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<document id="7755607E1634ED6311153601F718E384" ID-DOI="10.11646/zootaxa.4809.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3936752" ID-ZooBank="679EF63B-0051-4490-9380-BCD857326197" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594291765646" checkinUser="plazi" docAuthor="Naz, Saima, Najer, Tomas &amp; Gustafsson, Daniel R." docDate="2020" docId="03BC878AE860FFC27FE8FAC3FB55F834" docLanguage="en" docName="zootaxa.4809.3.1.pdf" docOrigin="Zootaxa 4809 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Picophilopterus tuktola Ansari 1947" docType="treatment" docVersion="5" lastPageNumber="435" masterDocId="FF85FFF2E842FFE07F7FFFA3FFFFFFA3" masterDocTitle="An annotated list of the species of lice (Insecta: Phthiraptera) described by Mohammad A. - R. Ansari" masterLastPageNumber="448" masterPageNumber="401" pageNumber="435" updateTime="1698779163507" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DC2B4F43396EF8168567C1317528F0B0">An annotated list of the species of lice (Insecta: Phthiraptera) described by Mohammad A. - R. Ansari</mods:title>
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<mods:name id="1728E8DCA66C5B52E8D6B8F4C8C33B4E" type="personal">
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<mods:namePart id="12F58AB7406070232007E967A2705D42">Naz, Saima</mods:namePart>
<mods:affiliation id="C3CCE07DE1E9C69926F17A8D4D348D99">Department of Veterinary Sciences, Faculty of Agrobiology, Food and Natural Resources, Czech University of Life Sciences, Kamycka 129, 165 00 Prague 6, Czechia.</mods:affiliation>
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<mods:namePart id="DB7B9F41859E88FE1C2D029DC640FA27">Najer, Tomas</mods:namePart>
<mods:affiliation id="F8C485FEACD27F3E8C73FBE02D7C0263">Department of Veterinary Sciences, Faculty of Agrobiology, Food and Natural Resources, Czech University of Life Sciences, Kamycka 129, 165 00 Prague 6, Czechia. &amp; najer @ af. czu. cz; https: // orcid. org / 0000 - 0003 - 4768 - 7906</mods:affiliation>
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<mods:namePart id="51AFF89457430A9C1AB6018CCE118C35">Gustafsson, Daniel R.</mods:namePart>
<mods:affiliation id="E73CB36E0893A1B7B5C0CA326D5F4910">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Guangdong Institute of Applied Biological Resources, 105 Xingang West Road, Haizhu District, Guangzhou 510260, China. https: // orcid. org / 0000 - 0001 - 8868 - 8310</mods:affiliation>
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<mods:date id="D15887F59F833B84640A156EE411AE91">2020</mods:date>
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<treatment id="03BC878AE860FFC27FE8FAC3FB55F834" ID-DOI="http://doi.org/10.5281/zenodo.4345154" ID-Zenodo-Dep="4345154" LSID="urn:lsid:plazi:treatment:03BC878AE860FFC27FE8FAC3FB55F834" httpUri="http://treatment.plazi.org/id/03BC878AE860FFC27FE8FAC3FB55F834" lastPageNumber="435" pageId="34" pageNumber="435">
<subSubSection id="C30F6517E860FFC27FE8FAC3FDBCFAD8" box="[151,579,1376,1403]" pageId="34" pageNumber="435" type="nomenclature">
<paragraph id="8BAA369CE860FFC27FE8FAC3FDBCFAD8" blockId="34.[151,1437,1376,1943]" box="[151,579,1376,1403]" pageId="34" pageNumber="435">
<heading id="D0E281F0E860FFC27FE8FAC3FDBCFAD8" bold="true" box="[151,579,1376,1403]" fontSize="11" level="1" pageId="34" pageNumber="435" reason="1">
<taxonomicName id="4C154D1FE860FFC27FE8FAC3FDBCFAD8" authority="Ansari, 1947" authorityName="Ansari" authorityYear="1947" box="[151,579,1376,1403]" class="Insecta" family="Philopteridae" genus="Picophilopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="tuktola">
<emphasis id="B961EA8EE860FFC27FE8FAC3FDBCFAD8" bold="true" box="[151,579,1376,1403]" pageId="34" pageNumber="435">
<emphasis id="B961EA8EE860FFC27FE8FAC3FE5AFAD8" bold="true" box="[151,421,1376,1403]" italics="true" pageId="34" pageNumber="435">Picophilopterus tuktola</emphasis>
<bibRefCitation id="EF844B6DE860FFC27ED3FAC3FDBCFAD8" author="Ansari, M. A. R." box="[428,579,1376,1403]" pageId="34" pageNumber="435" pagination="253 - 303" refId="ref29219" refString="Ansari, M. A. R. (1947) Mallophaga (Ischnocera) infesting birds in the Punjab (India). Proceedings of the National Institute of Science India, 13 (6), 253 - 303." type="journal article" year="1947">Ansari, 1947</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30F6517E860FFC27FB8FA26FC2FFA60" pageId="34" pageNumber="435" type="reference_group">
<paragraph id="8BAA369CE860FFC27FB8FA26FCE4FA3C" blockId="34.[151,1437,1376,1943]" box="[199,795,1412,1439]" pageId="34" pageNumber="435">
<treatmentCitationGroup id="AB0511B2E860FFC27FB8FA26FCE4FA3C" box="[199,795,1412,1439]" pageId="34" pageNumber="435">
<treatmentCitation id="0AB4108DE860FFC27FB8FA26FD5DFA3C" author="Ansari, M. A. R." box="[199,674,1412,1439]" page="265" pageId="34" pageNumber="435" year="1947">
<taxonomicName id="4C154D1FE860FFC27FB8FA26FD5DFA3C" authority="Ansari, 1947: 265" authorityName="Ansari" authorityPageNumber="265" authorityYear="1947" box="[199,674,1412,1439]" class="Insecta" family="Philopteridae" genus="Picophilopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="tuktola">
<emphasis id="B961EA8EE860FFC27FB8FA26FE30FA3C" box="[199,463,1413,1439]" italics="true" pageId="34" pageNumber="435">Picophilopterus tuktola</emphasis>
<bibRefCitation id="EF844B6DE860FFC27EA9FA26FD5DFA3C" author="Ansari, M. A. R." box="[470,674,1412,1439]" pageId="34" pageNumber="435" pagination="253 - 303" refId="ref29219" refString="Ansari, M. A. R. (1947) Mallophaga (Ischnocera) infesting birds in the Punjab (India). Proceedings of the National Institute of Science India, 13 (6), 253 - 303." type="journal article" year="1947">Ansari, 1947: 265</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 4ac.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA369CE860FFC27FE8FA0BFC2FFA60" blockId="34.[151,1437,1376,1943]" box="[151,976,1448,1475]" pageId="34" pageNumber="435">
<emphasis id="B961EA8EE860FFC27FE8FA0BFEB0FA60" bold="true" box="[151,335,1448,1475]" pageId="34" pageNumber="435">Current status:</emphasis>
<taxonomicName id="4C154D1FE860FFC27E29FA09FD21FA60" authority="(Ansari, 1947)" baseAuthorityName="Ansari" baseAuthorityYear="1947" box="[342,734,1449,1475]" class="Insecta" family="Philopteridae" genus="Penenirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="tuktola">
<emphasis id="B961EA8EE860FFC27E29FA09FDCEFA60" box="[342,561,1449,1475]" italics="true" pageId="34" pageNumber="435">Penenirmus tuktola</emphasis>
(
<bibRefCitation id="EF844B6DE860FFC27D40FA0AFD2AFA60" author="Ansari, M. A. R." box="[575,725,1449,1475]" pageId="34" pageNumber="435" pagination="253 - 303" refId="ref29219" refString="Ansari, M. A. R. (1947) Mallophaga (Ischnocera) infesting birds in the Punjab (India). Proceedings of the National Institute of Science India, 13 (6), 253 - 303." type="journal article" year="1947">Ansari, 1947</bibRefCitation>
)
</taxonomicName>
; see Remarks, below.
</paragraph>
</subSubSection>
<subSubSection id="C30F6517E860FFC27FE8FA6EFB1DF9F0" pageId="34" pageNumber="435" type="materials_examined">
<paragraph id="8BAA369CE860FFC27FE8FA6EFD50FA44" blockId="34.[151,1437,1376,1943]" box="[151,687,1485,1511]" pageId="34" pageNumber="435">
<emphasis id="B961EA8EE860FFC27FE8FA6EFEEBFA44" bold="true" box="[151,276,1485,1511]" pageId="34" pageNumber="435">
<typeStatus id="54AE883EE860FFC27FE8FA6EFF2DFA44" box="[151,210,1485,1511]" pageId="34" pageNumber="435">Type</typeStatus>
host:
</emphasis>
<taxonomicName id="4C154D1FE860FFC27E64FA6DFD50FA44" authority="squamatus Vigors." authorityName="Vigors." box="[283,687,1485,1511]" class="Aves" family="Picidae" genus="Picus" kingdom="Animalia" order="Piciformes" pageId="34" pageNumber="435" phylum="Chordata" rank="subSpecies" species="squamatus" subSpecies="squamatus">
<emphasis id="B961EA8EE860FFC27E64FA6DFDA8FA44" box="[283,599,1486,1511]" italics="true" pageId="34" pageNumber="435">Picus squamatus squamatus</emphasis>
Vigors.
</taxonomicName>
</paragraph>
<paragraph id="8BAA369CE860FFC27FE8FA52FDC0F9A8" blockId="34.[151,1437,1376,1943]" box="[151,575,1520,1547]" pageId="34" pageNumber="435">
<materialsCitation id="3B7D3CC1E860FFC27FE8FA52FDC4F9A8" box="[151,571,1520,1547]" country="India" location="Kullu" pageId="34" pageNumber="435" specimenCount="1" typeStatus="holotype">
<emphasis id="B961EA8EE860FFC27FE8FA52FEC6F9A8" bold="true" box="[151,313,1520,1547]" pageId="34" pageNumber="435">
<typeStatus id="54AE883EE860FFC27FE8FA52FF2DF9A8" box="[151,210,1521,1547]" pageId="34" pageNumber="435">Type</typeStatus>
locality:
</emphasis>
<location id="8ECA6047E860FFC27E3FFA52FE7FF9A8" LSID="urn:lsid:plazi:treatment:03BC878AE860FFC27FE8FAC3FB55F834:8ECA6047E860FFC27E3FFA52FE7FF9A8" box="[320,384,1521,1547]" country="India" name="Kullu" pageId="34" pageNumber="435">Kullu</location>
[as Kulu],
<collectingCountry id="F302760CE860FFC27E81FA52FDC4F9A8" box="[510,571,1521,1547]" name="India" pageId="34" pageNumber="435">India</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAA369CE860FFC27FE8F9B6FB1DF9F0" blockId="34.[151,1437,1376,1943]" pageId="34" pageNumber="435">
<emphasis id="B961EA8EE860FFC27FE8F9B6FEB8F98C" bold="true" box="[151,327,1556,1583]" pageId="34" pageNumber="435">Type material:</emphasis>
<materialsCitation id="3B7D3CC1E860FFC27E32F9B6FB21F9F0" collectionCode="NHML" pageId="34" pageNumber="435" specimenCode="NHML010684468" specimenCount="3" typeStatus="paratype">
Two
<typeStatus id="54AE883EE860FFC27EFAF9B6FE1AF98C" box="[389,485,1557,1583]" pageId="34" pageNumber="435" type="paratype">paratype</typeStatus>
females deposited in the
<collectionCode id="ED04AE59E860FFC27C7DF9B6FCACF98C" box="[770,851,1557,1583]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="34" pageNumber="435">NHML</collectionCode>
[slide
<specimenCode id="DBB39EE7E860FFC27CE0F9B7FB89F98C" box="[927,1142,1556,1583]" collectionCode="NHML" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="34" pageNumber="435">NHML010684468</specimenCode>
]. The
<typeStatus id="54AE883EE860FFC27BC7F9B6FAE5F98C" box="[1208,1306,1557,1583]" pageId="34" pageNumber="435" type="holotype">holotype</typeStatus>
female and the “Numerous males and females” recorded by
<bibRefCitation id="EF844B6DE860FFC27DCAF99AFC75F9F7" author="Ansari, M. A. R." box="[693,906,1593,1620]" pageId="34" pageNumber="435" pagination="253 - 303" refId="ref29219" refString="Ansari, M. A. R. (1947) Mallophaga (Ischnocera) infesting birds in the Punjab (India). Proceedings of the National Institute of Science India, 13 (6), 253 - 303." type="journal article" year="1947">Ansari (1947: 266)</bibRefCitation>
as
<typeStatus id="54AE883EE860FFC27CD0F99AFBE5F9F0" box="[943,1050,1593,1619]" pageId="34" pageNumber="435" type="paratype">paratypes</typeStatus>
appear to be lost
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F6517E860FFC27FE8F9FEFB55F834" pageId="34" pageNumber="435" type="discussion">
<paragraph id="8BAA369CE860FFC27FE8F9FEFB55F834" blockId="34.[151,1437,1376,1943]" pageId="34" pageNumber="435">
<emphasis id="B961EA8EE860FFC27FE8F9FEFEF2F9D4" bold="true" box="[151,269,1629,1655]" pageId="34" pageNumber="435">Remarks:</emphasis>
<bibRefCitation id="EF844B6DE860FFC27E6AF9FEFE00F9D4" author="Dalgleish, R. C." box="[277,511,1629,1655]" pageId="34" pageNumber="435" pagination="83 - 104" refId="ref31611" refString="Dalgleish, R. C. (1972) The Penenirmus (Mallophaga: Ischnocera) of the Picidae (Aves: Piciformes). Journal of the New York Entomological Society, 80, 83 - 104." type="journal article" year="1972">Dalgleish (1972: 98)</bibRefCitation>
regarded
<taxonomicName id="4C154D1FE860FFC27D0EF9FEFC85F9D4" authorityName="Ansari" authorityYear="1947" box="[625,890,1629,1655]" class="Insecta" family="Philopteridae" genus="Picophilopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="tuktola">
<emphasis id="B961EA8EE860FFC27D0EF9FEFC85F9D4" box="[625,890,1629,1655]" italics="true" pageId="34" pageNumber="435">Picophilopterus tuktola</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="4C154D1FE860FFC27BF0F9FDFEE9F938" authority="(Fabricius, 1798)" baseAuthorityName="Fabricius" baseAuthorityYear="1798" class="Insecta" family="Philopteridae" genus="Penenirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="pici">
<emphasis id="B961EA8EE860FFC27BF0F9FDFAB9F9D4" box="[1167,1350,1630,1655]" italics="true" pageId="34" pageNumber="435">Penenirmus pici</emphasis>
(
<bibRefCitation id="EF844B6DE860FFC27A29F9FEFEF1F938" author="Fabricius, J. C." pageId="34" pageNumber="435" pagination="570 - 571" refId="ref32015" refString="Fabricius, J. C. (1798) Pediculus. In: Supplementum entomologiae systematicae. Proft et Storch, Hafniae (Copenhagen), pp. 570 - 571." type="book chapter" year="1798">Fabricius, 1798</bibRefCitation>
)
</taxonomicName>
, stating that there were no significant differences between these two species. However,
<bibRefCitation id="EF844B6DE860FFC27B90F922FA67F938" author="Ansari, M. A. R." box="[1263,1432,1665,1691]" pageId="34" pageNumber="435" pagination="253 - 303" refId="ref29219" refString="Ansari, M. A. R. (1947) Mallophaga (Ischnocera) infesting birds in the Punjab (India). Proceedings of the National Institute of Science India, 13 (6), 253 - 303." type="journal article" year="1947">Ansaris (1947</bibRefCitation>
, fig. 4a) illustration of the female head of
<taxonomicName id="4C154D1FE860FFC27D18F905FD2EF91C" authorityName="Ansari" authorityYear="1947" box="[615,721,1701,1727]" class="Insecta" family="Philopteridae" genus="Picophilopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="tuktola">
<emphasis id="B961EA8EE860FFC27D18F905FD2EF91C" box="[615,721,1701,1727]" italics="true" pageId="34" pageNumber="435">P. tuktola</emphasis>
</taxonomicName>
shows the temporal setae as two elongate setae and one shorter seta, a feature that does not agree with
<taxonomicName id="4C154D1FE860FFC27D37F969FD72F940" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[584,653,1738,1763]" class="Insecta" family="Philopteridae" genus="Penenirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="pici">
<emphasis id="B961EA8EE860FFC27D37F969FD72F940" box="[584,653,1738,1763]" italics="true" pageId="34" pageNumber="435">P. pici</emphasis>
</taxonomicName>
, but that is closer to
<taxonomicName id="4C154D1FE860FFC27C0AF969FB6CF940" authority="Carriker, 1961" authorityName="Carriker" authorityYear="1961" box="[885,1171,1737,1763]" class="Insecta" family="Philopteridae" genus="Penenirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="arcticus">
<emphasis id="B961EA8EE860FFC27C0AF969FC16F940" box="[885,1001,1738,1763]" italics="true" pageId="34" pageNumber="435">P. arcticus</emphasis>
Carriker, 1961
</taxonomicName>
, using key to species of
<bibRefCitation id="EF844B6DE860FFC27FE8F94EFE7CF8A4" author="Dalgleish, R. C." box="[151,387,1772,1799]" pageId="34" pageNumber="435" pagination="83 - 104" refId="ref31611" refString="Dalgleish, R. C. (1972) The Penenirmus (Mallophaga: Ischnocera) of the Picidae (Aves: Piciformes). Journal of the New York Entomological Society, 80, 83 - 104." type="journal article" year="1972">Dalgleish (1972: 85)</bibRefCitation>
. Also, the shape of the head illustrated by
<bibRefCitation id="EF844B6DE860FFC27C21F94EFC05F8A4" author="Ansari, M. A. R." box="[862,1018,1773,1799]" pageId="34" pageNumber="435" pagination="253 - 303" refId="ref29219" refString="Ansari, M. A. R. (1947) Mallophaga (Ischnocera) infesting birds in the Punjab (India). Proceedings of the National Institute of Science India, 13 (6), 253 - 303." type="journal article" year="1947">Ansari (1947)</bibRefCitation>
differs from that of
<taxonomicName id="4C154D1FE860FFC27BA0F94DFADAF8A4" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[1247,1317,1774,1799]" class="Insecta" family="Philopteridae" genus="Penenirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="pici">
<emphasis id="B961EA8EE860FFC27BA0F94DFADAF8A4" box="[1247,1317,1774,1799]" italics="true" pageId="34" pageNumber="435">P. pici</emphasis>
</taxonomicName>
illustrated by
<bibRefCitation id="EF844B6DE860FFC27FC5F8B2FE8EF888" author="Dalgleish, R. C." box="[186,369,1809,1835]" pageId="34" pageNumber="435" pagination="83 - 104" refId="ref31611" refString="Dalgleish, R. C. (1972) The Penenirmus (Mallophaga: Ischnocera) of the Picidae (Aves: Piciformes). Journal of the New York Entomological Society, 80, 83 - 104." type="journal article" year="1972">Dalgleish (1972</bibRefCitation>
: fig. 12). Our examination of other species of
<taxonomicName id="4C154D1FE860FFC27C0EF8B1FC08F888" authorityName="Clay &amp; Meinertzhagen" authorityYear="1938" box="[881,1015,1810,1835]" class="Insecta" family="Philopteridae" genus="Penenirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA8EE860FFC27C0EF8B1FC08F888" box="[881,1015,1810,1835]" italics="true" pageId="34" pageNumber="435">Penenirmus</emphasis>
</taxonomicName>
from woodpeckers suggests that
<bibRefCitation id="EF844B6DE860FFC27A15F8B2FED3F8EC" author="Dalgleish, R. C." pageId="34" pageNumber="435" pagination="83 - 104" refId="ref31611" refString="Dalgleish, R. C. (1972) The Penenirmus (Mallophaga: Ischnocera) of the Picidae (Aves: Piciformes). Journal of the New York Entomological Society, 80, 83 - 104." type="journal article" year="1972">Dalgleish (1972)</bibRefCitation>
was very conservative in regard to accepting several nominal species as valid. Although we have not seen any specimen, we here tentatively resurrect
<taxonomicName id="4C154D1FE860FFC27DC2F8F9FC67F8D0" baseAuthorityName="Ansari" baseAuthorityYear="1947" box="[701,920,1881,1907]" class="Insecta" family="Philopteridae" genus="Penenirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="tuktola">
<emphasis id="B961EA8EE860FFC27DC2F8F9FC67F8D0" box="[701,920,1881,1907]" italics="true" pageId="34" pageNumber="435">Penenirmus tuktola</emphasis>
</taxonomicName>
as a separate species, until new material from the
<typeStatus id="54AE883EE860FFC27FBFF8DEFF0FF834" box="[192,240,1917,1943]" pageId="34" pageNumber="435">type</typeStatus>
host becomes available for study, redescription and comparison with related species.
</paragraph>
</subSubSection>
</treatment>
</document>