 3–4, ♂. 2, ♀. 1–2, Habitus. 3, Head in frontal view. 4, Head in lateral view." httpUri="https://zenodo.org/record/2614889/files/figure.png" pageId="3" pageNumber="510">Fig. 3</figureCitation>
) [clypeus longitudinally depressed and with two basal calli (
<figureCitation id="132E2A30FFD4C409FD17FBF311338C2F" box="[720,789,1049,1074]" captionStart="FIGURES 7–11" captionStartId="4.[151,264,1951,1974]" captionTargetBox="[178,1404,193,1922]" captionTargetId="figure@4.[178,1404,193,1922]" captionTargetPageId="4" captionText="FIGURES 7–11. Polistes (Gyrostoma) tenebricosus Lepeletier, 1836. 7, 11, ♀. 8–10, ♂. 7, subspecies leopoldi Bequaert, 1940. 8–11, typical form. 7, Habitus. 8, Head in frontal view. 9, Head in lateral view. 10, Apex of antenna in lateral view. 11, Propodeum in postero-dorsal view." httpUri="https://zenodo.org/record/2614893/files/figure.png" pageId="3" pageNumber="510">Fig. 8</figureCitation>
)], mandible simple [inner margin of mandible bulging in basal half], second tooth of mandible shorter than adjacent ones [three apical teeth of mandible of same length], gena in lateral view almost as broad as width of eye at ocular sinus and with a rounded but clearly visible angle on posterior margin (
<figureCitation id="132E2A30FFD4C409FEA2FB6F138D8C83" box="[357,427,1157,1182]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1982,2005]" captionTargetBox="[197,1377,206,1965]" captionTargetId="figure@2.[197,1377,193,1965]" captionTargetPageId="2" captionText="FIGURES 1–4. Polistes (Gyrostoma) nigrosericans Bequaert, 1940. 1, 3–4, ♂. 2, ♀. 1–2, Habitus. 3, Head in frontal view. 4, Head in lateral view." httpUri="https://zenodo.org/record/2614889/files/figure.png" pageId="3" pageNumber="510">Fig. 4</figureCitation>
) [gena in lateral view 1.5× as broad as width of eye at ocular sinus and with posterior margin regularly rounded (
<figureCitation id="132E2A30FFD4C409FE0EFB4310368CDF" box="[457,528,1193,1218]" captionStart="FIGURES 7–11" captionStartId="4.[151,264,1951,1974]" captionTargetBox="[178,1404,193,1922]" captionTargetId="figure@4.[178,1404,193,1922]" captionTargetPageId="4" captionText="FIGURES 7–11. Polistes (Gyrostoma) tenebricosus Lepeletier, 1836. 7, 11, ♀. 8–10, ♂. 7, subspecies leopoldi Bequaert, 1940. 8–11, typical form. 7, Habitus. 8, Head in frontal view. 9, Head in lateral view. 10, Apex of antenna in lateral view. 11, Propodeum in postero-dorsal view." httpUri="https://zenodo.org/record/2614893/files/figure.png" pageId="3" pageNumber="510">Fig. 9</figureCitation>
)], last flagellomere curved and tapering only at apex (
<figureCitation id="132E2A30FFD4C409FBB8FB4316E58CDF" box="[1151,1219,1193,1218]" captionStart="FIGURES 5–6" captionStartId="3.[151,264,699,722]" captionTargetBox="[151,1436,193,678]" captionTargetId="figure@3.[151,1436,193,678]" captionTargetPageId="3" captionText="FIGURES 5–6. Polistes (Gyrostoma) nigrosericans Bequaert, 1940. 5, ♂. 6, ♀. 5, Apex of antenna in lateral view. 6, Propodeum in postero-dorsal view." httpUri="https://zenodo.org/record/2614891/files/figure.png" pageId="3" pageNumber="510">Fig. 5</figureCitation>
) [last flagellomere straight and tapering from base to apex (
<figureCitation id="132E2A30FFD4C409FD9CFB2710898CFB" box="[603,687,1229,1254]" captionStart="FIGURES 7–11" captionStartId="4.[151,264,1951,1974]" captionTargetBox="[178,1404,193,1922]" captionTargetId="figure@4.[178,1404,193,1922]" captionTargetPageId="4" captionText="FIGURES 7–11. Polistes (Gyrostoma) tenebricosus Lepeletier, 1836. 7, 11, ♀. 8–10, ♂. 7, subspecies leopoldi Bequaert, 1940. 8–11, typical form. 7, Habitus. 8, Head in frontal view. 9, Head in lateral view. 10, Apex of antenna in lateral view. 11, Propodeum in postero-dorsal view." httpUri="https://zenodo.org/record/2614893/files/figure.png" pageId="3" pageNumber="510">Fig. 10</figureCitation>
)], T
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lateral view more strongly curved dorsally, mesosoma less densely punctured and propodeum with finer striae. Differences in females are less evident than in males, but can be summarized as follows: more robust species, anterior pronotal carina less elevated and not lamellate [anterior pronotal carina elevated and shortly lamellate], posterior face of propodeum broader than long and with a broad shallow depression (
<figureCitation id="132E2A30FFD4C409FE40FAB713F68D6B" box="[391,464,1373,1398]" captionStart="FIGURES 5–6" captionStartId="3.[151,264,699,722]" captionTargetBox="[151,1436,193,678]" captionTargetId="figure@3.[151,1436,193,678]" captionTargetPageId="3" captionText="FIGURES 5–6. Polistes (Gyrostoma) nigrosericans Bequaert, 1940. 5, ♂. 6, ♀. 5, Apex of antenna in lateral view. 6, Propodeum in postero-dorsal view." httpUri="https://zenodo.org/record/2614891/files/figure.png" pageId="3" pageNumber="510">Fig. 6</figureCitation>
) [posterior face of propodeum as long as broad and with a narrower and deeper depression (
<figureCitation id="132E2A30FFD4C409FED9FA6B13578D87" box="[286,369,1409,1434]" captionStart="FIGURES 7–11" captionStartId="4.[151,264,1951,1974]" captionTargetBox="[178,1404,193,1922]" captionTargetId="figure@4.[178,1404,193,1922]" captionTargetPageId="4" captionText="FIGURES 7–11. Polistes (Gyrostoma) tenebricosus Lepeletier, 1836. 7, 11, ♀. 8–10, ♂. 7, subspecies leopoldi Bequaert, 1940. 8–11, typical form. 7, Habitus. 8, Head in frontal view. 9, Head in lateral view. 10, Apex of antenna in lateral view. 11, Propodeum in postero-dorsal view." httpUri="https://zenodo.org/record/2614893/files/figure.png" pageId="3" pageNumber="510">Fig. 11</figureCitation>
)], T
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dorsal view as long as broad or broader than long [T1 longer than broad], T
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lateral view strongly curved dorsally [T
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lateral view weakly curved dorsally], dorsal plate of metapleura transversely striate [dorsal plate of metapleura punctate], clypeus and gena more coarsely punctured. Wings orange-yellow with orange venation [wings fuscous-hyaline with black-brown venation].
</paragraph>
</subSubSection>
</treatment>
</document>
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