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<document id="5ADB45EE4ECECA0A84CB78FF9A5AB6AC" ID-DOI="10.11646/zootaxa.5493.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="13330255" ID-ZooBank="16BD6CF7-E0FF-4A04-9ADE-C3DFBF92B7DC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1723803345649" checkinUser="plazi" docAuthor="Anichtchenko, Alexander &amp; Sciaky, Riccardo" docDate="2024" docId="03B3FE717728FFE474FD43E0A924EA65" docLanguage="en" docName="zootaxa.5493.4.2.pdf" docOrigin="Zootaxa 5493 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5493.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ophionea bhamoensis Bates 1892" docType="treatment" docVersion="2" lastPageNumber="338" masterDocId="FF8A86097720FFEE746A4027AA6FEB24" masterDocTitle="Contribution to the knowledge of the Philippines species of the tribe Odacanthini Laporte, 1834 (Coleoptera: Carabidae)" masterLastPageNumber="344" masterPageNumber="328" pageNumber="336" updateTime="1723804345995" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E9CB359725644BE8AAFF151B51260859">Contribution to the knowledge of the Philippines species of the tribe Odacanthini Laporte, 1834 (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="09FFA0C3D76DDBD652D2AA79C173066D">Anichtchenko, Alexander</mods:namePart>
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<mods:date id="E773EFBABEEE4C94A61A55C6F6806F33">2024</mods:date>
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<treatment id="03B3FE717728FFE474FD43E0A924EA65" ID-DOI="http://doi.org/10.5281/zenodo.13330225" ID-Zenodo-Dep="13330225" LSID="urn:lsid:plazi:treatment:03B3FE717728FFE474FD43E0A924EA65" httpUri="http://treatment.plazi.org/id/03B3FE717728FFE474FD43E0A924EA65" lastPageId="10" lastPageNumber="338" pageId="8" pageNumber="336">
<subSubSection id="C3001CEC7728FFE674FD43E0A8F9E8C6" box="[151,662,967,994]" pageId="8" pageNumber="336" type="nomenclature">
<paragraph id="8BA54F677728FFE674FD43E0A8F9E8C6" blockId="8.[151,662,967,1030]" box="[151,662,967,994]" pageId="8" pageNumber="336">
<heading id="D0EDF80B7728FFE674FD43E0A8F9E8C6" bold="true" box="[151,662,967,994]" fontSize="11" level="1" pageId="8" pageNumber="336" reason="1">
<taxonomicName id="4C1A34E47728FFE674FD43E0A8F9E8C6" ID-CoL="49YM4" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" box="[151,662,967,994]" class="Insecta" family="Carabidae" genus="Ophionea" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="336" phylum="Arthropoda" rank="species" species="bhamoensis">
<emphasis id="B96E93757728FFE674FD43E0A8F9E8C6" bold="true" box="[151,662,967,994]" pageId="8" pageNumber="336">
<emphasis id="B96E93757728FFE674FD43E0AB61E8C6" bold="true" box="[151,270,967,994]" italics="true" pageId="8" pageNumber="336">Ophionea</emphasis>
(s. str.)
<emphasis id="B96E93757728FFE6751B43E0A86FE8C6" bold="true" box="[369,512,967,994]" italics="true" pageId="8" pageNumber="336">bhamoensis</emphasis>
<bibRefCitation id="EF8B32967728FFE6766243EFA8F9E8C6" author="Bates, H. W." box="[520,662,968,994]" pageId="8" pageNumber="336" pagination="267 - 428" refId="ref9486" refString="Bates, H. W. (1892) List of Carabidae. Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. Annali del Museo Civico di Storia Naturale di Genova, 12 (2), 267 - 428." type="journal article" year="1892">Bates, 1892</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3001CEC7728FFE674FD43CBAB66EF22" box="[151,265,1004,1030]" pageId="8" pageNumber="336" type="description">
<paragraph id="8BA54F677728FFE674FD43CBAB66EF22" blockId="8.[151,662,967,1030]" box="[151,265,1004,1030]" pageId="8" pageNumber="336">
<heading id="D0EDF80B7728FFE674FD43CBAB66EF22" bold="true" box="[151,265,1004,1030]" fontSize="11" level="1" pageId="8" pageNumber="336" reason="1">
<figureCitation id="132153E27728FFE674FD43CBAB66EF22" box="[151,265,1004,1030]" captionStart="FIGURES 1116" captionStartId="9.[151,264,1759,1783]" captionTargetBox="[155,1426,185,1731]" captionTargetId="figure-20@9.[151,1435,181,1735]" captionTargetPageId="9" captionText="FIGURES 1116. Species of tribe Odacanthini of Philippines. (11)—Ophionea (s. str.) celebensis Baehr, 1996; (12)—Ophionea (s. str.) bhamoensis Bates, 1892; (13)—Ophionea (s.str.) ishiii Habu, 1961; (14)—Ophionea (s. str.) indica (Thunberg, 1784); (15)—Eucolliuris fuscipennis (Chaudoir, 1850); (16)—Polydamasium strandi Liebke, 1938." figureDoi="http://doi.org/10.5281/zenodo.13330261" httpUri="https://zenodo.org/record/13330261/files/figure.png" pageId="8" pageNumber="336">
<emphasis id="B96E93757728FFE674FD43CBAB66EF22" bold="true" box="[151,265,1004,1030]" pageId="8" pageNumber="336">Figure 12</emphasis>
</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3001CEC7728FFE674FD4417AFE8EF81" pageId="8" pageNumber="336" type="reference_group">
<paragraph id="8BA54F677728FFE674FD4417A9D8EF6C" blockId="8.[151,1415,1072,1189]" box="[151,951,1072,1097]" pageId="8" pageNumber="336">
<treatmentCitationGroup id="AB0A68497728FFE674FD4417A9D8EF6C" box="[151,951,1072,1097]" pageId="8" pageNumber="336">
<treatmentCitation id="0ABB69767728FFE674FD4417A816EF6D" author="Bates, H. W." box="[151,633,1072,1097]" page="380" pageId="8" pageNumber="336" year="1892">
<taxonomicName id="4C1A34E47728FFE674FD4417A816EF6D" ID-CoL="49YM4" authority="Bates, 1892: 380" authorityName="Bates" authorityPageNumber="380" authorityYear="1892" box="[151,633,1072,1097]" class="Insecta" family="Carabidae" genus="Ophionea" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="336" phylum="Arthropoda" rank="species" species="bhamoensis">
<emphasis id="B96E93757728FFE674FD4417AA94EF6C" box="[151,251,1072,1096]" italics="true" pageId="8" pageNumber="336">Ophionea</emphasis>
(s. str.)
<emphasis id="B96E93757728FFE675274417ABAAEF6C" box="[333,453,1072,1096]" italics="true" pageId="8" pageNumber="336">bhamoensis</emphasis>
<bibRefCitation id="EF8B32967728FFE675A14416A816EF6D" author="Bates, H. W." box="[459,633,1073,1097]" pageId="8" pageNumber="336" pagination="267 - 428" refId="ref9486" refString="Bates, H. W. (1892) List of Carabidae. Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. Annali del Museo Civico di Storia Naturale di Genova, 12 (2), 267 - 428." type="journal article" year="1892">Bates, 1892: 380</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Loc. Typ.: Bhamo (
<collectingCountry id="F30D0FF77728FFE677224417A9C3EF6C" box="[840,940,1072,1096]" name="Myanmar" pageId="8" pageNumber="336">Birmania</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA54F677728FFE674FD4468A960EF43" blockId="8.[151,1415,1072,1189]" box="[151,783,1103,1127]" pageId="8" pageNumber="336">
<taxonomicName id="4C1A34E47728FFE674FD4468A80BEF43" ID-CoL="49YM4" authority="sensu Maindron, 1910" authorityName="Maindron" authorityYear="1910" box="[151,612,1103,1127]" class="Insecta" family="Carabidae" genus="Ophionea" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="336" phylum="Arthropoda" rank="species" species="bhamoensis">
<emphasis id="B96E93757728FFE674FD4468AB16EF43" box="[151,377,1103,1127]" italics="true" pageId="8" pageNumber="336">Ophionea bhamoensis</emphasis>
sensu
<bibRefCitation id="EF8B32967728FFE675D44468A80BEF43" author="Maindron, M." box="[446,612,1103,1127]" pageId="8" pageNumber="336" pagination="34 - 35" refId="ref10046" refString="Maindron, M. (1910) Description d'une nouvelle espece du genre Ophionea Eschsch. (Col. Carabidae) propre a la Nouvelle- Guinee. Bulletin de la Societe entomologique de France, 1910, 34 - 35. https: // doi. org / 10.3406 / bsef. 1910.24624" type="journal article" year="1910">Maindron, 1910</bibRefCitation>
</taxonomicName>
, nec
<bibRefCitation id="EF8B32967728FFE676FD4477A960EF43" author="Bates, H. W." box="[663,783,1103,1127]" pageId="8" pageNumber="336" pagination="267 - 428" refId="ref9486" refString="Bates, H. W. (1892) List of Carabidae. Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. Annali del Museo Civico di Storia Naturale di Genova, 12 (2), 267 - 428." type="journal article" year="1892">Bates, 1892</bibRefCitation>
</paragraph>
<paragraph id="8BA54F677728FFE674FD4449AE2BEFA2" blockId="8.[151,1415,1072,1189]" box="[151,1092,1134,1158]" pageId="8" pageNumber="336">
<treatmentCitationGroup id="AB0A68497728FFE674FD4449AE2BEFA2" box="[151,1092,1134,1158]" pageId="8" pageNumber="336">
<treatmentCitation id="0ABB69767728FFE674FD4449A83CEFA2" author="Dupuis, P." box="[151,595,1134,1158]" page="270" pageId="8" pageNumber="336" year="1913">
<taxonomicName id="4C1A34E47728FFE674FD4449A83CEFA2" ID-CoL="49YM3" authority="Dupuis, 1913: 270" authorityName="Dupuis" authorityPageNumber="270" authorityYear="1913" box="[151,595,1134,1158]" class="Insecta" family="Carabidae" genus="Ophionea" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="336" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="B96E93757728FFE674FD4449AA94EFA2" box="[151,251,1134,1158]" italics="true" pageId="8" pageNumber="336">Ophionea</emphasis>
(s. str.)
<emphasis id="B96E93757728FFE675274449ABE1EFA2" box="[333,398,1134,1158]" italics="true" pageId="8" pageNumber="336">bakeri</emphasis>
<bibRefCitation id="EF8B32967728FFE675FE4449A83CEFA2" author="Dupuis, P." box="[404,595,1134,1158]" pageId="8" pageNumber="336" pagination="270 - 277" refId="ref9611" refString="Dupuis, P. (1913) Notes sur les Carabiques des iles Philippines. Annales de la Societe entomologique Belge, 57, 270 - 277." type="journal article" year="1913">Dupuis, 1913: 270</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Loc. Typ.: Los Banos, Iles
<collectingCountry id="F30D0FF77728FFE677054449A98EEFA2" box="[879,993,1134,1158]" name="Philippines" pageId="8" pageNumber="336">Philippines</collectingCountry>
<emphasis id="B96E93757728FFE6778D4449AE2BEFA2" bold="true" box="[999,1092,1134,1158]" pageId="8" pageNumber="336">
<taxonomicNameLabel id="A25D2E0E7728FFE6778D4449AE2BEFA2" box="[999,1092,1134,1158]" pageId="8" pageNumber="336">syn. nov.</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA54F677728FFE674FD44ABAFE8EF81" blockId="8.[151,1415,1072,1189]" box="[151,1415,1164,1189]" pageId="8" pageNumber="336">
<treatmentCitationGroup id="AB0A68497728FFE674FD44ABAFE8EF81" box="[151,1415,1164,1189]" pageId="8" pageNumber="336">
<treatmentCitation id="0ABB69767728FFE674FD44ABA881EF81" author="Terada, K. &amp; Wu, W." box="[151,750,1164,1189]" page="19" pageId="8" pageNumber="336" year="2014">
<taxonomicName id="4C1A34E47728FFE674FD44ABA881EF81" ID-CoL="8MK73" authority="Terada &amp; Wu, 2014: 19" authorityName="Terada &amp; Wu" authorityPageNumber="19" authorityYear="2014" box="[151,750,1164,1189]" class="Insecta" family="Carabidae" genus="Ophionea" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="336" phylum="Arthropoda" rank="subSpecies" species="bhamoensis" subSpecies="taiwanensis">
<emphasis id="B96E93757728FFE674FD44ABAB98EF80" box="[151,503,1164,1188]" italics="true" pageId="8" pageNumber="336">Ophionea bhamoensis taiwanensis</emphasis>
<bibRefCitation id="EF8B32967728FFE6759744ABA881EF81" author="Terada, K. &amp; Wu, W." box="[509,750,1164,1189]" pageId="8" pageNumber="336" pagination="15 - 41" refId="ref10136" refString="Terada, K. &amp; Wu, W. (2014) Notes on Taiwanese Caraboidea (Coleoptera) V. A Review of the Tribe Odacanthini (Carabidae) in Taiwan, with Description of Ophionea bhamoensis taiwanensis subsp. nov. Collection and Research, 27, 15 - 41." type="journal article" year="2014">Terada &amp; Wu, 2014: 19</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Loc. Typ.:
<collectingCountry id="F30D0FF77728FFE6770C44ABA9DCEF80" box="[870,947,1164,1188]" name="Taiwan" pageId="8" pageNumber="336">Taiwan</collectingCountry>
,
<collectingRegion id="49DE81857728FFE677D644ABAE2AEF81" box="[956,1093,1164,1189]" country="Taiwan" name="Yilan" pageId="8" pageNumber="336">Yilan County</collectingRegion>
, Nanao, Tawan River.
<emphasis id="B96E93757728FFE6714044AAAFE8EF81" bold="true" box="[1322,1415,1165,1189]" pageId="8" pageNumber="336">
<taxonomicNameLabel id="A25D2E0E7728FFE6714044AAAFE8EF81" box="[1322,1415,1165,1189]" pageId="8" pageNumber="336">syn. nov.</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3001CEC7728FFE674FD44F5A943EEE1" pageId="8" pageNumber="336" type="materials_examined">
<paragraph id="8BA54F677728FFE674FD44F5A943EEE1" blockId="8.[151,1437,1234,1945]" pageId="8" pageNumber="336">
<emphasis id="B96E93757728FFE674FD44F5AB66EFC8" bold="true" box="[151,265,1234,1260]" pageId="8" pageNumber="336">Material:</emphasis>
<materialsCitation id="3B72453A7728FFE6756544F4AA94EE7D" collectingDate="1885-07" collectingDateMax="2015-08" collectingDateMin="1885-07" collectionCode="BMNH, DUBC" collectorName="H. E. Andrewes" country="Philippines" county="Cotype" location="Fea" municipality="Bhamo" pageId="8" pageNumber="336" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Nueva Vizcaya">
1 ex: “
<collectingCounty id="62C437EB7728FFE6753244F4ABDFEFC9" box="[344,432,1235,1261]" pageId="8" pageNumber="336">Cotype</collectingCounty>
”, “
<collectingMunicipality id="6BC1D51D7728FFE675A544F4A84DEFC9" box="[463,546,1235,1261]" pageId="8" pageNumber="336">Bhamo</collectingMunicipality>
/ Birmania /
<location id="8EC519BC7728FFE676DF44F4A8B2EFC9" LSID="urn:lsid:plazi:treatment:03B3FE717728FFE474FD43E0A924EA65:8EC519BC7728FFE676DF44F4A8B2EFC9" box="[693,733,1235,1261]" country="Philippines" county="Cotype" municipality="Bhamo" name="Fea" pageId="8" pageNumber="336" stateProvince="Nueva Vizcaya">Fea</location>
<date id="FFA469A77728FFE6768944F4A922EFC9" box="[739,845,1234,1261]" pageId="8" pageNumber="336" value="1885-07">
<collectingDate id="EFE0904F7728FFE6768944F4A922EFC9" box="[739,845,1234,1261]" pageId="8" pageNumber="336" value="1885-07">VII 1885</collectingDate>
</date>
”, “
<location id="8EC519BC7728FFE6770444F4A9C8EFC9" LSID="urn:lsid:plazi:treatment:03B3FE717728FFE474FD43E0A924EA65:8EC519BC7728FFE6770444F4A9C8EFC9" box="[878,935,1235,1261]" country="Philippines" county="Cotype" municipality="Bhamo" name="Oph." pageId="8" pageNumber="336" stateProvince="Nueva Vizcaya">Oph.</location>
/
<location id="8EC519BC7728FFE677D644F4AE48EFC9" LSID="urn:lsid:plazi:treatment:03B3FE717728FFE474FD43E0A924EA65:8EC519BC7728FFE677D644F4AE48EFC9" box="[956,1063,1235,1261]" country="Philippines" county="Cotype" municipality="Bhamo" name="Bhamoen" pageId="8" pageNumber="336" stateProvince="Nueva Vizcaya">Bhamoen</location>
/ sis B. / n. sp.”, “
<collectorName id="26EF2AB17728FFE6709B44F4AFF2EFC9" box="[1265,1437,1235,1261]" pageId="8" pageNumber="336">H.E. Andrewes</collectorName>
coll. / B.M.1945-97” (
<collectionCode id="ED0BD7A27728FFE675F044D0AB9BEE35" box="[410,500,1271,1297]" pageId="8" pageNumber="336">BMNH</collectionCode>
);
<specimenCount id="9D1C84EE7728FFE6766144D0A853EE35" box="[523,572,1271,1297]" count="1" pageId="8" pageNumber="336" type="male">1 ♂</specimenCount>
: “
<collectingCountry id="F30D0FF77728FFE6763F44D0A8B7EE35" box="[597,728,1271,1297]" name="Philippines" pageId="8" pageNumber="336">Philippines</collectingCountry>
, Mindoro,
<location id="8EC519BC7728FFE6773044D0A98FEE35" LSID="urn:lsid:plazi:treatment:03B3FE717728FFE474FD43E0A924EA65:8EC519BC7728FFE6773044D0A98FEE35" box="[858,992,1271,1297]" country="Philippines" county="Cotype" municipality="Bhamo" name="Mt. Halcon" pageId="8" pageNumber="336" stateProvince="Nueva Vizcaya">Mt. Halcon</location>
,
<date id="FFA469A77728FFE6778544D0AE2BEE35" box="[1007,1092,1271,1297]" pageId="8" pageNumber="336" value="2013-10">
<collectingDate id="EFE0904F7728FFE6778544D0AE2BEE35" box="[1007,1092,1271,1297]" pageId="8" pageNumber="336" value="2013-10">X.2013</collectingDate>
</date>
” (
<collectionCode id="ED0BD7A27728FFE6700B44D0AEDAEE35" box="[1121,1205,1271,1297]" pageId="8" pageNumber="336">DUBC</collectionCode>
);
<specimenCount id="9D1C84EE7728FFE670A144D0AE94EE35" box="[1227,1275,1271,1297]" count="1" pageId="8" pageNumber="336" type="female">1 ♀</specimenCount>
: “
<collectingCountry id="F30D0FF77728FFE6717F44D0AFF6EE35" box="[1301,1433,1271,1297]" name="Philippines" pageId="8" pageNumber="336">Philippines</collectingCountry>
. N Luzon,
<collectingRegion id="49DE81857728FFE67565453CABD0EE11" box="[271,447,1307,1333]" country="Philippines" name="Nueva Vizcaya" pageId="8" pageNumber="336">Nueva Vizcaya</collectingRegion>
, Belance,
<date id="FFA469A77728FFE67653453DA8E9EE11" box="[569,646,1306,1333]" pageId="8" pageNumber="336" value="2015-05">
<collectingDate id="EFE0904F7728FFE67653453DA8E9EE11" box="[569,646,1306,1333]" pageId="8" pageNumber="336" value="2015-05">v.2015</collectingDate>
</date>
” (
<collectionCode id="ED0BD7A27728FFE676C9453CA898EE11" box="[675,759,1307,1333]" pageId="8" pageNumber="336">DUBC</collectionCode>
);
<specimenCount id="9D1C84EE7728FFE67764453CA952EE11" box="[782,829,1307,1333]" count="1" pageId="8" pageNumber="336" type="female">1 ♀</specimenCount>
: “
<collectingCountry id="F30D0FF77728FFE6773D453CA9B5EE11" box="[855,986,1307,1333]" name="Philippines" pageId="8" pageNumber="336">Philippines</collectingCountry>
, N Luzon,
<collectingRegion id="49DE81857728FFE6700B453CAEC2EE11" box="[1121,1197,1307,1333]" country="Philippines" name="Ifugao" pageId="8" pageNumber="336">Ifugao</collectingRegion>
, Banaue,
<date id="FFA469A77728FFE6714B453CAFFDEE11" box="[1313,1426,1306,1333]" pageId="8" pageNumber="336" value="2015-08">
<collectingDate id="EFE0904F7728FFE6714B453CAFFDEE11" box="[1313,1426,1306,1333]" pageId="8" pageNumber="336" value="2015-08">viii. 2015</collectingDate>
</date>
” (
<collectionCode id="ED0BD7A27728FFE674F54518AA9CEE7D" box="[159,243,1343,1369]" pageId="8" pageNumber="336">DUBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B72453A7728FFE675624518A9A3EE7D" box="[264,972,1343,1369]" collectingDate="2014-03" collectionCode="DUBC" country="Philippines" location="San Mariano" pageId="8" pageNumber="336" specimenCount="1" stateProvince="Isabela">
1 ex: “
<location id="8EC519BC7728FFE6753F4518AB83EE7D" LSID="urn:lsid:plazi:treatment:03B3FE717728FFE474FD43E0A924EA65:8EC519BC7728FFE6753F4518AB83EE7D" box="[341,492,1343,1369]" country="Philippines" name="San Mariano" pageId="8" pageNumber="336" stateProvince="Isabela">San Mariano</location>
,
<collectingRegion id="49DE81857728FFE675924518A825EE7D" box="[504,586,1343,1369]" country="Philippines" name="Isabela" pageId="8" pageNumber="336">Isabela</collectingRegion>
, E Luzon,
<date id="FFA469A77728FFE676A34518A93AEE7D" box="[713,853,1343,1369]" pageId="8" pageNumber="336" value="2014-03">
<collectingDate id="EFE0904F7728FFE676A34518A93AEE7D" box="[713,853,1343,1369]" pageId="8" pageNumber="336" value="2014-03">March 2014</collectingDate>
</date>
” (
<collectionCode id="ED0BD7A27728FFE6771A4518A9ABEE7D" box="[880,964,1343,1369]" pageId="8" pageNumber="336">DUBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B72453A7728FFE677B04518AB10EE59" collectingDate="2014-12" collectionCode="DUBC" country="Philippines" location="Sierra Madre" pageId="8" pageNumber="336" specimenCount="1" stateProvince="Quirino">
3 ex: “
<location id="8EC519BC7728FFE6704F4518AEAFEE7D" LSID="urn:lsid:plazi:treatment:03B3FE717728FFE474FD43E0A924EA65:8EC519BC7728FFE6704F4518AEAFEE7D" box="[1061,1216,1343,1369]" country="Philippines" name="Sierra Madre" pageId="8" pageNumber="336" stateProvince="Quirino">Sierra Madre</location>
,
<collectingRegion id="49DE81857728FFE670A74518AF48EE7D" box="[1229,1319,1343,1369]" country="Philippines" name="Quirino" pageId="8" pageNumber="336">Quirino</collectingRegion>
, E Luzon,
<date id="FFA469A77728FFE674FD4544AB64EE59" box="[151,267,1379,1405]" pageId="8" pageNumber="336" value="2014-12">
<collectingDate id="EFE0904F7728FFE674FD4544AB64EE59" box="[151,267,1379,1405]" pageId="8" pageNumber="336" value="2014-12">Dec. 2014</collectingDate>
</date>
” (
<collectionCode id="ED0BD7A27728FFE675494544AB18EE59" box="[291,375,1379,1405]" pageId="8" pageNumber="336">DUBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B72453A7728FFE675E34544AEF6EE59" box="[393,1177,1379,1405]" collectingDate="2014-12" collectionCode="DUBC" country="Philippines" location="Gutalac" pageId="8" pageNumber="336" specimenCount="1" stateProvince="Zamboanga del Norte">
3 ex: “
<location id="8EC519BC7728FFE675A54544A844EE59" LSID="urn:lsid:plazi:treatment:03B3FE717728FFE474FD43E0A924EA65:8EC519BC7728FFE675A54544A844EE59" box="[463,555,1379,1405]" country="Philippines" name="Gutalac" pageId="8" pageNumber="336" stateProvince="Zamboanga del Norte">Gutalac</location>
,
<collectingRegion id="49DE81857728FFE6765F4544A946EE59" box="[565,809,1379,1405]" country="Philippines" name="Zamboanga del Norte" pageId="8" pageNumber="336">Zamboanga del Norte</collectingRegion>
, Mindanao,
<date id="FFA469A77728FFE677DB4544AE4AEE59" box="[945,1061,1379,1405]" pageId="8" pageNumber="336" value="2014-12">
<collectingDate id="EFE0904F7728FFE677DB4544AE4AEE59" box="[945,1061,1379,1405]" pageId="8" pageNumber="336" value="2014-12">Dec. 2014</collectingDate>
</date>
” (
<collectionCode id="ED0BD7A27728FFE670574544AEFEEE59" box="[1085,1169,1379,1405]" pageId="8" pageNumber="336">DUBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B72453A7728FFE670C94544AB20EE85" collectingDate="2020-11" collectionCode="RSc" country="Philippines" location="Roxas" pageId="8" pageNumber="336" specimenCount="1" stateProvince="Palawan">
1 ex: “
<location id="8EC519BC7728FFE670834544AF5BEE59" LSID="urn:lsid:plazi:treatment:03B3FE717728FFE474FD43E0A924EA65:8EC519BC7728FFE670834544AF5BEE59" box="[1257,1332,1379,1405]" country="Philippines" name="Roxas" pageId="8" pageNumber="336" stateProvince="Palawan">Roxas</location>
,
<collectingRegion id="49DE81857728FFE671574544AFF3EE59" box="[1341,1436,1379,1405]" country="Philippines" name="Palawan" pageId="8" pageNumber="336">Palawan</collectingRegion>
<date id="FFA469A77728FFE674FD45A0AA99EE85" box="[151,246,1415,1441]" pageId="8" pageNumber="336" value="2020-11">
<collectingDate id="EFE0904F7728FFE674FD45A0AA99EE85" box="[151,246,1415,1441]" pageId="8" pageNumber="336" value="2020-11">XI.2020</collectingDate>
</date>
” (
<collectionCode id="ED0BD7A27728FFE6757945A0AB28EE85" box="[275,327,1415,1441]" pageId="8" pageNumber="336">RSc</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B72453A7728FFE6753545A0A8A9EE85" box="[351,710,1415,1441]" collectionCode="RSc" country="Philippines" location="Manila" pageId="8" pageNumber="336" specimenCount="1" stateProvince="Manila">
1 ex: “
<collectingRegion id="49DE81857728FFE675C545A0A86CEE85" box="[431,515,1415,1441]" country="Philippines" name="Manila" pageId="8" pageNumber="336">Manila</collectingRegion>
, Philipp.” (
<collectionCode id="ED0BD7A27728FFE676E045A0A8D1EE85" box="[650,702,1415,1441]" pageId="8" pageNumber="336">RSc</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B72453A7728FFE676BC45A0AF7FEE85" box="[726,1296,1415,1441]" collectingDate="2019-11" collectionCode="RSc" country="Philippines" location="Roxas" municipality="Magara" pageId="8" pageNumber="336" specimenCount="1" stateProvince="Palawan">
2 ex: “
<collectingMunicipality id="6BC1D51D7728FFE6774C45A0A9EEEE85" box="[806,897,1415,1441]" pageId="8" pageNumber="336">Magara</collectingMunicipality>
,
<location id="8EC519BC7728FFE677E545A0A9B7EE85" LSID="urn:lsid:plazi:treatment:03B3FE717728FFE474FD43E0A924EA65:8EC519BC7728FFE677E545A0A9B7EE85" box="[911,984,1415,1441]" country="Philippines" municipality="Magara" name="Roxas" pageId="8" pageNumber="336" stateProvince="Palawan">Roxas</location>
,
<collectingRegion id="49DE81857728FFE6778D45A0AE26EE85" box="[999,1097,1415,1441]" country="Philippines" name="Palawan" pageId="8" pageNumber="336">Palawan</collectingRegion>
,
<date id="FFA469A77728FFE6703245A0AED8EE85" box="[1112,1207,1415,1441]" pageId="8" pageNumber="336" value="2019-11">
<collectingDate id="EFE0904F7728FFE6703245A0AED8EE85" box="[1112,1207,1415,1441]" pageId="8" pageNumber="336" value="2019-11">XI.2019</collectingDate>
</date>
” (
<collectionCode id="ED0BD7A27728FFE670BE45A0AF67EE85" box="[1236,1288,1415,1441]" pageId="8" pageNumber="336">RSc</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B72453A7728FFE6714A45A0A87DEEE1" collectingDate="2019-10" collectionCode="RSc" country="Saint Vincent and the Grenadines" location="Palawan" pageId="8" pageNumber="336" specimenCount="1">
1 ex: “
<collectingCountry id="F30D0FF77728FFE6711A45A0AA80EEE1" name="Saint Vincent and the Grenadines" pageId="8" pageNumber="336">San Vicente</collectingCountry>
,
<location id="8EC519BC7728FFE67491458CAB32EEE1" LSID="urn:lsid:plazi:treatment:03B3FE717728FFE474FD43E0A924EA65:8EC519BC7728FFE67491458CAB32EEE1" box="[251,349,1451,1477]" country="Saint Vincent and the Grenadines" name="Palawan" pageId="8" pageNumber="336">Palawan</location>
,
<date id="FFA469A77728FFE6750D458CABD2EEE1" box="[359,445,1451,1477]" pageId="8" pageNumber="336" value="2019-10">
<collectingDate id="EFE0904F7728FFE6750D458CABD2EEE1" box="[359,445,1451,1477]" pageId="8" pageNumber="336" value="2019-10">X.2019</collectingDate>
</date>
” (
<collectionCode id="ED0BD7A27728FFE675BC458CA865EEE1" box="[470,522,1451,1477]" pageId="8" pageNumber="336">RSc</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B72453A7728FFE67674458CA947EEE1" box="[542,808,1451,1477]" collectingDate="2019-11" collectionCode="RSc" country="Saint Vincent and the Grenadines" pageId="8" pageNumber="336" specimenCount="1">
1 ex: id,
<date id="FFA469A77728FFE676EA458CA8B3EEE1" box="[640,732,1451,1477]" pageId="8" pageNumber="336" value="2019-11">
<collectingDate id="EFE0904F7728FFE676EA458CA8B3EEE1" box="[640,732,1451,1477]" pageId="8" pageNumber="336" value="2019-11">XI.2019</collectingDate>
</date>
(
<collectionCode id="ED0BD7A27728FFE67686458CA94FEEE1" box="[748,800,1451,1477]" pageId="8" pageNumber="336">RSc</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3001CEC7728FFE474AD45E9A924EA65" lastPageId="10" lastPageNumber="338" pageId="8" pageNumber="336" type="distribution">
<paragraph id="8BA54F677728FFE674AD45E9AA91ED29" blockId="8.[151,1437,1234,1945]" pageId="8" pageNumber="336">
<emphasis id="B96E93757728FFE674AD45E9AB0CEECC" bold="true" box="[199,355,1486,1512]" pageId="8" pageNumber="336">Distribution:</emphasis>
<collectingCountry id="F30D0FF77728FFE6750245E8ABCBEECD" box="[360,420,1487,1513]" name="India" pageId="8" pageNumber="336">India</collectingCountry>
,
<collectingCountry id="F30D0FF77728FFE675C545E8A871EECD" box="[431,542,1487,1513]" name="Myanmar" pageId="8" pageNumber="336">Myanmar</collectingCountry>
,
<collectingCountry id="F30D0FF77728FFE6764D45E8A8CAEECD" box="[551,677,1487,1513]" name="Philippines" pageId="8" pageNumber="336">Philippines</collectingCountry>
(Luzon, first citation for
<collectingRegion id="49DE81857728FFE677D145E8AE72EECD" box="[955,1053,1487,1513]" country="Philippines" name="Palawan" pageId="8" pageNumber="336">Palawan</collectingRegion>
, Mindoro and Mindanao),
<collectingCountry id="F30D0FF77728FFE6712945E8AFF7EECD" box="[1347,1432,1487,1513]" name="Taiwan" pageId="8" pageNumber="336">Taiwan</collectingCountry>
,
<collectingCountry id="F30D0FF77728FFE674FD45D4AA95ED29" box="[151,250,1523,1549]" name="Vietnam" pageId="8" pageNumber="336">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="8BA54F677728FFE674AD4631A94EEC2D" blockId="8.[151,1437,1234,1945]" pageId="8" pageNumber="336">
<emphasis id="B96E93757728FFE674AD4631ABFFED15" bold="true" box="[199,400,1558,1585]" pageId="8" pageNumber="336">Systematic notes</emphasis>
: The possible synonymy of
<taxonomicName id="4C1A34E47728FFE676B04630A986ED15" authority="Dupuis, 1913" authorityName="Dupuis" authorityYear="1913" box="[730,1001,1559,1585]" class="Insecta" family="Carabidae" genus="Ophionea" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="336" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="B96E93757728FFE676B04630A929ED15" box="[730,838,1559,1585]" italics="true" pageId="8" pageNumber="336">O. bakeri</emphasis>
<bibRefCitation id="EF8B32967728FFE677254630A986ED15" author="Dupuis, P." box="[847,1001,1559,1585]" pageId="8" pageNumber="336" pagination="270 - 277" refId="ref9611" refString="Dupuis, P. (1913) Notes sur les Carabiques des iles Philippines. Annales de la Societe entomologique Belge, 57, 270 - 277." type="journal article" year="1913">Dupuis, 1913</bibRefCitation>
</taxonomicName>
with
<taxonomicName id="4C1A34E47728FFE670474630AF06ED15" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" box="[1069,1385,1559,1585]" class="Insecta" family="Carabidae" genus="Ophionea" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="336" phylum="Arthropoda" rank="species" species="bhamoensis">
<emphasis id="B96E93757728FFE670474630AEB9ED15" box="[1069,1238,1559,1585]" italics="true" pageId="8" pageNumber="336">O. bhamoensis</emphasis>
<bibRefCitation id="EF8B32967728FFE6708A4630AF06ED15" author="Bates, H. W." box="[1248,1385,1559,1585]" pageId="8" pageNumber="336" pagination="267 - 428" refId="ref9486" refString="Bates, H. W. (1892) List of Carabidae. Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. Annali del Museo Civico di Storia Naturale di Genova, 12 (2), 267 - 428." type="journal article" year="1892">Bates, 1892</bibRefCitation>
</taxonomicName>
, has already been discussed by
<bibRefCitation id="EF8B32967728FFE675AF461CA835ED71" author="Baehr, M." box="[453,602,1595,1621]" pageId="8" pageNumber="336" pagination="1041 - 1084" refId="ref9337" refString="Baehr, M. (1996) The ground beetle genus Casnoidea Castelnau, Taxonomy, Phylogeny, Zoogeography (Insecta, Coleoptera, Carabidae, Odacanthinae). Invertebrate Taxonomy, 10, 1041 - 1084. https: // doi. org / 10.1071 / IT 9961041" type="journal article" year="1996">Baehr (1996)</bibRefCitation>
and
<bibRefCitation id="EF8B32967728FFE676FB461CA9FDED71" author="Terada, K. &amp; Wu, W." box="[657,914,1595,1621]" pageId="8" pageNumber="336" pagination="15 - 41" refId="ref10136" refString="Terada, K. &amp; Wu, W. (2014) Notes on Taiwanese Caraboidea (Coleoptera) V. A Review of the Tribe Odacanthini (Carabidae) in Taiwan, with Description of Ophionea bhamoensis taiwanensis subsp. nov. Collection and Research, 27, 15 - 41." type="journal article" year="2014">Terada and Wu (2014)</bibRefCitation>
. The study of numerous specimens has shown that the two features used to distinguish between these two species are extremely unstable.
<bibRefCitation id="EF8B32967728FFE670CD4678AF53ED5D" author="Baehr, M." box="[1191,1340,1631,1657]" pageId="8" pageNumber="336" pagination="1041 - 1084" refId="ref9337" refString="Baehr, M. (1996) The ground beetle genus Casnoidea Castelnau, Taxonomy, Phylogeny, Zoogeography (Insecta, Coleoptera, Carabidae, Odacanthinae). Invertebrate Taxonomy, 10, 1041 - 1084. https: // doi. org / 10.1071 / IT 9961041" type="journal article" year="1996">Baehr (1996)</bibRefCitation>
affirms, only on the basis of the original description (admitting that he has not directly seen any specimen of this species), that in
<taxonomicName id="4C1A34E47728FFE6748C4680ABE2EDE5" authorityName="Bates" authorityYear="1892" box="[230,397,1703,1729]" class="Insecta" family="Carabidae" genus="Ophionea" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="336" phylum="Arthropoda" rank="species" species="bhamoensis">
<emphasis id="B96E93757728FFE6748C4680ABE2EDE5" box="[230,397,1703,1729]" italics="true" pageId="8" pageNumber="336">O. bhamoensis</emphasis>
</taxonomicName>
the anterior white elytral spot is outside the dark fascia, while the posterior spot is included in fascia, while in
<taxonomicName id="4C1A34E47728FFE6750346ECABBAEDC1" authorityName="Dupuis" authorityYear="1913" box="[361,469,1739,1765]" class="Insecta" family="Carabidae" genus="Ophionea" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="336" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="B96E93757728FFE6750346ECABBAEDC1" box="[361,469,1739,1765]" italics="true" pageId="8" pageNumber="336">O. bakeri</emphasis>
</taxonomicName>
both spots are at least partly included in dark fascia. In the Cotype of
<taxonomicName id="4C1A34E47728FFE6709F46ECAFF3EDC1" authorityName="Bates" authorityYear="1892" box="[1269,1436,1739,1765]" class="Insecta" family="Carabidae" genus="Ophionea" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="336" phylum="Arthropoda" rank="species" species="bhamoensis">
<emphasis id="B96E93757728FFE6709F46ECAFF3EDC1" box="[1269,1436,1739,1765]" italics="true" pageId="8" pageNumber="336">O. bhamoensis</emphasis>
</taxonomicName>
examined by us, the posterior spot is only partly included.
</paragraph>
<paragraph id="8BA54F677728FFE674AD4734AE59ECBD" blockId="8.[151,1437,1234,1945]" pageId="8" pageNumber="336">
The number of discal pores in third interval of elytra should be
<quantity id="4CE2E2827728FFE677C34735A99CEC09" box="[937,1011,1810,1837]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.524" metricValueMin="1.27" pageId="8" pageNumber="336" unit="in" value="5.5" valueMax="6.0" valueMin="5.0">56 in</quantity>
<taxonomicName id="4C1A34E47728FFE677964734AECBEC09" authorityName="Bates" authorityYear="1892" box="[1020,1188,1811,1837]" class="Insecta" family="Carabidae" genus="Ophionea" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="336" phylum="Arthropoda" rank="species" species="bhamoensis">
<emphasis id="B96E93757728FFE677964734AECBEC09" box="[1020,1188,1811,1837]" italics="true" pageId="8" pageNumber="336">O. bhamoensis</emphasis>
</taxonomicName>
vs three in
<taxonomicName id="4C1A34E47728FFE6715A4734AB11EC75" authority="(Terada &amp; Wu 2014)" baseAuthorityName="Terada &amp; Wu" baseAuthorityYear="2014" class="Insecta" family="Carabidae" genus="Ophionea" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="336" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="B96E93757728FFE6715A4734AFF2EC09" box="[1328,1437,1811,1837]" italics="true" pageId="8" pageNumber="336">O. bakeri</emphasis>
(
<bibRefCitation id="EF8B32967728FFE674F44710AB19EC75" author="Terada, K. &amp; Wu, W." box="[158,374,1847,1873]" pageId="8" pageNumber="336" pagination="15 - 41" refId="ref10136" refString="Terada, K. &amp; Wu, W. (2014) Notes on Taiwanese Caraboidea (Coleoptera) V. A Review of the Tribe Odacanthini (Carabidae) in Taiwan, with Description of Ophionea bhamoensis taiwanensis subsp. nov. Collection and Research, 27, 15 - 41." type="journal article" year="2014">Terada &amp; Wu 2014</bibRefCitation>
)
</taxonomicName>
, yet, in
<specimenCount id="9D1C84EE7728FFE675B84710A805EC75" box="[466,618,1847,1873]" count="1" pageId="8" pageNumber="336" type="generic">one specimen</specimenCount>
from the
<collectingCountry id="F30D0FF77728FFE676B94710A93EEC75" box="[723,849,1847,1873]" name="Philippines" pageId="8" pageNumber="336">Philippines</collectingCountry>
that we examined has five discal pores.Additionally, the aedeagus is extremely similar in both species. Therefore, we propose here a new synonymy:
<taxonomicName id="4C1A34E47728FFE670B2477CABC8ECBD" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" class="Insecta" family="Carabidae" genus="Ophionea" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="336" phylum="Arthropoda" rank="species" species="bhamoensis">
<emphasis id="B96E93757728FFE670B2477CAF29EC51" box="[1240,1350,1883,1909]" italics="true" pageId="8" pageNumber="336">Ophionea</emphasis>
(s. str.)
<emphasis id="B96E93757728FFE674FD4758AB74ECBD" box="[151,283,1919,1945]" italics="true" pageId="8" pageNumber="336">bhamoensis</emphasis>
<bibRefCitation id="EF8B32967728FFE675484758ABC8ECBD" author="Bates, H. W." box="[290,423,1919,1945]" pageId="8" pageNumber="336" pagination="267 - 428" refId="ref9486" refString="Bates, H. W. (1892) List of Carabidae. Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. Annali del Museo Civico di Storia Naturale di Genova, 12 (2), 267 - 428." type="journal article" year="1892">Bates, 1892</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="4C1A34E47728FFE675AE4758A91CECBD" authority="Dupuis, 1913" authorityName="Dupuis" authorityYear="1913" box="[452,883,1919,1945]" class="Insecta" family="Carabidae" genus="Ophionea" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="336" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="B96E93757728FFE675AE4758A85DECBD" box="[452,562,1919,1945]" italics="true" pageId="8" pageNumber="336">Ophionea</emphasis>
(s. str.)
<emphasis id="B96E93757728FFE676E44758A8B9ECBD" box="[654,726,1919,1945]" italics="true" pageId="8" pageNumber="336">bakeri</emphasis>
<bibRefCitation id="EF8B32967728FFE676B64758A91CECBD" author="Dupuis, P." box="[732,883,1919,1945]" pageId="8" pageNumber="336" pagination="270 - 277" refId="ref9611" refString="Dupuis, P. (1913) Notes sur les Carabiques des iles Philippines. Annales de la Societe entomologique Belge, 57, 270 - 277." type="journal article" year="1913">Dupuis, 1913</bibRefCitation>
</taxonomicName>
<emphasis id="B96E93757728FFE677104759AE59ECBD" bold="true" box="[890,1078,1918,1945]" pageId="8" pageNumber="336">junior synonym</emphasis>
</paragraph>
<caption id="DF651FEF7729FFE774FD46F8AE2FEC1B" ID-DOI="http://doi.org/10.5281/zenodo.13330261" ID-Zenodo-Dep="13330261" httpUri="https://zenodo.org/record/13330261/files/figure.png" pageId="9" pageNumber="337" startId="9.[151,264,1759,1783]" targetBox="[155,1426,185,1731]" targetPageId="9" targetType="figure">
<paragraph id="8BA54F677729FFE774FD46F8AE2FEC1B" blockId="9.[151,1436,1759,1855]" pageId="9" pageNumber="337">
<emphasis id="B96E93757729FFE774FD46F8AB3EEDD3" bold="true" box="[151,337,1759,1783]" pageId="9" pageNumber="337">FIGURES 1116.</emphasis>
Species of tribe
<taxonomicName id="4C1A34E47729FFE7759C46F8A81DEDD3" authorityName="Laporte" authorityYear="1834" box="[502,626,1759,1783]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="337" phylum="Arthropoda" rank="tribe" tribe="Odacanthini">Odacanthini</taxonomicName>
of Philippines. (11)—
<taxonomicName id="4C1A34E47729FFE7773A46F8AE83EDD3" authority="Baehr, 1996" authorityName="Baehr" authorityYear="1996" box="[848,1260,1759,1783]" class="Insecta" family="Carabidae" genus="Ophionea" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="337" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis id="B96E93757729FFE7773A46F8A9DBEDD3" box="[848,948,1759,1783]" italics="true" pageId="9" pageNumber="337">Ophionea</emphasis>
(s. str.)
<emphasis id="B96E93757729FFE7706846F8AE05EDD3" box="[1026,1130,1759,1783]" italics="true" pageId="9" pageNumber="337">celebensis</emphasis>
<bibRefCitation id="EF8B32967729FFE7700546F8AE83EDD3" author="Baehr, M." box="[1135,1260,1759,1783]" pageId="9" pageNumber="337" pagination="1041 - 1084" refId="ref9337" refString="Baehr, M. (1996) The ground beetle genus Casnoidea Castelnau, Taxonomy, Phylogeny, Zoogeography (Insecta, Coleoptera, Carabidae, Odacanthinae). Invertebrate Taxonomy, 10, 1041 - 1084. https: // doi. org / 10.1071 / IT 9961041" type="journal article" year="1996">Baehr, 1996</bibRefCitation>
</taxonomicName>
; (12)—
<taxonomicName id="4C1A34E47729FFE7715246F8AB8FEC3F" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" class="Insecta" family="Carabidae" genus="Ophionea" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="337" phylum="Arthropoda" rank="species" species="bhamoensis">
<emphasis id="B96E93757729FFE7715246F8AFF3EDD3" box="[1336,1436,1759,1783]" italics="true" pageId="9" pageNumber="337">Ophionea</emphasis>
(s. str.)
<emphasis id="B96E93757729FFE7748F4724AB32EC3F" box="[229,349,1795,1819]" italics="true" pageId="9" pageNumber="337">bhamoensis</emphasis>
<bibRefCitation id="EF8B32967729FFE7750E4724AB8FEC3F" author="Bates, H. W." box="[356,480,1795,1819]" pageId="9" pageNumber="337" pagination="267 - 428" refId="ref9486" refString="Bates, H. W. (1892) List of Carabidae. Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. Annali del Museo Civico di Storia Naturale di Genova, 12 (2), 267 - 428." type="journal article" year="1892">Bates, 1892</bibRefCitation>
</taxonomicName>
; (13)—
<taxonomicName id="4C1A34E47729FFE776444724A9FBEC3F" authority="Habu, 1961" authorityName="Habu" authorityYear="1961" box="[558,916,1795,1819]" class="Insecta" family="Carabidae" genus="Ophionea" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="337" phylum="Arthropoda" rank="species" species="ishiii">
<emphasis id="B96E93757729FFE776444724A8FDEC3F" box="[558,658,1795,1819]" italics="true" pageId="9" pageNumber="337">Ophionea</emphasis>
(s.str.)
<emphasis id="B96E93757729FFE7768A4724A97CEC3F" box="[736,787,1795,1819]" italics="true" pageId="9" pageNumber="337">ishiii</emphasis>
<bibRefCitation id="EF8B32967729FFE777704724A9FBEC3F" author="Habu, A." box="[794,916,1795,1819]" pageId="9" pageNumber="337" pagination="91 - 126" refId="ref9639" refString="Habu, A. (1961) Odacanthini of Japan (Coleoptera, Carabidae). Bulletin of the National Institute of Agricultural Sciences, C (13), 91 - 126." type="journal article" year="1961">Habu, 1961</bibRefCitation>
</taxonomicName>
; (14)—
<taxonomicName id="4C1A34E47729FFE777894724AFF6EC3F" authority="(Thunberg, 1784)" baseAuthorityName="Thunberg" baseAuthorityYear="1784" box="[995,1433,1795,1819]" class="Insecta" family="Carabidae" genus="Ophionea" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="337" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B96E93757729FFE777894724AE28EC3F" box="[995,1095,1795,1819]" italics="true" pageId="9" pageNumber="337">Ophionea</emphasis>
(s. str.)
<emphasis id="B96E93757729FFE770F64724AEB4EC3F" box="[1180,1243,1795,1819]" italics="true" pageId="9" pageNumber="337">indica</emphasis>
(Thunberg, 1784)
</taxonomicName>
; (15)—
<taxonomicName id="4C1A34E47729FFE774B14700A815EC1B" authority="(Chaudoir, 1850)" baseAuthorityName="Chaudoir" baseAuthorityYear="1850" box="[219,634,1831,1855]" class="Insecta" family="Carabidae" genus="Eucolliuris" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="337" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="B96E93757729FFE774B14700ABABEC1B" box="[219,452,1831,1855]" italics="true" pageId="9" pageNumber="337">Eucolliuris fuscipennis</emphasis>
(Chaudoir, 1850)
</taxonomicName>
; (16)—
<taxonomicName id="4C1A34E47729FFE776A24700AE53EC1B" authority="Liebke, 1938" authorityName="Liebke" authorityYear="1938" box="[712,1084,1831,1855]" class="Insecta" family="Carabidae" genus="Polydamasium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="337" phylum="Arthropoda" rank="species" species="strandi">
<emphasis id="B96E93757729FFE776A24700A9C2EC1B" box="[712,941,1831,1855]" italics="true" pageId="9" pageNumber="337">Polydamasium strandi</emphasis>
<bibRefCitation id="EF8B32967729FFE777D94700AE53EC1B" author="Liebke, M." box="[947,1084,1831,1855]" pageId="9" pageNumber="337" pagination="37 - 141" refId="ref9967" refString="Liebke, M. (1938) Denkschrift uber die Carabiden-Tribus Colliurini. Festschrift zum 60. Geburtstage von Professor Dr. Embrik Strand, 4, 37 - 141." type="journal article" year="1938">Liebke, 1938</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BA54F677729FFE774AD47B6AED6ECD7" blockId="9.[151,1436,1937,2036]" pageId="9" pageNumber="337">
Concerning the
<taxonomicName id="4C1A34E47729FFE7751D47B5A843EC8F" authorityName="Terada &amp; Wu" authorityYear="2014" box="[375,556,1938,1964]" class="Insecta" family="Carabidae" genus="Ophionea" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="337" phylum="Arthropoda" rank="subSpecies" species="bhamoensis" subSpecies="taiwanensis">
ssp.
<emphasis id="B96E93757729FFE775C247B5A843EC8F" box="[424,556,1938,1963]" italics="true" pageId="9" pageNumber="337">taiwanensis</emphasis>
</taxonomicName>
, the characters adopted for a subspecific distinction are extremely weak and lay largely within the limits of the intraspecific variation, so we propose the synonymy:
<taxonomicName id="4C1A34E47729FFE770274792AB74ECD7" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" class="Insecta" family="Carabidae" genus="Ophionea" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="337" phylum="Arthropoda" rank="species" species="bhamoensis">
<emphasis id="B96E93757729FFE770274792AED4ECEB" box="[1101,1211,1973,1999]" italics="true" pageId="9" pageNumber="337">Ophionea</emphasis>
(s. str.)
<emphasis id="B96E93757729FFE771724792AFF3ECEB" box="[1304,1436,1973,1999]" italics="true" pageId="9" pageNumber="337">bhamoensis</emphasis>
<bibRefCitation id="EF8B32967729FFE774FD47FDAB74ECD7" author="Bates, H. W." box="[151,283,2009,2036]" pageId="9" pageNumber="337" pagination="267 - 428" refId="ref9486" refString="Bates, H. W. (1892) List of Carabidae. Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. Annali del Museo Civico di Storia Naturale di Genova, 12 (2), 267 - 428." type="journal article" year="1892">Bates, 1892</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="4C1A34E47729FFE7755347FEA99AECD7" authority="Terada &amp; Wu, 2014" authorityName="Terada &amp; Wu" authorityYear="2014" box="[313,1013,2009,2036]" class="Insecta" family="Carabidae" genus="Ophionea" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="337" phylum="Arthropoda" rank="subSpecies" species="bhamoensis" subSpecies="taiwanensis">
<emphasis id="B96E93757729FFE7755347FEA85CECD7" box="[313,563,2009,2035]" italics="true" pageId="9" pageNumber="337">Ophionea bhamoensis</emphasis>
(s. str.)
<emphasis id="B96E93757729FFE776E447FDA97DECD7" box="[654,786,2010,2035]" italics="true" pageId="9" pageNumber="337">taiwanensis</emphasis>
<bibRefCitation id="EF8B32967729FFE7777347FEA99AECD7" author="Terada, K. &amp; Wu, W." box="[793,1013,2009,2036]" pageId="9" pageNumber="337" pagination="15 - 41" refId="ref10136" refString="Terada, K. &amp; Wu, W. (2014) Notes on Taiwanese Caraboidea (Coleoptera) V. A Review of the Tribe Odacanthini (Carabidae) in Taiwan, with Description of Ophionea bhamoensis taiwanensis subsp. nov. Collection and Research, 27, 15 - 41." type="journal article" year="2014">Terada &amp; Wu, 2014</bibRefCitation>
</taxonomicName>
<emphasis id="B96E93757729FFE7779647FEAED6ECD7" bold="true" box="[1020,1209,2009,2035]" pageId="9" pageNumber="337">junior synonym</emphasis>
</paragraph>
<paragraph id="8BA54F67772AFFE474AD40B0A924EA65" blockId="10.[151,1437,151,321]" pageId="10" pageNumber="338">
As mentioned by
<bibRefCitation id="EF8B3296772AFFE475EE40B0A8FCEB95" author="Anagha, V. S. S. &amp; Sabu, T. K." box="[388,659,151,177]" pageId="10" pageNumber="338" pagination="523 - 536" refId="ref9157" refString="Anagha, V. S. S. &amp; Sabu, T. K. (2024) Additions to the taxonomy of the genus Ophionea Klug, 1821 (Coleoptera: Carabidae: Lebiinae) and new species from India. Zootaxa, 5437 (4), 523 - 536. https: // doi. org / 10.11646 / zootaxa. 5437.4.5" type="journal article" year="2024">Anagha and Sabu (2024)</bibRefCitation>
for
<taxonomicName id="4C1A34E4772AFFE476D640B0AE0DEB95" authority="Schmidt-Gobel, 1846" authorityName="Schmidt-Gobel" authorityYear="1846" box="[700,1122,151,177]" class="Insecta" family="Carabidae" genus="Ophionea" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="338" phylum="Arthropoda" rank="species" species="nigrofasciata">
<emphasis id="B96E9375772AFFE476D640B0A900EB95" box="[700,879,151,177]" italics="true" pageId="10" pageNumber="338">O. nigrofasciata</emphasis>
Schmidt-Göbel, 1846
</taxonomicName>
as present in the
<collectingCountry id="F30D0FF7772AFFE4717D40B0AFF7EB95" box="[1303,1432,151,177]" name="Philippines" pageId="10" pageNumber="338">Philippines</collectingCountry>
, in our opinion, should be referred to
<taxonomicName id="4C1A34E4772AFFE47628409CA885EBF1" authorityName="Bates" authorityYear="1892" box="[578,746,187,213]" class="Insecta" family="Carabidae" genus="Ophionea" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="338" phylum="Arthropoda" rank="species" species="bhamoensis">
<emphasis id="B96E9375772AFFE47628409CA885EBF1" box="[578,746,187,213]" italics="true" pageId="10" pageNumber="338">O. bhamoensis</emphasis>
</taxonomicName>
: the distribution listed by these authors for
<taxonomicName id="4C1A34E4772AFFE4708E409CAFF3EBF1" authorityName="Schmidt-Gobel" authorityYear="1846" box="[1252,1436,187,213]" class="Insecta" family="Carabidae" genus="Ophionea" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="338" phylum="Arthropoda" rank="species" species="nigrofasciata">
<emphasis id="B96E9375772AFFE4708E409CAFF3EBF1" box="[1252,1436,187,213]" italics="true" pageId="10" pageNumber="338">O. nigrofasciata</emphasis>
</taxonomicName>
as extremely widespread, not corresponding at all from that given by
<bibRefCitation id="EF8B3296772AFFE477DA40F8AE26EBDD" author="Baehr, M." box="[944,1097,223,249]" pageId="10" pageNumber="338" pagination="1041 - 1084" refId="ref9337" refString="Baehr, M. (1996) The ground beetle genus Casnoidea Castelnau, Taxonomy, Phylogeny, Zoogeography (Insecta, Coleoptera, Carabidae, Odacanthinae). Invertebrate Taxonomy, 10, 1041 - 1084. https: // doi. org / 10.1071 / IT 9961041" type="journal article" year="1996">Baehr (1996)</bibRefCitation>
. We conclude that this data is extremely doubtful. Thus we redefine
<taxonomicName id="4C1A34E4772AFFE476264124A89BEA39" authorityName="Bates" authorityYear="1892" box="[588,756,259,285]" class="Insecta" family="Carabidae" genus="Ophionea" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="338" phylum="Arthropoda" rank="species" species="bhamoensis">
<emphasis id="B96E9375772AFFE476264124A89BEA39" box="[588,756,259,285]" italics="true" pageId="10" pageNumber="338">O. bhamoensis</emphasis>
</taxonomicName>
as certainly distributed only in
<collectingCountry id="F30D0FF7772AFFE4700A4124AEA0EA39" box="[1120,1231,259,285]" name="Myanmar" pageId="10" pageNumber="338">Myanmar</collectingCountry>
,
<collectingCountry id="F30D0FF7772AFFE470B14124AF7AEA39" box="[1243,1301,259,285]" name="India" pageId="10" pageNumber="338">India</collectingCountry>
(
<bibRefCitation id="EF8B3296772AFFE4714F4124AB7DEA65" author="Anagha, V. S. S. &amp; Sabu, T. K." pageId="10" pageNumber="338" pagination="523 - 536" refId="ref9157" refString="Anagha, V. S. S. &amp; Sabu, T. K. (2024) Additions to the taxonomy of the genus Ophionea Klug, 1821 (Coleoptera: Carabidae: Lebiinae) and new species from India. Zootaxa, 5437 (4), 523 - 536. https: // doi. org / 10.11646 / zootaxa. 5437.4.5" type="journal article" year="2024">Anagha &amp; Sabu 2024</bibRefCitation>
),
<collectingCountry id="F30D0FF7772AFFE4754F4100AB18EA65" box="[293,375,295,321]" name="Taiwan" pageId="10" pageNumber="338">Taiwan</collectingCountry>
(
<bibRefCitation id="EF8B3296772AFFE475EF4100A830EA65" author="Terada, K. &amp; Wu, W." box="[389,607,295,321]" pageId="10" pageNumber="338" pagination="15 - 41" refId="ref10136" refString="Terada, K. &amp; Wu, W. (2014) Notes on Taiwanese Caraboidea (Coleoptera) V. A Review of the Tribe Odacanthini (Carabidae) in Taiwan, with Description of Ophionea bhamoensis taiwanensis subsp. nov. Collection and Research, 27, 15 - 41." type="journal article" year="2014">Terada &amp; Wu 2014</bibRefCitation>
) and the
<collectingCountry id="F30D0FF7772AFFE476AC4100A927EA65" box="[710,840,295,321]" name="Philippines" pageId="10" pageNumber="338">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>