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<document id="F360FDE3E461116846290EF8DB9105DE" ID-CLB-Dataset="287311" ID-DOI="https://doi.org/10.5852/ejt.2023.881.2175" ID-GBIF-Dataset="225ee515-e526-4cc0-b86b-85e8f2e4c83a" ID-ISSN="2118-9773" ID-Zenodo-Dep="8178280" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="admin" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="admin" checkinTime="1690201937279" checkinUser="felipe" docAuthor="Moonlight, Peter. W., Jara-Muñoz, Orlando A., Purvis, David A., Delves, Jay, Allen, Josh P. &amp; Reynel, Carlos" docDate="2023" docId="03B26B4BFF94FFCDFE37F9B3AFE3EE11" docLanguage="en" docName="ejt-881_1-334.imf" docOrigin="European Journal of Taxonomy 881" docSource="http://dx.doi.org/10.5852/ejt.2023.881.2175" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia condorensis Jara &amp; Moonlight 2023, sp. nov." docType="treatment" docVersion="13" lastPageNumber="53" masterDocId="FF8B1333FFA5FFF9FFD9FFBFAD27EA79" masterDocTitle="The genus Begonia (Begoniaceae) in Peru" masterLastPageNumber="334" masterPageNumber="1" pageNumber="50" updateTime="1732576113334" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="2805185DB74E2165BC628CBFE6E743C1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D10BA862D19EDAC0EB4713DFEADB276D">
<mods:title id="9CBCBC3182ED347B4BC11A354C7826C6">The genus Begonia (Begoniaceae) in Peru</mods:title>
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<mods:name id="5EC1379BCB8BC31959D3E28551857B27" type="personal">
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<mods:namePart id="8DD33C2E8F1BA54AE8F4117AF5B2A08D">Moonlight, Peter. W.</mods:namePart>
<mods:affiliation id="D867688B7F9DA00BD11A99E5FCCC0053">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK. &amp; Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland.</mods:affiliation>
<mods:nameIdentifier id="0E6AD4D364821D279B0E341438459384" type="email">moonligp@tcd.ie</mods:nameIdentifier>
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<mods:namePart id="C6933BB32746438D5FA707F83560C608">Jara-Muñoz, Orlando A.</mods:namePart>
<mods:affiliation id="9C03FAEFDDDFB4825A6BBCCC78901907">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá D. C., Colombia.</mods:affiliation>
<mods:nameIdentifier id="9D448A8BD11213AE4B89CBEC5EB54271" type="email">oajaram@unal.edu.co</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="8C1C99066A711875B0EF4E8F3E5B2949">Purvis, David A.</mods:namePart>
<mods:affiliation id="CCF52158DAEB2141ADE7F61E5326DD41">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="D28E5329173C92CF3958E602FA8269F1" type="email">dpurvis@rbge.org.uk</mods:nameIdentifier>
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<mods:namePart id="E6CEEE3AEF56F796EA1420C1DCB9E5C4">Delves, Jay</mods:namePart>
<mods:affiliation id="380BFA4341743170A6B75FE91C0A602F">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="D82F8245D36E46370E8CEFA069E34A72" type="email">j.delves@westernsydney.edu.au</mods:nameIdentifier>
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<mods:roleTerm id="B5AD55AE6287928903CED36BBE63FBBF">Author</mods:roleTerm>
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<mods:namePart id="62A943B397926EEE67FA3168081F2649">Allen, Josh P.</mods:namePart>
<mods:affiliation id="85EFECCA988E589A1CAF6F5917F6DE92">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="DE91AE7F8A58BDD4076B79DDB0E18F62" type="email">myrmeciaman@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F7C4A018AA682FADAF7F83B293EF1AEF">Reynel, Carlos</mods:namePart>
<mods:affiliation id="3723ECA47C700135C290F49F5861F84D">Facultad de Ciencias Forestales, Universidad Nacional Agraria-La Molina, Lima 12, Peru.</mods:affiliation>
<mods:nameIdentifier id="D6E8DD46C8595BDC6D60103A4CEC621E" type="email">reynel@lamolina.edu.pe</mods:nameIdentifier>
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<mods:title id="493038DA46951BAE1E85D9F622B7AB00">European Journal of Taxonomy</mods:title>
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<mods:date id="E4D99C3A61037C855B92C20CA4BB1DCD">2023</mods:date>
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<mods:number id="E0E57996B775E7AAD64B3E803719B24F">2023-07-18</mods:number>
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<mods:classification id="31EEDE39EF1DFA30126F015402CBD570">journal article</mods:classification>
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<mods:identifier id="D21D3969C96652D2CCE05D032DD70BA3" type="DOI">https://doi.org/10.5852/ejt.2023.881.2175</mods:identifier>
<mods:identifier id="C9300A81127EB5B289EC89851385B0F5" type="GBIF-Dataset">225ee515-e526-4cc0-b86b-85e8f2e4c83a</mods:identifier>
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<treatment id="03B26B4BFF94FFCDFE37F9B3AFE3EE11" ID-DOI="http://doi.org/10.5281/zenodo.10559122" ID-GBIF-Taxon="221057976" ID-Zenodo-Dep="10559122" LSID="urn:lsid:plazi:treatment:03B26B4BFF94FFCDFE37F9B3AFE3EE11" httpUri="http://treatment.plazi.org/id/03B26B4BFF94FFCDFE37F9B3AFE3EE11" lastPageId="52" lastPageNumber="53" pageId="49" pageNumber="50" scope_class="Equisetopsida" scope_family="Begoniaceae" scope_order="Cucurbitales" scope_subClass="Magnoliidae" scope_superOrder="Rosanae">
<subSubSection id="C30189D6FF94FFC8FE37F9B3A961EC5F" box="[494,1094,1547,1574]" pageId="49" pageNumber="50" type="nomenclature">
<paragraph id="8BA4DA5DFF94FFC8FE37F9B3A961EC5F" blockId="49.[494,1094,1547,1647]" box="[494,1094,1547,1574]" pageId="49" pageNumber="50">
6.
<taxonomicName id="4C1BA1DEFF94FFC8FDD5F9B3AEC3EC5F" ID-CoL="CFGSL" authority="Jara &amp; Moonlight" authorityName="Jara &amp; Moonlight" authorityYear="2023" box="[524,996,1547,1574]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B96F064FFF94FFC8FDD5F9B3AE2BEC5F" bold="true" box="[524,780,1547,1574]" italics="true" pageId="49" pageNumber="50">Begonia condorensis</emphasis>
Jara &amp; Moonlight
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FF94FFC8FC32F9B3A961EC5F" box="[1003,1094,1548,1574]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF94FFC8FD9EF98FAE41EC16" pageId="49" pageNumber="50" type="description">
<paragraph id="8BA4DA5DFF94FFC8FD9EF98FAECAEC33" blockId="49.[494,1094,1547,1647]" box="[583,1005,1584,1610]" pageId="49" pageNumber="50">urn:lsid:ipni.org:names:77323287-1</paragraph>
<paragraph id="8BA4DA5DFF94FFC8FD17F9EAAE41EC16" blockId="49.[494,1094,1547,1647]" box="[718,870,1621,1647]" pageId="49" pageNumber="50">
<figureCitation id="1320C6D8FF94FFC8FD17F9EAAE0FEC16" box="[718,808,1621,1647]" captionStart="Fig" captionStartId="51.[189,232,1928,1954]" captionTargetBox="[227,1334,414,1820]" captionTargetId="figure-12@51.[227,1361,347,1877]" captionTargetPageId="51" captionText="Fig. 20. Begonia condorensis Jara &amp; Moonlight sp. nov. A. Habit. B. Staminate flower, front view. C. Staminate flower, side. D. Pistillate flower, front view. E. Style and stigma. Illustration by Anna Dorward from E. Rodríguez 2996 (US)." figureDoi="http://doi.org/10.5281/zenodo.8178327" httpUri="https://zenodo.org/record/8178327/files/figure.png" pageId="49" pageNumber="50">Figs 20</figureCitation>
,
<figureCitation id="1320C6D8FF94FFC8FCEDF9EAAE41EC16" box="[820,870,1621,1647]" captionStart="Fig" captionStartId="52.[189,232,1892,1918]" captionTargetBox="[194,1392,1224,1843]" captionTargetId="figure-267@52.[189,1398,1214,1849]" captionTargetPageId="52" captionText="Fig. 21. Distribution of Begonia sect. Casparya (Klotzsch) Warb. in Peru and surrounding countries. A. B. condorensis Jara &amp; Moonlight sp. nov. (red) and B. hirta (Klotzsch) L.B.Sm. &amp; B.G.Schub. (blue). B. B. urticae L.f. (red). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178331" httpUri="https://zenodo.org/record/8178331/files/figure.png" pageId="49" pageNumber="50">21A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF94FFC8FF64F927AE53ED29" pageId="49" pageNumber="50" type="diagnosis">
<paragraph id="8BA4DA5DFF94FFC8FF64F927AC10ECCB" blockId="49.[189,311,1688,1714]" box="[189,311,1688,1714]" pageId="49" pageNumber="50">
<heading id="D0EC6D31FF94FFC8FF64F927AC10ECCB" bold="true" box="[189,311,1688,1714]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B96F064FFF94FFC8FF64F927AC10ECCB" bold="true" box="[189,311,1688,1714]" pageId="49" pageNumber="50">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF94FFC8FF64F976AE53ED29" blockId="49.[189,1399,1736,1873]" pageId="49" pageNumber="50">
<taxonomicName id="4C1BA1DEFF94FFC8FF64F976AC9EEC9B" authorityName="Moonlight" authorityYear="2023" box="[189,441,1736,1762]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B96F064FFF94FFC8FF64F976AC9EEC9B" box="[189,441,1736,1762]" italics="true" pageId="49" pageNumber="50">Begonia condorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FF94FFC8FE1DF976AF05EC9A" box="[452,546,1737,1763]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to
<taxonomicName id="4C1BA1DEFF94FFC8FCD5F976A98EEC9B" authority="L. B. Sm. &amp; B. G. Schub." authorityName="L. B. Sm. &amp; B. G. Schub." box="[780,1193,1736,1762]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="valvata">
<emphasis id="B96F064FFF94FFC8FCD5F976AEA0EC9B" box="[780,903,1736,1762]" italics="true" pageId="49" pageNumber="50">B. valvata</emphasis>
L.B.Sm. &amp; B.G.Schub.
</taxonomicName>
but differs in its number of stamens (4–6 vs 4); the apices of the tepals of the staminate flowers (rounded to truncate vs retuse); the shape of its leaf laminae (lanceolate to oblanceolate vs oblong); and the indumentum on the underside of the leaves (sparsely pilose vs densely villous).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF94FFC8FF64F8C5AE25ED90" pageId="49" pageNumber="50" type="etymology">
<paragraph id="8BA4DA5DFF94FFC8FF64F8C5AC62EDED" blockId="49.[189,325,1914,1940]" box="[189,325,1914,1940]" pageId="49" pageNumber="50">
<heading id="D0EC6D31FF94FFC8FF64F8C5AC62EDED" bold="true" box="[189,325,1914,1940]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B96F064FFF94FFC8FF64F8C5AC62EDED" bold="true" box="[189,325,1914,1940]" pageId="49" pageNumber="50">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF94FFC8FF64F815AE25ED90" blockId="49.[189,1399,1961,2025]" pageId="49" pageNumber="50">
<taxonomicName id="4C1BA1DEFF94FFC8FF64F815AC92EDBA" authorityName="Moonlight" authorityYear="2023" box="[189,437,1961,1987]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B96F064FFF94FFC8FF64F815AC92EDBA" box="[189,437,1961,1987]" italics="true" pageId="49" pageNumber="50">Begonia condorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FF94FFC8FE67F815AF3FEDBD" box="[446,536,1962,1988]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
is endemic to the Cordillera del Cóndor, which spans the borders of
<collectingCountry id="F30C9ACDFF94FFC8FA98F815A850EDBD" box="[1345,1399,1962,1988]" name="Peru" pageId="49" pageNumber="50">Peru</collectingCountry>
and
<collectingCountry id="F30C9ACDFF94FFC8FF36F871AC74ED91" box="[239,339,1998,2024]" name="Ecuador" pageId="49" pageNumber="50">Ecuador</collectingCountry>
. It is named for this mountain range.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF97FFCBFF64FEB6AF19E90C" pageId="50" pageNumber="51" type="materials_examined">
<paragraph id="8BA4DA5DFF97FFCBFF64FEB6ADDCEB5A" blockId="50.[189,251,265,291]" box="[189,251,265,291]" pageId="50" pageNumber="51">
<heading id="D0EC6D31FF97FFCBFF64FEB6ADDCEB5A" bold="true" box="[189,251,265,291]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
<emphasis id="B96F064FFF97FFCBFF64FEB6ADDCEB5A" bold="true" box="[189,251,265,291]" pageId="50" pageNumber="51">
<typeStatus id="54A064FFFF97FFCBFF64FEB6ADDCEB5A" box="[189,251,265,291]" pageId="50" pageNumber="51">Type</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF97FFCBFF64FE86A994EBB9" blockId="50.[189,1399,312,448]" pageId="50" pageNumber="51">
<materialsCitation id="3B73D000FF97FFCBFF64FE86A822EBE2" ID-GBIF-Occurrence="4517461798" collectingDate="2006-04-24" collectionCode="HUT" collectorName="E. Rodriguez &amp; S. Orroyo &amp; J. Nunez &amp; L. Campos" country="Peru" county="Huarango" elevation="2060" latitude="-5.271389" location="Cordillera Huarango" longLatPrecision="21" longitude="-78.67027" municipality="San Ignacio" pageId="50" pageNumber="51" specimenCode="HUT44721" specimenCount="1" stateProvince="Cajamarca Region" typeStatus="holotype">
<collectingCountry id="F30C9ACDFF97FFCBFF64FE86AC2FEB2A" box="[189,264,313,339]" name="Peru" pageId="50" pageNumber="51">PERU</collectingCountry>
–
<emphasis id="B96F064FFF97FFCBFEFDFE87AE27EB2B" bold="true" box="[292,768,312,338]" pageId="50" pageNumber="51">
<collectingRegion id="49DF14BFFF97FFCBFEFDFE87AF32EB2B" box="[292,533,312,338]" country="Peru" name="Cajamarca" pageId="50" pageNumber="51">Cajamarca Region</collectingRegion>
: Prov.
<collectingMunicipality id="6BC04027FF97FFCBFDB2FE87AE27EB2B" box="[619,768,312,338]" pageId="50" pageNumber="51">San Ignacio</collectingMunicipality>
</emphasis>
• Dist.
<collectingCounty id="62C5A2D1FF97FFCBFC81FE86AEE8EB2A" box="[856,975,313,339]" pageId="50" pageNumber="51">Huarango</collectingCounty>
,
<location id="8EC48C86FF97FFCBFC00FE87A9ECEB2A" LSID="urn:lsid:plazi:treatment:03B26B4BFF94FFCDFE37F9B3AFE3EE11:8EC48C86FF97FFCBFC00FE87A9ECEB2A" box="[985,1227,312,339]" country="Peru" county="Huarango" latitude="-5.271389" longLatPrecision="21" longitude="-78.67027" municipality="San Ignacio" name="Cordillera Huarango" pageId="50" pageNumber="51" stateProvince="Cajamarca Region">Cordillera Huarango</location>
(prolongación al sur de la Cordillera del Cóndor),
<location id="8EC48C86FF97FFCBFD8FFEE2AE64EB0E" LSID="urn:lsid:plazi:treatment:03B26B4BFF94FFCDFE37F9B3AFE3EE11:8EC48C86FF97FFCBFD8FFEE2AE64EB0E" box="[598,835,349,375]" country="Peru" county="Huarango" latitude="-5.271389" longLatPrecision="21" longitude="-78.67027" municipality="San Ignacio" name="Sector El Romerillo" pageId="50" pageNumber="51" stateProvince="Cajamarca Region">Sector El Romerillo</location>
;
<geoCoordinate id="EE2FBC9AFF97FFCBFC94FEE4AEEAEB0E" box="[845,973,347,375]" degrees="5" direction="south" minutes="16" orientation="latitude" pageId="50" pageNumber="51" precision="15" seconds="17" value="-5.271389">5°16′17″ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF97FFCBFC00FEE4A954EB0E" box="[985,1139,347,375]" degrees="78" direction="west" minutes="40" orientation="longitude" pageId="50" pageNumber="51" precision="15" seconds="13" value="-78.67027">78°40′13″ W</geoCoordinate>
;
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<quantity id="4CE377B8FF97FFCBFBA7FEE2A9F1EB0E" box="[1150,1238,349,375]" metricMagnitude="3" metricUnit="m" metricValue="2.06" pageId="50" pageNumber="51" unit="m" value="2060.0">2060 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF97FFCBFAF8FEE2ADDCEBE2" pageId="50" pageNumber="51" value="2006-04-24">
<collectingDate id="EFE10575FF97FFCBFAF8FEE2ADDCEBE2" pageId="50" pageNumber="51" value="2006-04-24">24 Apr. 2006</collectingDate>
</date>
;
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<collectorName id="26EEBF8BFF97FFCBFED0FE3DAC8EEBE2" box="[265,425,385,411]" pageId="50" pageNumber="51">E. Rodríguez</collectorName>
,
<collectorName id="26EEBF8BFF97FFCBFE6FFE3DAF08EBE5" box="[438,559,386,412]" pageId="50" pageNumber="51">S. Orroyo</collectorName>
,
<collectorName id="26EEBF8BFF97FFCBFDE4FE3DAF82EBE2" box="[573,677,386,411]" pageId="50" pageNumber="51">J. Nuñez</collectorName>
&amp;
<collectorName id="26EEBF8BFF97FFCBFD16FE3DAE76EBE5" box="[719,849,386,412]" pageId="50" pageNumber="51">L. Campos</collectorName>
2996
</emphasis>
;
<typeStatus id="54A064FFFF97FFCBFC7EFE3EA935EBE2" box="[935,1042,385,411]" pageId="50" pageNumber="51">holotype</typeStatus>
:
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[
<specimenCode id="DBBD7226FF97FFCBFBB2FE3EA9DDEBE2" box="[1131,1274,385,411]" collectionCode="HUT" country="Japan" httpUri="http://grbio.org/cool/gtk4-xv5b" name="HUT Culture Collection" pageId="50" pageNumber="51">HUT44721</specimenCode>
];
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<materialsCitation id="3B73D000FF97FFCBFAD7FE3EA994EBB9" ID-GBIF-Occurrence="4517461927" collectionCode="HUT, MO, US, QCNE" pageId="50" pageNumber="51" specimenCode="HUT44721, MO-2194463, US00951218, QCNE0220802" specimenCount="1" typeStatus="isotypes">
<typeStatus id="54A064FFFF97FFCBFAD7FE3EA854EBE2" box="[1294,1395,385,411]" pageId="50" pageNumber="51">isotypes</typeStatus>
:
<collectionCode id="ED0A4298FF97FFCBFF66FE19ADDCEBB9" box="[191,251,422,448]" collectionName="HUT" country="Japan" httpUri="http://grbio.org/cool/gtk4-xv5b" name="HUT Culture Collection" pageId="50" pageNumber="51">HUT</collectionCode>
[
<specimenCode id="DBBD7226FF97FFCBFED0FE19ACB0EBB9" box="[265,407,422,448]" collectionCode="HUT" country="Japan" httpUri="http://grbio.org/cool/gtk4-xv5b" name="HUT Culture Collection" pageId="50" pageNumber="51">HUT44721</specimenCode>
],
<collectionCode id="ED0A4298FF97FFCBFE73FE19ACFEEBB9" box="[426,473,422,448]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="50" pageNumber="51" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FF97FFCBFE31FE19AFAFEBB9" box="[488,648,422,448]" pageId="50" pageNumber="51">MO-2194463</specimenCode>
],
<collectionCode id="ED0A4298FF97FFCBFD79FE19AFE2EBB9" box="[672,709,422,448]" pageId="50" pageNumber="51">US</collectionCode>
[
<specimenCode id="DBBD7226FF97FFCBFD0EFE1AAE55EBB9" box="[727,882,421,448]" pageId="50" pageNumber="51">US00951218</specimenCode>
],
<collectionCode id="ED0A4298FF97FFCBFC53FE19AEFDEBB9" box="[906,986,422,448]" collectionName="QCNE" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13634" lsid="urn:lsid:biocol.org:col:13634" name="Museo Ecuatoriano de Ciencias Naturales" pageId="50" pageNumber="51" type="Herbarium">QCNE</collectionCode>
[
<specimenCode id="DBBD7226FF97FFCBFC31FE19A98FEBB9" box="[1000,1192,422,448]" collectionCode="QCNE" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13634" lsid="urn:lsid:biocol.org:col:13634" name="Museo Ecuatoriano de Ciencias Naturales" pageId="50" pageNumber="51" type="Herbarium">QCNE0220802</specimenCode>
].
</materialsCitation>
</paragraph>
<paragraph id="8BA4DA5DFF97FFCBFF64FE57ACE3E87B" blockId="50.[189,452,488,514]" box="[189,452,488,514]" pageId="50" pageNumber="51">
<heading id="D0EC6D31FF97FFCBFF64FE57ACE3E87B" bold="true" box="[189,452,488,514]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
<emphasis id="B96F064FFF97FFCBFF64FE57ACE3E87B" bold="true" box="[189,452,488,514]" pageId="50" pageNumber="51">Specimens examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF97FFCBFF64FDA7AC89E8D9" blockId="50.[189,1399,536,672]" pageId="50" pageNumber="51">
<materialsCitation id="3B73D000FF97FFCBFF64FDA7AC89E8D9" ID-GBIF-Occurrence="4517461321" collectingDate="2005-03-13" collectionCode="HUT, MO" collectorName="W. Quizhpe &amp; B. Medina &amp; C. Aguirre &amp; M. Prado" country="Ecuador" elevation="1800" latitude="-4.7169447" location="Region de la Cordillera del Condor" longLatPrecision="21" longitude="-78.96305" municipality="Palanda" pageId="50" pageNumber="51" specimenCode="HUT45696, MO-2135430" specimenCount="1" stateProvince="Prov. Santiago - Zamora">
<collectingCountry id="F30C9ACDFF97FFCBFF64FDA7AC6CE84B" box="[189,331,536,562]" name="Ecuador" pageId="50" pageNumber="51">ECUADOR</collectingCountry>
–
<emphasis id="B96F064FFF97FFCBFEBEFDA7AFA8E84B" bold="true" box="[359,655,536,563]" pageId="50" pageNumber="51">
<collectingRegion id="49DF14BFFF97FFCBFEBEFDA7AFA8E84B" box="[359,655,536,563]" pageId="50" pageNumber="51">Prov. Santiago -Zamora</collectingRegion>
</emphasis>
• Dist.
<collectingMunicipality id="6BC04027FF97FFCBFD3EFDA7AE60E84B" box="[743,839,536,562]" pageId="50" pageNumber="51">Palanda</collectingMunicipality>
,
<location id="8EC48C86FF97FFCBFC8BFDA7A9C8E84B" LSID="urn:lsid:plazi:treatment:03B26B4BFF94FFCDFE37F9B3AFE3EE11:8EC48C86FF97FFCBFC8BFDA7A9C8E84B" box="[850,1263,536,562]" country="Ecuador" latitude="-4.7169447" longLatPrecision="21" longitude="-78.96305" municipality="Palanda" name="Region de la Cordillera del Condor" pageId="50" pageNumber="51" stateProvince="Prov. Santiago - Zamora">Región de la Cordillera del Cóndor</location>
,
<location id="8EC48C86FF97FFCBFB23FDA6A855E84B" LSID="urn:lsid:plazi:treatment:03B26B4BFF94FFCDFE37F9B3AFE3EE11:8EC48C86FF97FFCBFB23FDA6A855E84B" box="[1274,1394,536,563]" country="Ecuador" latitude="-4.7169447" longLatPrecision="21" longitude="-78.96305" municipality="Palanda" name="sector Sur" pageId="50" pageNumber="51" stateProvince="Prov. Santiago - Zamora">sector Sur</location>
,
<location id="8EC48C86FF97FFCBFF64FD83AF75E82F" LSID="urn:lsid:plazi:treatment:03B26B4BFF94FFCDFE37F9B3AFE3EE11:8EC48C86FF97FFCBFF64FD83AF75E82F" box="[189,594,572,599]" country="Ecuador" county="Region de la Cordillera del Condor" latitude="-4.7169447" longLatPrecision="21" longitude="-78.96305" municipality="Palanda" name="Parroquia San Francisco de Vergel" pageId="50" pageNumber="51" stateProvince="Prov. Santiago - Zamora">Parroquia San Francisco de Vergel</location>
,
<location id="8EC48C86FF97FFCBFD84FD82AEF3E82F" LSID="urn:lsid:plazi:treatment:03B26B4BFF94FFCDFE37F9B3AFE3EE11:8EC48C86FF97FFCBFD84FD82AEF3E82F" box="[605,980,572,599]" country="Ecuador" latitude="-4.7169447" longLatPrecision="21" longitude="-78.96305" municipality="Palanda" name="Margen izqueirdo del rio Vergel" pageId="50" pageNumber="51" stateProvince="Prov. Santiago - Zamora">Margen izqueirdo del río Vergel</location>
, pantano;
<geoCoordinate id="EE2FBC9AFF97FFCBFB93FD84A9ECE82E" box="[1098,1227,571,599]" degrees="4" direction="south" minutes="43" orientation="latitude" pageId="50" pageNumber="51" precision="15" seconds="01" value="-4.7169447">4°43′01″ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF97FFCBFB0EFD84A855E82E" box="[1239,1394,571,599]" degrees="78" direction="west" minutes="57" orientation="longitude" pageId="50" pageNumber="51" precision="15" seconds="47" value="-78.96305">78°57′47″ W</geoCoordinate>
;
<elevation id="00363D6EFF97FFCBFF64FDDEAC72E802" box="[189,341,609,636]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="50" pageNumber="51" unit="m" value="1800.0">
<quantity id="4CE377B8FF97FFCBFF64FDDEAC31E805" box="[189,278,609,636]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="50" pageNumber="51" unit="m" value="1800.0">1800 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF97FFCBFEBBFDDEAF25E805" box="[354,514,609,636]" pageId="50" pageNumber="51" value="2005-03-13">
<collectingDate id="EFE10575FF97FFCBFEBBFDDEAF25E805" box="[354,514,609,636]" pageId="50" pageNumber="51" value="2005-03-13">13 Mar. 2005</collectingDate>
</date>
;
<emphasis id="B96F064FFF97FFCBFDD7FDDDA9A3E802" box="[526,1156,609,635]" italics="true" pageId="50" pageNumber="51">
<collectorName id="26EEBF8BFF97FFCBFDD7FDDDAFBFE802" box="[526,664,609,635]" pageId="50" pageNumber="51">W. Quizhpe</collectorName>
,
<collectorName id="26EEBF8BFF97FFCBFD7DFDDDAE06E802" box="[676,801,609,635]" pageId="50" pageNumber="51">B. Medina</collectorName>
,
<collectorName id="26EEBF8BFF97FFCBFCF7FDDEAE8DE802" box="[814,938,609,635]" pageId="50" pageNumber="51">C. Aguirre</collectorName>
&amp;
<collectorName id="26EEBF8BFF97FFCBFC09FDDDA966E802" box="[976,1089,609,635]" pageId="50" pageNumber="51">M. Prado</collectorName>
1013
</emphasis>
;
<collectionCode id="ED0A4298FF97FFCBFB4AFDDDA9E8E805" box="[1171,1231,610,636]" country="Japan" httpUri="http://grbio.org/cool/gtk4-xv5b" name="HUT Culture Collection" pageId="50" pageNumber="51">HUT</collectionCode>
[
<specimenCode id="DBBD7226FF97FFCBFB04FDDEA84CE805" box="[1245,1387,609,636]" collectionCode="HUT" country="Japan" httpUri="http://grbio.org/cool/gtk4-xv5b" name="HUT Culture Collection" pageId="50" pageNumber="51">HUT45696</specimenCode>
],
<collectionCode id="ED0A4298FF97FFCBFF64FD39ADCBE8D9" box="[189,236,646,672]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="50" pageNumber="51" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FF97FFCBFF24FD3AACBAE8D9" box="[253,413,645,672]" pageId="50" pageNumber="51">MO-2135430</specimenCode>
].
</materialsCitation>
</paragraph>
<paragraph id="8BA4DA5DFF97FFCBFF64FD76AF19E90C" blockId="50.[187,1399,712,885]" pageId="50" pageNumber="51">
<materialsCitation id="3B73D000FF97FFCBFF64FD76AE91E955" ID-GBIF-Occurrence="4517461865" collectionCode="HUT, MO, US" collectorName="E. Rodriguez R." country="Peru" county="Huarango" elevation="1700" latitude="-5.1666665" location="Nuevo Mundo" longLatPrecision="1305" longitude="-78.53333" municipality="San Ignacio" pageId="50" pageNumber="51" specimenCode="MO-1100939, US00843967" specimenCount="1" stateProvince="Cajamarca Region">
<collectingCountry id="F30C9ACDFF97FFCBFF64FD76AC2FE89A" box="[189,264,713,739]" name="Peru" pageId="50" pageNumber="51">PERU</collectingCountry>
–
<emphasis id="B96F064FFF97FFCBFEFCFD77AE26E89B" bold="true" box="[293,769,712,738]" pageId="50" pageNumber="51">
<collectingRegion id="49DF14BFFF97FFCBFEFCFD77AF31E89B" box="[293,534,712,738]" country="Peru" name="Cajamarca" pageId="50" pageNumber="51">Cajamarca Region</collectingRegion>
: Prov.
<collectingMunicipality id="6BC04027FF97FFCBFDB5FD77AE26E89B" box="[620,769,712,738]" pageId="50" pageNumber="51">San Ignacio</collectingMunicipality>
</emphasis>
• Dist.
<collectingCounty id="62C5A2D1FF97FFCBFC80FD76AEF7E89A" box="[857,976,713,739]" pageId="50" pageNumber="51">Huarango</collectingCounty>
,
<location id="8EC48C86FF97FFCBFC02FD76A9AFE89B" LSID="urn:lsid:plazi:treatment:03B26B4BFF94FFCDFE37F9B3AFE3EE11:8EC48C86FF97FFCBFC02FD76A9AFE89B" box="[987,1160,712,739]" country="Peru" county="Huarango" latitude="-5.1666665" longLatPrecision="1305" longitude="-78.53333" municipality="San Ignacio" name="Nuevo Mundo" pageId="50" pageNumber="51" stateProvince="Cajamarca Region">Nuevo Mundo</location>
,
<location id="8EC48C86FF97FFCBFB4DFD77A879E89B" LSID="urn:lsid:plazi:treatment:03B26B4BFF94FFCDFE37F9B3AFE3EE11:8EC48C86FF97FFCBFB4DFD77A879E89B" box="[1172,1374,712,738]" country="Peru" county="Huarango" latitude="-5.1666665" longLatPrecision="1305" longitude="-78.53333" municipality="San Ignacio" name="Caserio Pisaguas" pageId="50" pageNumber="51" stateProvince="Cajamarca Region">Caserío Pisaguas</location>
,
<locationDeviation id="9A16C37FFF97FFCBFAB0FD76AEF4E97E" pageId="50" pageNumber="51">a 2 horas del poblado y al norte, margen derecha quebrada Santa Rosa</locationDeviation>
;
<geoCoordinate id="EE2FBC9AFF97FFCBFC07FD54A913E97E" box="[990,1076,747,775]" degrees="5" direction="south" minutes="10" orientation="latitude" pageId="50" pageNumber="51" precision="925" value="-5.1666665">5°10′ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF97FFCBFB99FD54A997E97E" box="[1088,1200,747,775]" degrees="78" direction="west" minutes="32" orientation="longitude" pageId="50" pageNumber="51" precision="925" value="-78.53333">78°32′ W</geoCoordinate>
;
<elevation id="00363D6EFF97FFCBFB62FD52A876E97E" box="[1211,1361,749,775]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="50" pageNumber="51" unit="m" value="1700.0">
<quantity id="4CE377B8FF97FFCBFB62FD52A834E97E" box="[1211,1299,749,775]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="50" pageNumber="51" unit="m" value="1700.0">1700 m</quantity>
a.s.l.
</elevation>
;
<emphasis id="B96F064FFF97FFCBFA84FD52ACB1E952" italics="true" pageId="50" pageNumber="51">
<collectorName id="26EEBF8BFF97FFCBFA84FD52AC72E952" pageId="50" pageNumber="51">E. Rodríguez R.</collectorName>
1900
</emphasis>
;
<collectionCode id="ED0A4298FF97FFCBFE7AFCADACC7E955" box="[419,480,786,812]" collectionName="HUT" country="Japan" httpUri="http://grbio.org/cool/gtk4-xv5b" name="HUT Culture Collection" pageId="50" pageNumber="51">HUT</collectionCode>
,
<collectionCode id="ED0A4298FF97FFCBFE33FCAEAF3EE952" box="[490,537,785,811]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="50" pageNumber="51" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FF97FFCBFDF1FCAEAFE0E952" box="[552,711,785,811]" pageId="50" pageNumber="51">MO-1100939</specimenCode>
],
<collectionCode id="ED0A4298FF97FFCBFD05FCAEAE26E952" box="[732,769,785,811]" pageId="50" pageNumber="51">US</collectionCode>
[
<specimenCode id="DBBD7226FF97FFCBFCC8FCAEAE8BE952" box="[785,940,785,811]" pageId="50" pageNumber="51">US00843967</specimenCode>
]
</materialsCitation>
<materialsCitation id="3B73D000FF97FFCBFC63FCADAF19E90C" ID-GBIF-Occurrence="4517461431" collectionCode="HUT" collectorName="S. Arroyo A &amp; E. Rodriguez R." county="Huarango" elevation="2062" latitude="-5.2714167" location="Cordillera Huarango (El Romerillo)" longLatPrecision="1" longitude="-78.67042" pageId="50" pageNumber="51" specimenCount="1">
• Dist.
<collectingCounty id="62C5A2D1FF97FFCBFBDEFCADA959E955" box="[1031,1150,786,812]" pageId="50" pageNumber="51">Huarango</collectingCounty>
,
<location id="8EC48C86FF97FFCBFB5EFCAEAC4FE936" LSID="urn:lsid:plazi:treatment:03B26B4BFF94FFCDFE37F9B3AFE3EE11:8EC48C86FF97FFCBFB5EFCAEAC4FE936" county="Huarango" latitude="-5.2714167" longLatPrecision="1" longitude="-78.67042" name="Cordillera Huarango (El Romerillo)" pageId="50" pageNumber="51">Cordillera Huarango (El Romerillo)</location>
,
<locationDeviation id="9A16C37FFF97FFCBFEAAFC89AFDCE936" box="[371,763,821,848]" pageId="50" pageNumber="51">Base cordillera en el lado oriental</locationDeviation>
;
<geoCoordinate id="EE2FBC9AFF97FFCBFCDFFC8BAEB9E929" box="[774,926,820,848]" degrees="5" direction="south" minutes="16" orientation="latitude" pageId="50" pageNumber="51" precision="1" seconds="17.1" value="-5.2714167">5°16′17.1″ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF97FFCBFC73FC8BA97DE929" box="[938,1114,820,848]" degrees="78" direction="west" minutes="40" orientation="longitude" pageId="50" pageNumber="51" precision="1" seconds="13.5" value="-78.67042">78°40′13.5″ W</geoCoordinate>
;
<elevation id="00363D6EFF97FFCBFBBCFC8AA9DCE936" box="[1125,1275,821,848]" metricMagnitude="3" metricUnit="m" metricValue="2.062" pageId="50" pageNumber="51" unit="m" value="2062.0">
<quantity id="4CE377B8FF97FFCBFBBCFC8AA99AE929" box="[1125,1213,821,848]" metricMagnitude="3" metricUnit="m" metricValue="2.062" pageId="50" pageNumber="51" unit="m" value="2062.0">2062 m</quantity>
a.s.l.
</elevation>
;
<emphasis id="B96F064FFF97FFCBFAD1FC89ACCDE90D" italics="true" pageId="50" pageNumber="51">
<collectorName id="26EEBF8BFF97FFCBFAD1FC89ADF6E90D" pageId="50" pageNumber="51">S. Arroyo A</collectorName>
. &amp;
<collectorName id="26EEBF8BFF97FFCBFF22FCE4AC90E90D" box="[251,439,858,884]" pageId="50" pageNumber="51">E. Rodríguez R.</collectorName>
171
</emphasis>
;
<collectionCode id="ED0A4298FF97FFCBFE25FCE4AF1DE90C" box="[508,570,859,885]" collectionName="HUT" country="Japan" httpUri="http://grbio.org/cool/gtk4-xv5b" name="HUT Culture Collection" pageId="50" pageNumber="51">HUT</collectionCode>
.
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</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF97FFCBFF64FC23AEF3ED71" pageId="50" pageNumber="51" type="description">
<paragraph id="8BA4DA5DFF97FFCBFF64FC23AC68E9CF" blockId="50.[189,335,924,950]" box="[189,335,924,950]" pageId="50" pageNumber="51">
<heading id="D0EC6D31FF97FFCBFF64FC23AC68E9CF" bold="true" box="[189,335,924,950]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
<emphasis id="B96F064FFF97FFCBFF64FC23AC68E9CF" bold="true" box="[189,335,924,950]" pageId="50" pageNumber="51">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF97FFCBFF64FC73AEF3ED71" blockId="50.[187,1401,972,1801]" pageId="50" pageNumber="51">
Caulescent herb, to
<quantity id="4CE377B8FF97FFCBFE6AFC73AF27E99E" box="[435,512,972,999]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="50" pageNumber="51" unit="cm" value="50.0">50 cm</quantity>
high.
<emphasis id="B96F064FFF97FFCBFD8AFC72AFABE99E" box="[595,652,973,999]" italics="true" pageId="50" pageNumber="51">Stem</emphasis>
erect, branching; internodes to
<quantity id="4CE377B8FF97FFCBFBCEFC73A94CE99E" box="[1047,1131,972,999]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="50" pageNumber="51" unit="cm" value="5.5">5.5 cm</quantity>
long, to
<quantity id="4CE377B8FF97FFCBFB38FC72A80EE99E" box="[1249,1321,973,999]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="50" pageNumber="51" unit="mm" value="3.0">3 mm</quantity>
thick, succulent, green, sparsely to densely pilose.
<emphasis id="B96F064FFF97FFCBFD10FC4EAE00EE72" box="[713,807,1009,1035]" italics="true" pageId="50" pageNumber="51">Stipules</emphasis>
deciduous, lanceolate to elliptic, 5–20 ×
<quantity id="4CE377B8FF97FFCBFAD7FC4EA855EE72" box="[1294,1394,1009,1036]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="50" pageNumber="51" unit="mm" value="3.0" valueMax="5.0" valueMin="1.0">1–5 mm</quantity>
, apex acute, mucronate, translucent, brown, glabrous, margin entire, aciliate.
<emphasis id="B96F064FFF97FFCBFBBAFBA9A992EE56" box="[1123,1205,1046,1071]" italics="true" pageId="50" pageNumber="51">Leaves</emphasis>
5–10 per stem, alternate, basifixed; petiole
<quantity id="4CE377B8FF97FFCBFDD0FB85AFA8EE2D" box="[521,655,1082,1109]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="15.0" metricValueMin="2.0" pageId="50" pageNumber="51" unit="cm" value="0.85" valueMax="1.5" valueMin="0.2">0.2–1.5 cm</quantity>
long, green, sparsely to densely villous; blade subsymmetric, lanceolate to oblanceolate, to 13 ×
<quantity id="4CE377B8FF97FFCBFD8CFBE1AF8FEE00" box="[597,680,1118,1145]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="50" pageNumber="51" unit="cm" value="4.5">4.5 cm</quantity>
, membranaceous, apex acuminate, base dimidiate, the largest side rounded, the narrow side cuneate and up to
<quantity id="4CE377B8FF97FFCBFD21FB3CAE1AEEE4" box="[760,829,1155,1182]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="50" pageNumber="51" unit="mm" value="5.0">5 mm</quantity>
further along the petiole, margin serrate, ciliate, upper surface green, sparsely pilose, lower surface green, glabrous to sparsely pilose, sparsely pilose on the veins, veins pinnate, with 6–10 secondary veins on the larger side, 5–7 secondary veins on the smaller side.
<emphasis id="B96F064FFF97FFCBFE87FB4FAF21EF73" box="[350,518,1264,1290]" italics="true" pageId="50" pageNumber="51">Inflorescences</emphasis>
1 per stem, bisexual, axillary, erect, cymose, with 4 branches, bearing up to 4 staminate flowers and 1 pistillate flower, protandrous; peduncle to
<quantity id="4CE377B8FF97FFCBFBCEFAABA940EF56" box="[1047,1127,1300,1327]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="50" pageNumber="51" unit="cm" value="3.5">3.5 cm</quantity>
long, sparsely villous, bracts deciduous, elliptic to ovate, 3–10 ×
<quantity id="4CE377B8FF97FFCBFD68FA86AE31EF2D" box="[689,790,1337,1364]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="50" pageNumber="51" unit="mm" value="2.5" valueMax="4.0" valueMin="1.0">1–4 mm</quantity>
, translucent, red to orange, glabrous, apex rounded, margin entire to serrate at the apex, aciliate to ciliate at the apex.
<emphasis id="B96F064FFF97FFCBFC62FAE1A9A9EF0E" box="[955,1166,1373,1400]" italics="true" pageId="50" pageNumber="51">Staminate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FF97FFCBFAFDFAE1A850EF01" box="[1316,1399,1374,1400]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="50" pageNumber="51" unit="mm" value="17.0">17 mm</quantity>
long, densely villous; tepals 4, projecting, outer 2 ovate, 8–13 ×
<quantity id="4CE377B8FF97FFCBFC1CFA3DA91DEFE5" box="[965,1082,1410,1436]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="50" pageNumber="51" unit="mm" value="7.0" valueMax="10.0" valueMin="4.0">4–10 mm</quantity>
, apex rounded to truncate, orange to red, glabrous, margin entire, aciliate, inner 2 elliptic, 7–17 ×
<quantity id="4CE377B8FF97FFCBFBD9FA19A943EFB8" box="[1024,1124,1446,1473]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="50" pageNumber="51" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6 mm</quantity>
, apex acute to truncate, orange to red, glabrous, margin entire, aciliate; stamens 4–6, projecting, white, filaments
<quantity id="4CE377B8FF97FFCBFB0AFA74A813EF9C" box="[1235,1332,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="50" pageNumber="51" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
long, free, anthers ellipsoid, ca 3 ×
<quantity id="4CE377B8FF97FFCBFDE8FA50AFB5EC73" box="[561,658,1519,1546]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="50" pageNumber="51" unit="mm" value="0.5">0.5 mm</quantity>
, dehiscing via lateral slits, connectives extended to
<quantity id="4CE377B8FF97FFCBFAC8FA50A855EC73" box="[1297,1394,1519,1546]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="50" pageNumber="51" unit="mm" value="0.2">0.2 mm</quantity>
, symmetrically basifixed.
<emphasis id="B96F064FFF97FFCBFE38F9ABAF80EC57" box="[481,679,1556,1582]" italics="true" pageId="50" pageNumber="51">Pistillate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FF97FFCBFCE1F9ABAE5DEC57" box="[824,890,1556,1582]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="50" pageNumber="51" unit="mm" value="8.0">8 mm</quantity>
long; bracteoles lacking; tepals 5, subequal, persistence unknown, projecting, the largest ovate, 9–17 ×
<quantity id="4CE377B8FF97FFCBFCB1F987AEFBEC2A" box="[872,988,1592,1619]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="50" pageNumber="51" unit="mm" value="7.0" valueMax="10.0" valueMin="4.0">4–10 mm</quantity>
, apex acute rounded, orange to red, glabrous, margin entire, aciliate, the smallest elliptic to ovate, 9–14 ×
<quantity id="4CE377B8FF97FFCBFC30F9E3A944EC0E" box="[1001,1123,1628,1655]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="5.0" metricValueMin="1.5" pageId="50" pageNumber="51" unit="mm" value="3.25" valueMax="5.0" valueMin="1.5">1.5–5 mm</quantity>
, apex acute to rounded, orange to red, glabrous, margin entire, aciliate; ovary body obdeltoid, 3–5 ×
<quantity id="4CE377B8FF97FFCBFB82F93EA9E6ECE5" box="[1115,1217,1665,1692]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="50" pageNumber="51" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
, orange to red, sparsely villous, equally 3-winged, wings horn-shaped, ca 2 ×
<quantity id="4CE377B8FF97FFCBFC6EF919A926ECB9" box="[951,1025,1702,1728]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="50" pageNumber="51" unit="mm" value="8.0">8 mm</quantity>
; 3-locular, placentae unknown; styles 3, orange to red, fused at the base,
<quantity id="4CE377B8FF97FFCBFD42F975AFDBEC9C" box="[667,764,1738,1765]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="50" pageNumber="51" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
long, irregularly 5 times divided, stigmatic papillae in a spirally-twisted band.
<emphasis id="B96F064FFF97FFCBFE0EF950AFB0ED71" box="[471,663,1774,1800]" italics="true" pageId="50" pageNumber="51">Fruiting pedicel</emphasis>
unknown.
<emphasis id="B96F064FFF97FFCBFCC3F950AE70ED71" box="[794,855,1775,1800]" italics="true" pageId="50" pageNumber="51">Fruit</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF97FFCDFF64F88EA9C1EB33" lastPageId="52" lastPageNumber="53" pageId="50" pageNumber="51" type="conservation">
<paragraph id="8BA4DA5DFF97FFCBFF64F88EAF4AED32" blockId="50.[189,621,1841,1867]" box="[189,621,1841,1867]" pageId="50" pageNumber="51">
<heading id="D0EC6D31FF97FFCBFF64F88EAF4AED32" bold="true" box="[189,621,1841,1867]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
<emphasis id="B96F064FFF97FFCBFF64F88EAF4AED32" bold="true" box="[189,621,1841,1867]" pageId="50" pageNumber="51">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF97FFCDFF64F8DEA9C1EB33" blockId="50.[189,1400,1888,2025]" lastBlockId="52.[189,1398,265,331]" lastPageId="52" lastPageNumber="53" pageId="50" pageNumber="51">
Known from three localities in the Cordillera del Cóndor and with an EOO of
<quantity id="4CE377B8FF97FFCBFB83F8DEA995ED02" box="[1114,1202,1889,1915]" metricMagnitude="5" metricUnit="m" metricValue="7.2" pageId="50" pageNumber="51" unit="km" value="720.0">720 km</quantity>
<superScript id="7C6E7715FF97FFCBFB6AF8DFA99BED17" attach="right" box="[1203,1212,1888,1902]" fontSize="6" pageId="50" pageNumber="51">2</superScript>
. The Cordillera is partially deforested on its Western flank in
<collectingCountry id="F30C9ACDFF97FFCBFD14F839AE24EDD9" box="[717,771,1926,1952]" name="Peru" pageId="50" pageNumber="51">Peru</collectingCountry>
and
<collectingCountry id="F30C9ACDFF97FFCBFCE2F83AAEB8EDE6" box="[827,927,1925,1951]" name="Ecuador" pageId="50" pageNumber="51">Ecuador</collectingCountry>
, but extensive forest remains elsewhere. It is likely that
<taxonomicName id="4C1BA1DEFF97FFCBFEAEF814AF00EDBD" authorityName="Moonlight" authorityYear="2023" box="[375,551,1962,1988]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B96F064FFF97FFCBFEAEF814AF00EDBD" box="[375,551,1962,1988]" italics="true" pageId="50" pageNumber="51">B. condorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FF97FFCBFDE8F815AFAAEDBD" box="[561,653,1962,1988]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
is found in much of this habitat. We were unable to locate the species at its
<typeStatus id="54A064FFFF97FFCBFEBCF870ACBFED90" box="[357,408,1999,2025]" pageId="50" pageNumber="51">type</typeStatus>
locality during fieldwork in 2016 but this is likely because it is common only in specific microhabitats. Given its small range, few known localities, and the deforestation on the west of the Cordillera del Cóndor, we assess
<taxonomicName id="4C1BA1DEFF91FFCDFDA9FE8EAE38EB33" authorityName="Moonlight" authorityYear="2023" box="[624,799,304,330]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B96F064FFF91FFCDFDA9FE8EAE38EB33" box="[624,799,304,330]" italics="true" pageId="52" pageNumber="53">B. condorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FF91FFCDFCFEFE8EAEA6EB32" box="[807,897,305,331]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
as Vulnerable (VU B2ab(iii)).
</paragraph>
</subSubSection>
<caption id="DF648AD5FF96FFCAFF64F837AFB4ED91" ID-DOI="http://doi.org/10.5281/zenodo.8178327" ID-Zenodo-Dep="8178327" httpUri="https://zenodo.org/record/8178327/files/figure.png" pageId="51" pageNumber="52" startId="51.[189,232,1928,1954]" targetBox="[227,1334,414,1820]" targetPageId="51">
<paragraph id="8BA4DA5DFF96FFCAFF64F837AFB4ED91" blockId="51.[189,1399,1928,2024]" pageId="51" pageNumber="52">
<emphasis id="B96F064FFF96FFCAFF64F837AC39EDDB" bold="true" box="[189,286,1928,1954]" pageId="51" pageNumber="52">Fig. 20.</emphasis>
<taxonomicName id="4C1BA1DEFF96FFCAFEF1F836AE21EDDB" authority="Jara &amp; Moonlight" authorityName="Jara &amp; Moonlight" authorityYear="2023" box="[296,774,1928,1954]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="51" pageNumber="52" phylum="Tracheophyta" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B96F064FFF96FFCAFEF1F836AF03EDDB" box="[296,548,1928,1954]" italics="true" pageId="51" pageNumber="52">Begonia condorensis</emphasis>
Jara &amp; Moonlight
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FF96FFCAFCC9F837AE49EDDB" box="[784,878,1928,1954]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B96F064FFF96FFCAFCA0F837AEA9EDDB" bold="true" box="[889,910,1928,1954]" pageId="51" pageNumber="52">A</emphasis>
. Habit.
<emphasis id="B96F064FFF96FFCAFC2AF837A920EDDB" bold="true" box="[1011,1031,1928,1954]" pageId="51" pageNumber="52">B</emphasis>
. Staminate flower, front view.
<emphasis id="B96F064FFF96FFCAFF64F814ADF5EDBC" bold="true" box="[189,210,1963,1989]" pageId="51" pageNumber="52">C</emphasis>
. Staminate flower, side.
<emphasis id="B96F064FFF96FFCAFDD8F814AF31EDBC" bold="true" box="[513,534,1963,1989]" pageId="51" pageNumber="52">D</emphasis>
. Pistillate flower, front view.
<emphasis id="B96F064FFF96FFCAFC5AF814AEB0EDBC" bold="true" box="[899,919,1963,1989]" pageId="51" pageNumber="52">E</emphasis>
. Style and stigma. Illustration by Anna Dorward from
<emphasis id="B96F064FFF96FFCAFEB4F870AF6CED91" box="[365,587,1998,2024]" italics="true" pageId="51" pageNumber="52">E. Rodríguez 2996</emphasis>
(US).
</paragraph>
</caption>
<subSubSection id="C30189D6FF91FFCDFF64FEC9AF00E928" pageId="52" pageNumber="53" type="discussion">
<paragraph id="8BA4DA5DFF91FFCDFF64FEC9AC97EBE8" blockId="52.[189,432,374,401]" box="[189,432,374,401]" pageId="52" pageNumber="53">
<heading id="D0EC6D31FF91FFCDFF64FEC9AC97EBE8" bold="true" box="[189,432,374,401]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="B96F064FFF91FFCDFF64FEC9AC97EBE8" bold="true" box="[189,432,374,401]" pageId="52" pageNumber="53">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF91FFCDFF64FE16A96AE843" blockId="52.[189,1399,425,571]" pageId="52" pageNumber="53">
Within
<collectingCountry id="F30C9ACDFF91FFCDFEC2FE15AC73EBBD" box="[283,340,426,452]" name="Peru" pageId="52" pageNumber="53">Peru</collectingCountry>
,
<taxonomicName id="4C1BA1DEFF91FFCDFEBFFE15AF3BEBBA" authorityName="Moonlight" authorityYear="2023" box="[358,540,425,451]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B96F064FFF91FFCDFEBFFE15AF3BEBBA" box="[358,540,425,451]" italics="true" pageId="52" pageNumber="53">B. condorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FF91FFCDFDF3FE15AFABEBBD" box="[554,652,426,452]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
is the only species with pinnately veined leaves and &lt;10 stamens. It superficially resembles
<taxonomicName id="4C1BA1DEFF91FFCDFD82FE6DAFF6EB92" box="[603,721,466,491]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="urticae">
<emphasis id="B96F064FFF91FFCDFD82FE6DAFF6EB92" box="[603,721,466,491]" italics="true" pageId="52" pageNumber="53">B. urticae</emphasis>
</taxonomicName>
, especially relatively glabrous individuals with relatively membranaceous leaves,
<taxonomicName id="4C1BA1DEFF91FFCDFE3BFE46AF7CE86B" box="[482,603,505,530]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="urticae">
<emphasis id="B96F064FFF91FFCDFE3BFE46AF7CE86B" box="[482,603,505,530]" italics="true" pageId="52" pageNumber="53">B. urticae</emphasis>
</taxonomicName>
differs in its much smaller staminate (up to
<quantity id="4CE377B8FF91FFCDFB58FE46A9EFE86A" box="[1153,1224,505,531]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="52" pageNumber="53" unit="mm" value="8.0">8 mm</quantity>
long vs up to
<quantity id="4CE377B8FF91FFCDFF64FD9FAC37E842" box="[189,272,544,571]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="52" pageNumber="53" unit="mm" value="17.0">17 mm</quantity>
long) and pistillate tepals (up to
<quantity id="4CE377B8FF91FFCDFD4BFD9FAFF1E842" box="[658,726,544,571]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="52" pageNumber="53" unit="mm" value="6.0">6 mm</quantity>
long versus up to
<quantity id="4CE377B8FF91FFCDFC77FD9FA926E842" box="[942,1025,544,571]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="52" pageNumber="53" unit="mm" value="17.0">17 mm</quantity>
long).
</paragraph>
<paragraph id="8BA4DA5DFF91FFCDFF64FDD9AC23E8F9" blockId="52.[189,260,614,640]" box="[189,260,614,640]" pageId="52" pageNumber="53">
<heading id="D0EC6D31FF91FFCDFF64FDD9AC23E8F9" bold="true" box="[189,260,614,640]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="B96F064FFF91FFCDFF64FDD9AC23E8F9" bold="true" box="[189,260,614,640]" pageId="52" pageNumber="53">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF91FFCDFF64FD26AF00E928" blockId="52.[188,1399,665,849]" pageId="52" pageNumber="53">
The fruits of
<taxonomicName id="4C1BA1DEFF91FFCDFE8FFD25AF22E8CA" authorityName="Moonlight" authorityYear="2023" box="[342,517,665,691]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B96F064FFF91FFCDFE8FFD25AF22E8CA" box="[342,517,665,691]" italics="true" pageId="52" pageNumber="53">B. condorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FF91FFCDFDD4FD26AF40E8CA" box="[525,615,665,691]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
remain unknown. A single duplicate of the specimen
<emphasis id="B96F064FFF91FFCDFB05FD25AC3FE8A2" italics="true" pageId="52" pageNumber="53">E. Rodríguez R. 1900</emphasis>
(MO [MO-1100939]) has a relatively mature pistillate flower with a well-developed ovary. This suggests that the fruit is similar to that of
<taxonomicName id="4C1BA1DEFF91FFCDFD7AFD56AE3CE97B" box="[675,795,744,770]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="valvata">
<emphasis id="B96F064FFF91FFCDFD7AFD56AE3CE97B" box="[675,795,744,770]" italics="true" pageId="52" pageNumber="53">B. valvata</emphasis>
</taxonomicName>
, which is a large, three-horned capsule with a large apical column. In contrast to the fruits of
<taxonomicName id="4C1BA1DEFF91FFCDFD44FCAEAE33E953" box="[669,788,784,810]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="valvata">
<emphasis id="B96F064FFF91FFCDFD44FCAEAE33E953" box="[669,788,784,810]" italics="true" pageId="52" pageNumber="53">B. valvata</emphasis>
</taxonomicName>
, the horns of
<taxonomicName id="4C1BA1DEFF91FFCDFC6DFCAEA945E953" authorityName="Moonlight" authorityYear="2023" box="[948,1122,784,810]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B96F064FFF91FFCDFC6DFCAEA945E953" box="[948,1122,784,810]" italics="true" pageId="52" pageNumber="53">B. condorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FF91FFCDFBB3FCAFA9E4E953" box="[1130,1219,784,810]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
are most likely projecting rather than reflexed.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF91FFCDFF64FCC2AFE3EE11" pageId="52" pageNumber="53" type="distribution">
<paragraph id="8BA4DA5DFF91FFCDFF64FCC2ACD2E9EE" blockId="52.[189,501,893,919]" box="[189,501,893,919]" pageId="52" pageNumber="53">
<heading id="D0EC6D31FF91FFCDFF64FCC2ACD2E9EE" bold="true" box="[189,501,893,919]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="B96F064FFF91FFCDFF64FCC2ACD2E9EE" bold="true" box="[189,501,893,919]" pageId="52" pageNumber="53">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF91FFCDFF64FC0EAFE3EE11" blockId="52.[187,1400,944,1129]" pageId="52" pageNumber="53">
Known from
<collectingCountry id="F30C9ACDFF91FFCDFE87FC0FACE7E9B3" box="[350,448,944,970]" name="Ecuador" pageId="52" pageNumber="53">Ecuador</collectingCountry>
and
<collectingCountry id="F30C9ACDFF91FFCDFE22FC0EAF16E9B2" box="[507,561,945,971]" name="Peru" pageId="52" pageNumber="53">Peru</collectingCountry>
and collected in the Ecuadorian Province of Santiago-Zamora and the border of
<collectingRegion id="49DF14BFFF91FFCDFEF7FC67AC8CE98B" box="[302,427,984,1010]" country="Peru" name="Cajamarca" pageId="52" pageNumber="53">Cajamarca</collectingRegion>
and Amazonas Regions in
<collectingCountry id="F30C9ACDFF91FFCDFD3BFC67AE3FE98B" box="[738,792,984,1010]" name="Peru" pageId="52" pageNumber="53">Peru</collectingCountry>
(
<figureCitation id="1320C6D8FF91FFCDFCFFFC67AEB6E98B" box="[806,913,984,1010]" captionStart="Fig" captionStartId="52.[189,232,1892,1918]" captionTargetBox="[194,1392,1224,1843]" captionTargetId="figure-267@52.[189,1398,1214,1849]" captionTargetPageId="52" captionText="Fig. 21. Distribution of Begonia sect. Casparya (Klotzsch) Warb. in Peru and surrounding countries. A. B. condorensis Jara &amp; Moonlight sp. nov. (red) and B. hirta (Klotzsch) L.B.Sm. &amp; B.G.Schub. (blue). B. B. urticae L.f. (red). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178331" httpUri="https://zenodo.org/record/8178331/files/figure.png" pageId="52" pageNumber="53">Fig. 21A</figureCitation>
). Found within lower and middle montane forest at an elevation of
<quantity id="4CE377B8FF91FFCDFE3CFC40AFADEE63" box="[485,650,1023,1050]" metricMagnitude="3" metricUnit="m" metricValue="1.8809999999999998" metricValueMax="2.062" metricValueMin="1.7" pageId="52" pageNumber="53" unit="m" value="1881.0" valueMax="2062.0" valueMin="1700.0">1700–2062 m</quantity>
a.s.l. Like other species in
<emphasis id="B96F064FFF91FFCDFC04FBBFAED1EE60" box="[989,1014,1024,1049]" italics="true" pageId="52" pageNumber="53">B.</emphasis>
sect.
<taxonomicName id="4C1BA1DEFF91FFCDFB99FBBFA994EE63" box="[1088,1203,1024,1050]" class="Magnoliopsida" family="Begoniaceae" genus="Casparya" kingdom="Plantae" order="Cucurbitales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F064FFF91FFCDFB99FBBFA994EE63" box="[1088,1203,1024,1050]" italics="true" pageId="52" pageNumber="53">Casparya</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1BA1DEFF91FFCDFB1DFBBFA851EE60" authorityName="Moonlight" authorityYear="2023" box="[1220,1398,1023,1049]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="condorensis" status="sp. nov.">
<emphasis id="B96F064FFF91FFCDFB1DFBBFA851EE60" box="[1220,1398,1023,1049]" italics="true" pageId="52" pageNumber="53">B. condorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FF91FFCDFF64FB98AC3BEE38" box="[189,284,1063,1089]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
is found in shaded humid microhabitats, including around waterfalls, and its horned fruits appear adapted for dispersal by water drops.
</paragraph>
</subSubSection>
</treatment>
</document>
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