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<mods:title id="1C958A5DFC36091F3920623A87F20124">The genus Begonia (Begoniaceae) in Peru</mods:title>
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75. 
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74: 610 (
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], 
<collectionCode id="ED0A4298FE8CFED0FB41FDC0A98AE8E0" box="[1176,1197,639,665]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15513" name="Nationaal Herbarium Nederland" pageId="297" pageNumber="298" type="Herbarium">U</collectionCode>
[
<specimenCode id="DBBD7226FE8CFED0FB62FDC0A867E8E0" box="[1211,1344,639,665]" collectionCode="U" country="Netherlands" lsid="urn:lsid:biocol.org:col:15513" name="Nationaal Herbarium Nederland" pageId="297" pageNumber="298" type="Herbarium">U0139626</specimenCode>
], 
<collectionCode id="ED0A4298FE8CFED0FA88FDC0A851E8E0" box="[1361,1398,639,665]" pageId="297" pageNumber="298">US</collectionCode>
[
<specimenCode id="DBBD7226FE8CFED0FF2DFD1CACA9E8C7" box="[244,398,675,702]" pageId="297" pageNumber="298">US00115468</specimenCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE8CFED0FF64FD78A813E898" box="[189,1332,711,738]" pageId="297" pageNumber="298" type="reference_group">
<paragraph id="8BA4DA5DFE8CFED0FF64FD78A813E898" blockId="297.[189,1399,493,738]" box="[189,1332,711,738]" pageId="297" pageNumber="298">
<bibRefCitation id="EF8AA7ACFE8CFED0FF64FD78AF01E89B" author="Brako &amp; Zarucchi" box="[189,550,711,738]" firstAuthor="Brako" page="195" pageId="297" pageNumber="298" refId="ref225797" refString="Brako L. &amp; Zarucchi L. 1993. Catalogue of the Flowering Plants and Gymnosperms of Peru. Monographs in Systematic Botany from the Missouri Botanical Garden 45, Missouri Botanical Garden, St Louis." type="book" year="1993">Brako &amp; Zarucchi (1993: 195)</bibRefCitation>
; 
<bibRefCitation id="EF8AA7ACFE8CFED0FDEAFD78AEBAE898" author="Leon &amp; Monsalve" box="[563,925,711,737]" firstAuthor="Leon" page="169" pageId="297" pageNumber="298" pagination="165 - 170" refId="ref229226" refString="Leon B. &amp; Monsalve C. 2006. Begoniaceae endemicas del Peru. Revista Peruana de Biologia 13 (2): 165 - 170. https: // doi. org / 10.15381 / rpb. v 13 i 2.1810" type="journal article" year="2006">
<collectingRegion id="49DF14BFFE8CFED0FDEAFD78AF48E898" box="[563,623,711,737]" country="Nicaragua" name="Leon" pageId="297" pageNumber="298">León</collectingRegion>
&amp; Monsalve (2006: 169)
</bibRefCitation>
; 
<bibRefCitation id="EF8AA7ACFE8CFED0FC70FD78A817E898" author="Moonlight &amp; Tebbitt" box="[937,1328,711,737]" firstAuthor="Moonlight" page="119" pageId="297" pageNumber="298" pagination="111 - 122" refId="ref229847" refString="Moonlight P. W. &amp; Tebbitt M. C. 2016. Two new Peruvian species of Begonia (Begoniaceae) and an amended description of Begonia thyrsoidea. Edinburgh Journal of Botany 74 (2): 111 - 122. https: // doi. org / 10.1017 / S 0960428616000299" type="journal article" year="2016">Moonlight &amp; Tebbitt (2016: 119)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE8CFED0FF64FCB5ACCAE901" pageId="297" pageNumber="298" type="etymology">
<paragraph id="8BA4DA5DFE8CFED0FF64FCB5AC62E95D" blockId="297.[189,325,778,804]" box="[189,325,778,804]" pageId="297" pageNumber="298">
<heading id="D0EC6D31FE8CFED0FF64FCB5AC62E95D" bold="true" box="[189,325,778,804]" fontSize="11" level="3" pageId="297" pageNumber="298" reason="3">
<emphasis id="B96F064FFE8CFED0FF64FCB5AC62E95D" bold="true" box="[189,325,778,804]" pageId="297" pageNumber="298">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFE8CFED0FF64FC86ACCAE901" blockId="297.[189,1398,825,888]" pageId="297" pageNumber="298">Named for the resemblance of the unusual androecium of the species to a thyrsus, a wand or staff from ancient Greek mythology.</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE8CFED0FF64FC1FAE82EE4A" pageId="297" pageNumber="298" type="materials_examined">
<paragraph id="8BA4DA5DFE8CFED0FF64FC1FACE3E9C3" blockId="297.[189,452,928,954]" box="[189,452,928,954]" pageId="297" pageNumber="298">
<heading id="D0EC6D31FE8CFED0FF64FC1FACE3E9C3" bold="true" box="[189,452,928,954]" fontSize="11" level="3" pageId="297" pageNumber="298" reason="3">
<emphasis id="B96F064FFE8CFED0FF64FC1FACE3E9C3" bold="true" box="[189,452,928,954]" pageId="297" pageNumber="298">Specimens examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFE8CFED0FF64FC6EAE82EE4A" blockId="297.[189,1399,976,1076]" pageId="297" pageNumber="298">
<collectingCountry id="F30C9ACDFE8CFED0FF64FC6EAC2FE992" box="[189,264,977,1003]" name="Peru" pageId="297" pageNumber="298">PERU</collectingCountry>
– 
<emphasis id="B96F064FFE8CFED0FEF5FC6FAF97E993" bold="true" box="[300,688,976,1002]" pageId="297" pageNumber="298">
<collectingRegion id="49DF14BFFE8CFED0FEF5FC6FACF2E993" box="[300,469,976,1002]" country="Peru" name="Puno" pageId="297" pageNumber="298">
Puno 
<materialsCitation id="3B73D000FE8CFED0FEA1FC6FAE82EE4A" collectingDate="2015-01-09" collectionCode="E, MOL" collectorName="M. C. Tebbitt &amp; A. Daza" county="Prov. Carabaya" elevation="1503" latitude="-13.633333" location="Dist. San Gaban" longLatPrecision="1290" longitude="-70.45" municipality="Dist. San Gaban" pageId="297" pageNumber="298" specimenCode="E01059318" specimenCount="1">
Region: 
<collectingCounty id="62C5A2D1FE8CFED0FE3FFC6FAF97E993" box="[486,688,976,1002]" pageId="297" pageNumber="298">Prov. Carabaya</collectingCounty>
• 
<collectingMunicipality id="6BC04027FE8CFED0FD09FC6FAEBEE993" box="[720,921,976,1002]" pageId="297" pageNumber="298">Dist. San Gabán</collectingMunicipality>
, 
<locationDeviation id="9A16C37FFE8CFED0FC71FC6FA865E993" box="[936,1346,976,1002]" firstLocation="Ollachea" pageId="297" pageNumber="298" secondLocation="San Gaban">between Ollachea and San Gabán</locationDeviation>
, 
<locationDeviation id="9A16C37FFE8CFED0FA88FC6FAE53EE76" pageId="297" pageNumber="298">km 259+200, below the hydroelectric substation San Gabán II</locationDeviation>
; 
<geoCoordinate id="EE2FBC9AFE8CFED0FC58FC4CAECFEE76" box="[897,1000,1011,1039]" degrees="13" direction="south" minutes="38" orientation="latitude" pageId="297" pageNumber="298" precision="925" value="-13.633333">13°38′ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFE8CFED0FC2FFC4CA94FEE76" box="[1014,1128,1011,1039]" degrees="70" direction="west" minutes="27" orientation="longitude" pageId="297" pageNumber="298" precision="925" value="-70.45">70°27′ W</geoCoordinate>
; 
<elevation id="00363D6EFE8CFED0FBACFC4BA87EEE77" box="[1141,1369,1012,1039]" metricMagnitude="3" metricUnit="m" metricValue="1.5025" metricValueMax="1.555" metricValueMin="1.45" pageId="297" pageNumber="298" unit="m" value="1502.5" valueMax="1555.0" valueMin="1450.0">
<quantity id="4CE377B8FE8CFED0FBACFC4BA83EEE76" box="[1141,1305,1012,1039]" metricMagnitude="3" metricUnit="m" metricValue="1.5025" metricValueMax="1.555" metricValueMin="1.45" pageId="297" pageNumber="298" unit="m" value="1502.5" valueMax="1555.0" valueMin="1450.0">1450–1555 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFE8CFED0FAB1FC4AAC16EE4D" pageId="297" pageNumber="298" value="2015-01-09">
<collectingDate id="EFE10575FE8CFED0FAB1FC4AAC16EE4D" pageId="297" pageNumber="298" value="2015-01-09">9 Jan. 2015</collectingDate>
</date>
; 
<emphasis id="B96F064FFE8CFED0FEE4FBA6AFADEE4A" box="[317,650,1049,1075]" italics="true" pageId="297" pageNumber="298">
<collectorName id="26EEBF8BFE8CFED0FEE4FBA6ACF6EE4A" box="[317,465,1049,1075]" pageId="297" pageNumber="298">M.C. Tebbitt</collectorName>
&amp; 
<collectorName id="26EEBF8BFE8CFED0FE2FFBA6AF71EE4A" box="[502,598,1049,1075]" pageId="297" pageNumber="298">A. Daza</collectorName>
809
</emphasis>
; 
<collectionCode id="ED0A4298FE8CFED0FD40FBA6AF8CEE4A" box="[665,683,1049,1075]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="297" pageNumber="298" type="Herbarium">E</collectionCode>
[
<specimenCode id="DBBD7226FE8CFED0FD65FBA6AE63EE4D" box="[700,836,1049,1076]" collectionCode="E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="297" pageNumber="298" type="Herbarium">E01059318</specimenCode>
], 
<collectionCode id="ED0A4298FE8CFED0FC85FBA6AE86EE4A" box="[860,929,1049,1075]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="297" pageNumber="298" type="Herbarium">MOL</collectionCode>
.
</materialsCitation>
</collectingRegion>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE8CFED3FF64FBE4AEAEEB3E" lastPageId="298" lastPageNumber="299" pageId="297" pageNumber="298" type="description">
<paragraph id="8BA4DA5DFE8CFED0FF64FBE4AC68EE0C" blockId="297.[189,335,1115,1141]" box="[189,335,1115,1141]" pageId="297" pageNumber="298">
<heading id="D0EC6D31FE8CFED0FF64FBE4AC68EE0C" bold="true" box="[189,335,1115,1141]" fontSize="11" level="3" pageId="297" pageNumber="298" reason="3">
<emphasis id="B96F064FFE8CFED0FF64FBE4AC68EE0C" bold="true" box="[189,335,1115,1141]" pageId="297" pageNumber="298">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFE8CFED3FF64FB34AEAEEB3E" blockId="297.[187,1401,1163,2025]" lastBlockId="298.[189,1399,265,327]" lastPageId="298" lastPageNumber="299" pageId="297" pageNumber="298">
Caulescent, tuberous herb, to 
<quantity id="4CE377B8FE8CFED0FDF2FB34AF5FEEDC" box="[555,632,1163,1190]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="297" pageNumber="298" unit="cm" value="15.0">15 cm</quantity>
high. 
<emphasis id="B96F064FFE8CFED0FD12FB34AE37EEDC" box="[715,784,1163,1189]" italics="true" pageId="297" pageNumber="298">Tuber</emphasis>
ellipsoid to sub-globose, 1–3 × 
<quantity id="4CE377B8FE8CFED0FB7AFB34A83FEEDC" box="[1187,1304,1163,1190]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="297" pageNumber="298" unit="cm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 cm</quantity>
, with 1 growing point. 
<emphasis id="B96F064FFE8CFED0FEA0FB0FAC99EEB3" box="[377,446,1200,1226]" italics="true" pageId="297" pageNumber="298">Stems</emphasis>
1–5, erect, unbranched or few-branched; internodes to 
<quantity id="4CE377B8FE8CFED0FBBEFB10A99CEEB3" box="[1127,1211,1199,1226]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="297" pageNumber="298" unit="cm" value="7.5">7.5 cm</quantity>
long, to 
<quantity id="4CE377B8FE8CFED0FAF6FB10A850EEB3" box="[1327,1399,1199,1226]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="297" pageNumber="298" unit="mm" value="1.0">1 mm</quantity>
thick, wiry, pale red, glabrous to sparsely tomentose. 
<emphasis id="B96F064FFE8CFED0FCF2FB6CAEAEEE94" box="[811,905,1235,1261]" italics="true" pageId="297" pageNumber="298">Stipules</emphasis>
persistent, ovate to lanceolate, 1–3 × 
<quantity id="4CE377B8FE8CFED0FA9AFB6CAC23EF6B" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="297" pageNumber="298" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5– 1 mm</quantity>
, apex acute, aristate, translucent, pale green, glabrous, margin irregularly-lacerate, ciliate. 
<emphasis id="B96F064FFE8CFED0FAFDFB46A851EF6B" box="[1316,1398,1273,1298]" italics="true" pageId="297" pageNumber="298">Leaves</emphasis>
3–7 per stem, alternate, basifixed; petiole 
<quantity id="4CE377B8FE8CFED0FD6FFAA3AE1BEF4E" box="[694,828,1308,1335]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="4.5" metricValueMin="0.5" pageId="297" pageNumber="298" unit="cm" value="2.5" valueMax="4.5" valueMin="0.5">0.5–4.5 cm</quantity>
long, pale red, glabrous to sparsely tomentose; blade asymmetrical, broadly lanceolate to ovate, to 7 × 
<quantity id="4CE377B8FE8CFED0FC84FAFEAEBBEF22" box="[861,924,1345,1371]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="297" pageNumber="298" unit="cm" value="7.0">7 cm</quantity>
, succulent, apex acute, base truncate to cordate, basal lobes not overlapping, sinus to 
<quantity id="4CE377B8FE8CFED0FD37FADAAE64EF06" box="[750,835,1381,1407]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="297" pageNumber="298" unit="mm" value="18.0">18 mm</quantity>
deep, margin crenate-lobed, the lobes dentate, ciliate, upper surface dark green, sparsely to moderately pilose, lower surface pale green flushed purple, sparsely pilose to densely pilose on the veins, veins palmate-pinnate, 6–9 veined from the base, with 2–4 secondary veins on the larger side, 1–3 secondary veins on the smaller side. 
<emphasis id="B96F064FFE8CFED0FBF3FA6DA9F5EF95" box="[1066,1234,1490,1516]" italics="true" pageId="297" pageNumber="298">Inflorescences</emphasis>
1–6 per stem, bisexual, axillary, erect, an asymmetrical cyme, with 2–3 branches, bearing up to 2 staminate flowers and 1 pistillate flower, protandrous; peduncle to 
<quantity id="4CE377B8FE8CFED0FD22F9A5AE6CEC4C" box="[763,843,1562,1589]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="297" pageNumber="298" unit="cm" value="7.5">7.5 cm</quantity>
long, pale red, glabrous to sparsely pubescent, bracts persistent, oblong to narrowly ovate, 2–4 × 
<quantity id="4CE377B8FE8CFED0FCF5F980AEB3EC20" box="[812,916,1599,1625]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="297" pageNumber="298" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
, translucent, pale green, glabrous, apex acute, margin irregularly-lacerate, ciliate. 
<emphasis id="B96F064FFE8CFED0FD7DF9DCAE50EC04" box="[676,887,1635,1661]" italics="true" pageId="297" pageNumber="298">Staminate flowers</emphasis>
: pedicels to 
<quantity id="4CE377B8FE8CFED0FBD0F9DCA97CEC07" box="[1033,1115,1635,1662]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="297" pageNumber="298" unit="mm" value="20.0">20 mm</quantity>
long, glabrous; tepals 4, spreading, outer 2 ovate, 4–9 × 
<quantity id="4CE377B8FE8CFED0FDFDF937AFA1ECDB" box="[548,646,1672,1698]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="297" pageNumber="298" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
, apex rounded, white, tinged red outside, glabrous, margin entire, aciliate, inner 2 elliptic to oblong, 3–7 × 
<quantity id="4CE377B8FE8CFED0FD4BF913AE2CECBF" box="[658,779,1708,1735]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="297" pageNumber="298" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.5–4 mm</quantity>
, apex obtuse, white, glabrous, margin entire, aciliate; stamens 15–20, projecting, yellow, filaments 
<quantity id="4CE377B8FE8CFED0FD0EF96FAE40EC92" box="[727,871,1744,1771]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="297" pageNumber="298" unit="mm" value="1.0" valueMax="1.5" valueMin="0.5">0.5–1.5 mm</quantity>
long, fused into a 
<quantity id="4CE377B8FE8CFED0FB93F96FA9F6EC92" box="[1098,1233,1744,1771]" metricMagnitude="-3" metricUnit="m" metricValue="2.375" metricValueMax="2.75" metricValueMin="2.0" pageId="297" pageNumber="298" unit="mm" value="2.375" valueMax="2.75" valueMin="2.0">2–2.75 mm</quantity>
long column, anthers cuboid, ca 0.75 × 
<quantity id="4CE377B8FE8CFED0FE2BF94BAF77ED76" box="[498,592,1780,1807]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="297" pageNumber="298" unit="mm" value="0.5">0.5 mm</quantity>
, dehiscing via lateral slits, connectives not extended, symmetrically basifixed. 
<emphasis id="B96F064FFE8CFED0FEECF8A6ACDBED4A" box="[309,508,1817,1843]" italics="true" pageId="297" pageNumber="298">Pistillate flowers</emphasis>
: pedicels to 
<quantity id="4CE377B8FE8CFED0FD49F8A6AFC5ED4A" box="[656,738,1817,1843]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="297" pageNumber="298" unit="mm" value="17.0">17 mm</quantity>
long; bracteoles absent; tepals 5, subequal, deciduous in fruit, spreading, narrowly elliptic to obovate, 4–5.5 × 
<quantity id="4CE377B8FE8CFED0FCEDF882AE88ED21" box="[820,943,1853,1880]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="297" pageNumber="298" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 mm</quantity>
, apex obtuse, white tinged red outside, glabrous, margin entire, aciliate; ovary body ellipsoid to obovoid, 2–4 × 
<quantity id="4CE377B8FE8CFED0FBC0F8DEA959ED05" box="[1049,1150,1889,1916]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="297" pageNumber="298" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
, green tinged purple, glabrous, unequally 3-winged, wings triangular, largest 2–5 × 
<quantity id="4CE377B8FE8CFED0FC73F83AA900EDD9" box="[938,1063,1925,1952]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="297" pageNumber="298" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
, smallest 3–5 × ca 
<quantity id="4CE377B8FE8CFED0FACBF83AA855EDD9" box="[1298,1394,1925,1952]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="297" pageNumber="298" unit="mm" value="0.5">0.5 mm</quantity>
; 3-locular, placentae branches divided, bearing ovules on both surfaces; styles 3, yellow, fused at the base, 
<quantity id="4CE377B8FE8CFED0FED8F871ACA8ED90" box="[257,399,1998,2025]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="297" pageNumber="298" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.5–3.5 mm</quantity>
long, once-divided, stigmatic papillae in a spirally-twisted band. 
<emphasis id="B96F064FFE8CFED0FB4EF870A87EED91" box="[1175,1369,1998,2024]" italics="true" pageId="297" pageNumber="298">Fruiting pedicel</emphasis>
to 
<quantity id="4CE377B8FE8FFED3FF64FEB6AC3FEB5D" box="[189,280,265,292]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="298" pageNumber="299" unit="mm" value="1.7">1.7 mm</quantity>
long. 
<emphasis id="B96F064FFE8FFED3FEBDFEB5ACC4EB5A" box="[356,483,265,291]" italics="true" pageId="298" pageNumber="299">Fruit body</emphasis>
ovoid, to 7 × 
<quantity id="4CE377B8FE8FFED3FD55FEB6AFF4EB5D" box="[652,723,265,292]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="298" pageNumber="299" unit="mm" value="4.0">4 mm</quantity>
, drying brown, wings same shape as in ovary, the largest expanding to 9 × 
<quantity id="4CE377B8FE8FFED3FE52FE92ACF5EB3E" box="[395,466,301,327]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="298" pageNumber="299" unit="mm" value="7.0">7 mm</quantity>
, the smallest expanding to 8 × 
<quantity id="4CE377B8FE8FFED3FCE7FE93AEA2EB3E" box="[830,901,300,327]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="298" pageNumber="299" unit="mm" value="3.0">3 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE8FFED3FF64FED1AE08E859" pageId="298" pageNumber="299" type="conservation">
<paragraph id="8BA4DA5DFE8FFED3FF64FED1AF4AEBF1" blockId="298.[189,621,365,392]" box="[189,621,365,392]" pageId="298" pageNumber="299">
<heading id="D0EC6D31FE8FFED3FF64FED1AF4AEBF1" bold="true" box="[189,621,365,392]" fontSize="11" level="3" pageId="298" pageNumber="299" reason="3">
<emphasis id="B96F064FFE8FFED3FF64FED1AF4AEBF1" bold="true" box="[189,621,365,392]" pageId="298" pageNumber="299">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFE8FFED3FF64FE23AE08E859" blockId="298.[189,1399,412,544]" pageId="298" pageNumber="299">
Assessed by 
<bibRefCitation id="EF8AA7ACFE8FFED3FE89FE23AF52EBCF" author="Leon &amp; Monsalve" box="[336,629,412,438]" firstAuthor="Leon" pageId="298" pageNumber="299" pagination="165 - 170" refId="ref229226" refString="Leon B. &amp; Monsalve C. 2006. Begoniaceae endemicas del Peru. Revista Peruana de Biologia 13 (2): 165 - 170. https: // doi. org / 10.15381 / rpb. v 13 i 2.1810" type="journal article" year="2006">
<collectingRegion id="49DF14BFFE8FFED3FE89FE23ACABEBCF" box="[336,396,412,438]" country="Nicaragua" name="Leon" pageId="298" pageNumber="299">León</collectingRegion>
&amp; Monsalve (2006)
</bibRefCitation>
and 
<bibRefCitation id="EF8AA7ACFE8FFED3FD73FE23AECAEBCF" author="Moonlight &amp; Tebbitt" box="[682,1005,412,438]" firstAuthor="Moonlight" pageId="298" pageNumber="299" pagination="111 - 122" refId="ref229847" refString="Moonlight P. W. &amp; Tebbitt M. C. 2016. Two new Peruvian species of Begonia (Begoniaceae) and an amended description of Begonia thyrsoidea. Edinburgh Journal of Botany 74 (2): 111 - 122. https: // doi. org / 10.1017 / S 0960428616000299" type="journal article" year="2016">Moonlight &amp; Tebbitt (2016)</bibRefCitation>
as Data Deficient (DD). The latter authors noted that they were unable to locate the species at its 
<typeStatus id="54A064FFFE8FFED3FC73FE7FAEFAEBA3" box="[938,989,448,474]" pageId="298" pageNumber="299">type</typeStatus>
locality during fieldwork in either 2014 or 2015, and that the area has undergone widespread deforestation. We consider it appropriate to reassess 
<taxonomicName id="4C1BA1DEFE8FFED3FEFAFDB8ACE6E859" box="[291,449,518,544]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="298" pageNumber="299" phylum="Tracheophyta" rank="species" species="thyrsoidea">
<emphasis id="B96F064FFE8FFED3FEFAFDB8ACE6E859" box="[291,449,518,544]" italics="true" pageId="298" pageNumber="299">B. thyrsoidea</emphasis>
</taxonomicName>
as Endangered (EN B1ab(iii)).
</paragraph>
</subSubSection>
<caption id="DF648AD5FE8FFED3FF64F814A9D7ED91" ID-DOI="http://doi.org/10.5281/zenodo.8178505" ID-Zenodo-Dep="8178505" httpUri="https://zenodo.org/record/8178505/files/figure.png" pageId="298" pageNumber="299" startId="298.[189,232,1963,1989]" targetBox="[236,1329,655,1889]" targetPageId="298">
<paragraph id="8BA4DA5DFE8FFED3FF64F814A9D7ED91" blockId="298.[189,1398,1963,2025]" pageId="298" pageNumber="299">
<emphasis id="B96F064FFE8FFED3FF64F814AC3CEDBC" bold="true" box="[189,283,1963,1989]" pageId="298" pageNumber="299">Fig. 94.</emphasis>
<taxonomicName id="4C1BA1DEFE8FFED3FEFAF813AE59EDBC" authority="Irmsch. A. Habit. B. Staminate" box="[291,894,1963,1990]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="298" pageNumber="299" phylum="Tracheophyta" rank="species" species="thyrsoidea">
<emphasis id="B96F064FFE8FFED3FEFAF813AF2DEDBC" box="[291,522,1963,1989]" italics="true" pageId="298" pageNumber="299">Begonia thyrsoidea</emphasis>
Irmsch. 
<emphasis id="B96F064FFE8FFED3FDABF814AFA0EDBC" bold="true" box="[626,647,1963,1989]" pageId="298" pageNumber="299">A</emphasis>
. Habit. 
<emphasis id="B96F064FFE8FFED3FD3EF814AFDDEDBC" bold="true" box="[743,762,1963,1989]" pageId="298" pageNumber="299">B</emphasis>
. Staminate
</taxonomicName>
flower, side view. 
<emphasis id="B96F064FFE8FFED3FB85F814A956EDBC" bold="true" box="[1116,1137,1963,1989]" pageId="298" pageNumber="299">C</emphasis>
. Pistillate flower, side view. Illustration by Peter Moonlight from photographs of 
<emphasis id="B96F064FFE8FFED3FCB6F870A99CED91" box="[879,1211,1998,2025]" italics="true" pageId="298" pageNumber="299">M.C. Tebbitt &amp; A. Daza 809</emphasis>
(E).
</paragraph>
</caption>
<subSubSection id="C30189D6FE8EFED2FF64FEB7AE2FEBC4" pageId="299" pageNumber="300" type="discussion">
<paragraph id="8BA4DA5DFE8EFED2FF64FEB7AC97EB5A" blockId="299.[189,432,264,291]" box="[189,432,264,291]" pageId="299" pageNumber="300">
<heading id="D0EC6D31FE8EFED2FF64FEB7AC97EB5A" bold="true" box="[189,432,264,291]" fontSize="11" level="3" pageId="299" pageNumber="300" reason="3">
<emphasis id="B96F064FFE8EFED2FF64FEB7AC97EB5A" bold="true" box="[189,432,264,291]" pageId="299" pageNumber="300">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFE8EFED2FF64FE87AE2FEBC4" blockId="299.[189,1398,312,446]" pageId="299" pageNumber="300">
Unique among small tuberous begonias in 
<collectingCountry id="F30C9ACDFE8EFED2FD19FE87AFD1EB2B" box="[704,758,312,338]" name="Peru" pageId="299" pageNumber="300">Peru</collectingCountry>
in its staminate flowers with its anthers united into a&gt; 
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long column. It is also unusual in its asymmetrical inflorescences, which are only found amongst tuberous Peruvian begonias in members of the octopetala group of 
<emphasis id="B96F064FFE8EFED2FC08FE3FAECDEBE0" box="[977,1002,384,409]" italics="true" pageId="299" pageNumber="300">B.</emphasis>
sect. 
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<emphasis id="B96F064FFE8EFED2FBF7FE3FA989EBE3" box="[1070,1198,384,410]" italics="true" pageId="299" pageNumber="300">Eupetalum</emphasis>
</taxonomicName>
with many more tepals on both the staminate and pistillate flowers.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE8EFED2FF64FE5AA91EE80E" pageId="299" pageNumber="300" type="distribution">
<paragraph id="8BA4DA5DFE8EFED2FF64FE5AACD2E879" blockId="299.[189,501,485,512]" box="[189,501,485,512]" pageId="299" pageNumber="300">
<heading id="D0EC6D31FE8EFED2FF64FE5AACD2E879" bold="true" box="[189,501,485,512]" fontSize="11" level="3" pageId="299" pageNumber="300" reason="3">
<emphasis id="B96F064FFE8EFED2FF64FE5AACD2E879" bold="true" box="[189,501,485,512]" pageId="299" pageNumber="300">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFE8EFED2FF64FDAAA91EE80E" blockId="299.[188,1399,533,631]" pageId="299" pageNumber="300">
Endemic to 
<collectingCountry id="F30C9ACDFE8EFED2FE93FDAAACA7E856" box="[330,384,533,559]" name="Peru" pageId="299" pageNumber="300">Peru</collectingCountry>
and known from 
<collectingRegion id="49DF14BFFE8EFED2FD97FDAAAFBEE856" box="[590,665,533,559]" country="Peru" name="Cusco" pageId="299" pageNumber="300">Cuzco</collectingRegion>
and 
<collectingRegion id="49DF14BFFE8EFED2FD09FDAAAE2BE856" box="[720,780,533,559]" country="Peru" name="Puno" pageId="299" pageNumber="300">Puno</collectingRegion>
Regions (
<figureCitation id="1320C6D8FE8EFED2FC5AFDAAAED7E856" box="[899,1008,533,559]" captionStart="Fig" captionStartId="288.[189,232,1857,1883]" captionTargetBox="[194,1390,1193,1812]" captionTargetId="figure-313@288.[189,1398,1183,1818]" captionTargetPageId="288" captionText="Fig. 90. Distribution of the Begonia sect. Ignota in Peru and surrounding countries. A. B. bifurcata L.B.Sm. &amp; B.G.Schub. (red), B. erythrothrix Tebbitt &amp; Moonlight (red), and B. occultata J.P.Allen &amp; Moonlight sp. nov. (green). B. B. speculum Moonlight &amp; Tebbitt (red), B. thyrsoidea Irmsch., and B. urubambensis Tebbitt (green). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178495" httpUri="https://zenodo.org/record/8178495/files/figure.png" pageId="299" pageNumber="300">Fig. 90A</figureCitation>
). Found in lower montane forests at 
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a.s.l. on moist, shaded cliffs. As a tuberous herb, 
<taxonomicName id="4C1BA1DEFE8EFED2FC61FD85A972E82A" box="[952,1109,569,595]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="299" pageNumber="300" phylum="Tracheophyta" rank="species" species="thyrsoidea">
<emphasis id="B96F064FFE8EFED2FC61FD85A972E82A" box="[952,1109,569,595]" italics="true" pageId="299" pageNumber="300">B. thyrsoidea</emphasis>
</taxonomicName>
presumably dies back to its tuber in the dry season and flowers in the wet season (January to March).
</paragraph>
</subSubSection>
</treatment>
</document>