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<document id="367322FB3E5F8538B78A07EF2218A0AB" ID-DOI="https://doi.org/10.5852/ejt.2023.881.2175" ID-ISSN="2118-9773" ID-Zenodo-Dep="8178280" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690201937279" checkinUser="felipe" docAuthor="Moonlight, Peter. W., Jara-Muñoz, Orlando A., Purvis, David A., Delves, Jay, Allen, Josh P. &amp; Reynel, Carlos" docDate="2023" docId="03B26B4BFE8AFED6FDE7FB92A928ED91" docLanguage="en" docName="ejt-881_1-334.imf" docOrigin="European Journal of Taxonomy 881" docSource="http://dx.doi.org/10.5852/ejt.2023.881.2175" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia diversifolia var. nana Walp." docType="treatment" docVersion="7" lastPageNumber="304" masterDocId="FF8B1333FFA5FFF9FFD9FFBFAD27EA79" masterDocTitle="The genus Begonia (Begoniaceae) in Peru" masterLastPageNumber="334" masterPageNumber="1" pageNumber="304" updateTime="1707123162844" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="D2EB902B4F2E6CBD9D2467EBE095F6B0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BE302A781B50B3E1E6A48D17E7AC5E80">
<mods:title id="9AE4F8CF9C716F8A317B1A471848E70F">The genus Begonia (Begoniaceae) in Peru</mods:title>
</mods:titleInfo>
<mods:name id="30E232AE35B3EB5880C43B2666909192" type="personal">
<mods:role id="6BA17A19D73FF473C3E5351FE5B9C06B">
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<mods:namePart id="003423066DDB54251CB1BA9DAEF788B3">Moonlight, Peter. W.</mods:namePart>
<mods:affiliation id="EFFCB0B669295CD999A7ED04A9DAB4D8">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK. &amp; Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland.</mods:affiliation>
<mods:nameIdentifier id="BD6ADACFD62060FB1D5645045F5E3A2C" type="email">moonligp@tcd.ie</mods:nameIdentifier>
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<mods:role id="3465CD4713BE2B1BCA06A10503E699A1">
<mods:roleTerm id="46ACDAE206766959EA567C969AF8A6A3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BC79D722714847A31AC3F351649E2CDA">Jara-Muñoz, Orlando A.</mods:namePart>
<mods:affiliation id="27321C70C7EB767963F9267D029618F1">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá D. C., Colombia.</mods:affiliation>
<mods:nameIdentifier id="7D48D72FED3787F67CFA9E5A2C490E76" type="email">oajaram@unal.edu.co</mods:nameIdentifier>
</mods:name>
<mods:name id="4A97816D6D62881C0F5F3697C868D02A" type="personal">
<mods:role id="822C098C1BC24E112E9E5D476F1A9C0E">
<mods:roleTerm id="EDD1BECAC5B6B45131D843E43E844EDC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AFDCD1CE7EA6EAFADBCC4924A19E3BB2">Purvis, David A.</mods:namePart>
<mods:affiliation id="8C2F12AEE32A84C1C0F331BCE7A9A617">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="B55A991393CAB44127EEBD3A50BDB5B3" type="email">dpurvis@rbge.org.uk</mods:nameIdentifier>
</mods:name>
<mods:name id="AB4F2E6111E96C9834BEC3AA82348C46" type="personal">
<mods:role id="5124A5161BCDBD990B694FA52BEEDFCD">
<mods:roleTerm id="BFC7C3BD9EDDD103BB1817F905C01BDC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="83A60D2F5974637D2EC4531A327E9698">Delves, Jay</mods:namePart>
<mods:affiliation id="E3A57B004FF4714914FC1DF3846AD6D6">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="87F732464AF913F3AC6096E0150D4255" type="email">j.delves@westernsydney.edu.au</mods:nameIdentifier>
</mods:name>
<mods:name id="57A25622AEC33B6EA6D85CD420B6CC0F" type="personal">
<mods:role id="ABD05EBD1E49025AE1AA59FE1C216410">
<mods:roleTerm id="0827C694868FDA7084E3A0738915FB67">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5FD3734E611BB34F38D4A6A224E2E80A">Allen, Josh P.</mods:namePart>
<mods:affiliation id="8F09FA7A36BC8B19847860A3AA1A55C3">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="190AC5DFADEF8DAC14158AAF84FB3A8A" type="email">myrmeciaman@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm id="6656D02387A4806216A200E172876EE3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8F46C4EDCFC5F687ADDCAB306DB2712B">Reynel, Carlos</mods:namePart>
<mods:affiliation id="642B60F2A30832F070156880FE424DFD">Facultad de Ciencias Forestales, Universidad Nacional Agraria-La Molina, Lima 12, Peru.</mods:affiliation>
<mods:nameIdentifier id="5FD49BB4D72861B81744102FB70F8D73" type="email">reynel@lamolina.edu.pe</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="100F81E88FB4BAAACE9F15727E222B8C">text</mods:typeOfResource>
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<mods:title id="DD9BE3378576715738E7A9ED1BCE4378">European Journal of Taxonomy</mods:title>
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<mods:part id="4EF27FA723E40307B9865AEF421EFA66">
<mods:date id="6349F3ABEAF7C130404FCE11C97769A0">2023</mods:date>
<mods:detail id="C5B278CACD41893C202B91CE536CD1FB" type="pubDate">
<mods:number id="645548B17EC6BB1048387F2DE38F247E">2023-07-18</mods:number>
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<mods:detail id="450854FFA24D7C08EE6B49EECB3CD5FE" type="volume">
<mods:number id="81A05346EA6F9E1F177F6078523D05B7">881</mods:number>
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<mods:url id="D53FBE0B8029E7EC7369A5F33528C6DC">http://dx.doi.org/10.5852/ejt.2023.881.2175</mods:url>
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<mods:classification id="A7731277B3D1AA722AE19DBB84E8A313">journal article</mods:classification>
<mods:identifier id="ACD712ACC122AACEE0B639849BC0F43A" type="DOI">https://doi.org/10.5852/ejt.2023.881.2175</mods:identifier>
<mods:identifier id="373B2C5470EB971D0D065B5F29DE1C3D" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="BF84EBDE7BDCA60CFB5C2927651501F9" type="Zenodo-Dep">8178280</mods:identifier>
</mods:mods>
<treatment id="03B26B4BFE8AFED6FDE7FB92A928ED91" LSID="urn:lsid:plazi:treatment:03B26B4BFE8AFED6FDE7FB92A928ED91" httpUri="http://treatment.plazi.org/id/03B26B4BFE8AFED6FDE7FB92A928ED91" lastPageNumber="304" pageId="303" pageNumber="304">
<subSubSection id="C30189D6FE8AFED6FDE7FB92AED1EE3E" box="[574,1014,1069,1096]" pageId="303" pageNumber="304" type="nomenclature">
<paragraph id="8BA4DA5DFE8AFED6FDE7FB92AED1EE3E" blockId="303.[574,1014,1069,1096]" box="[574,1014,1069,1096]" pageId="303" pageNumber="304">
<heading id="D0EC6D31FE8AFED6FDE7FB92AED1EE3E" box="[574,1014,1069,1096]" centered="true" fontSize="11" level="2" pageId="303" pageNumber="304" reason="2">
<taxonomicName id="4C1BA1DEFE8AFED6FDE7FB92AED1EE3E" ID-CoL="87YY7" ID-ENA="234383" authority="Walp." box="[574,1014,1069,1096]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="303" pageNumber="304" phylum="Tracheophyta" rank="variety" species="diversifolia" variety="nana">
<emphasis id="B96F064FFE8AFED6FDE7FB92AE15EE31" bold="true" box="[574,818,1069,1096]" italics="true" pageId="303" pageNumber="304">Begonia diversifolia</emphasis>
var.
<emphasis id="B96F064FFE8AFED6FCB5FB91AE8DEE31" bold="true" box="[876,938,1070,1096]" italics="true" pageId="303" pageNumber="304">nana</emphasis>
Walp.
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE8AFED6FF64FBCCAC80EEA3" pageId="303" pageNumber="304" type="reference_group">
<paragraph id="8BA4DA5DFE8AFED6FF64FBCCAC80EEA3" blockId="303.[189,1399,1139,1281]" pageId="303" pageNumber="304">
<emphasis id="B96F064FFE8AFED6FF64FBCCA9EEEEF4" box="[189,1225,1139,1165]" italics="true" pageId="303" pageNumber="304">Novorum Actorum Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum</emphasis>
19 (suppl. 1): 408 (
<bibRefCitation id="EF8AA7ACFE8AFED6FEFCFB26ACFBEECA" author="Walpers" box="[293,476,1177,1203]" firstAuthor="Walpers" pageId="303" pageNumber="304" pagination="408 - 409" refId="ref233350" refString="Walpers W. G. 1843 b. Begoniaceae R. Br. Novorum Actorum Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum 19 (Suppl. 1): 408 - 409." type="journal article" year="1843" yearSuffix="b">Walpers 1843b</bibRefCitation>
). –
<treatmentCitationGroup id="AB0BFD73FE8AFED6FDDCFB25AC80EEA3" pageId="303" pageNumber="304">
<taxonomicName id="4C1BA1DEFE8AFED6FDDCFB25AECBEECD" ID-CoL="7YMJJ" authority="(Walp.) A. DC." authorityName="A. DC." baseAuthorityName="Walp." box="[517,1004,1177,1204]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="303" pageNumber="304" phylum="Tracheophyta" rank="variety" species="hirtella" variety="nana">
<emphasis id="B96F064FFE8AFED6FDDCFB25AFE3EECA" box="[517,708,1177,1203]" italics="true" pageId="303" pageNumber="304">Begonia hirtella</emphasis>
var.
<emphasis id="B96F064FFE8AFED6FD27FB25AE1EEECA" box="[766,825,1178,1203]" italics="true" pageId="303" pageNumber="304">nana</emphasis>
(Walp.) A.DC.
</taxonomicName>
,
<treatmentCitation id="0ABAFC4CFE8AFED6FC2FFB26AC84EEA3" author="de Candolle" page="345" pageId="303" pageNumber="304" year="1861">
<emphasis id="B96F064FFE8AFED6FC2FFB26A9EAEECA" box="[1014,1229,1177,1203]" italics="true" pageId="303" pageNumber="304">Flora Brasiliensis</emphasis>
4 (1): 345 (
<bibRefCitation id="EF8AA7ACFE8AFED6FA80FB26ACBDEEA3" author="de Candolle" firstAuthor="de Candolle" pageId="303" pageNumber="304" pagination="337 - 396" refId="ref226005" refString="de Candolle A. P. 1861. Begoniaceae. In: Martius C. P. F. (ed.) Flora Brasiliensis 4 (1): 337 - 396. Frid. Fleischer, Munich &amp; Leipzig." type="journal article" year="1861">de Candolle 1861</bibRefCitation>
)
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE8AFED6FE77FB7EACC4EF78" pageId="303" pageNumber="304" type="materials_examined">
<paragraph id="8BA4DA5DFE8AFED6FE77FB7EA867EEA3" blockId="303.[189,1399,1139,1281]" box="[430,1344,1216,1243]" pageId="303" pageNumber="304">
–
<emphasis id="B96F064FFE8AFED6FE1CFB7FAF24EEA3" bold="true" box="[453,515,1216,1242]" pageId="303" pageNumber="304">
<typeStatus id="54A064FFFE8AFED6FE1CFB7FAF24EEA3" box="[453,515,1216,1242]" pageId="303" pageNumber="304">Type</typeStatus>
</emphasis>
:
<materialsCitation id="3B73D000FE8AFED6FDCBFB7EA867EEA3" box="[530,1344,1216,1243]" collectorName="F. J. F. Meyen" country="Peru" location="ad Altos de Toledo" pageId="303" pageNumber="304" specimenCount="1">
<collectingCountry id="F30C9ACDFE8AFED6FDCBFB7EAF7AEEA2" box="[530,605,1217,1243]" name="Peru" pageId="303" pageNumber="304">PERU</collectingCountry>
•
<location id="8EC48C86FE8AFED6FDAFFB7FAE72EEA3" LSID="urn:lsid:plazi:treatment:03B26B4BFE8AFED6FDE7FB92A928ED91:8EC48C86FE8AFED6FDAFFB7FAE72EEA3" box="[630,853,1216,1242]" country="Peru" name="ad Altos de Toledo" pageId="303" pageNumber="304">ad Altos de Toledo</location>
,
<emphasis id="B96F064FFE8AFED6FCB8FB7EAED0EEA3" box="[865,1015,1217,1242]" italics="true" pageId="303" pageNumber="304">
<collectorName id="26EEBF8BFE8AFED6FCB8FB7EAED0EEA3" box="[865,1015,1217,1242]" pageId="303" pageNumber="304">F.J.F. Meyen</collectorName>
</emphasis>
[n.v., herbarium unknown].
</materialsCitation>
</paragraph>
<paragraph id="8BA4DA5DFE8AFED6FF64FB58ACC4EF78" blockId="303.[189,1399,1139,1281]" box="[189,483,1255,1281]" pageId="303" pageNumber="304">
<bibRefCitation id="EF8AA7ACFE8AFED6FF64FB58ACF8EF78" author="de Candolle" box="[189,479,1255,1281]" firstAuthor="de Candolle" page="299" pageId="303" pageNumber="304" pagination="266 - 408" refId="ref226047" refString="de Candolle A. P. 1864. Begoniaceae. In: de Candolle A. P. (ed.) Prodromus Systematis Naturalis Regni Vegetabilis 15 (1): 266 - 408. V. Masson &amp; fils, Paris." type="journal article" year="1864">de Candolle (1864: 299)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE8AFED6FF64FA91A928ED91" pageId="303" pageNumber="304" type="discussion">
<paragraph id="8BA4DA5DFE8AFED6FF64FA91AC23EF31" blockId="303.[189,260,1326,1352]" box="[189,260,1326,1352]" pageId="303" pageNumber="304">
<heading id="D0EC6D31FE8AFED6FF64FA91AC23EF31" bold="true" box="[189,260,1326,1352]" fontSize="11" level="3" pageId="303" pageNumber="304" reason="3">
<emphasis id="B96F064FFE8AFED6FF64FA91AC23EF31" bold="true" box="[189,260,1326,1352]" pageId="303" pageNumber="304">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFE8AFED6FF64FADFA928ED91" blockId="303.[187,1400,1376,2025]" pageId="303" pageNumber="304">
Walpers described
<taxonomicName id="4C1BA1DEFE8AFED6FE4DFADEAFD0EF03" authority="Walp." authorityName="Walp." box="[404,759,1376,1403]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="303" pageNumber="304" phylum="Tracheophyta" rank="variety" species="diversifolia" variety="nana">
<emphasis id="B96F064FFE8AFED6FE4DFADEAF1DEF03" box="[404,570,1376,1402]" italics="true" pageId="303" pageNumber="304">B. diversifolia</emphasis>
var.
<emphasis id="B96F064FFE8AFED6FDABFADEAF8AEF03" box="[626,685,1377,1402]" italics="true" pageId="303" pageNumber="304">nana</emphasis>
Walp.
</taxonomicName>
based upon a collection made by
<emphasis id="B96F064FFE8AFED6FBABFADEA821EF03" box="[1138,1286,1377,1402]" italics="true" pageId="303" pageNumber="304">F.J.F. Meyen</emphasis>
(
<bibRefCitation id="EF8AA7ACFE8AFED6FACAFADFAC6EEFD8" author="Walpers" firstAuthor="Walpers" page="409" pageId="303" pageNumber="304" pagination="408 - 409" refId="ref233350" refString="Walpers W. G. 1843 b. Begoniaceae R. Br. Novorum Actorum Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum 19 (Suppl. 1): 408 - 409." type="journal article" year="1843" yearSuffix="b">Walpers 1843b: 409</bibRefCitation>
). Meyen’s collection was made in Altos de Toledo, a pass between the cities of Puno and Arequipa (
<bibRefCitation id="EF8AA7ACFE8AFED6FEE0FA11AF2AEFB1" author="Weberbauer" box="[313,525,1454,1480]" firstAuthor="Weberbauer" pageId="303" pageNumber="304" pagination="293 - 305" refId="ref233535" refString="Weberbauer A. 1936. Historia de la exploracion botanica del Peru. Revista de la Universidad Catolica del Peru 6: 293 - 305." type="journal article" year="1936">Weberbauer 1936</bibRefCitation>
). Almost without exception, the full length of all passes between these cities are above
<quantity id="4CE377B8FE8AFED6FEE1FA6AACB4EF96" box="[312,403,1493,1519]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="303" pageNumber="304" unit="m" value="4000.0">4000 m</quantity>
in elevation and are dry, high elevation grasslands unsuitable for
<taxonomicName id="4C1BA1DEFE8AFED6FB69FA69A850EF96" box="[1200,1399,1493,1519]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="303" pageNumber="304" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B96F064FFE8AFED6FB69FA69A834EF96" box="[1200,1299,1494,1519]" italics="true" pageId="303" pageNumber="304">Begonia</emphasis>
species.
</taxonomicName>
This is a variety of
<taxonomicName id="4C1BA1DEFE8AFED6FE74FA42AFE4EC6F" authority="Graham" authorityName="Graham" box="[429,707,1532,1558]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="303" pageNumber="304" phylum="Tracheophyta" rank="species" species="diversifolia">
<emphasis id="B96F064FFE8AFED6FE74FA42AF70EC6F" box="[429,599,1532,1558]" italics="true" pageId="303" pageNumber="304">B. diversifolia</emphasis>
Graham
</taxonomicName>
, which is a member of
<emphasis id="B96F064FFE8AFED6FC39FA42AEDDEC6F" box="[992,1018,1533,1558]" italics="true" pageId="303" pageNumber="304">B.</emphasis>
sect.
<taxonomicName id="4C1BA1DEFE8AFED6FB99FA43A850EC6F" authority="Ziesenh." authorityName="Ziesenh." box="[1088,1399,1532,1558]" class="Magnoliopsida" family="Begoniaceae" genus="Quadriperigonia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cucurbitales" pageId="303" pageNumber="304" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F064FFE8AFED6FB99FA43A82EEC6F" box="[1088,1289,1532,1558]" italics="true" pageId="303" pageNumber="304">Quadriperigonia</emphasis>
Ziesenh.
</taxonomicName>
and only distantly related to any Peruvian
<taxonomicName id="4C1BA1DEFE8AFED6FD1BF99CAEA3EC44" box="[706,900,1571,1597]" pageId="303" pageNumber="304">
<emphasis id="B96F064FFE8AFED6FD1BF99CAE02EC45" box="[706,805,1571,1596]" italics="true" pageId="303" pageNumber="304">Begonia</emphasis>
species
</taxonomicName>
(
<bibRefCitation id="EF8AA7ACFE8AFED6FC4FF99CA980EC44" author="Moonlight P. W. &amp; Reynel C." box="[918,1191,1571,1597]" journalOrPublisher="Phytotaxa" pageId="303" pageNumber="304" pagination="116 - 126" part="381" refId="ref229806" refString="Moonlight P. W. &amp; Reynel C. 2018. Two new species of Begonia from Peru. Phytotaxa 381 (1): 116 - 126. https: // doi. org / 10.11646 / phytotaxa. 381.1.15" title="Two new species of Begonia from Peru" type="journal article" year="2018">
Moonlight
<emphasis id="B96F064FFE8AFED6FBC7F99CA97BEC45" box="[1054,1116,1571,1597]" italics="true" pageId="303" pageNumber="304">et al.</emphasis>
2018
</bibRefCitation>
). The description provided by Walpers is insufficient to identify this specimen to any known Peruvian species. Alphonse Pyramus de Candolle moved this taxon to a variety of
<taxonomicName id="4C1BA1DEFE8AFED6FCEAF9CEAE8CECF3" authorityName=", F. Sellow" authorityYear="2192" box="[819,939,1648,1674]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="303" pageNumber="304" phylum="Tracheophyta" rank="species" species="hirtella">
<emphasis id="B96F064FFE8AFED6FCEAF9CEAE8CECF3" box="[819,939,1648,1674]" italics="true" pageId="303" pageNumber="304">B. hirtella</emphasis>
</taxonomicName>
in his Flora of
<collectingCountry id="F30C9ACDFE8AFED6FB85F9CFA983ECF3" box="[1116,1188,1648,1674]" name="Brazil" pageId="303" pageNumber="304">Brazil</collectingCountry>
account and cited several Brazilian specimens of
<taxonomicName id="4C1BA1DEFE8AFED6FDEEF927AF96ECC8" authorityName=", F. Sellow" authorityYear="2192" box="[567,689,1687,1713]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="303" pageNumber="304" phylum="Tracheophyta" rank="species" species="hirtella">
<emphasis id="B96F064FFE8AFED6FDEEF927AF96ECC8" box="[567,689,1687,1713]" italics="true" pageId="303" pageNumber="304">B. hirtella</emphasis>
</taxonomicName>
as the same variety (
<bibRefCitation id="EF8AA7ACFE8AFED6FC6EF928A9A9ECC8" author="de Candolle" box="[951,1166,1687,1713]" firstAuthor="de Candolle" pageId="303" pageNumber="304" pagination="337 - 396" refId="ref226005" refString="de Candolle A. P. 1861. Begoniaceae. In: Martius C. P. F. (ed.) Flora Brasiliensis 4 (1): 337 - 396. Frid. Fleischer, Munich &amp; Leipzig." type="journal article" year="1861">de Candolle 1861</bibRefCitation>
) and later repeated this citation (
<bibRefCitation id="EF8AA7ACFE8AFED6FE80F901AF17ECA1" author="de Candolle" box="[345,560,1726,1752]" firstAuthor="de Candolle" pageId="303" pageNumber="304" pagination="266 - 408" refId="ref226047" refString="de Candolle A. P. 1864. Begoniaceae. In: de Candolle A. P. (ed.) Prodromus Systematis Naturalis Regni Vegetabilis 15 (1): 266 - 408. V. Masson &amp; fils, Paris." type="journal article" year="1864">de Candolle 1864</bibRefCitation>
). It is unclear whether de Candolle saw the type of
<taxonomicName id="4C1BA1DEFE8AFED6FB43F900ADDFEC86" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="303" pageNumber="304" phylum="Tracheophyta" rank="variety" species="diversifolia" variety="nana">
<emphasis id="B96F064FFE8AFED6FB43F900A865ECA1" box="[1178,1346,1726,1752]" italics="true" pageId="303" pageNumber="304">B. diversifolia</emphasis>
var.
<emphasis id="B96F064FFE8AFED6FF64F959ADDFEC86" box="[189,248,1766,1791]" italics="true" pageId="303" pageNumber="304">nana</emphasis>
</taxonomicName>
. He also cited a specimen collected by Meyen in
<collectingRegion id="49DF14BFFE8AFED6FC92F95AAEDCEC86" box="[843,1019,1765,1791]" country="Brazil" name="Rio de Janeiro" pageId="303" pageNumber="304">Rio de Janeiro</collectingRegion>
, which is a
<typeStatus id="54A064FFFE8AFED6FB50F959A9C3ED79" box="[1161,1252,1766,1792]" pageId="303" pageNumber="304">syntype</typeStatus>
of Walper’s
<taxonomicName id="4C1BA1DEFE8AFED6FF67F8B2AC94ED5F" authority="Walp." authorityName="Walp." box="[190,435,1804,1830]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="303" pageNumber="304" phylum="Tracheophyta" rank="species" species="meyeniana">
<emphasis id="B96F064FFE8AFED6FF67F8B2AC44ED5F" box="[190,355,1805,1830]" italics="true" pageId="303" pageNumber="304">B. meyeniana</emphasis>
Walp.
</taxonomicName>
(a taxon of uncertain application;
<bibRefCitation id="EF8AA7ACFE8AFED6FCB9F8B3A9BCED5E" author="Jacques &amp; Mamede" box="[864,1179,1804,1831]" firstAuthor="Jacques" pageId="303" pageNumber="304" pagination="52 - 65" refId="ref228655" refString="Jacques E. L. &amp; Mamede M. C. H. 2005. Notas nomenclaturais em Begonia L. (Begoniaceae). Revista Brasileira de Botanica 28 (3): 579 - 588. https: // doi. org / 10.1590 / S 0100 - 84042005000300014 Jara-Munoz A., Richardson J. E. &amp; Madrinan S. 2019. Character evolution and recircumscription of the northern Andean Begonia section Casparya (Begoniaceae). Systematic Botany 44 (1): 52 - 65. https: // doi. org / 10.1600 / 036364419 X 697895" type="journal article" year="2005">Jacques &amp; Mamede 2005</bibRefCitation>
). It is possible de Candolle confused these specimens or also that Walpers confused the type locality of the
<specimenCount id="9D1D11D4FE8AFED6FB1AF88CA854ED34" box="[1219,1395,1843,1869]" pageId="303" pageNumber="304" type="generic">two specimens</specimenCount>
. Walper’s herbarium was sold upon his death and the location of his specimens are unknown. We have been unable to locate any specimens matching his description. Given the considerable uncertainty surrounding the type locality, protologue description, and classification of this species, we suggest that if the type specimen cannot be located that this name should be rejected.
</paragraph>
</subSubSection>
</treatment>
</document>
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