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<mods:title id="2956E7EEBCAB54D92D9284B397031044">The fast-running flies (Diptera, Hybotidae, Tachydromiinae) of Singapore and adjacent regions</mods:title>
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Genus
<taxonomicName id="4C154D375D3DFFE6FD11BE6A35CC8078" ID-CoL="7NQ5" ID-ENA="626653" authority="Loew, 1859" authorityName="Loew" authorityYear="1859" box="[719,951,267,294]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA65D3DFFE6FD11BE6A355E8078" bold="true" box="[719,805,267,294]" italics="true" pageId="123" pageNumber="124">Stilpon</emphasis>
<bibRefCitation id="EF844B455D3DFFE6FCF3BE6D35CC8078" author="Loew" box="[813,951,267,294]" firstAuthor="Loew" pageId="123" pageNumber="124" pagination="1 - 50" refId="ref69704" refString="Loew H. 1859. Neue Beitrage zur Kenntniss der Dipteren. Sechster Beitrag. Programm der Koniglichen Realschule zu Meseritz 1859: 1 - 50." type="journal article" year="1859">Loew, 1859</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C30F653F5D3DFFE6FF63BE31372883BD" pageId="123" pageNumber="124" type="reference_group">
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<treatmentCitationGroup id="AB05119A5D3DFFE6FF63BE31331580E6" pageId="123" pageNumber="124">
<treatmentCitation id="0AB410A55D3DFFE6FF63BE31346E8034" author="Meigen J. W." box="[189,533,336,362]" page="343" pageId="123" pageNumber="124" year="1830">
<taxonomicName id="4C154D375D3DFFE6FF63BE31346E8034" authority="Meigen, 1830: 343" authorityName="Meigen" authorityPageNumber="343" authorityYear="1830" box="[189,533,336,362]" class="Insecta" family="Hybotidae" genus="Agatachys" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA65D3DFFE6FF63BE31374C8034" box="[189,311,336,362]" italics="true" pageId="123" pageNumber="124">Agatachys</emphasis>
<bibRefCitation id="EF844B455D3DFFE6FEE2BE31346E8034" author="Meigen" box="[316,533,336,362]" firstAuthor="Meigen" pageId="123" pageNumber="124" refId="ref69871" refString="Meigen J. W. 1830. Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Sechster Theil. Schulz, Hamm." type="book" year="1830">Meigen, 1830: 343</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(as MS name '
<taxonomicName id="4C154D375D3DFFE6FD66BE3135EA8034" authorityName="Meigen" authorityYear="1830" box="[696,913,336,362]" class="Insecta" family="Hybotidae" genus="Agatachys" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="B961EAA65D3DFFE6FD66BE3135EA8034" box="[696,913,336,362]" italics="true" pageId="123" pageNumber="124">Agatachys flavipes</emphasis>
</taxonomicName>
' of Winthem).Type-species:
<taxonomicName id="4C154D375D3DFFE6FB00BE3137A580CF" ID-CoL="8FC9J" authority="Meigen, 1830" authorityName="Meigen" authorityYear="1830" class="Insecta" family="Hybotidae" genus="Tachydromia" kingdom="Animalia" order="Diptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="celeripes">
<emphasis id="B961EAA65D3DFFE6FB00BE31375C80CF" italics="true" pageId="123" pageNumber="124">Tachydromia celeripes</emphasis>
<bibRefCitation id="EF844B455D3DFFE6FEEABE1637A580CF" author="Meigen" box="[308,478,375,401]" firstAuthor="Meigen" pageId="123" pageNumber="124" refId="ref69871" refString="Meigen J. W. 1830. Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Sechster Theil. Schulz, Hamm." type="book" year="1830">Meigen, 1830</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="4C154D375D3DFFE6FDCCBE19352E80CF" ID-CoL="8FCBL" authority="Fallen, 1815" authorityName="Fallen" authorityYear="1815" box="[530,853,374,401]" class="Insecta" family="Hybotidae" genus="Tachydromia" kingdom="Animalia" order="Diptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="graminum">
<emphasis id="B961EAA65D3DFFE6FDCCBE1934D580CF" box="[530,686,376,401]" italics="true" pageId="123" pageNumber="124">T. graminum</emphasis>
<bibRefCitation id="EF844B455D3DFFE6FD65BE16352E80CF" author="Fallen" box="[699,853,374,401]" firstAuthor="Fallen" pageId="123" pageNumber="124" refId="ref68744" refString="Fallen C. F 1815. Empididae Sveciae. Lundae. 16 pp." type="book" year="1815">Fallén, 1815</bibRefCitation>
</taxonomicName>
) [designation by
<treatmentCitation id="0AB410A55D3DFFE6FBEABE16333B80CF" author="Coquillett D. W." box="[1076,1344,374,401]" page="504" pageId="123" pageNumber="124" year="1910">
<bibRefCitation id="EF844B455D3DFFE6FBEABE16333B80CF" author="Coquillett" box="[1076,1344,374,401]" firstAuthor="Coquillett" pageId="123" pageNumber="124" pagination="124 - 131" refId="ref68240" refString="Coquillett D. W. 1910. New genera and species of North American Diptera. Proceedings of the Entomological Society of Washington 12: 124 - 131." type="journal article" year="1910">Coquillett, 1910: 504</bibRefCitation>
</treatmentCitation>
], by monotypy. Suppressed by the I.C.Z.N. (1997: 200 [Opinion 1881]), see
<bibRefCitation id="EF844B455D3DFFE6FBD7BEFC331180E6" author="Cumming &amp; Evenhuis" box="[1033,1386,413,440]" firstAuthor="Cumming" pageId="123" pageNumber="124" pagination="104 - 105" refId="ref68421" refString="Cumming J. M. &amp; Evenhuis N. L. 1996. Stilpon Loew, 1859 (Insecta, Diptera): proposed conservation. Bulletin of Zoological Nomenclature 53: 104 - 105." type="journal article" year="1996">Cumming &amp; Evenhuis (1996)</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA36B45D3DFFE6FF63BEA53486835B" blockId="123.[189,1399,336,671]" pageId="123" pageNumber="124">
<treatmentCitationGroup id="AB05119A5D3DFFE6FF63BEA53486835B" pageId="123" pageNumber="124">
<treatmentCitation id="0AB410A55D3DFFE6FF63BEA537B28080" author="Loew H." box="[189,457,452,479]" page="34" pageId="123" pageNumber="124" year="1859">
<taxonomicName id="4C154D375D3DFFE6FF63BEA537B28080" ID-CoL="7NQ5" authority="Loew, 1859: 34" authorityName="Loew" authorityPageNumber="34" authorityYear="1859" box="[189,457,452,479]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA65D3DFFE6FF63BEA5376B8080" box="[189,272,452,478]" italics="true" pageId="123" pageNumber="124">Stilpon</emphasis>
<bibRefCitation id="EF844B455D3DFFE6FEC8BEA537B28080" author="Loew" box="[278,457,452,479]" firstAuthor="Loew" pageId="123" pageNumber="124" pagination="1 - 50" refId="ref69704" refString="Loew H. 1859. Neue Beitrage zur Kenntniss der Dipteren. Sechster Beitrag. Programm der Koniglichen Realschule zu Meseritz 1859: 1 - 50." type="journal article" year="1859">Loew, 1859: 34</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(as subgenus of
<taxonomicName id="4C154D375D3DFFE6FD59BEA435298081" ID-CoL="486C" authority="Meigen" authorityName="Meigen" authorityYear="1822" box="[647,850,452,479]" class="Insecta" family="Hybotidae" genus="Drapetis" kingdom="Animalia" order="Diptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA65D3DFFE6FD59BEA434958080" box="[647,750,453,478]" italics="true" pageId="123" pageNumber="124">Drapetis</emphasis>
Meigen
</taxonomicName>
). Type-species:
<taxonomicName id="4C154D375D3DFFE6FBD7BEA53681835B" ID-CoL="8FCBL" authority="Fallen, 1815" authorityName="Fallen" authorityYear="1815" class="Insecta" family="Hybotidae" genus="Tachydromia" kingdom="Animalia" order="Diptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="graminum">
<emphasis id="B961EAA65D3DFFE6FBD7BEA5335A8080" box="[1033,1313,452,478]" italics="true" pageId="123" pageNumber="124">Tachydromia graminum</emphasis>
<bibRefCitation id="EF844B455D3DFFE6FAF8BEA53681835B" author="Fallen" firstAuthor="Fallen" pageId="123" pageNumber="124" refId="ref68744" refString="Fallen C. F 1815. Empididae Sveciae. Lundae. 16 pp." type="book" year="1815">Fallén, 1815</bibRefCitation>
</taxonomicName>
, by subsequent designation
<treatmentCitation id="0AB410A55D3DFFE6FD9DBE8A3495835B" author="Loew H." box="[579,750,490,517]" page="5" pageId="123" pageNumber="124" year="1864">
<bibRefCitation id="EF844B455D3DFFE6FD9DBE8A3495835B" author="Loew" box="[579,750,490,517]" firstAuthor="Loew" pageId="123" pageNumber="124" pagination="1 - 60" refId="ref69731" refString="Loew H. 1864. Ueber die schlesischen Arten der Gattungen Tachypeza Meig. (Tachypeza, Tachista, Dysaletria) und Microphorus Macq. (Trichina und Microphorus). Zeitschrift fur Entomologie 14: 1 - 60." type="journal article" year="1864">Loew, 1864: 5</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA36B45D3DFFE6FF63BD703724830C" blockId="123.[189,1399,336,671]" pageId="123" pageNumber="124">
<treatmentCitationGroup id="AB05119A5D3DFFE6FF63BD703724830C" pageId="123" pageNumber="124">
<treatmentCitation id="0AB410A55D3DFFE6FF63BD7034658375" author="Dahl F." box="[189,542,529,555]" page="362" pageId="123" pageNumber="124" year="1909">
<taxonomicName id="4C154D375D3DFFE6FF63BD7034658375" authority="Dahl, 1909: 362" authorityName="Dahl" authorityPageNumber="362" authorityYear="1909" box="[189,542,529,555]" class="Insecta" family="Hybotidae" genus="Tetraneurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA65D3DFFE6FF63BD7037208375" box="[189,347,529,555]" italics="true" pageId="123" pageNumber="124">Tetraneurella</emphasis>
<bibRefCitation id="EF844B455D3DFFE6FEBEBD7034658375" author="Dahl" box="[352,542,529,555]" firstAuthor="Dahl" pageId="123" pageNumber="124" pagination="360 - 377" refId="ref68596" refString="Dahl F. 1909. Die Gattung Limosina und die bioconotische Forschung. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin 1909: 360 - 377." type="journal article" year="1909">Dahl, 1909: 362</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Type-species:
<taxonomicName id="4C154D375D3DFFE6FD13BD7335BE8375" authority="Dahl, 1909" authorityName="Dahl" authorityYear="1909" box="[717,965,529,555]" class="Insecta" family="Hybotidae" genus="Tetraneurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="beckeri">
<emphasis id="B961EAA65D3DFFE6FD13BD7335458375" box="[717,830,529,555]" italics="true" pageId="123" pageNumber="124">T. beckeri</emphasis>
<bibRefCitation id="EF844B455D3DFFE6FC9ABD7035BE8375" author="Dahl" box="[836,965,529,555]" firstAuthor="Dahl" pageId="123" pageNumber="124" pagination="360 - 377" refId="ref68596" refString="Dahl F. 1909. Die Gattung Limosina und die bioconotische Forschung. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin 1909: 360 - 377." type="journal article" year="1909">Dahl, 1909</bibRefCitation>
</taxonomicName>
[=
<taxonomicName id="4C154D375D3DFFE6FC34BD7033118375" ID-CoL="8F6GR" authority="(Fallen, 1915)" baseAuthorityName="Fallen" baseAuthorityYear="1915" box="[1002,1386,528,555]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="graminum">
<emphasis id="B961EAA65D3DFFE6FC34BD7032C78375" box="[1002,1212,529,555]" italics="true" pageId="123" pageNumber="124">Stilpon graminum</emphasis>
(Fallén, 1915)
</taxonomicName>
], by monotypy.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA36B45D3DFFE6FF63BD3F373783C1" blockId="123.[189,1399,336,671]" pageId="123" pageNumber="124">
<treatmentCitationGroup id="AB05119A5D3DFFE6FF63BD3F373783C1" pageId="123" pageNumber="124">
<treatmentCitation id="0AB410A55D3DFFE6FF63BD3F34308326" author="Seguy E." box="[189,587,606,633]" page="83" pageId="123" pageNumber="124" year="1950">
<taxonomicName id="4C154D375D3DFFE6FF63BD3F34308326" authority="Seguy, 1950: 83" authorityName="Seguy" authorityPageNumber="83" authorityYear="1950" box="[189,587,606,633]" class="Insecta" family="Hybotidae" genus="Pseudostilpon" kingdom="Animalia" order="Diptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA65D3DFFE6FF63BD3F37188326" box="[189,355,606,632]" italics="true" pageId="123" pageNumber="124">Pseudostilpon</emphasis>
<bibRefCitation id="EF844B455D3DFFE6FEABBD3F34308326" author="Seguy" box="[373,587,606,633]" firstAuthor="Seguy" pageId="123" pageNumber="124" pagination="83 - 87" refId="ref70338" refString="Seguy E. 1950. Un nouveau genre de Corynetine du Midi de la France (Dipt., Empididae). Vie et milieu 1: 83 - 87." type="journal article" year="1950">Séguy, 1950: 83</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Type-species:
<taxonomicName id="4C154D375D3DFFE6FCCDBD3F32AE8327" authority="Perris, 1852" authorityName="Perris" authorityYear="1852" box="[787,1237,606,633]" class="Insecta" family="Hybotidae" genus="Tachydromia" kingdom="Animalia" order="Diptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="paludosa">
<emphasis id="B961EAA65D3DFFE6FCCDBD3F32528326" box="[787,1065,606,632]" italics="true" pageId="123" pageNumber="124">Tachydromia paludosa</emphasis>
Perris, 1852
</taxonomicName>
, by original designation.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA36B45D3DFFE6FF63BDA8372883BD" blockId="123.[189,339,713,739]" box="[189,339,713,739]" pageId="123" pageNumber="124">
<heading id="D0E281D85D3DFFE6FF63BDA8372883BD" bold="true" box="[189,339,713,739]" fontSize="11" level="3" pageId="123" pageNumber="124" reason="3">
<emphasis id="B961EAA65D3DFFE6FF63BDA8372883BD" bold="true" box="[189,339,713,739]" pageId="123" pageNumber="124">Recognition</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30F653F5D3DFFE6FF63BD9A375482EE" pageId="123" pageNumber="124" type="description">
<paragraph id="8BAA36B45D3DFFE6FF63BD9A375482EE" blockId="123.[189,1398,763,944]" pageId="123" pageNumber="124">
<taxonomicName id="4C154D375D3DFFE6FF63BD9A376B824B" authorityName="Loew" authorityYear="1859" box="[189,272,763,789]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA65D3DFFE6FF63BD9A376B824B" box="[189,272,763,789]" italics="true" pageId="123" pageNumber="124">Stilpon</emphasis>
</taxonomicName>
is recognized from other drapetine genera that occur in the Oriental Region by the following combination of features: eyes contiguous on face, frons with sides nearly parallel; antenna with dorsoapical arista-like stylus; wing with cell br much shorter than cell bm, A
<subScript id="179134F15D3DFFE6FBB8BC3B32148236" attach="left" box="[1126,1135,858,872]" fontSize="6" pageId="123" pageNumber="124">1</subScript>
very weak or absent; abdominal tergites lacking squamiform setae; male terminalia with single rod-shaped ejaculatory apodeme.
</paragraph>
</subSubSection>
<subSubSection id="C30F653F5D3DFFE6FF63BCBB344286B5" pageId="123" pageNumber="124" type="diagnosis">
<paragraph id="8BAA36B45D3DFFE6FF63BCBB374D82AA" blockId="123.[189,310,986,1012]" box="[189,310,986,1012]" pageId="123" pageNumber="124">
<heading id="D0E281D85D3DFFE6FF63BCBB374D82AA" bold="true" box="[189,310,986,1012]" fontSize="11" level="3" pageId="123" pageNumber="124" reason="3">
<emphasis id="B961EAA65D3DFFE6FF63BCBB374D82AA" bold="true" box="[189,310,986,1012]" pageId="123" pageNumber="124">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAA36B45D3DFFE6FF63BB6C344286B5" blockId="123.[189,1399,1036,2027]" pageId="123" pageNumber="124">
Very small flies, usually 1.0-
<quantity id="4CED9B515D3DFFE6FDD3BB6D34138579" box="[525,616,1036,1063]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="123" pageNumber="124" unit="mm" value="1.5">1.5 mm</quantity>
(rarely 2.0-
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). Eyes contiguous on face. Ommatidia slightly enlarged below antennae. Frons linear to sublinear, narrow to fairly wide. Face strongly convex. Gena barely extended below eye.Ocellar tubercle with 2pairs of bristles.One pair of prominent inclinate vertical bristles. Antennae with pedicel large and globose, bearing long ventral preapical seta; postpedicel small, ovate, with very prominent dorsoapical extension; stylus dorsoapical, long. Palpus elongate-ovate, with distinct apical seta. Thorax blackish brown to yellow in ground-colour. Scutum shiney or tomentose; anepisternum largely shiney. Thoracic bristles mostly only slightly prominent. Metaepimeron large. Halter with knob yellow to black, rarely absent. Wing normally developed or sometimes shortened; nearly hyaline or with distinct pattern; Rs originating halfway along R
<subScript id="179134F15D3DFFE6FC3EBA323596843F" attach="left" box="[992,1005,1363,1377]" fontSize="6" pageId="123" pageNumber="124">1;</subScript>
vein R
<subScript id="179134F15D3DFFE6FBE0BA323222843F" attach="left" box="[1086,1113,1363,1377]" fontSize="6" pageId="123" pageNumber="124">2+3</subScript>
complete or incomplete; cell br short, distinctly less than length of cell bm; crossvein bm-cu nearly transverse; A
<subScript id="179134F15D3DFFE6FB18BA1832B484D9" attach="left" box="[1222,1231,1401,1415]" fontSize="6" pageId="123" pageNumber="124">1</subScript>
and crossvein CuA
<subScript id="179134F15D3DFFE6FF2ABAC1368684F0" attach="left" box="[244,253,1440,1454]" fontSize="6" pageId="123" pageNumber="124">2</subScript>
absent. Legs short, often with distinct colour pattern; fore femur thickened; mid femur slender to barely thickened, usually armed with bristles and spinules or spines arranged in specific patterns; fore tibia more or less spindle-like; mid tibia often armed with ventral spinules; hind tibia slender, lacking prominent bristles, rarely with modified posterior apical comb. Abdomen with segments 1-7 lightly sclerotised, rarely segments 1-2 modified; squamiform setae absent. Gland-like intersegmental structures present or absent. Male terminalia asymmetrical, rotated 90° to right. Epandrium completely divided. Left epandrial lamella small and fused to hypandrium. Left surstylus divided into 3 (or 4) lobes; upper lobe with or without surstylar comb. Right epandrial lamella usually large, positioned ventrally. Right surstylus large to moderately large, undivided, sometimes with apical spines. Cerci, including subepandrial sclerite, fused together basally, rarely fused completely into one large lobe, sometimes greatly reduced and bearing apical spines. Phallus elongate, well sclerotised, hair-like or, rarely, straight or very short and rather weakly sclerotised; single rod-shaped ejaculatory apodeme present. Female similar to male except ordinary setation on mid legs and unmodified abdominal segments 1-2; abdomen always without gland-like structures; terminalia short to elongate; tergite 8 not fused laterally with sternite 8; sternite 8 entire, or with apex hinged and partly or completely separated from base. Cercus elongate-ovate or broadly ovate.
</paragraph>
</subSubSection>
<subSubSection id="C30F653F5D3AFFE1FF63BE6A35D2857E" pageId="124" pageNumber="125" type="discussion">
<paragraph id="8BAA36B45D3AFFE1FF63BE6A3754807B" blockId="124.[189,303,267,293]" box="[189,303,267,293]" pageId="124" pageNumber="125">
<heading id="D0E281D85D3AFFE1FF63BE6A3754807B" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="124" pageNumber="125" reason="3">
<emphasis id="B961EAA65D3AFFE1FF63BE6A3754807B" bold="true" box="[189,303,267,293]" pageId="124" pageNumber="125">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BAA36B45D3AFFE1FF63BE5A348A80FE" blockId="124.[189,1399,315,416]" pageId="124" pageNumber="125">
The species of the genus
<taxonomicName id="4C154D375D3AFFE1FE30BE5A343A800B" authorityName="Loew" authorityYear="1859" box="[494,577,315,341]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA65D3AFFE1FE30BE5A343A800B" box="[494,577,315,341]" italics="true" pageId="124" pageNumber="125">Stilpon</emphasis>
</taxonomicName>
are distributed in the Afrotropical (2 species), Nearctic (13 species), Oriental (26 species) and Palaearctic (12 species) regions, with 54 currently known species (including 5 new species recognised herein from
<collectingCountry id="F30276245D3AFFE1FDB7BEE4349E80FE" box="[617,741,389,416]" name="Singapore" pageId="124" pageNumber="125">Singapore</collectingCountry>
).
</paragraph>
<paragraph id="8BAA36B45D3AFFE1FF63BEA835C183C3" blockId="124.[189,1399,456,669]" pageId="124" pageNumber="125">
<bibRefCitation id="EF844B455D3AFFE1FF63BEA8372680BD" author="Smith" box="[189,349,456,483]" firstAuthor="Smith" pageId="124" pageNumber="125" pagination="63 - 112" refId="ref71017" refString="Smith K. G. V. 1965. Diptera from Nepal. Empididae. Bulletin of the British Museum (Natural History), Entomology 17: 63 - 112." type="journal article" year="1965">Smith (1965)</bibRefCitation>
described the first Oriental species
<taxonomicName id="4C154D375D3AFFE1FCC8BEA8359D80BD" authorityName="Smith" authorityYear="1965" box="[790,998,457,483]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="divergens">
<emphasis id="B961EAA65D3AFFE1FCC8BEA8359D80BD" box="[790,998,457,483]" italics="true" pageId="124" pageNumber="125">Stilpon divergens</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF844B455D3AFFE1FC26BEA8330D80BA" author="Shamshev &amp; Grootaert" box="[1016,1398,457,484]" firstAuthor="Shamshev" pageId="124" pageNumber="125" pagination="315 - 346" refId="ref70430" refString="Shamshev I. V. &amp; Grootaert P. 2004 b. A review of the genus Stilpon Loew (Diptera, Empidoidea, Hybotidae) from the Oriental region. The Raffles Bulletin of Zoology 52: 315 - 346." type="journal article" year="2004" yearSuffix="b">Shamshev &amp; Grootaert (2004b)</bibRefCitation>
published 15 new species of
<taxonomicName id="4C154D375D3AFFE1FDC4BE8F34168356" authorityName="Loew" authorityYear="1859" box="[538,621,494,520]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA65D3AFFE1FDC4BE8F34168356" box="[538,621,494,520]" italics="true" pageId="124" pageNumber="125">Stilpon</emphasis>
</taxonomicName>
found in northeastern
<collectingCountry id="F30276245D3AFFE1FCA1BE8F359C8356" box="[895,999,494,520]" name="Thailand" pageId="124" pageNumber="125">Thailand</collectingCountry>
above
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altitude. Two short papers followed: first with two new species from
<collectingCountry id="F30276245D3AFFE1FCDBBD7235058373" box="[773,894,531,557]" name="Cambodia" pageId="124" pageNumber="125">Cambodia</collectingCountry>
(
<bibRefCitation id="EF844B455D3AFFE1FC53BD7232988370" author="Shamshev &amp; Grootaert" box="[909,1251,531,558]" firstAuthor="Shamshev" pageId="124" pageNumber="125" pagination="4031 - 4065" refId="ref69126" refString="Grootaert P. &amp; Shamshev I. 2006. The genus Platypalpus Macquart (Diptera: Hybotidae) from Northeast Thailand with comments on the species groups in the Oriental region. Journal of Natural History 39 (47): 4031 - 4065. http: // dx. doi. org / 10.1080 / 00222930500533781" type="journal article" year="2006">Shamshev &amp; Grootaert 2006</bibRefCitation>
) and second with two new species from southern
<collectingCountry id="F30276245D3AFFE1FD84BD5934DB830C" box="[602,672,568,594]" name="China" pageId="124" pageNumber="125">China</collectingCountry>
(
<bibRefCitation id="EF844B455D3AFFE1FD72BD5935DD830D" author="Shamshev I. V. &amp; Grootaert P." box="[684,934,568,595]" journalOrPublisher="European Journal of Entomology" pageId="124" pageNumber="125" pagination="107 - 118" part="102" refId="ref70472" refString="Shamshev I. V. &amp; Grootaert P. 2005 a. Eothalassius, a new genus of parathalasiine flies (Diptera, Empidoidea, Dolichopodidae) from Southeast Asia and Papua New Guinea. European Journal of Entomology 102: 107 - 118." title="Eothalassius, a new genus of parathalasiine flies (Diptera, Empidoidea, Dolichopodidae) from Southeast Asia and Papua New Guinea" type="journal article" year="2005">
Shamshev
<emphasis id="B961EAA65D3AFFE1FCF4BD583519830C" box="[810,866,568,594]" italics="true" pageId="124" pageNumber="125">et al.</emphasis>
2005
</bibRefCitation>
).
<bibRefCitation id="EF844B455D3AFFE1FC68BD593288830C" author="Cumming &amp; Cooper" box="[950,1267,568,594]" firstAuthor="Cumming" pageId="124" pageNumber="125" pagination="951 - 998" refId="ref68318" refString="Cumming J. M. &amp; Cooper B. E. 1992. A revision of the Nearctic species of the Tachydromiine fly genus Stilpon Loew (Diptera: Empidoidea). The Canadian Entomologist 124 (6): 951 - 998. http: // dx. doi. org / 10.4039 / Ent 124951 - 6" type="journal article" year="1992">Cumming &amp; Cooper (1992)</bibRefCitation>
recognised three informal species groups of
<taxonomicName id="4C154D375D3AFFE1FDE2BD3C34F48329" authorityName="Loew" authorityYear="1859" box="[572,655,605,631]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA65D3AFFE1FDE2BD3C34F48329" box="[572,655,605,631]" italics="true" pageId="124" pageNumber="125">Stilpon</emphasis>
</taxonomicName>
(
<taxonomicName id="4C154D375D3AFFE1FD41BD3C35698329" authorityName="Loew" authorityYear="1862" box="[671,786,605,631]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="varipes">
<emphasis id="B961EAA65D3AFFE1FD41BD3C35698329" box="[671,786,605,631]" italics="true" pageId="124" pageNumber="125">S. varipes</emphasis>
</taxonomicName>
group,
<taxonomicName id="4C154D375D3AFFE1FCB2BD3C32798329" baseAuthorityName="Fallen" baseAuthorityYear="1915" box="[876,1026,605,631]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="graminum">
<emphasis id="B961EAA65D3AFFE1FCB2BD3C32798329" box="[876,1026,605,631]" italics="true" pageId="124" pageNumber="125">S. graminum</emphasis>
</taxonomicName>
group, and
<taxonomicName id="4C154D375D3AFFE1FB52BD3C33618329" authorityName="Smith" authorityYear="1965" box="[1164,1306,605,631]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="divergens">
<emphasis id="B961EAA65D3AFFE1FB52BD3C33618329" box="[1164,1306,605,631]" italics="true" pageId="124" pageNumber="125">S. divergens</emphasis>
</taxonomicName>
group), and
<bibRefCitation id="EF844B455D3AFFE1FF31BDE3341A83C3" author="Shamshev &amp; Grootaert" box="[239,609,642,669]" firstAuthor="Shamshev" pageId="124" pageNumber="125" pagination="315 - 346" refId="ref70430" refString="Shamshev I. V. &amp; Grootaert P. 2004 b. A review of the genus Stilpon Loew (Diptera, Empidoidea, Hybotidae) from the Oriental region. The Raffles Bulletin of Zoology 52: 315 - 346." type="journal article" year="2004" yearSuffix="b">Shamshev &amp; Grootaert (2004b)</bibRefCitation>
added the
<taxonomicName id="4C154D375D3AFFE1FD3FBDE2351D83C2" authorityName="Shamshev &amp; Grootaert" authorityYear="2004" box="[737,870,642,669]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="seeluang">
<emphasis id="B961EAA65D3AFFE1FD3FBDE2351D83C2" box="[737,870,642,669]" italics="true" pageId="124" pageNumber="125">S. seeluang</emphasis>
</taxonomicName>
group.
</paragraph>
<paragraph id="8BAA36B45D3AFFE1FF63BDA435428211" blockId="124.[189,1398,709,847]" pageId="124" pageNumber="125">
Species of
<taxonomicName id="4C154D375D3AFFE1FEE0BDA437EA8381" authorityName="Loew" authorityYear="1859" box="[318,401,709,735]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA65D3AFFE1FEE0BDA437EA8381" box="[318,401,709,735]" italics="true" pageId="124" pageNumber="125">Stilpon</emphasis>
</taxonomicName>
are quite rare at low altitudes in the tropics as is demonstrated here too. On the other hand they are quite common at higher altitudes as is shown by the material collected during the TIGER project in
<collectingCountry id="F30276245D3AFFE1FEE9BC6E37D88277" box="[311,419,783,809]" name="Thailand" pageId="124" pageNumber="125">Thailand</collectingCountry>
. In the latter survey project,
<taxonomicName id="4C154D375D3AFFE1FD25BC7135358274" authorityName="Loew" authorityYear="1859" box="[763,846,784,810]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA65D3AFFE1FD25BC7135358274" box="[763,846,784,810]" italics="true" pageId="124" pageNumber="125">Stilpon</emphasis>
</taxonomicName>
is as common and diverse as
<taxonomicName id="4C154D375D3AFFE1FB69BC7133388274" authorityName="Macquart" authorityYear="1827" box="[1207,1347,784,810]" class="Insecta" family="Hybotidae" genus="Platypalpus" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA65D3AFFE1FB69BC7133388274" box="[1207,1347,784,810]" italics="true" pageId="124" pageNumber="125">Platypalpus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C154D375D3AFFE1FF63BC5437298211" box="[189,338,821,847]" class="Insecta" family="Hybotidae" genus="Elaphropeza" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA65D3AFFE1FF63BC5437298211" box="[189,338,821,847]" italics="true" pageId="124" pageNumber="125">Elaphropeza</emphasis>
</taxonomicName>
and many new species await description.
</paragraph>
<paragraph id="8BAA36B45D3AFFE1FF63BC1932D582E9" blockId="124.[189,1399,888,1056]" pageId="124" pageNumber="125">
Species of this genus inhabit different biotopes but usually occurring in the low-lying vegetation zones (
<bibRefCitation id="EF844B455D3AFFE1FF1BBCFC372E82E9" author="Collin" box="[197,341,925,951]" firstAuthor="Collin" pageId="124" pageNumber="125" pagination="1 - 782" refId="ref68182" refString="Collin J. E. 1961. Empididae. In: British Flies 6: 1 - 782. University Press, Cambridge." type="journal article" year="1961">Collin 1961</bibRefCitation>
;
<bibRefCitation id="EF844B455D3AFFE1FEBCBCFC378182E6" author="Chvala" box="[354,506,925,952]" firstAuthor="Chvala" pageId="124" pageNumber="125" pagination="1 - 336" refId="ref68000" refString="Chvala M. 1975. The Tachydromiinae (Dipt. Empididae) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica 3: 1 - 336." type="journal article" year="1975">Chvála 1975</bibRefCitation>
;
<bibRefCitation id="EF844B455D3AFFE1FDD8BCFC354682E9" author="Cumming &amp; Cooper" box="[518,829,925,951]" firstAuthor="Cumming" pageId="124" pageNumber="125" pagination="951 - 998" refId="ref68318" refString="Cumming J. M. &amp; Cooper B. E. 1992. A revision of the Nearctic species of the Tachydromiine fly genus Stilpon Loew (Diptera: Empidoidea). The Canadian Entomologist 124 (6): 951 - 998. http: // dx. doi. org / 10.4039 / Ent 124951 - 6" type="journal article" year="1992">Cumming &amp; Cooper 1992</bibRefCitation>
;
<bibRefCitation id="EF844B455D3AFFE1FC97BCFC32D982E9" author="Przhiboro &amp; Shamshev" box="[841,1186,925,951]" firstAuthor="Przhiboro" pageId="124" pageNumber="125" pagination="110 - 116" refId="ref70216" refString="Przhiboro A. &amp; Shamshev I. 2007. Shore habitats of larval Stilpon graminum (Fallen, 1815) in Northwestern Russia (Diptera Hybotidae). Bulletin de la Societe royale belge d'Entomologie 143: 110 - 116." type="journal article" year="2007">Przhiboro &amp; Shamshev 2007</bibRefCitation>
).
</paragraph>
<paragraph id="8BAA36B45D3AFFE1FF63BC8135D2857E" blockId="124.[189,1399,888,1056]" pageId="124" pageNumber="125">
The key given below is limited to
<taxonomicName id="4C154D375D3AFFE1FD90BC8134DA82A4" authorityName="Loew" authorityYear="1859" box="[590,673,992,1018]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA65D3AFFE1FD90BC8134DA82A4" box="[590,673,992,1018]" italics="true" pageId="124" pageNumber="125">Stilpon</emphasis>
</taxonomicName>
from low altitudes including only
<taxonomicName id="4C154D375D3AFFE1FBE7BC8032D082A4" authorityName=", Shamshev &amp; Grootaert" authorityYear="2004" box="[1081,1195,992,1019]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="laawae">
<emphasis id="B961EAA65D3AFFE1FBE7BC8032D082A4" box="[1081,1195,992,1019]" italics="true" pageId="124" pageNumber="125">S. laawae</emphasis>
</taxonomicName>
of the 15 species known from northern
<collectingCountry id="F30276245D3AFFE1FE61BB64345C8541" box="[447,551,1029,1055]" name="Thailand" pageId="124" pageNumber="125">Thailand</collectingCountry>
(
<bibRefCitation id="EF844B455D3AFFE1FDE8BB6435E7857E" author="Shamshev &amp; Grootaert" box="[566,924,1029,1056]" firstAuthor="Shamshev" pageId="124" pageNumber="125" pagination="315 - 346" refId="ref70430" refString="Shamshev I. V. &amp; Grootaert P. 2004 b. A review of the genus Stilpon Loew (Diptera, Empidoidea, Hybotidae) from the Oriental region. The Raffles Bulletin of Zoology 52: 315 - 346." type="journal article" year="2004" yearSuffix="b">Shamshev &amp; Grootaert 2004b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30F653F5D3AFFE0FF63BB28330C8364" lastPageId="125" lastPageNumber="126" pageId="124" pageNumber="125" type="key">
<paragraph id="8BAA36B45D3AFFE1FF63BB2834CE853D" blockId="124.[189,1399,1096,2027]" box="[189,693,1096,1123]" pageId="124" pageNumber="125">
<heading id="D0E281D85D3AFFE1FF63BB2834CE853D" bold="true" box="[189,693,1096,1123]" fontSize="11" level="3" pageId="124" pageNumber="125" reason="3">
<emphasis id="B961EAA65D3AFFE1FF63BB2834CE853D" bold="true" box="[189,693,1096,1123]" pageId="124" pageNumber="125">
Key to species of
<taxonomicName id="4C154D375D3AFFE1FE4ABB293791853D" ID-CoL="7NQ5" authorityName="Loew" authorityYear="1859" box="[404,490,1096,1123]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA65D3AFFE1FE4ABB293791853D" bold="true" box="[404,490,1096,1123]" italics="true" pageId="124" pageNumber="125">Stilpon</emphasis>
</taxonomicName>
from
<collectingCountry id="F30276245D3AFFE1FDE8BB2834CE853D" box="[566,693,1097,1123]" name="Singapore" pageId="124" pageNumber="125">Singapore</collectingCountry>
</emphasis>
</heading>
</paragraph>
<key id="FF8461855D3AFFE0FF63BB18330C8364" lastPageId="125" lastPageNumber="126" pageId="124" pageNumber="125">
<keyStep id="30E12E115D3AFFE1FF63BB18330D85E7" pageId="124" pageNumber="125">
<paragraph id="8BAA36B45D3AFFE1FF63BB18330D85CD" blockId="124.[189,1399,1096,2027]" box="[189,1398,1145,1171]" pageId="124" pageNumber="125">
<keyLead id="30E495815D3AFFE1FF63BB18330D85CD" box="[189,1398,1145,1171]" pageId="124" pageNumber="125">1. Thorax entirely black.............................................................................................................................2</keyLead>
</paragraph>
<paragraph id="8BAA36B45D3AFFE1FF1ABBFE330D85E7" blockId="124.[189,1399,1096,2027]" box="[196,1398,1182,1209]" pageId="124" pageNumber="125">
<keyLead id="30E495815D3AFFE1FF1ABBFE330D85E7" box="[196,1398,1182,1209]" pageId="124" pageNumber="125"> Thorax at least with yellow pleura........................................................................................................5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E12E115D3AFFE1FF63BB83330C847E" pageId="124" pageNumber="125">
<paragraph id="8BAA36B45D3AFFE1FF63BB83330C85A2" blockId="124.[189,1399,1096,2027]" box="[189,1399,1249,1281]" pageId="124" pageNumber="125">
<keyLead id="30E495815D3AFFE1FF63BB83330C85A2" box="[189,1399,1249,1281]" pageId="124" pageNumber="125">
2. Wing lacking R
<subScript id="179134F15D3AFFE1FE7EBB9237C0845F" attach="left" box="[416,443,1267,1281]" fontSize="6" pageId="124" pageNumber="125">2+3</subScript>
. Palpus brown...................................................................
<taxonomicName id="4C154D375D3AFFE1FB80BB80337785A5" ID-CoL="TY7K" authority="Grootaert &amp; Shamshev, 2012" authorityName="Grootaert &amp; Shamshev" authorityYear="2012" box="[1118,1292,1249,1275]" class="Insecta" family="Hybotidae" genus="Chersodromia" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="nigripennis" status="sp. nov.">
<emphasis id="B961EAA65D3AFFE1FB80BB80337785A5" bold="true" box="[1118,1292,1249,1275]" italics="true" pageId="124" pageNumber="125">S. nigripennis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257DD5D3AFFE1FAC6BB83330C85A2" box="[1304,1399,1250,1276]" pageId="124" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAA36B45D3AFFE1FF1BBA66330C847E" blockId="124.[189,1399,1096,2027]" box="[197,1399,1286,1318]" pageId="124" pageNumber="125">
<keyLead id="30E495815D3AFFE1FF1BBA66330C847E" box="[197,1399,1286,1318]" pageId="124" pageNumber="125">
Wing with R
<subScript id="179134F15D3AFFE1FEADBA7937F58478" attach="left" box="[371,398,1304,1318]" fontSize="6" pageId="124" pageNumber="125">2+3</subScript>
. Palpus yellow..............................................................................................................3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E12E115D3AFFE1FF63BA2B330C8743" pageId="124" pageNumber="125">
<paragraph id="8BAA36B45D3AFFE1FF63BA2B330C848A" blockId="124.[189,1399,1096,2027]" pageId="124" pageNumber="125">
<keyLead id="30E495815D3AFFE1FF63BA2B330C848A" pageId="124" pageNumber="125">
3. R
<subScript id="179134F15D3AFFE1FF2CBA3D37768434" attach="left" box="[242,269,1372,1386]" fontSize="6" pageId="124" pageNumber="125">4+5</subScript>
strongly curved toward costa in apical part; distance between apices of R
<subScript id="179134F15D3AFFE1FB7DBA3D32C58434" attach="left" box="[1187,1214,1372,1386]" fontSize="6" pageId="124" pageNumber="125">2+3</subScript>
and R
<subScript id="179134F15D3AFFE1FAD1BA3D33528434" attach="left" box="[1295,1321,1372,1386]" fontSize="6" pageId="124" pageNumber="125">4+5</subScript>
about 1.5 times shorter than distance between apices of R
<subScript id="179134F15D3AFFE1FC87BAE0351984D1" attach="left" box="[857,866,1409,1423]" fontSize="6" pageId="124" pageNumber="125">1</subScript>
and R
<subScript id="179134F15D3AFFE1FC6DBAE035B584D1" attach="left" box="[947,974,1409,1423]" fontSize="6" pageId="124" pageNumber="125">2+3</subScript>
. Hind femur (except narrow basal portion) brownish. Male: abdominal tergite 1 produced laterally into small corner-like projection bearing 3 short bristles................................................................................
<taxonomicName id="4C154D375D3AFFE1FB9FBAD83370848D" ID-CoL="TY7X" authority="Grootaert &amp; Shamshev, 2012" authorityName="Grootaert &amp; Shamshev" authorityYear="2012" box="[1089,1291,1465,1491]" class="Insecta" family="Hybotidae" genus="Chersodromia" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="singaporensis" status="sp. nov.">
<emphasis id="B961EAA65D3AFFE1FB9FBAD83370848D" bold="true" box="[1089,1291,1465,1491]" italics="true" pageId="124" pageNumber="125">S. singaporensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257DD5D3AFFE1FAC9BADB330C848A" box="[1303,1399,1466,1492]" pageId="124" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAA36B45D3AFFE1FF1BBABE330C8743" blockId="124.[189,1399,1096,2027]" pageId="124" pageNumber="125">
<keyLead id="30E495815D3AFFE1FF1BBABE330C8743" pageId="124" pageNumber="125">
R
<subScript id="179134F15D3AFFE1FF39BA91377984A0" attach="left" box="[231,258,1520,1534]" fontSize="6" pageId="124" pageNumber="125">4+5</subScript>
evenly arcuate; distance between apices of R
<subScript id="179134F15D3AFFE1FCF6BA91353884A0" attach="left" box="[808,835,1520,1534]" fontSize="6" pageId="124" pageNumber="125">2+3</subScript>
and R
<subScript id="179134F15D3AFFE1FC4BBA9135CB84A0" attach="left" box="[917,944,1520,1534]" fontSize="6" pageId="124" pageNumber="125">4+5</subScript>
at least subequal to distance between apices of R
<subScript id="179134F15D3AFFE1FEBBB9743715877D" attach="left" box="[357,366,1557,1571]" fontSize="6" pageId="124" pageNumber="125">1</subScript>
and R
<subScript id="179134F15D3AFFE1FE61B974379B877D" attach="left" box="[447,480,1557,1571]" fontSize="6" pageId="124" pageNumber="125">2+3.</subScript>
Hind femur wholly yellow. Abdominal tergite 1 unmodified........................4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E12E115D3AFFE1FF63B926330C87AB" pageId="124" pageNumber="125">
<paragraph id="8BAA36B45D3AFFE1FF63B926330C87DB" blockId="124.[189,1399,1096,2027]" pageId="124" pageNumber="125">
<keyLead id="30E495815D3AFFE1FF63B926330C87DB" pageId="124" pageNumber="125">
4. Only tarsomere 5 of all legs black. Male: mid femur with 3 moderately long, ventral, brownish bristles in basal 1/3 (
<figureCitation id="132E2A315D3AFFE1FE11B90D344D87D8" box="[463,566,1644,1670]" captionStart="Figs 199-203" captionStartId="125.[189,241,1895,1921]" captionTargetBox="[221,1379,845,1842]" captionTargetId="figure@125.[189,1398,819,1855]" captionText="Figs 199-203. Stilpon, ♂. 199-202. S. angkorensis Shamshev &amp; Grootaert, 2006. 199. mid leg, lateral view. 200. right surstylus, dorsal view. 201. epandrium with cerci, dorsal view. 202. left surstylus, lateral view. 203. S. goesi Shamshev &amp; Grootaert, 2006, mid leg, anterior view. (after Shamshev &amp; Grootaert 2006)." figureDoi="http://doi.org/10.5281/zenodo.3723772" httpUri="https://zenodo.org/record/3723772/files/figure.png" pageId="124" pageNumber="125">Fig. 203</figureCitation>
). (
<collectingCountry id="F30276245D3AFFE1FD8CB90D34B787D9" box="[594,716,1644,1671]" name="Cambodia" pageId="124" pageNumber="125">Cambodia</collectingCountry>
)...............................
<taxonomicName id="4C154D375D3AFFE1FC64B90D330C87DB" ID-CoL="52QKG" authority="Shamshev &amp; Grootaert, 2006" authorityName="Shamshev &amp; Grootaert" authorityYear="2006" box="[954,1399,1643,1670]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="goesi">
<emphasis id="B961EAA65D3AFFE1FC64B90D326C87D8" bold="true" box="[954,1047,1644,1670]" italics="true" pageId="124" pageNumber="125">S. goesi</emphasis>
<bibRefCitation id="EF844B455D3AFFE1FBC0B90D330C87DB" author="Shamshev &amp; Grootaert" box="[1054,1399,1643,1670]" firstAuthor="Shamshev" pageId="124" pageNumber="125" pagination="4031 - 4065" refId="ref69126" refString="Grootaert P. &amp; Shamshev I. 2006. The genus Platypalpus Macquart (Diptera: Hybotidae) from Northeast Thailand with comments on the species groups in the Oriental region. Journal of Natural History 39 (47): 4031 - 4065. http: // dx. doi. org / 10.1080 / 00222930500533781" type="journal article" year="2006">Shamshev &amp; Grootaert, 2006</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAA36B45D3AFFE1FF1BB9F0330C87AB" blockId="124.[189,1399,1096,2027]" pageId="124" pageNumber="125">
<keyLead id="30E495815D3AFFE1FF1BB9F0330C87AB" pageId="124" pageNumber="125">
Tarsomere 5 of all legs and tarsomere 1 of hind leg brownish. Male: mid femur with 1 black, posteroventral spine beyond middle and row of 7 spine-like bristles in basal part (
<figureCitation id="132E2A315D3AFFE1FB25B9D73312878E" box="[1275,1385,1718,1744]" captionStart="Figs 199-203" captionStartId="125.[189,241,1895,1921]" captionTargetBox="[221,1379,845,1842]" captionTargetId="figure@125.[189,1398,819,1855]" captionText="Figs 199-203. Stilpon, ♂. 199-202. S. angkorensis Shamshev &amp; Grootaert, 2006. 199. mid leg, lateral view. 200. right surstylus, dorsal view. 201. epandrium with cerci, dorsal view. 202. left surstylus, lateral view. 203. S. goesi Shamshev &amp; Grootaert, 2006, mid leg, anterior view. (after Shamshev &amp; Grootaert 2006)." figureDoi="http://doi.org/10.5281/zenodo.3723772" httpUri="https://zenodo.org/record/3723772/files/figure.png" pageId="124" pageNumber="125">Fig. 199</figureCitation>
). (
<collectingCountry id="F30276245D3AFFE1FF3DB9BA372787A8" box="[227,348,1755,1782]" name="Cambodia" pageId="124" pageNumber="125">Cambodia</collectingCountry>
)....................................................................
<taxonomicName id="4C154D375D3AFFE1FC84B9BA330C87AB" ID-CoL="52QK5" authority="Shamshev &amp; Grootaert, 2006" authorityName="Shamshev &amp; Grootaert" authorityYear="2006" box="[858,1399,1754,1781]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="angkorensis">
<emphasis id="B961EAA65D3AFFE1FC84B9BA327587AB" bold="true" box="[858,1038,1754,1781]" italics="true" pageId="124" pageNumber="125">S. angkorensis</emphasis>
<bibRefCitation id="EF844B455D3AFFE1FBC6B9BA330C87AB" author="Shamshev &amp; Grootaert" box="[1048,1399,1755,1781]" firstAuthor="Shamshev" pageId="124" pageNumber="125" pagination="4031 - 4065" refId="ref69126" refString="Grootaert P. &amp; Shamshev I. 2006. The genus Platypalpus Macquart (Diptera: Hybotidae) from Northeast Thailand with comments on the species groups in the Oriental region. Journal of Natural History 39 (47): 4031 - 4065. http: // dx. doi. org / 10.1080 / 00222930500533781" type="journal article" year="2006">Shamshev &amp; Grootaert, 2006</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E12E115D3AFFE1FF63B87F330C8603" pageId="124" pageNumber="125">
<paragraph id="8BAA36B45D3AFFE1FF63B87F330C8667" blockId="124.[189,1399,1096,2027]" box="[189,1399,1822,1849]" pageId="124" pageNumber="125">
<keyLead id="30E495815D3AFFE1FF63B87F330C8667" box="[189,1399,1822,1849]" pageId="124" pageNumber="125">
5. Hind tibia strongly curved. Halter yellow..........................................................
<taxonomicName id="4C154D375D3AFFE1FBA6B87F33748667" ID-CoL="BDDV2" authority="Grootaert &amp; Shamshev, 2012" authorityName="Grootaert &amp; Shamshev" authorityYear="2012" box="[1144,1295,1822,1849]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="arcuatum" status="sp. nov.">
<emphasis id="B961EAA65D3AFFE1FBA6B87F33748667" bold="true" box="[1144,1295,1822,1849]" italics="true" pageId="124" pageNumber="125">S. arcuatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257DD5D3AFFE1FAC7B87E330C8667" box="[1305,1399,1823,1849]" pageId="124" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAA36B45D3AFFE1FF1BB825330C8603" blockId="124.[189,1399,1096,2027]" box="[197,1399,1859,1886]" pageId="124" pageNumber="125">
<keyLead id="30E495815D3AFFE1FF1BB825330C8603" box="[197,1399,1859,1886]" pageId="124" pageNumber="125"> Hind tibia unmodified. Halter brownish or black.................................................................................6</keyLead>
</paragraph>
</keyStep>
<key id="FF8461855D3AFFE0FF63B8E7330C8364" lastPageId="125" lastPageNumber="126" pageId="124" pageNumber="125">
<keyStep id="30E12E115D3AFFE1FF63B8E7330C86B5" pageId="124" pageNumber="125">
<paragraph id="8BAA36B45D3AFFE1FF63B8E7330C86FF" blockId="124.[189,1399,1096,2027]" box="[189,1399,1926,1953]" pageId="124" pageNumber="125">
<keyLead id="30E495815D3AFFE1FF63B8E7330C86FF" box="[189,1399,1926,1953]" pageId="124" pageNumber="125">
6. Scutum entirely brown. Hind femur largely brownish......
<taxonomicName id="4C154D375D3AFFE1FCAAB8E6330C86FF" ID-CoL="393DC" authority="Shamshev &amp; Grootaert, 2004" authorityName="Shamshev &amp; Grootaert" authorityYear="2004" box="[884,1399,1926,1953]" class="Insecta" family="Hybotidae" genus="Elaphropeza" kingdom="Animalia" order="Diptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B961EAA65D3AFFE1FCAAB8E6326286FF" bold="true" box="[884,1049,1926,1953]" italics="true" pageId="124" pageNumber="125">S. malayensis</emphasis>
<bibRefCitation id="EF844B455D3AFFE1FBFEB8E7330C86FF" author="Shamshev I. V. &amp; Grootaert P." box="[1056,1399,1926,1953]" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="124" pageNumber="125" pagination="45 - 58" part="52" refId="ref70369" refString="Shamshev I. V. &amp; Grootaert P. 2004 a. Descriptions of four new species of the genus Microphorella Becker (Diptera: Empidoidea, Microphoridae, Parathalassiini) from Southeast Asia and New Guinea, with notes on the relationships within the genus. The Raffles Bulletin of Zoology 52 (1): 45 - 58." title="Descriptions of four new species of the genus Microphorella Becker (Diptera: Empidoidea, Microphoridae, Parathalassiini) from Southeast Asia and New Guinea, with notes on the relationships within the genus" type="journal article" year="2004">Shamshev &amp; Grootaert, 2004</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAA36B45D3AFFE1FF1BB8CD330C86B5" blockId="124.[189,1399,1096,2027]" pageId="124" pageNumber="125">
<keyLead id="30E495815D3AFFE1FF1BB8CD330C86B5" pageId="124" pageNumber="125"> Scutum largely yellow, with brownish patch above wing base. Hind femur at most somewhat brownish in apical part.........................................................................................................................................7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E12E115D3BFFE0FF63BE6D330C8032" pageId="125" pageNumber="126">
<paragraph id="8BAA36B45D3BFFE0FF63BE6D330C8017" blockId="125.[189,1399,267,571]" pageId="125" pageNumber="126">
<keyLead id="30E495815D3BFFE0FF63BE6D330C8017" pageId="125" pageNumber="126">
7. Wing finely infuscate; R
<subScript id="179134F15D3BFFE0FE23BE7C34638075" attach="left" box="[509,536,285,299]" fontSize="6" pageId="125" pageNumber="126">4+5</subScript>
slightly arcuate in apical part. Male: upper lobe of left surstylus lacking surstylar comb (
<figureCitation id="132E2A315D3BFFE0FE47BE4F347B8017" box="[409,512,302,329]" captionStart="Figs 210-213" captionStartId="130.[189,241,1930,1956]" captionTargetBox="[199,1363,1132,1862]" captionTargetId="figure@130.[189,1398,1085,1889]" captionText="Figs 210-213. Stilpon laawae, Shamshev &amp; Grootaert, 2004, ♂ terminalia (Material Singapore). 210. right surstylus, dorsal view. 211. epandrium with cerci, dorsal view. 212. left surstylus, lateral view. 213. cerci, right lateral view. Scale bar= 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3723778" httpUri="https://zenodo.org/record/3723778/files/figure.png" pageId="125" pageNumber="126">Fig. 211</figureCitation>
). (
<collectingCountry id="F30276245D3BFFE0FDC2BE4F34F18016" box="[540,650,302,328]" name="Thailand" pageId="125" pageNumber="126">Thailand</collectingCountry>
;
<collectingCountry id="F30276245D3BFFE0FD48BE4F35758017" box="[662,782,302,329]" name="Singapore" pageId="125" pageNumber="126">Singapore</collectingCountry>
)...................
<taxonomicName id="4C154D375D3BFFE0FC7CBE4E330C8017" ID-CoL="52QKN" authority="Shamshev &amp; Grootaert, 2004" authorityName=", Shamshev &amp; Grootaert" authorityYear="2004" box="[930,1399,302,329]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="laawae">
<emphasis id="B961EAA65D3BFFE0FC7CBE4E326D8017" bold="true" box="[930,1046,302,329]" italics="true" pageId="125" pageNumber="126">S. laawae</emphasis>
<bibRefCitation id="EF844B455D3BFFE0FBC0BE4F330C8017" author="Shamshev I. V. &amp; Grootaert P." box="[1054,1399,302,329]" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="125" pageNumber="126" pagination="45 - 58" part="52" refId="ref70369" refString="Shamshev I. V. &amp; Grootaert P. 2004 a. Descriptions of four new species of the genus Microphorella Becker (Diptera: Empidoidea, Microphoridae, Parathalassiini) from Southeast Asia and New Guinea, with notes on the relationships within the genus. The Raffles Bulletin of Zoology 52 (1): 45 - 58." title="Descriptions of four new species of the genus Microphorella Becker (Diptera: Empidoidea, Microphoridae, Parathalassiini) from Southeast Asia and New Guinea, with notes on the relationships within the genus" type="journal article" year="2004">Shamshev &amp; Grootaert, 2004</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAA36B45D3BFFE0FF1BBE33330C8032" blockId="125.[189,1399,267,571]" box="[197,1399,338,364]" pageId="125" pageNumber="126">
<keyLead id="30E495815D3BFFE0FF1BBE33330C8032" box="[197,1399,338,364]" pageId="125" pageNumber="126"> Wing maculate or darker along some veins........................................................................................8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E12E115D3BFFE0FF63BEF2330C8364" pageId="125" pageNumber="126">
<paragraph id="8BAA36B45D3BFFE0FF63BEF2330C808F" blockId="125.[189,1399,267,571]" pageId="125" pageNumber="126">
<keyLead id="30E495815D3BFFE0FF63BEF2330C808F" pageId="125" pageNumber="126">
8. Wing maculate; vein R
<subScript id="179134F15D3BFFE0FE3FBEC4378780ED" attach="left" box="[481,508,421,435]" fontSize="6" pageId="125" pageNumber="126">4+5</subScript>
strongly curved toward costa in apical part. Male: upper lobe of left surstylus with markedly developed surstylar comb (
<figureCitation id="132E2A315D3BFFE0FD11BED7354F808E" box="[719,820,438,464]" captionStart="Figs 220-224" captionStartId="134.[189,241,1930,1956]" captionTargetBox="[198,1378,420,1867]" captionTargetId="figure@134.[189,1398,400,1890]" captionText="Figs 220-224. Stilpon neesoonensis sp. nov., ♂. 220. head lateral, lateral view. 221. mid leg, anterior view. 222. right epandrial lamella, lateral view. 223. epandrium with cerci, dorsal view. 224. left surstylus and cercus, dorso-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3723782" httpUri="https://zenodo.org/record/3723782/files/figure.png" pageId="125" pageNumber="126">Fig. 223</figureCitation>
) (
<collectingCountry id="F30276245D3BFFE0FC93BED735BD808F" box="[845,966,438,465]" name="Singapore" pageId="125" pageNumber="126">Singapore</collectingCountry>
)..................
<taxonomicName id="4C154D375D3BFFE0FB8DBED73368808E" ID-CoL="BDDV3" authority="Grootaert &amp; Shamshev, 2012" authorityName="Grootaert &amp; Shamshev" authorityYear="2012" box="[1107,1299,438,464]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="neesoonensis" status="sp. nov.">
<emphasis id="B961EAA65D3BFFE0FB8DBED73368808E" bold="true" box="[1107,1299,438,464]" italics="true" pageId="125" pageNumber="126">S. neesoonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257DD5D3BFFE0FAC5BED6330C808F" box="[1307,1399,439,465]" pageId="125" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAA36B45D3BFFE0FF1BBEBB330C8364" blockId="125.[189,1399,267,571]" pageId="125" pageNumber="126">
<keyLead id="30E495815D3BFFE0FF1BBEBB330C8364" pageId="125" pageNumber="126">
Wing broadly brownish infuscate along R
<subScript id="179134F15D3BFFE0FD0DBE8A349580A7" attach="left" box="[723,750,491,505]" fontSize="6" pageId="125" pageNumber="126">2+3</subScript>
and CuA
<subScript id="179134F15D3BFFE0FCBDBE8A351780A7" attach="left" box="[867,876,491,505]" fontSize="6" pageId="125" pageNumber="126">1</subScript>
(except subapical part). R
<subScript id="179134F15D3BFFE0FB74BE8A32BE80A7" attach="left" box="[1194,1221,491,505]" fontSize="6" pageId="125" pageNumber="126">4+5</subScript>
evenly curved toward costa in apical part. Male: upper lobe of left surstylus lacking surstylar comb (
<figureCitation id="132E2A315D3BFFE0FADBBE9C33158349" box="[1285,1390,509,535]" captionStart="Figs 232-235" captionStartId="140.[189,241,1965,1991]" captionTargetBox="[193,1381,1084,1904]" captionTargetId="figure@140.[189,1398,1040,1925]" captionText="Figs 232-235. Stilpon weilingae sp. nov., ♂. 232. right surstylus, lateral view. 233. epandrium with cerci, dorsal view. 234. left surstylus, lateral view. 235. right epandrial lamella." figureDoi="http://doi.org/10.5281/zenodo.3723788" httpUri="https://zenodo.org/record/3723788/files/figure.png" pageId="125" pageNumber="126">Fig. 233</figureCitation>
) (
<collectingCountry id="F30276245D3BFFE0FF3DBD4137238365" box="[227,344,544,571]" name="Singapore" pageId="125" pageNumber="126">Singapore</collectingCountry>
)...........................................................................................................
<taxonomicName id="4C154D375D3BFFE0FBA8BD4133768365" ID-CoL="BDDV6" authority="Grootaert &amp; Shamshev, 2012" authorityName="Grootaert &amp; Shamshev" authorityYear="2012" box="[1142,1293,544,571]" class="Insecta" family="Hybotidae" genus="Stilpon" kingdom="Animalia" order="Diptera" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="weilingae" status="sp. nov.">
<emphasis id="B961EAA65D3BFFE0FBA8BD4133768365" bold="true" box="[1142,1293,544,571]" italics="true" pageId="125" pageNumber="126">S. weilingae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257DD5D3BFFE0FAC6BD41330C8364" box="[1304,1399,544,570]" pageId="125" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</key>
</subSubSection>
</treatment>
</document>