treatments-xml/data/03/B3/87/03B387F5FF9EC16142C6FADDFF6BF831.xml

<document id="6F78D4E0062504E7864FF335519BE247" ID-CLB-Dataset="9462" ID-DOI="10.11646/zootaxa.4890.2.5" ID-GBIF-Dataset="768fbdbe-ad7e-415f-8e22-52c0a52c71f6" ID-ISSN="1175-5326" ID-Zenodo-Dep="4301869" ID-ZooBank="82A5AFC2-8E8D-472D-97B5-9FFD5CB8CC31" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1606905221522" checkinUser="plazi" docAuthor="Scheers, Kevin &amp; Hájek, Jiří" docDate="2020" docId="03B387F5FF9EC16142C6FADDFF6BF831" docLanguage="en" docName="zootaxa.4890.2.5.pdf" docOrigin="Zootaxa 4890 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neoclypeodytes balkei Scheers &amp; Hájek 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="248" masterDocId="FF8AFF8DFF9FC1624251FFF2FF99FFC5" masterDocTitle="Neoclypeodytes Young from Panama, with description of two new species (Coleoptera: Dytiscidae: Hydroporinae: Bidessini)" masterLastPageNumber="256" masterPageNumber="245" pageNumber="246" updateTime="1698847084457" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D48B775CAF4576AAB0E06E9BA111EB7C">Neoclypeodytes Young from Panama, with description of two new species (Coleoptera: Dytiscidae: Hydroporinae: Bidessini)</mods:title>
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<mods:namePart id="AA032745A6B97FE37F252EF00DE832AE">Scheers, Kevin</mods:namePart>
<mods:affiliation id="9435966DBE35B9F842EF50B6D2B26200">Research Institute for Nature and Forest (INBO), Havenlaan 88 bus 73, 1000 Brussels, Belgium.</mods:affiliation>
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<mods:namePart id="265DF4924B8C191CB27720B686C750BD">Hájek, Jiří</mods:namePart>
<mods:affiliation id="CB76AC6FC9ABFC0CF1750AE0CA23ED18">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic. jiri. hajek @ nm. cz; https: // orcid. org / 0000 - 0001 - 5779 - 1542</mods:affiliation>
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<mods:title id="3CC4CB4DD6EE9EA3F062C86C3CE6E1F7">Zootaxa</mods:title>
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<mods:date id="49C93750479AC4134DF9D7CB9D790FBD">2020</mods:date>
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<treatment id="03B387F5FF9EC16142C6FADDFF6BF831" ID-DOI="http://doi.org/10.5281/zenodo.4327975" ID-GBIF-Taxon="170604211" ID-Zenodo-Dep="4327975" LSID="urn:lsid:plazi:treatment:03B387F5FF9EC16142C6FADDFF6BF831" httpUri="http://treatment.plazi.org/id/03B387F5FF9EC16142C6FADDFF6BF831" lastPageId="3" lastPageNumber="248" pageId="1" pageNumber="246">
<subSubSection id="C3006568FF9EC16342C6FADDFD9FFA8F" box="[151,518,1327,1354]" pageId="1" pageNumber="246" type="nomenclature">
<paragraph id="8BA536E3FF9EC16342C6FADDFD9FFA8F" blockId="1.[151,518,1327,1390]" box="[151,518,1327,1354]" pageId="1" pageNumber="246">
<heading id="D0ED818FFF9EC16342C6FADDFD9FFA8F" bold="true" box="[151,518,1327,1354]" fontSize="11" level="1" pageId="1" pageNumber="246" reason="1">
<emphasis id="B96EEAF1FF9EC16342C6FADDFD9FFA8F" bold="true" box="[151,518,1327,1354]" pageId="1" pageNumber="246">
<taxonomicName id="4C1A4D60FF9EC16342C6FADDFE38FA8F" ID-CoL="8LZDK" authority="Scheers &amp; Hájek, 2020" authorityName="Scheers &amp; Hájek" authorityYear="2020" box="[151,417,1327,1354]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="species" species="balkei" status="sp. nov.">
<emphasis id="B96EEAF1FF9EC16342C6FADDFE38FA8F" bold="true" box="[151,417,1327,1354]" italics="true" pageId="1" pageNumber="246">Neoclypeodytes balkei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D578AFF9EC16343F8FAC2FD9FFA8F" box="[425,518,1328,1354]" pageId="1" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3006568FF9EC16342C6FAA6FF72FAAB" box="[151,235,1364,1390]" pageId="1" pageNumber="246" type="description">
<paragraph id="8BA536E3FF9EC16342C6FAA6FF72FAAB" blockId="1.[151,518,1327,1390]" box="[151,235,1364,1390]" pageId="1" pageNumber="246">
(
<figureCitation id="13212A66FF9EC16342F1FAA6FF79FAAB" box="[160,224,1364,1390]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1939]" captionTargetBox="[169,1422,191,1875]" captionTargetId="figure-19@2.[155,1433,179,1890]" captionTargetPageId="2" captionText="FIGURE 1. Neoclypeodytes balkei sp. nov. A. dorsal habitus; B. ventral habitus; C. median lobe of aedeagus in ventral view; D. median lobe of aedeagus in lateral view; E. lateral lobe (paramere). Scale bars: A–B = 1 mm; C–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301871" httpUri="https://zenodo.org/record/4301871/files/figure.png" pageId="1" pageNumber="246">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3006568FF9EC16342C6FA6EFB77F9E7" pageId="1" pageNumber="246" type="materials_examined">
<paragraph id="8BA536E3FF9EC16342C6FA6EFB77F9E7" blockId="1.[151,1437,1436,1967]" pageId="1" pageNumber="246">
<emphasis id="B96EEAF1FF9EC16342C6FA6EFED1FA73" bold="true" box="[151,328,1436,1462]" pageId="1" pageNumber="246">Type material.</emphasis>
<materialsCitation id="3B723CBEFF9EC1634303FA6EFCBEFA3B" ID-GBIF-Occurrence="2986026308" collectingDate="2007-06-19" collectionCode="ZSMG" country="Panama" determinerName="K. Scheers &amp; J. Hajek" elevation="270" location="Campana" municipality="Anam" pageId="1" pageNumber="246" specimenCount="2" typeStatus="holotype">
<typeStatus id="54A18841FF9EC1634303FA6EFE26FA73" box="[338,447,1436,1462]" pageId="1" pageNumber="246" type="holotype">Holotype</typeStatus>
: m#, “ 
<collectingCountry id="F30D7673FF9EC163405BFA6EFDF3FA73" box="[522,618,1436,1462]" name="Panama" pageId="1" pageNumber="246">Panama</collectingCountry>
: 
<collectingCountry id="F30D7673FF9EC1634026FA6FFD4AFA72" box="[631,723,1437,1463]" name="Panama" pageId="1" pageNumber="246">Panama</collectingCountry>
, 
<location id="8EC56038FF9EC16340B0FA6EFCD7FA73" LSID="urn:lsid:plazi:treatment:03B387F5FF9EC16142C6FADDFF6BF831:8EC56038FF9EC16340B0FA6EFCD7FA73" box="[737,846,1436,1462]" country="Panama" municipality="Anam" name="Campana" pageId="1" pageNumber="246">Campana</location>
, / cascade below 
<collectingMunicipality id="6BC1AC99FF9EC163464AFA6EFBC6FA73" box="[1051,1119,1436,1462]" pageId="1" pageNumber="246">Anam</collectingMunicipality>
office, / 
<quantity id="4CE29B06FF9EC163469CFA6EFA96FA73" box="[1229,1295,1436,1462]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="246" unit="m" value="270.0">
<elevation id="0037D1D0FF9EC163469CFA6EFA96FA73" box="[1229,1295,1436,1462]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="246" unit="m" value="270.0">270m</elevation>
</quantity>
, 
<date id="FFA41023FF9EC163474CFA6EFA0EFA73" box="[1309,1431,1436,1462]" pageId="1" pageNumber="246" value="2007-06-19">
<collectingDate id="EFE0E9CBFF9EC163474CFA6EFA0EFA73" box="[1309,1431,1436,1462]" pageId="1" pageNumber="246" value="2007-06-19">19.vi.2007</collectingDate>
</date>
, / 08.40.818N 079.54.075W, / Balke (PAN 07) [p] // 
<typeStatus id="54A18841FF9EC16340A9FA32FC15FA1F" box="[760,908,1472,1498]" pageId="1" pageNumber="246" type="holotype">HOLOTYPE</typeStatus>
m# / 
<taxonomicName id="4C1A4D60FF9EC1634184FA32FB5EFA1F" authorityName="Young" authorityYear="1967" box="[981,1223,1472,1498]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEAF1FF9EC1634184FA32FB5EFA1F" box="[981,1223,1472,1498]" italics="true" pageId="1" pageNumber="246">NEOCLYPEODYTES</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C1A4D60FF9EC16346B2FA32FABEFA1F" authority="Scheers &amp; Hájek, 2020" authorityName="Scheers &amp; Hájek" authorityYear="2020" box="[1251,1319,1472,1498]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="species" species="balkei" status="sp. nov.">
<emphasis id="B96EEAF1FF9EC16346B2FA32FABEFA1F" box="[1251,1319,1472,1498]" italics="true" pageId="1" pageNumber="246">balkei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D578AFF9EC1634760FA33FA12FA1E" box="[1329,1419,1473,1499]" pageId="1" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
/ 
<determinerName id="7E79DA27FF9EC16342C6FA17FE8AFA3B" box="[151,275,1508,1535]" pageId="1" pageNumber="246">K. Scheers</determinerName>
&amp; 
<determinerName id="7E79DA27FF9EC163436DFA17FE03FA3B" box="[316,410,1508,1535]" pageId="1" pageNumber="246">J. Hájek</determinerName>
det. 2020 [p, red label]” (
<collectionCode id="ED0BAE26FF9EC163409AFA16FC86FA3B" box="[715,799,1508,1534]" pageId="1" pageNumber="246">ZSMG</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B723CBEFF9EC1634165FA17FB73F9E7" ID-GBIF-Occurrence="2986026310" collectingDate="2007-06-19" collectionCode="BMNH, CKSB, NMPC, ZSMG" country="Panama" determinerName="K. Scheers &amp; J. Hajek" elevation="270" location="Campana" municipality="Anam" pageId="1" pageNumber="246" specimenCount="46" typeStatus="paratype">
<typeStatus id="54A18841FF9EC1634165FA17FC3CFA3A" box="[820,933,1509,1535]" pageId="1" pageNumber="246" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D1CFD6AFF9EC16341E3FA16FBC9FA3B" box="[946,1104,1508,1534]" pageId="1" pageNumber="246" type="generic">46 specimens</specimenCount>
; same label data as holotype (
<collectionCode id="ED0BAE26FF9EC16342CEF9FAFF6EF9E7" box="[159,247,1544,1570]" pageId="1" pageNumber="246">BMNH</collectionCode>
, 
<collectionCode id="ED0BAE26FF9EC1634352F9FAFED6F9E7" box="[259,335,1544,1570]" pageId="1" pageNumber="246">CKSB</collectionCode>
, 
<collectionCode id="ED0BAE26FF9EC163430BF9FAFE35F9E7" box="[346,428,1544,1570]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="246">NMPC</collectionCode>
, 
<collectionCode id="ED0BAE26FF9EC16343E6F9FAFD93F9E7" box="[439,522,1544,1570]" pageId="1" pageNumber="246">ZSMG</collectionCode>
). Each 
<typeStatus id="54A18841FF9EC163400AF9FBFD22F9E6" box="[603,699,1545,1571]" pageId="1" pageNumber="246" type="paratype">paratype</typeStatus>
is provided with the appropriate red printed label
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3006568FF9EC1614296F9DEFC88FA89" lastPageId="3" lastPageNumber="248" pageId="1" pageNumber="246" type="description">
<paragraph id="8BA536E3FF9EC1634296F9DEFA08F9AF" blockId="1.[151,1437,1436,1967]" pageId="1" pageNumber="246">
<emphasis id="B96EEAF1FF9EC1634296F9DEFD10F983" bold="true" box="[199,649,1579,1606]" pageId="1" pageNumber="246">
Description of male 
<typeStatus id="54A18841FF9EC16343E9F9DEFDB8F983" box="[440,545,1580,1606]" pageId="1" pageNumber="246" type="holotype">holotype</typeStatus>
. 
<emphasis id="B96EEAF1FF9EC163407CF9D9FD10F983" bold="true" box="[557,649,1579,1606]" italics="true" pageId="1" pageNumber="246">Habitus</emphasis>
</emphasis>
(
<figureCitation id="13212A66FF9EC16340C9F9DEFD77F983" box="[664,750,1580,1606]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1939]" captionTargetBox="[169,1422,191,1875]" captionTargetId="figure-19@2.[155,1433,179,1890]" captionTargetPageId="2" captionText="FIGURE 1. Neoclypeodytes balkei sp. nov. A. dorsal habitus; B. ventral habitus; C. median lobe of aedeagus in ventral view; D. median lobe of aedeagus in lateral view; E. lateral lobe (paramere). Scale bars: A–B = 1 mm; C–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301871" httpUri="https://zenodo.org/record/4301871/files/figure.png" pageId="1" pageNumber="246">Fig. 1A</figureCitation>
). Ovoid-shaped, slightly attenuate anteriorly and strongly attenuate posteriorly; lateral outline nearly continuous between pronotum and elytra in dorsal aspect. Surface shiny.
</paragraph>
<paragraph id="8BA536E3FF9EC16342C6F981FA12F86B" blockId="1.[151,1437,1436,1967]" pageId="1" pageNumber="246">
<emphasis id="B96EEAF1FF9EC16342C6F981FE83F94B" bold="true" box="[151,282,1651,1678]" italics="true" pageId="1" pageNumber="246">Coloration.</emphasis>
Head piceous. Antennae with antennomeres I–IV uniformly pale testaceous, antennomeres V–XI slightly infuscated apically. Palpi pale testaceous. Disc of pronotum piceous, medially slightly lightened; lateral parts of pronotum pale testaceous, broadly reaching inwards. Elytron dark piceous with two testaceous maculae; anterior (subbasal) macula subquadrangular, extending from lateral margin to one fifth of elytral width to suture, anterior margin straight with small emargination sublaterally, posterior margin largely concave with posteriorly protruding tooth near suture; posterior macula not reaching suture, anteriorly not reaching lateral margin, with concave emargination towards suture, posteriorly reaching lateral margin and apex of elytron (
<figureCitation id="13212A66FF9EC163467FF8BEFB10F8A3" box="[1070,1161,1868,1894]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1939]" captionTargetBox="[169,1422,191,1875]" captionTargetId="figure-19@2.[155,1433,179,1890]" captionTargetPageId="2" captionText="FIGURE 1. Neoclypeodytes balkei sp. nov. A. dorsal habitus; B. ventral habitus; C. median lobe of aedeagus in ventral view; D. median lobe of aedeagus in lateral view; E. lateral lobe (paramere). Scale bars: A–B = 1 mm; C–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301871" httpUri="https://zenodo.org/record/4301871/files/figure.png" pageId="1" pageNumber="246">Fig. 1A</figureCitation>
); epipleuron completely testaceous. Legs including tarsi testaceous. Prosternum, mesepisterna and mesepimera rufotestaceous; metaventrite, metepisterna and metacoxae rufotestaceous, somewhat darkened at lateral portions. Abdominal ventrites piceous.
</paragraph>
<caption id="DF65666BFF9DC16042C6F889FA91F872" ID-DOI="http://doi.org/10.5281/zenodo.4301871" ID-Zenodo-Dep="4301871" httpUri="https://zenodo.org/record/4301871/files/figure.png" pageId="2" pageNumber="247" startId="2.[151,250,1915,1939]" targetBox="[169,1422,191,1875]" targetPageId="2">
<paragraph id="8BA536E3FF9DC16042C6F889FA91F872" blockId="2.[151,1436,1915,1976]" pageId="2" pageNumber="247">
<emphasis id="B96EEAF1FF9DC16042C6F889FE8DF851" bold="true" box="[151,276,1915,1940]" pageId="2" pageNumber="247">FIGURE 1.</emphasis>
<taxonomicName id="4C1A4D60FF9DC160434AF88EFE67F851" authority="Scheers &amp; Hájek, 2020" authorityName="Scheers &amp; Hájek" authorityYear="2020" box="[283,510,1916,1940]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="247" phylum="Arthropoda" rank="species" species="balkei" status="sp. nov.">
<emphasis id="B96EEAF1FF9DC160434AF88EFE67F851" box="[283,510,1916,1940]" italics="true" pageId="2" pageNumber="247">Neoclypeodytes balkei</emphasis>
</taxonomicName>
<emphasis id="B96EEAF1FF9DC1604054F88EFDCFF851" bold="true" box="[517,598,1916,1940]" pageId="2" pageNumber="247">
<taxonomicNameLabel id="A25D578AFF9DC1604054F88EFDCFF851" box="[517,598,1916,1940]" pageId="2" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. dorsal habitus; B. ventral habitus; C. median lobe of aedeagus in ventral view; D. median lobe of aedeagus in lateral view; E. lateral lobe (paramere). Scale bars: A–B = 1 mm; C–E = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BA536E3FF9CC1614296FF6AFD38FF3D" blockId="3.[151,1437,152,2036]" pageId="3" pageNumber="248">
<emphasis id="B96EEAF1FF9CC1614296FF6AFE9FFF75" bold="true" box="[199,262,152,176]" pageId="3" pageNumber="248">Head</emphasis>
with fine but distinct microreticulation anterior of occipital line; punctation consisting of fine punctures near eyes and coarser punctures on clypeus; anterior clypeal margin medially distinctly flattened; occipital line narrowly separated from posterior margin of eyes.
</paragraph>
<paragraph id="8BA536E3FF9CC1614296FEF6FB35FE42" blockId="3.[151,1437,152,2036]" pageId="3" pageNumber="248">
<emphasis id="B96EEAF1FF9CC1614296FEF6FEA6FED9" bold="true" box="[199,319,260,284]" pageId="3" pageNumber="248">Pronotum</emphasis>
widest at base; lateral bead distinct; lateral margin evenly rounded; microreticulation absent; punctation consisting of punctures separated by 2–3× puncture diameter, punctation somewhat sparser laterally and distinctly denser near anterior and posterior margins; basal striae sinuated, distinctly curved inwards near base and slightly outwards anteriorly, length about 0.55× length of pronotum measured along same line as stria.
</paragraph>
<paragraph id="8BA536E3FF9CC1614296FE61FB2EFE37" blockId="3.[151,1437,152,2036]" pageId="3" pageNumber="248">
<emphasis id="B96EEAF1FF9CC1614296FE61FE8BFE6E" bold="true" box="[199,274,403,427]" pageId="3" pageNumber="248">Elytra</emphasis>
with basal striae straight, about same length than those on pronotum; sutural striae absent; microreticulation absent; punctation similar to that of pronotal disc, but with punctures slightly larger; epipleura sparsely punctate, with moderately oblique humeral carina; lateral epipleural rim anteriorly visible in dorsal view.
</paragraph>
<paragraph id="8BA536E3FF9CC1614296FE0CFC63FDFC" blockId="3.[151,1437,152,2036]" pageId="3" pageNumber="248">
<emphasis id="B96EEAF1FF9CC1614296FE0CFE9FFDD0" bold="true" box="[199,262,510,534]" pageId="3" pageNumber="248">Legs.</emphasis>
Pro- and mesotibiae club shaped. Pro- and mesotarsomeres 1–3 distinctly broadened, ventrally with adhesive setae. Metatarsomeres with well-developed natatorial setae on both sides.
</paragraph>
<paragraph id="8BA536E3FF9CC1614296FDB7FD1DFCBE" blockId="3.[151,1437,152,2036]" pageId="3" pageNumber="248">
<emphasis id="B96EEAF1FF9CC1614296FDB7FE77FD98" box="[199,494,581,605]" inLineHeading="true" pageId="3" pageNumber="248" reason="1">
<emphasis id="B96EEAF1FF9CC1614296FDB7FEE5FD98" bold="true" box="[199,380,581,605]" pageId="3" pageNumber="248">Ventral surface</emphasis>
(
<figureCitation id="13212A66FF9CC16143DBFDB7FE44FD98" box="[394,477,581,605]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1939]" captionTargetBox="[169,1422,191,1875]" captionTargetId="figure-19@2.[155,1433,179,1890]" captionTargetPageId="2" captionText="FIGURE 1. Neoclypeodytes balkei sp. nov. A. dorsal habitus; B. ventral habitus; C. median lobe of aedeagus in ventral view; D. median lobe of aedeagus in lateral view; E. lateral lobe (paramere). Scale bars: A–B = 1 mm; C–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301871" httpUri="https://zenodo.org/record/4301871/files/figure.png" pageId="3" pageNumber="248">Fig. 1B</figureCitation>
).
</emphasis>
Prosternum medially narrowly convex, coarsely sculptured; prosternal process rather broad, subovate with apex broadly obtuse, laterally margined, slightly constricted medially. Metaventrite distinctly punctate over entire surface, punctures separated by 2–4× puncture diameter. Intralinear space of metacoxa broad (ML/MW = 2.40), sparsely and irregularly punctate with fine punctures, punctures separated by 1–4× puncture diameter; metacoxal plates distinctly punctate with coarse punctures, punctures separated by 1–3× puncture diameter, plates narrowly impunctate along anterior and posterior margins. Abdominal ventrite I coarsely punctate along margins, ventrite II sparsely and finely punctate; succeeding ventrites smooth with a row of setae bearing punctures along posterior margin and some sparse and fine punctures scattered over surface; ventrites IV and V with a tuft of long setae medially; ventrite VI unmodified.
</paragraph>
<paragraph id="8BA536E3FF9CC1614296FC75FDE4FB94" blockId="3.[151,1437,152,2036]" pageId="3" pageNumber="248">
<emphasis id="B96EEAF1FF9CC1614296FC75FEEEFC5B" bold="true" box="[199,375,902,927]" inLineHeading="true" pageId="3" pageNumber="248" reason="1">Male genitalia.</emphasis>
Median lobe in ventral view rather slender, parallel over whole length; apex distinctly truncate (
<figureCitation id="13212A66FF9CC16142CEFC58FF6FFC06" box="[159,246,938,963]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1939]" captionTargetBox="[169,1422,191,1875]" captionTargetId="figure-19@2.[155,1433,179,1890]" captionTargetPageId="2" captionText="FIGURE 1. Neoclypeodytes balkei sp. nov. A. dorsal habitus; B. ventral habitus; C. median lobe of aedeagus in ventral view; D. median lobe of aedeagus in lateral view; E. lateral lobe (paramere). Scale bars: A–B = 1 mm; C–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301871" httpUri="https://zenodo.org/record/4301871/files/figure.png" pageId="3" pageNumber="248">Fig. 1C</figureCitation>
); ventral surface in apical part lacking scale-like setae (see under 
<taxonomicName id="4C1A4D60FF9CC161418BFC59FBEFFC07" authority="Scheers &amp; Hájek, 2020" authorityName="Scheers &amp; Hájek" authorityYear="2020" box="[986,1142,938,962]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="fortunensis" status="sp. nov.">
<emphasis id="B96EEAF1FF9CC161418BFC59FBEFFC07" box="[986,1142,938,962]" italics="true" pageId="3" pageNumber="248">N. fortunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D578AFF9CC161462FFC59FB4EFC07" box="[1150,1239,939,962]" pageId="3" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
, and cf. 
<figureCitation id="13212A66FF9CC1614767FC58FA09FC06" box="[1334,1424,938,963]" captionStart="FIGURE 3" captionStartId="7.[151,250,1817,1841]" captionTargetBox="[211,1374,195,1777]" captionTargetId="figure-17@7.[199,1389,181,1793]" captionTargetPageId="7" captionText="FIGURE 3. Neoclypeodytes fortunensis sp. nov. A. dorsal habitus; B. ventral habitus; C. median lobe of aedeagus in ventral view; D. detail of apex of median lobe with scale-like setae. E. median lobe of aedeagus in lateral view; F. lateral lobe (paramere). Scale bars: A–B = 1 mm; C, E–F = 0.2 mm; D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4301875" httpUri="https://zenodo.org/record/4301875/files/figure.png" pageId="3" pageNumber="248">Fig. 3D</figureCitation>
). Median lobe in lateral view broadest in apical third; ventral outline almost evenly curved; dorsal outline almost straight medially, distinctly curved inwards near apex (
<figureCitation id="13212A66FF9CC16140AFFC00FCCFFBCF" box="[766,854,1010,1034]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1939]" captionTargetBox="[169,1422,191,1875]" captionTargetId="figure-19@2.[155,1433,179,1890]" captionTargetPageId="2" captionText="FIGURE 1. Neoclypeodytes balkei sp. nov. A. dorsal habitus; B. ventral habitus; C. median lobe of aedeagus in ventral view; D. median lobe of aedeagus in lateral view; E. lateral lobe (paramere). Scale bars: A–B = 1 mm; C–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301871" httpUri="https://zenodo.org/record/4301871/files/figure.png" pageId="3" pageNumber="248">Fig. 1D</figureCitation>
). Lateral lobe (paramere) with apical segment elongated, nearly straight; in apical half on inner side with several long setae; apex distinctly curved but not protruding, subapical incision not perceptible (
<figureCitation id="13212A66FF9CC161404BFBCBFDF6FB94" box="[538,623,1081,1105]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1939]" captionTargetBox="[169,1422,191,1875]" captionTargetId="figure-19@2.[155,1433,179,1890]" captionTargetPageId="2" captionText="FIGURE 1. Neoclypeodytes balkei sp. nov. A. dorsal habitus; B. ventral habitus; C. median lobe of aedeagus in ventral view; D. median lobe of aedeagus in lateral view; E. lateral lobe (paramere). Scale bars: A–B = 1 mm; C–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301871" httpUri="https://zenodo.org/record/4301871/files/figure.png" pageId="3" pageNumber="248">Fig. 1E</figureCitation>
).
</paragraph>
<paragraph id="8BA536E3FF9CC1614296FBAFFCF0FB5C" blockId="3.[151,1437,152,2036]" pageId="3" pageNumber="248">
<emphasis id="B96EEAF1FF9CC1614296FBAFFED5FBB0" bold="true" box="[199,332,1117,1141]" inLineHeading="true" pageId="3" pageNumber="248" reason="1">Variability.</emphasis>
With the exception of small variability in extent and shape of elytral color pattern, there is no significant variation present within the specimens of the 
<typeStatus id="54A18841FF9CC16140BAFB73FC82FB5C" box="[747,795,1153,1177]" pageId="3" pageNumber="248">type</typeStatus>
series.
</paragraph>
<paragraph id="8BA536E3FF9CC1614296FB56FB0DFB25" blockId="3.[151,1437,152,2036]" pageId="3" pageNumber="248">
<emphasis id="B96EEAF1FF9CC1614296FB56FE2DFB79" bold="true" box="[199,436,1188,1212]" inLineHeading="true" pageId="3" pageNumber="248" reason="1">Sexual dimorphism.</emphasis>
The females have the dorsal surface completely reticulated with irregular polygonal meshes and thus they appear submatt; pro- and mesotarsomeres are less broadened than in males.
</paragraph>
<paragraph id="8BA536E3FF9CC1614296FB1EFC88FA89" blockId="3.[151,1437,152,2036]" pageId="3" pageNumber="248">
<emphasis id="B96EEAF1FF9CC1614296FB1EFEE5FAC6" bold="true" box="[199,380,1260,1284]" inLineHeading="true" pageId="3" pageNumber="248" reason="1">Measurements.</emphasis>
TL: 
<quantity id="4CE29B06FF9CC16143E7FB1EFDC0FAC6" box="[438,601,1260,1284]" metricMagnitude="-3" metricUnit="m" metricValue="2.0999999999999996" metricValueMax="2.15" metricValueMin="2.05" pageId="3" pageNumber="248" unit="mm" value="2.0999999999999996" valueMax="2.15" valueMin="2.05">2.05–2.15 mm</quantity>
(mean value: 2.10 ± 
<quantity id="4CE29B06FF9CC161411CFB1EFC2FFAC1" box="[845,950,1260,1284]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="248" unit="mm" value="0.03">0.03 mm</quantity>
); 
<typeStatus id="54A18841FF9CC161419AFB1EFBA9FAC1" box="[971,1072,1260,1284]" pageId="3" pageNumber="248" type="holotype">holotype</typeStatus>
: 
<quantity id="4CE29B06FF9CC161466CFB1EFB3CFAC6" box="[1085,1189,1260,1284]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="3" pageNumber="248" unit="mm" value="2.15">2.15 mm</quantity>
. TL-h: 
<quantity id="4CE29B06FF9CC16146A8FB1EFA05FAC6" box="[1273,1436,1260,1284]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="1.95" metricValueMin="1.85" pageId="3" pageNumber="248" unit="mm" value="1.9" valueMax="1.95" valueMin="1.85">1.85–1.95 mm</quantity>
(mean value: 1.85 ± 
<quantity id="4CE29B06FF9CC1614327FAE2FE45FAED" box="[374,476,1296,1320]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="248" unit="mm" value="0.04">0.04 mm</quantity>
); 
<typeStatus id="54A18841FF9CC16143BFFAE2FDCAFAED" box="[494,595,1296,1320]" pageId="3" pageNumber="248" type="holotype">holotype</typeStatus>
: 
<quantity id="4CE29B06FF9CC161400DFAE2FD5BFAE2" box="[604,706,1296,1320]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="3" pageNumber="248" unit="mm" value="1.95">1.95 mm</quantity>
. TW: 
<quantity id="4CE29B06FF9CC1614150FAE2FC38FAE2" box="[769,929,1296,1320]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="3" pageNumber="248" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.10–1.20 mm</quantity>
(mean value: 1.15 ± 
<quantity id="4CE29B06FF9CC16146D4FAE2FB75FAED" box="[1157,1260,1296,1320]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="248" unit="mm" value="0.01">0.01 mm</quantity>
); 
<typeStatus id="54A18841FF9CC16146ACFAE2FAFBFAED" box="[1277,1378,1296,1320]" pageId="3" pageNumber="248" type="holotype">holotype</typeStatus>
: 
<quantity id="4CE29B06FF9CC161473DFAE2FF5FFA8E" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="248" unit="mm" value="1.2">1.20 mm</quantity>
. Ratio TL/TW: 1.73–1.82 (mean value: 1.80 ± 0.02).
</paragraph>
</subSubSection>
<subSubSection id="C3006568FF9CC1614296FAA5FE37F93D" pageId="3" pageNumber="248" type="diagnosis">
<paragraph id="8BA536E3FF9CC1614296FAA5FD31F9E7" blockId="3.[151,1437,152,2036]" pageId="3" pageNumber="248">
<emphasis id="B96EEAF1FF9CC1614296FAA5FE55FAAA" bold="true" box="[199,460,1367,1391]" inLineHeading="true" pageId="3" pageNumber="248" reason="1">Differential diagnosis.</emphasis>
<taxonomicName id="4C1A4D60FF9CC1614382FAA5FD55FAAA" authority="Scheers &amp; Hájek, 2020" authorityName="Scheers &amp; Hájek" authorityYear="2020" box="[467,716,1367,1391]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="balkei" status="sp. nov.">
<emphasis id="B96EEAF1FF9CC1614382FAA5FD55FAAA" box="[467,716,1367,1391]" italics="true" pageId="3" pageNumber="248">Neoclypeodytes balkei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D578AFF9CC1614082FAAAFCB0FAAA" box="[723,809,1368,1391]" pageId="3" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
is, together with 
<taxonomicName id="4C1A4D60FF9CC16141BAFAAAFBFFFAAA" baseAuthorityName="Sharp" baseAuthorityYear="1887" box="[1003,1126,1367,1391]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="curtulus">
<emphasis id="B96EEAF1FF9CC16141BAFAAAFBFFFAAA" box="[1003,1126,1367,1391]" italics="true" pageId="3" pageNumber="248">N. curtulus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1A4D60FF9CC16146CCFAAAFAA1FAAA" authority="Scheers &amp; Hájek, 2020" authorityName="Scheers &amp; Hájek" authorityYear="2020" box="[1181,1336,1367,1391]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="fortunensis" status="sp. nov.">
<emphasis id="B96EEAF1FF9CC16146CCFAAAFAA1FAAA" box="[1181,1336,1367,1391]" italics="true" pageId="3" pageNumber="248">N. fortunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D578AFF9CC1614711FAAAFA01FAAA" box="[1344,1432,1368,1391]" pageId="3" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
, one of only three representatives of the genus known south of 
<collectingCountry id="F30D7673FF9CC161411AFA89FC5EFA56" box="[843,967,1403,1427]" name="Guatemala" pageId="3" pageNumber="248">Guatemala</collectingCountry>
. The three species are easily distinguished from each other by the color pattern (cf. 
<figureCitation id="13212A66FF9CC1614003FA6DFD32FA72" box="[594,683,1439,1463]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1939]" captionTargetBox="[169,1422,191,1875]" captionTargetId="figure-19@2.[155,1433,179,1890]" captionTargetPageId="2" captionText="FIGURE 1. Neoclypeodytes balkei sp. nov. A. dorsal habitus; B. ventral habitus; C. median lobe of aedeagus in ventral view; D. median lobe of aedeagus in lateral view; E. lateral lobe (paramere). Scale bars: A–B = 1 mm; C–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301871" httpUri="https://zenodo.org/record/4301871/files/figure.png" pageId="3" pageNumber="248">Figs 1A</figureCitation>
, 
<figureCitation id="13212A66FF9CC16140E6FA6DFD5DFA72" box="[695,708,1439,1463]" captionStart="FIGURE 2" captionStartId="5.[151,250,1914,1938]" captionTargetBox="[164,1419,198,1882]" captionTargetId="figure-17@5.[151,1435,181,1890]" captionTargetPageId="5" captionText="FIGURE 2. Neoclypeodytes curtulus (Sharp, 1887) A. dorsal habitus of the lectotype; B. dorsal habitus of specimen from Campana; C. ventral habitus; D. median lobe of aedeagus in ventral view; E. median lobe of aedeagus in lateral view; F. lateral lobe (paramere). Scale bars: A–C = 1 mm; D–F = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301873" httpUri="https://zenodo.org/record/4301873/files/figure.png" pageId="3" pageNumber="248">2</figureCitation>
A–B and 3A), and the male genitalia with the median lobe parallel in ventral view, not medially expanded in lateral view (
<figureCitation id="13212A66FF9CC1614153FA30FCDDFA1E" box="[770,836,1474,1499]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1939]" captionTargetBox="[169,1422,191,1875]" captionTargetId="figure-19@2.[155,1433,179,1890]" captionTargetPageId="2" captionText="FIGURE 1. Neoclypeodytes balkei sp. nov. A. dorsal habitus; B. ventral habitus; C. median lobe of aedeagus in ventral view; D. median lobe of aedeagus in lateral view; E. lateral lobe (paramere). Scale bars: A–B = 1 mm; C–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301871" httpUri="https://zenodo.org/record/4301871/files/figure.png" pageId="3" pageNumber="248">Figs 1</figureCitation>
C–D) and lacking distinct scale-like setae at the apex (which are very distinct in both 
<taxonomicName id="4C1A4D60FF9CC16143AEFA15FDE5FA3B" baseAuthorityName="Sharp" baseAuthorityYear="1887" box="[511,636,1510,1534]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="curtulus">
<emphasis id="B96EEAF1FF9CC16143AEFA15FDE5FA3B" box="[511,636,1510,1534]" italics="true" pageId="3" pageNumber="248">N. curtulus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1A4D60FF9CC16140E2FA15FCC9FA3B" authority="Scheers &amp; Hájek, 2020" authorityName="Scheers &amp; Hájek" authorityYear="2020" box="[691,848,1510,1534]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="fortunensis" status="sp. nov.">
<emphasis id="B96EEAF1FF9CC16140E2FA15FCC9FA3B" box="[691,848,1510,1534]" italics="true" pageId="3" pageNumber="248">N. fortunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D578AFF9CC1614109FA15FC2BFA3B" box="[856,946,1510,1534]" pageId="3" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
), and the apical part of the lateral lobe with appendage not protruding (
<figureCitation id="13212A66FF9CC1614392F9F8FD85F9E7" box="[451,540,1546,1570]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1939]" captionTargetBox="[169,1422,191,1875]" captionTargetId="figure-19@2.[155,1433,179,1890]" captionTargetPageId="2" captionText="FIGURE 1. Neoclypeodytes balkei sp. nov. A. dorsal habitus; B. ventral habitus; C. median lobe of aedeagus in ventral view; D. median lobe of aedeagus in lateral view; E. lateral lobe (paramere). Scale bars: A–B = 1 mm; C–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301871" httpUri="https://zenodo.org/record/4301871/files/figure.png" pageId="3" pageNumber="248">Figs 1E</figureCitation>
, 
<figureCitation id="13212A66FF9CC1614079F9F8FDDCF9E7" box="[552,581,1546,1570]" captionStart="FIGURE 2" captionStartId="5.[151,250,1914,1938]" captionTargetBox="[164,1419,198,1882]" captionTargetId="figure-17@5.[151,1435,181,1890]" captionTargetPageId="5" captionText="FIGURE 2. Neoclypeodytes curtulus (Sharp, 1887) A. dorsal habitus of the lectotype; B. dorsal habitus of specimen from Campana; C. ventral habitus; D. median lobe of aedeagus in ventral view; E. median lobe of aedeagus in lateral view; F. lateral lobe (paramere). Scale bars: A–C = 1 mm; D–F = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301873" httpUri="https://zenodo.org/record/4301873/files/figure.png" pageId="3" pageNumber="248">2F</figureCitation>
and 
<figureCitation id="13212A66FF9CC161402AF9F8FD05F9E7" box="[635,668,1546,1570]" captionStart="FIGURE 3" captionStartId="7.[151,250,1817,1841]" captionTargetBox="[211,1374,195,1777]" captionTargetId="figure-17@7.[199,1389,181,1793]" captionTargetPageId="7" captionText="FIGURE 3. Neoclypeodytes fortunensis sp. nov. A. dorsal habitus; B. ventral habitus; C. median lobe of aedeagus in ventral view; D. detail of apex of median lobe with scale-like setae. E. median lobe of aedeagus in lateral view; F. lateral lobe (paramere). Scale bars: A–B = 1 mm; C, E–F = 0.2 mm; D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4301875" httpUri="https://zenodo.org/record/4301875/files/figure.png" pageId="3" pageNumber="248">3F</figureCitation>
).
</paragraph>
<paragraph id="8BA536E3FF9CC1614296F9DCFE37F93D" blockId="3.[151,1437,152,2036]" pageId="3" pageNumber="248">
The new species differs from the other species in the genus by the combination of the distinctive color pattern, the absence of sutural striae, the shape of the basal pronotal and elytral striae, and the male genitalia. In the identification key of 
<bibRefCitation id="EF8B4B12FF9CC16143D1F987FD81F948" author="Miller, K. B." box="[384,536,1653,1677]" pageId="3" pageNumber="248" pagination="87 - 123" refId="ref5642" refString="Miller, K. B. (2001) Revision and phylogenetic analysis of the New World genus Neoclypeodytes Young (Coleoptera: Dytiscidae: Hydroporinae: Bidessini). Systematic Entomology, 26, 87 - 123. https: // doi. org / 10.1046 / j. 1365 - 3113.2001.00144. x" type="journal article" year="2001">Miller (2001)</bibRefCitation>
, the new species falls into couplet 24, together with 
<taxonomicName id="4C1A4D60FF9CC1614630F984FAD4F948" authority="(Clark, 1862)" baseAuthorityName="Clark" baseAuthorityYear="1862" box="[1121,1357,1653,1677]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="fryii">
<emphasis id="B96EEAF1FF9CC1614630F984FB36F948" box="[1121,1199,1653,1677]" italics="true" pageId="3" pageNumber="248">N. fryii</emphasis>
(Clark, 1862)
</taxonomicName>
and 
<taxonomicName id="4C1A4D60FF9CC16147D2F984FE4DF974" authority="(Sharp, 1882)" baseAuthorityName="Sharp" baseAuthorityYear="1882" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="quadrinotatus">
<emphasis id="B96EEAF1FF9CC16147D2F984FEAAF974" italics="true" pageId="3" pageNumber="248">N. quadrinotatus</emphasis>
(Sharp, 1882)
</taxonomicName>
; from both these species, 
<taxonomicName id="4C1A4D60FF9CC16140BEF968FCCBF974" authority="Scheers &amp; Hájek, 2020" authorityName="Scheers &amp; Hájek" authorityYear="2020" box="[751,850,1689,1713]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="balkei" status="sp. nov.">
<emphasis id="B96EEAF1FF9CC16140BEF968FCCBF974" box="[751,850,1689,1713]" italics="true" pageId="3" pageNumber="248">N. balkei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D578AFF9CC1614109F96BFC34F975" box="[856,941,1689,1712]" pageId="3" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
differs by the posteriorly strongly attenuated habitus, more strongly punctured metacoxal plates, and by the male genitalia (cf. 
<figureCitation id="13212A66FF9CC1614661F94EFBE8F910" box="[1072,1137,1724,1749]" captionStart="FIGURE 1" captionStartId="2.[151,250,1915,1939]" captionTargetBox="[169,1422,191,1875]" captionTargetId="figure-19@2.[155,1433,179,1890]" captionTargetPageId="2" captionText="FIGURE 1. Neoclypeodytes balkei sp. nov. A. dorsal habitus; B. ventral habitus; C. median lobe of aedeagus in ventral view; D. median lobe of aedeagus in lateral view; E. lateral lobe (paramere). Scale bars: A–B = 1 mm; C–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301871" httpUri="https://zenodo.org/record/4301871/files/figure.png" pageId="3" pageNumber="248">Figs 1</figureCitation>
C–E, and Figs 20C–E and 21C–E in 
<bibRefCitation id="EF8B4B12FF9CC1614359F912FE32F93D" author="Miller, K. B." box="[264,427,1760,1784]" pageId="3" pageNumber="248" pagination="87 - 123" refId="ref5642" refString="Miller, K. B. (2001) Revision and phylogenetic analysis of the New World genus Neoclypeodytes Young (Coleoptera: Dytiscidae: Hydroporinae: Bidessini). Systematic Entomology, 26, 87 - 123. https: // doi. org / 10.1046 / j. 1365 - 3113.2001.00144. x" type="journal article" year="2001">Miller (2001))</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3006568FF9CC1614296F8F6FF4DF86F" pageId="3" pageNumber="248" type="etymology">
<paragraph id="8BA536E3FF9CC1614296F8F6FD16F8FA" blockId="3.[151,1437,152,2036]" pageId="3" pageNumber="248">
<emphasis id="B96EEAF1FF9CC1614296F8F6FED4F8D9" bold="true" box="[199,333,1796,1820]" inLineHeading="true" pageId="3" pageNumber="248" reason="1">Etymology.</emphasis>
The new species is named after Michael Balke, who collected the 
<typeStatus id="54A18841FF9CC1614613F8F6FBEBF8D9" box="[1090,1138,1796,1820]" pageId="3" pageNumber="248">type</typeStatus>
series of this species. The specific epithet is a noun in the genitive case.
</paragraph>
<paragraph id="8BA536E3FF9CC1614296F8B9FF4DF86F" blockId="3.[151,1437,152,2036]" pageId="3" pageNumber="248">
<emphasis id="B96EEAF1FF9CC1614296F8B9FE6CF8A6" bold="true" box="[199,501,1867,1891]" inLineHeading="true" pageId="3" pageNumber="248" reason="1">Collection circumstances.</emphasis>
The specimens were collected in small water holes in the bedrock around a cascade of a forest stream (M. Balke, pers. comm. 2020) (
<figureCitation id="13212A66FF9CC1614093F89DFC9EF842" box="[706,775,1903,1927]" captionStart="FIGURE 4" captionStartId="8.[151,250,1770,1794]" captionTargetBox="[151,1437,885,1747]" captionTargetId="figure-345@8.[151,1437,885,1747]" captionTargetPageId="8" captionText="FIGURE 4. Habitat of Neoclypeodytes balkei sp. nov. and N. curtulus (Sharp, 1887) at Campana; small cascade of forest stream below ANAM office (Photo M. Balke)." figureDoi="http://doi.org/10.5281/zenodo.4301877" httpUri="https://zenodo.org/record/4301877/files/figure.png" pageId="3" pageNumber="248">Fig. 4</figureCitation>
). At this locality, the species was co-occurring with 
<taxonomicName id="4C1A4D60FF9CC161471FF882FF54F86E" baseAuthorityName="Sharp" baseAuthorityYear="1887" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="curtulus">
<emphasis id="B96EEAF1FF9CC161471FF882FF54F86E" italics="true" pageId="3" pageNumber="248">N. curtulus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3006568FF9CC1614296F844FF6BF831" pageId="3" pageNumber="248" type="distribution">
<paragraph id="8BA536E3FF9CC1614296F844FF6BF831" blockId="3.[151,1437,152,2036]" pageId="3" pageNumber="248">
<emphasis id="B96EEAF1FF9CC1614296F844FEC6F80B" bold="true" box="[199,351,1974,1998]" inLineHeading="true" pageId="3" pageNumber="248" reason="1">Distribution.</emphasis>
To date the species is only known from the 
<typeStatus id="54A18841FF9CC1614101F845FC19F80A" box="[848,896,1975,1999]" pageId="3" pageNumber="248">type</typeStatus>
locality at Campana, 
<collectingRegion id="49DEF801FF9CC1614624F845FAA0F80A" box="[1141,1337,1975,1999]" country="Panama" name="Panama" pageId="3" pageNumber="248">Panama province</collectingRegion>
, 
<collectingCountry id="F30D7673FF9CC1614715F845FA05F80B" box="[1348,1436,1975,1998]" name="Panama" pageId="3" pageNumber="248">Panama</collectingCountry>
(
<figureCitation id="13212A66FF9CC16142F1F82BFF7DF831" box="[160,228,2009,2036]" captionStart="FIGURE 6" captionStartId="10.[151,250,852,876]" captionTargetBox="[151,1435,181,829]" captionTargetId="figure-19@10.[151,1435,181,829]" captionTargetPageId="10" captionText="FIGURE 6. Map of distribution of Neoclypeodytes species in Panama." figureDoi="http://doi.org/10.5281/zenodo.4301881" httpUri="https://zenodo.org/record/4301881/files/figure.png" pageId="3" pageNumber="248">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>