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<document id="D7B9593EB1436D4E8CA19E7F17509DBA" ID-CLB-Dataset="304901" ID-DOI="10.1649/0010-065X(2001)055[0515:EAANGA]2.0.CO;2" ID-GBIF-Dataset="0647875f-6005-4b9f-9388-2318633c573f" ID-ISSN="1938-4394" ID-Zenodo-Dep="10102874" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1630156293176" checkinUser="tatiana" docAuthor="Brown, Kirby W. &amp; Triplehorn, Charles A." docDate="2001" docId="03A95D4BFFEA5021F3B9FB1568849F62" docLanguage="en" docName="TheColeopBull.55.4.515-521.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (4)" docSource="http://dx.doi.org/10.1649/0010-065x(2001)055[0515:eaanga]2.0.co;2" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Epitragosoma Brown and Triplehorn, new genus" docType="treatment" docVersion="2" lastPageNumber="519" masterDocId="FF902533FFEA5025F22BFF9369029D4C" masterDocTitle="EPITRAGOSOMA ARENARIA, A New Genus and Species from Texas (Coleoptera: Tenebrionidae)" masterLastPageNumber="521" masterPageNumber="515" pageNumber="515" updateTime="1730314773783" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="14BADFA6FE9ACA7989E284C37BB9550E">EPITRAGOSOMA ARENARIA, A New Genus and Species from Texas (Coleoptera: Tenebrionidae)</mods:title>
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<mods:namePart id="758A3372582759F013AA637CDA023DF0">Brown, Kirby W.</mods:namePart>
<mods:affiliation id="1CF810162C20F20EF919752953868838">P. O. Box 1842 Paradise, CA 95967, U. S. A.</mods:affiliation>
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<mods:namePart id="6FEF077AE6D17E0495F864C1961CB5CB">Triplehorn, Charles A.</mods:namePart>
<mods:affiliation id="58EBBC8D546F42B256025FE00BE88B21">Museum of Biological Diversity The Ohio State University 1315 Kinnear Road Columbus, OH 43212 ­ 1192, U. S. A.</mods:affiliation>
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<mods:title id="59B40AE6A89A626D8A7E27B9ACAC129B">The Coleopterists Bulletin</mods:title>
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<mods:date id="DB80EA85265AABBC0FD4C973232E1D48">2001</mods:date>
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<subSubSection id="C31ABFD6FFEA5025F3B9FB156AA399D7" box="[402,929,1158,1179]" pageId="0" pageNumber="515" type="nomenclature">
<paragraph id="8BBFEC5DFFEA5025F3B9FB156AA399D7" blockId="0.[402,929,1158,1206]" box="[402,929,1158,1179]" pageId="0" pageNumber="515">
<heading id="D0F75B31FFEA5025F3B9FB156AA399D7" box="[402,929,1158,1179]" pageId="0" pageNumber="515">
<taxonomicName id="4C0097DEFFEA5025F3B9FB156A1C99D7" authority="Brown and Triplehorn" authorityName="Brown and Triplehorn" box="[402,798,1158,1179]" class="Insecta" family="Tenebrionidae" genus="Epitragosoma" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFEA5025F3B9FB156B2499D7" box="[402,550,1158,1179]" italics="true" pageId="0" pageNumber="515">Epitragosoma</emphasis>
Brown and Triplehorn
</taxonomicName>
,
<emphasis id="B974304FFFEA5025F100FB156AA399D7" bold="true" box="[811,929,1158,1179]" pageId="0" pageNumber="515">
<taxonomicNameLabel id="A2478D34FFEA5025F100FB156AA399D7" box="[811,929,1158,1179]" pageId="0" pageNumber="515" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C31ABFD6FFEA5027F04CFB32683799F5" lastPageId="2" lastPageNumber="517" pageId="0" pageNumber="515" type="description">
<paragraph id="8BBFEC5DFFEA5025F04CFB326BC999FA" blockId="0.[402,929,1158,1206]" box="[615,715,1185,1206]" pageId="0" pageNumber="515">
<figureCitation id="133BF0D8FFEA5025F04CFB326BC999FA" box="[615,715,1185,1206]" captionStart-0="Fig" captionStart-1="Figs" captionStart-2="Figs" captionStartId-0="1.[269,303,1483,1502]" captionStartId-1="2.[272,315,931,950]" captionStartId-2="4.[272,315,1608,1627]" captionTargetBox-0="[242,1078,269,1455]" captionTargetBox-1="[252,1080,268,903]" captionTargetId-0="figure-11@1.[239,1084,265,1462]" captionTargetId-1="figure-12@2.[248,1083,265,910]" captionTargetId-2="figure-10@3.[246,1078,265,1644]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Fig. 1. Epitragosoma arenaria dorsal habitus, female." captionText-1="Figs. 23. Epitragosoma arenaria. 2) apical abdominal tergites, female; 3) apical abdominal tergites, male." captionText-2="Figs. 48. Epitragosoma arenaria. 4) prosternal process, ventral view; 5) head, dorsal; 6) antenna; 7) metatarsus; 8) metathoracic wing." figureDoi-0="http://doi.org/10.5281/zenodo.10102876" figureDoi-1="http://doi.org/10.5281/zenodo.10102878" figureDoi-2="http://doi.org/10.5281/zenodo.14014557" httpUri-0="https://zenodo.org/record/10102876/files/figure.png" httpUri-1="https://zenodo.org/record/10102878/files/figure.png" httpUri-2="https://zenodo.org/record/14014557/files/figure.png" pageId="0" pageNumber="515">Figs. 18</figureCitation>
</paragraph>
<paragraph id="8BBFEC5DFFEA5027F33BFB5B683799F5" blockId="0.[249,1083,1224,1652]" lastBlockId="2.[249,1083,1022,1209]" lastPageId="2" lastPageNumber="517" pageId="0" pageNumber="515">
<emphasis id="B974304FFFEA5025F33BFB5B688B9997" bold="true" box="[272,393,1224,1243]" pageId="0" pageNumber="515">Description.</emphasis>
Body (
<figureCitation id="133BF0D8FFEA5025F3FDFB5A6B129990" box="[470,528,1225,1244]" captionStart="Fig" captionStartId="1.[269,303,1483,1502]" captionTargetBox="[242,1078,269,1455]" captionTargetId="figure-11@1.[239,1084,265,1462]" captionTargetPageId="1" captionText="Fig. 1. Epitragosoma arenaria dorsal habitus, female." figureDoi="http://doi.org/10.5281/zenodo.10102876" httpUri="https://zenodo.org/record/10102876/files/figure.png" pageId="0" pageNumber="515">Fig. 1</figureCitation>
) elongate­elliptical, convex, clothed with short but conspicuous pubescence.
<emphasis id="B974304FFFEA5025F3FAFB726B0999B8" box="[465,523,1249,1268]" italics="true" pageId="0" pageNumber="515">Head:</emphasis>
(
<figureCitation id="133BF0D8FFEA5025F036FB726B5B99B8" box="[541,601,1249,1268]" captionStart="Figs" captionStartId="4.[272,315,1608,1627]" captionTargetId="figure-10@3.[246,1078,265,1644]" captionTargetPageId="3" captionText="Figs. 48. Epitragosoma arenaria. 4) prosternal process, ventral view; 5) head, dorsal; 6) antenna; 7) metatarsus; 8) metathoracic wing." figureDoi="http://doi.org/10.5281/zenodo.14014557" httpUri="https://zenodo.org/record/14014557/files/figure.png" pageId="0" pageNumber="515">Fig. 5</figureCitation>
) convex; epistoma with median lobe prolonged, moderately acute; lateral lobes obtusely rounded, not prominent. Ventral post­genal process prolonged, narrow, sharp and slightly everted. Eyes large, weakly convex and not protruding, coarsely faceted; supraorbital carina absent to extremely weak. Maxillary palpus with apical segment securiform. Antenna (
<figureCitation id="133BF0D8FFEA5025F0F7FAD26A159818" box="[732,791,1345,1364]" captionStart="Figs" captionStartId="4.[272,315,1608,1627]" captionTargetId="figure-10@3.[246,1078,265,1644]" captionTargetPageId="3" captionText="Figs. 48. Epitragosoma arenaria. 4) prosternal process, ventral view; 5) head, dorsal; 6) antenna; 7) metatarsus; 8) metathoracic wing." figureDoi="http://doi.org/10.5281/zenodo.14014557" httpUri="https://zenodo.org/record/14014557/files/figure.png" pageId="0" pageNumber="515">Fig. 6</figureCitation>
) not reaching hind margin of pronotum when extended caudad, basal six segments cylindrical with third and fourth segments longest and subequal, width one­half length, segments 710 somewhat flattened, subequal in length but each slightly broader than preceding segment, each with a tomentose sensory area, eleventh segment two­thirds as broad as tenth, rounded apically with apical half occupied with a tomentose sensory area.
<emphasis id="B974304FFFEA5025F116FA2A6AA39880" box="[829,929,1465,1484]" italics="true" pageId="0" pageNumber="515">Pronotum:</emphasis>
convex, lateral margin rounded, lateral marginal bead scarcely defined and not reaching posterior angle; lateral margin arcuate in dorsal view, widest at middle or at hind angles; apical and basal angles acute; apical margin broadly and shallowly concave; basal margin strongly bisinuate.
<emphasis id="B974304FFFEA5025F314F98A68839B60" box="[319,385,1561,1580]" italics="true" pageId="0" pageNumber="515">Elytra:</emphasis>
fused along suture, convex, broadly oval; pseudopleural crest faintly carinate from apex to humeri, interval narrow at apex, gradually widening towards humeri.
<emphasis id="B974304FFFEA5025F2D2F9DA68BE9B10" box="[249,444,1609,1628]" italics="true" pageId="0" pageNumber="515">Metathoracic wings:</emphasis>
(
<figureCitation id="133BF0D8FFEA5025F3E1F9DA6B069B10" box="[458,516,1609,1628]" captionStart="Figs" captionStartId="4.[272,315,1608,1627]" captionTargetId="figure-10@3.[246,1078,265,1644]" captionTargetPageId="3" captionText="Figs. 48. Epitragosoma arenaria. 4) prosternal process, ventral view; 5) head, dorsal; 6) antenna; 7) metatarsus; 8) metathoracic wing." figureDoi="http://doi.org/10.5281/zenodo.14014557" httpUri="https://zenodo.org/record/14014557/files/figure.png" pageId="0" pageNumber="515">Fig. 8</figureCitation>
) reduced and non­functional, venation vestigial.
<emphasis id="B974304FFFEA5025F1FDF9DA6D3B9B10" box="[982,1081,1609,1628]" italics="true" pageId="0" pageNumber="515">Prosternal</emphasis>
<emphasis id="B974304FFFEA5025F2D2F9F268409B38" box="[249,322,1633,1652]" italics="true" pageId="0" pageNumber="515">process</emphasis>
<emphasis id="B974304FFFEA5025F369F9F268459B38" box="[322,327,1633,1652]" italics="true" pageId="0" pageNumber="515">:</emphasis>
(
<figureCitation id="133BF0D8FFEA5025F373F9F268909B38" box="[344,402,1633,1652]" captionStart="Figs" captionStartId="4.[272,315,1608,1627]" captionTargetId="figure-10@3.[246,1078,265,1644]" captionTargetPageId="3" captionText="Figs. 48. Epitragosoma arenaria. 4) prosternal process, ventral view; 5) head, dorsal; 6) antenna; 7) metatarsus; 8) metathoracic wing." figureDoi="http://doi.org/10.5281/zenodo.14014557" httpUri="https://zenodo.org/record/14014557/files/figure.png" pageId="0" pageNumber="515">Fig. 4</figureCitation>
) horizontally produced behind coxae.
<emphasis id="B974304FFFEA5025F123F9F26A8F9B38" box="[776,909,1633,1652]" italics="true" pageId="0" pageNumber="515">Mesosternum:</emphasis>
distinctly but not deeply excavate.
<emphasis id="B974304FFFE85027F38BFC6D6B20995D" box="[416,546,1022,1041]" italics="true" pageId="2" pageNumber="517">Metasternum:</emphasis>
short, strongly bifurcate posteriorly.
<emphasis id="B974304FFFE85027F1A2FC6D6ABE995D" box="[905,956,1022,1041]" italics="true" pageId="2" pageNumber="517">Legs:</emphasis>
stout; profemur weakly expanded apically and not acute; protarsus and mesotarsus with dense pads of golden setae on plantar surfaces of basal four segments, with occasional coarse, stout setae; metatarsus (
<figureCitation id="133BF0D8FFE85027F383FBD568E39915" box="[424,481,1094,1113]" captionStart="Figs" captionStartId="4.[272,315,1608,1627]" captionTargetId="figure-10@3.[246,1078,265,1644]" captionTargetPageId="3" captionText="Figs. 48. Epitragosoma arenaria. 4) prosternal process, ventral view; 5) head, dorsal; 6) antenna; 7) metatarsus; 8) metathoracic wing." figureDoi="http://doi.org/10.5281/zenodo.14014557" httpUri="https://zenodo.org/record/14014557/files/figure.png" pageId="2" pageNumber="517">Fig. 7</figureCitation>
) with pads more poorly developed and stout setae more prominent.
<emphasis id="B974304FFFE85027F31EFBCD68FB993D" box="[309,505,1118,1137]" italics="true" pageId="2" pageNumber="517">Abdominal tergum 7</emphasis>
of female (
<figureCitation id="133BF0D8FFE85027F041FBCD6BA6993D" box="[618,676,1118,1137]" captionStart="Figs" captionStartId="2.[272,315,931,950]" captionTargetBox="[252,1080,268,903]" captionTargetId="figure-12@2.[248,1083,265,910]" captionTargetPageId="2" captionText="Figs. 23. Epitragosoma arenaria. 2) apical abdominal tergites, female; 3) apical abdominal tergites, male." figureDoi="http://doi.org/10.5281/zenodo.10102878" httpUri="https://zenodo.org/record/10102878/files/figure.png" pageId="2" pageNumber="517">Fig. 2</figureCitation>
) highly modified, sharply bifurcate with a deep central sinus; of male (
<figureCitation id="133BF0D8FFE85027F039FBE56B4F99C5" box="[530,589,1142,1161]" captionStart="Figs" captionStartId="2.[272,315,931,950]" captionTargetBox="[252,1080,268,903]" captionTargetId="figure-12@2.[248,1083,265,910]" captionTargetPageId="2" captionText="Figs. 23. Epitragosoma arenaria. 2) apical abdominal tergites, female; 3) apical abdominal tergites, male." figureDoi="http://doi.org/10.5281/zenodo.10102878" httpUri="https://zenodo.org/record/10102878/files/figure.png" pageId="2" pageNumber="517">Fig. 3</figureCitation>
) weakly bifurcate with a shallow V shaped sinus.
<emphasis id="B974304FFFE85027F2D2FB1D68C099ED" box="[249,450,1166,1185]" italics="true" pageId="2" pageNumber="517">Abdominal tergum 8</emphasis>
of female trapezoidal, wider apically; of male normal lunate in shape.
</paragraph>
</subSubSection>
<caption id="DF7FBCD5FFEB5024F326FA586A1D9892" ID-DOI="http://doi.org/10.5281/zenodo.10102876" ID-Zenodo-Dep="10102876" box="[269,799,1483,1502]" httpUri="https://zenodo.org/record/10102876/files/figure.png" pageId="1" pageNumber="516" startId="1.[269,303,1483,1502]" targetBox="[242,1078,269,1455]" targetPageId="1">
<paragraph id="8BBFEC5DFFEB5024F326FA586A1D9892" blockId="1.[269,799,1483,1502]" box="[269,799,1483,1502]" pageId="1" pageNumber="516">
<emphasis id="B974304FFFEB5024F326FA58684E9892" bold="true" box="[269,332,1483,1502]" pageId="1" pageNumber="516">Fig. 1.</emphasis>
<taxonomicName id="4C0097DEFFEB5024F348FA586B429892" box="[355,576,1483,1502]" class="Insecta" family="Tenebrionidae" genus="Epitragosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="516" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis id="B974304FFFEB5024F348FA586B429892" box="[355,576,1483,1502]" italics="true" pageId="1" pageNumber="516">Epitragosoma arenaria</emphasis>
</taxonomicName>
dorsal habitus, female.
</paragraph>
</caption>
<caption id="DF7FBCD5FFE85027F33BFC3068EB9E82" ID-DOI="http://doi.org/10.5281/zenodo.10102878" ID-Zenodo-Dep="10102878" httpUri="https://zenodo.org/record/10102878/files/figure.png" pageId="2" pageNumber="517" startId="2.[272,315,931,950]" targetBox="[252,1080,268,903]" targetPageId="2">
<paragraph id="8BBFEC5DFFE85027F33BFC3068EB9E82" blockId="2.[249,1081,931,974]" pageId="2" pageNumber="517">
<emphasis id="B974304FFFE85027F33BFC3068709EFA" bold="true" box="[272,370,931,950]" pageId="2" pageNumber="517">Figs. 23.</emphasis>
<taxonomicName id="4C0097DEFFE85027F3A2FC306B6A9EFA" box="[393,616,931,950]" class="Insecta" family="Tenebrionidae" genus="Epitragosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis id="B974304FFFE85027F3A2FC306B6A9EFA" box="[393,616,931,950]" italics="true" pageId="2" pageNumber="517">Epitragosoma arenaria</emphasis>
</taxonomicName>
.
<emphasis id="B974304FFFE85027F05CFC306B899EFA" bold="true" box="[631,651,931,950]" pageId="2" pageNumber="517">2)</emphasis>
apical abdominal tergites, female;
<emphasis id="B974304FFFE85027F1CEFC306AFA9EFA" bold="true" box="[997,1016,931,950]" pageId="2" pageNumber="517">3)</emphasis>
apical abdominal tergites, male.
</paragraph>
</caption>
<subSubSection id="C31ABFD6FFE85027F338FB5D684299B2" pageId="2" pageNumber="517" type="etymology">
<paragraph id="8BBFEC5DFFE85027F338FB5D684299B2" blockId="2.[249,1082,1230,1652]" pageId="2" pageNumber="517">
<emphasis id="B974304FFFE85027F338FB5D689199AF" bold="true" box="[275,403,1230,1251]" pageId="2" pageNumber="517">Etymology.</emphasis>
The name of this genus refers to its resemblance to
<taxonomicName id="4C0097DEFFE85027F19FFB5C684299B2" authority="Casey." authorityName="Casey." class="Insecta" family="Tenebrionidae" genus="Epitragopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFE85027F19FFB5C6D3899A8" box="[948,1082,1231,1252]" italics="true" pageId="2" pageNumber="517">Epitragopsis</emphasis>
Casey.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C31ABFD6FFE85027F338FA976D2C9855" box="[275,1070,1284,1305]" pageId="2" pageNumber="517" type="type_taxon">
<paragraph id="8BBFEC5DFFE85027F338FA976D2C9855" blockId="2.[249,1082,1230,1652]" box="[275,1070,1284,1305]" pageId="2" pageNumber="517">
<emphasis id="B974304FFFE85027F338FA9768AB9855" bold="true" box="[275,425,1284,1305]" pageId="2" pageNumber="517">
<typeStatus id="54BB52FFFFE85027F338FA97684E9855" box="[275,332,1284,1305]" pageId="2" pageNumber="517">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C0097DEFFE85027F39AFA976A9A9855" authority="Brown and Triplehorn" authorityName="Brown and Triplehorn" box="[433,920,1284,1305]" class="Insecta" family="Tenebrionidae" genus="Epitragosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="species" species="arenaria" status="sp. nov.">Epitragosoma arenaria Brown and Triplehorn</taxonomicName>
,
<emphasis id="B974304FFFE85027F18FFA976D2C9855" bold="true" box="[932,1070,1284,1305]" pageId="2" pageNumber="517">
<taxonomicNameLabel id="A2478D34FFE85027F18FFA976D2B9855" box="[932,1065,1284,1305]" pageId="2" pageNumber="517" rank="species">new species</taxonomicNameLabel>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31ABFD6FFE85021F338FA8D68849F62" lastPageId="4" lastPageNumber="519" pageId="2" pageNumber="517" type="diagnosis">
<paragraph id="8BBFEC5DFFE85027F338FA8D6BF29B38" blockId="2.[249,1082,1230,1652]" pageId="2" pageNumber="517">
<emphasis id="B974304FFFE85027F338FA8D68A6987F" bold="true" box="[275,420,1310,1331]" pageId="2" pageNumber="517">Comparison.</emphasis>
This new genus resembles
<taxonomicName id="4C0097DEFFE85027F0FCFA8C6AB39878" authority="Casey" authorityName="Casey" authorityYear="1907" box="[727,945,1311,1332]" class="Insecta" family="Tenebrionidae" genus="Lobometopon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFE85027F0FCFA8C6A659878" box="[727,871,1311,1332]" italics="true" pageId="2" pageNumber="517">Lobometopon</emphasis>
Casey
</taxonomicName>
and keys to that genus in Arnett 1960 (it is included in Arnett and Thomas 2001). It differs in the supraorbital carina usually totally absent, third antennal segment subequal to fourth (third much longer than fourth in
<taxonomicName id="4C0097DEFFE85027F0DDFAFC6A8498C8" authorityName="Casey" authorityYear="1907" box="[758,902,1391,1412]" class="Insecta" family="Tenebrionidae" genus="Lobometopon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFE85027F0DDFAFC6A8498C8" box="[758,902,1391,1412]" italics="true" pageId="2" pageNumber="517">Lobometopon</emphasis>
</taxonomicName>
), weak to absent lateral marginal ridge on pronotum almost never reaching posterior angles, and brachypterous wings with fused elytra and short metasternum. Only two other genera of
<taxonomicName id="4C0097DEFFE85027F343FA2C68D39898" authorityName="Blanchard" authorityYear="1845" box="[360,465,1471,1492]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="tribe" tribe="Epitragini">Epitragini</taxonomicName>
are brachypterous,
<taxonomicName id="4C0097DEFFE85027F082FA2C6A539898" authority="Horn" authorityName="Horn" authorityYear="1885" box="[681,849,1471,1492]" class="Insecta" family="Tenebrionidae" genus="Conoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFE85027F082FA2C6A129898" box="[681,784,1471,1492]" italics="true" pageId="2" pageNumber="517">Conoecus</emphasis>
Horn
</taxonomicName>
and
<taxonomicName id="4C0097DEFFE85027F1A1FA2C686998A2" authority="Champion. Both" authorityName="Champion. Both" authorityYear="1884" class="Insecta" family="Tenebrionidae" genus="Tydeolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFE85027F1A1FA2C6AEA9898" box="[906,1000,1471,1492]" italics="true" pageId="2" pageNumber="517">Tydeolus</emphasis>
Champion. Both
</taxonomicName>
have the pronotum widest anterior to the midpoint.
<taxonomicName id="4C0097DEFFE85027F18DFA496D0F98A3" authorityName="Horn" authorityYear="1885" box="[934,1037,1498,1519]" class="Insecta" family="Tenebrionidae" genus="Conoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFE85027F18DFA496D0F98A3" box="[934,1037,1498,1519]" italics="true" pageId="2" pageNumber="517">Conoecus</emphasis>
</taxonomicName>
has a deflexed prosternal process, convex rather than excavate mesosternum, and rounded posterior pronotal angles.
<taxonomicName id="4C0097DEFFE85027F044F99C6BCF9B68" authorityName="Champion. Both" authorityYear="1884" box="[623,717,1551,1572]" class="Insecta" family="Tenebrionidae" genus="Tydeolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFE85027F044F99C6BCF9B68" box="[623,717,1551,1572]" italics="true" pageId="2" pageNumber="517">Tydeolus</emphasis>
</taxonomicName>
known from one species in
<collectingCountry id="F317ACCDFFE85027F62AF99C681B9B72" name="Mexico" pageId="2" pageNumber="517">Mexico</collectingCountry>
has a flattened rather than convex head.
<taxonomicName id="4C0097DEFFE85027F0FBF9BA6A639B72" box="[720,865,1577,1598]" class="Insecta" family="Tenebrionidae" genus="Epitragosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="519" phylum="Arthropoda" rank="genus">Epitragosoma</taxonomicName>
superficially resembles
<taxonomicName id="4C0097DEFFE85027F305F9D768FC9B15" authority="Casey" authorityName="Casey" box="[302,510,1604,1625]" class="Insecta" family="Tenebrionidae" genus="Epitragopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFE85027F305F9D768B69B15" box="[302,436,1604,1625]" italics="true" pageId="2" pageNumber="517">Epitragopsis</emphasis>
Casey
</taxonomicName>
which has a weak prosternal process, weakly excavated mesosternum, and more protruding eyes.
</paragraph>
<paragraph id="8BBFEC5DFFEE5021F338FE9C68849F62" blockId="4.[249,1082,271,559]" pageId="4" pageNumber="519">
The structure of the seventh tergite (
<figureCitation id="133BF0D8FFEE5021F08AFE9C6A0B9C68" box="[673,777,271,292]" captionStart="Figs" captionStartId="2.[272,315,931,950]" captionTargetBox="[252,1080,268,903]" captionTargetId="figure-12@2.[248,1083,265,910]" captionTargetPageId="2" captionText="Figs. 23. Epitragosoma arenaria. 2) apical abdominal tergites, female; 3) apical abdominal tergites, male." figureDoi="http://doi.org/10.5281/zenodo.10102878" httpUri="https://zenodo.org/record/10102878/files/figure.png" pageId="4" pageNumber="519">Figs. 2, 3</figureCitation>
) is a remarkable and previously overlooked character in
<taxonomicName id="4C0097DEFFEE5021F065FEBA6BB89C72" authorityName="Blanchard" authorityYear="1845" box="[590,698,297,318]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="519" phylum="Arthropoda" rank="tribe" tribe="Epitragini">Epitragini</taxonomicName>
. It is sexually dimorphic in many genera and more highly modified in females than in males. The following discussion pertains to females. The seventh tergite in
<taxonomicName id="4C0097DEFFEE5021F117FECC6AF19C38" authority="Casey" authorityName="Casey" authorityYear="1907" box="[828,1011,351,372]" class="Insecta" family="Tenebrionidae" genus="Cyrtomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="519" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFEE5021F117FECC6AAB9C38" box="[828,937,351,372]" italics="true" pageId="4" pageNumber="519">Cyrtomius</emphasis>
Casey
</taxonomicName>
,
<taxonomicName id="4C0097DEFFEE5021F1D4FECC687B9CC2" authority="Casey" authorityName="Casey" authorityYear="1907" class="Insecta" family="Tenebrionidae" genus="Bothrotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="519" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFEE5021F1D4FECC682F9CC3" italics="true" pageId="4" pageNumber="519">Bothrotes</emphasis>
Casey
</taxonomicName>
and most
<taxonomicName id="4C0097DEFFEE5021F3DDFEE96BAB9CC2" authority="Casey" authorityName="Casey" authorityYear="1907" box="[502,681,377,399]" class="Insecta" family="Tenebrionidae" genus="Pechalius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="519" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFEE5021F3DDFEE96B5F9CC3" box="[502,605,378,399]" italics="true" pageId="4" pageNumber="519">Pechalius</emphasis>
Casey
</taxonomicName>
has an apical cavity with a central projection. In
<taxonomicName id="4C0097DEFFEE5021F3A6FE076B239CE5" box="[397,545,404,425]" class="Insecta" family="Tenebrionidae" genus="Epitragosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="519" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFEE5021F3A6FE076B239CE5" box="[397,545,404,425]" italics="true" pageId="4" pageNumber="519">Epitragosoma</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0097DEFFEE5021F005FE076BBC9CE5" authorityName="Casey" authorityYear="1907" box="[558,702,404,425]" class="Insecta" family="Tenebrionidae" genus="Lobometopon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="519" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFEE5021F005FE076BBC9CE5" box="[558,702,404,425]" italics="true" pageId="4" pageNumber="519">Lobometopon</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0097DEFFEE5021F0E0FE076ADF9CE5" authority="(Horn)" baseAuthorityName="Horn" baseAuthorityYear="1870" box="[715,989,404,425]" class="Insecta" family="Tenebrionidae" genus="Pechalius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="519" phylum="Arthropoda" rank="species" species="dentiger">
<emphasis id="B974304FFFEE5021F0E0FE076A8D9CE5" box="[715,911,404,425]" italics="true" pageId="4" pageNumber="519">Pechalius dentiger</emphasis>
(Horn)
</taxonomicName>
and
<taxonomicName id="4C0097DEFFEE5021F624FE0768A29C88" authority="Casey" authorityName="Casey" authorityYear="1907" class="Insecta" family="Tenebrionidae" genus="Metopoloba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="519" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFEE5021F624FE0768549C88" italics="true" pageId="4" pageNumber="519">Metopoloba</emphasis>
Casey
</taxonomicName>
the apical cavity lacks a central projection. In
<emphasis id="B974304FFFEE5021F188FE3C6D3B9C88" box="[931,1081,431,452]" italics="true" pageId="4" pageNumber="519">
<taxonomicName id="4C0097DEFFEE5021F188FE3C6D379C88" authorityName="Casey" authorityYear="1907" box="[931,1077,431,452]" class="Insecta" family="Tenebrionidae" genus="Lobometopon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="519" phylum="Arthropoda" rank="genus">Lobometopon</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C0097DEFFEE5021F300FE5968BD9C93" box="[299,447,458,479]" class="Insecta" family="Tenebrionidae" genus="Epitragosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="519" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFEE5021F300FE5968BD9C93" box="[299,447,458,479]" italics="true" pageId="4" pageNumber="519">Epitragosoma</emphasis>
</taxonomicName>
the cavity is deep and located between two pronounced projections that in
<taxonomicName id="4C0097DEFFEE5021F3EFFE776B5A9CB5" box="[452,600,484,505]" class="Insecta" family="Tenebrionidae" genus="Epitragosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="519" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFEE5021F3EFFE776B5A9CB5" box="[452,600,484,505]" italics="true" pageId="4" pageNumber="519">Epitragosoma</emphasis>
</taxonomicName>
are sharp and narrow. The seventh tergite is only weakly emarginate in
<taxonomicName id="4C0097DEFFEE5021F009FE6C6BF99F58" authority="LeConte" authorityName="LeConte" authorityYear="1866" box="[546,763,511,532]" class="Insecta" family="Tenebrionidae" genus="Schoenicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="519" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFEE5021F009FE6C6B9A9F58" box="[546,664,511,532]" italics="true" pageId="4" pageNumber="519">Schoenicus</emphasis>
LeConte
</taxonomicName>
and is unmodified in
<taxonomicName id="4C0097DEFFEE5021F1C5FE6C68849F62" authority="Casey." authorityName="Casey." class="Insecta" family="Tenebrionidae" genus="Epitragodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="519" phylum="Arthropoda" rank="genus">
<emphasis id="B974304FFFEE5021F1C5FE6C68349F63" italics="true" pageId="4" pageNumber="519">Epitragodes</emphasis>
Casey.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>