117 lines
16 KiB
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117 lines
16 KiB
XML
<document id="8F1B6D6D00EBC4B20307B568149FA765" ID-DOI="10.1080/00222930500140629" ID-ISSN="1464-5262" ID-Zenodo-Dep="5219994" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629350369910" checkinUser="felipe" docAuthor="Bálint, Zsolt, Vértesy, Zofia & Biró, László P." docDate="2005" docId="03A4CD19FF830D15FDD7FB977A47FB08" docLanguage="en" docName="JNatHist.39.31.2935-2952.pdf" docOrigin="Journal of Natural History 39 (31)" docSource="http://dx.doi.org/10.1080/00222930500140629" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Penaincisalia perezi Balint 2001" docType="treatment" docVersion="3" lastPageNumber="2940" masterDocId="FF9DB561FF850D12FFC2FFC47B7EFFD7" masterDocTitle="Microstructures and nanostructures of high Andean Penaincisalia lycaenid butterfly scales (Lepidoptera: Lycaenidae): descriptions and interpretations" masterLastPageNumber="2952" masterPageNumber="2935" pageNumber="2939" updateTime="1699069084240" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="0F97E9D48D32BD86ED19A85A677241BA">Microstructures and nanostructures of high Andean Penaincisalia lycaenid butterfly scales (Lepidoptera: Lycaenidae): descriptions and interpretations</mods:title>
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<mods:namePart id="75E7535242DDDDE4757F35341584D688">Bálint, Zsolt</mods:namePart>
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<mods:namePart id="E9D5F0D2E244241F34B997E9E6429D38">Vértesy, Zofia</mods:namePart>
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<mods:namePart id="1CB55B4673B1538C1B9497C68D8C115F">Biró, László P.</mods:namePart>
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<mods:typeOfResource id="7964DE3E20D519BF23B805C3734EDF15">text</mods:typeOfResource>
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<mods:title id="93E56EA8D0A67CA772028D3767200AF6">Journal of Natural History</mods:title>
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<mods:date id="B3F558E1E3882B46FB47C0EFD267B27E">2005</mods:date>
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<mods:number id="44557FBDD36F9DA06141F1B0C5DA9614">2005-09-15</mods:number>
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<mods:number id="CFD83EA101A824A13A5A142716A33D0A">39</mods:number>
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<treatment id="03A4CD19FF830D15FDD7FB977A47FB08" ID-DOI="http://doi.org/10.5281/zenodo.5220012" ID-Zenodo-Dep="5220012" LSID="urn:lsid:plazi:treatment:03A4CD19FF830D15FDD7FB977A47FB08" httpUri="http://treatment.plazi.org/id/03A4CD19FF830D15FDD7FB977A47FB08" lastPageId="7" lastPageNumber="2940" pageId="6" pageNumber="2939">
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<subSubSection id="C3172F84FF830D14FDD7FB977868FBBC" box="[533,790,1107,1131]" pageId="6" pageNumber="2939" type="nomenclature">
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<paragraph id="8BB27C0FFF830D14FDD7FB977868FBBC" blockId="6.[533,790,1107,1163]" box="[533,790,1107,1131]" pageId="6" pageNumber="2939">
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<heading id="D0FACB63FF830D14FDD7FB977868FBBC" box="[533,790,1107,1131]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="2939" reason="2">
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<taxonomicName id="4C0D078CFF830D14FDD7FB977868FBBC" authorityName="Balint" authorityYear="2001" box="[533,790,1107,1131]" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="2939" phylum="Arthropoda" rank="species" species="perezi">
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<emphasis id="B979A01DFF830D14FDD7FB977868FBBC" bold="true" box="[533,790,1107,1131]" italics="true" pageId="6" pageNumber="2939">Penaincisalia perezi</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3172F84FF830D15FD82FBB77A47FB08" lastPageId="7" lastPageNumber="2940" pageId="6" pageNumber="2939" type="description">
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<paragraph id="8BB27C0FFF830D14FD82FBB77995FB5C" blockId="6.[533,790,1107,1163]" box="[576,747,1139,1163]" pageId="6" pageNumber="2939">
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(
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<figureCitation id="1336608AFF830D14FD85FBB779B8FB5C" box="[583,710,1139,1163]" captionStart="Figure 1" captionStartId="5.[125,185,1025,1044]" captionTargetBox="[128,1173,163,986]" captionTargetId="figure-110@5.[125,1173,133,989]" captionTargetPageId="5" captionText="Figure 1. Lycaenid butterfly species investigated showing male dorsal wing surfaces. (A) Penaincisalia aurulenta Johnson, 1990 (Peru: PN Huascarán, Quebrada Demanda; HNHM); (B) Penaincisalia perezi Bálint, 2001 (Peru: PN Husacarán; Llanganuco; HNHM paratype); (C) Penaincisalia culminicola (Staudinger, 1894) (Peru: PNB Huascarán, Llanganuco; HNHM); (D) Thecloxurina loxurina (Felder and Felder, 1865) (Ecuador: Carchi, Res. Forest. Golondrinas; HNHM); (E) Thecloxurina atymna (Hewitson, 1870) (Ecuador: Zamora, Cord. Lagunillas; HNHM); (F) Thecloxurina atymna (Hewitson, 1870) (Ecuador: Azuay, Cuenca; HNHM)." figureDoi="http://doi.org/10.5281/zenodo.5219996" httpUri="https://zenodo.org/record/5219996/files/figure.png" pageId="6" pageNumber="2939">Figures 1B</figureCitation>
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,
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<figureCitation id="1336608AFF830D14FD10FBB7799FFB5C" box="[722,737,1139,1163]" captionStart="Figure 3" captionStartId="8.[138,198,910,929]" captionTargetBox="[138,1186,133,874]" captionTargetId="figure-209@8.[138,1186,133,874]" captionTargetPageId="8" captionText="Figure 3. Scanning electron micrographs of Penaincisalia perezi scale microstructures. (A) Tile- and shovel-shaped scales with dentate anterior margins; (B) low longitudinal ridges with flutes, straight cross ribs, open microcells, deep-laying pepper-pot structures; (C) moderate longitudinal ridges, curved cross-ribs, pepper-pot layers; (D) high longitudinal ridges, v-shaped cross ribs, small circular opening in microcell abyss." figureDoi="http://doi.org/10.5281/zenodo.5220000" httpUri="https://zenodo.org/record/5220000/files/figure.png" pageId="6" pageNumber="2939">3</figureCitation>
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)
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</paragraph>
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<paragraph id="8BB27C0FFF830D14FF67FB717930FA58" blockId="6.[138,1185,1205,1423]" pageId="6" pageNumber="2939">
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The dorsal colour is vivid Magenta Orange with Ecclesiastic Purple iridescence (
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<figureCitation id="1336608AFF830D14FF50FB117A76FB3A" box="[146,264,1237,1261]" captionStart="Figure 1" captionStartId="5.[125,185,1025,1044]" captionTargetBox="[128,1173,163,986]" captionTargetId="figure-110@5.[125,1173,133,989]" captionTargetPageId="5" captionText="Figure 1. Lycaenid butterfly species investigated showing male dorsal wing surfaces. (A) Penaincisalia aurulenta Johnson, 1990 (Peru: PN Huascarán, Quebrada Demanda; HNHM); (B) Penaincisalia perezi Bálint, 2001 (Peru: PN Husacarán; Llanganuco; HNHM paratype); (C) Penaincisalia culminicola (Staudinger, 1894) (Peru: PNB Huascarán, Llanganuco; HNHM); (D) Thecloxurina loxurina (Felder and Felder, 1865) (Ecuador: Carchi, Res. Forest. Golondrinas; HNHM); (E) Thecloxurina atymna (Hewitson, 1870) (Ecuador: Zamora, Cord. Lagunillas; HNHM); (F) Thecloxurina atymna (Hewitson, 1870) (Ecuador: Azuay, Cuenca; HNHM)." figureDoi="http://doi.org/10.5281/zenodo.5219996" httpUri="https://zenodo.org/record/5219996/files/figure.png" pageId="6" pageNumber="2939">Figure 1B</figureCitation>
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). The scales are tile- or shovel-shaped with a uniformly toothed anterior margin. The length/width ratio of the tile-shaped scales is 3/1 (150/
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<quantity id="4CF5D1EAFF830D14FC7BFB317F78FADA" box="[953,1030,1267,1293]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="2939" unit="mm" value="50.0">50 mm</quantity>
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); the shovelshaped scales are shorter, the length/width ratio is 2/1 (100/
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<quantity id="4CF5D1EAFF830D14FC8BFAD278E9FAF9" box="[841,919,1300,1326]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="2939" unit="mm" value="50.0">50 mm</quantity>
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). The dentation of the tile-shaped scales is more pronounced while the dentation of the shovel-shaped scales is far less prominent (
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<figureCitation id="1336608AFF830D14FE80FA937AC5FAB8" box="[322,443,1367,1391]" captionStart="Figure 3" captionStartId="8.[138,198,910,929]" captionTargetBox="[138,1186,133,874]" captionTargetId="figure-209@8.[138,1186,133,874]" captionTargetPageId="8" captionText="Figure 3. Scanning electron micrographs of Penaincisalia perezi scale microstructures. (A) Tile- and shovel-shaped scales with dentate anterior margins; (B) low longitudinal ridges with flutes, straight cross ribs, open microcells, deep-laying pepper-pot structures; (C) moderate longitudinal ridges, curved cross-ribs, pepper-pot layers; (D) high longitudinal ridges, v-shaped cross ribs, small circular opening in microcell abyss." figureDoi="http://doi.org/10.5281/zenodo.5220000" httpUri="https://zenodo.org/record/5220000/files/figure.png" pageId="6" pageNumber="2939">Figure 3A</figureCitation>
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). We found
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<specimenCount id="9D0BB786FF830D14FD8BFA9379BBFAB8" box="[585,709,1367,1391]" pageId="6" pageNumber="2939" type="generic" typeStatus="types">three types</specimenCount>
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of scale microstructures, which suggest a transformation series (see Discussion):
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</paragraph>
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<paragraph id="8BB27C0FFF830D14FF48FA7C7AAFF985" blockId="6.[138,1185,1464,1684]" pageId="6" pageNumber="2939">
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1. The longitudinal ridges are low, showing flutes on the longitudinal ridges, the cross ribs are straight. The space between the cross ribs is approximately
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; but highly variable, there are larger and smaller ratios. The widely open microcells are rectangular with fragmented nanostructures, the trabeculae and the lower scale membrane can be clearly seen (
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<figureCitation id="1336608AFF830D14FE8EF9FE7ABAF985" box="[332,452,1594,1618]" captionStart="Figure 2" captionStartId="7.[125,185,910,929]" captionTargetBox="[125,1173,133,874]" captionTargetId="figure-186@7.[125,1173,133,874]" captionTargetPageId="7" captionText="Figure 2. Scanning electron micrographs of Penaincisalia aurulenta scale microstructures. (A) Tile- and shovelshaped scales with dentate anterior margins; (B) longitudinal ridges with flutes, straight cross ribs; (C) widely open microcells, wispy internal structures; (D) widely open microcells, deep-laying pepper-pot structures." figureDoi="http://doi.org/10.5281/zenodo.5219998" httpUri="https://zenodo.org/record/5219998/files/figure.png" pageId="6" pageNumber="2939">Figure 2B</figureCitation>
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).
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</paragraph>
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<paragraph id="8BB27C0FFF830D15FF48F998788DFBEC" blockId="6.[138,1185,1464,1684]" lastBlockId="7.[125,1172,1025,1247]" lastPageId="7" lastPageNumber="2940" pageId="6" pageNumber="2939">
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2. The longitudinal ridges are moderate with flutes, which with curved cross ribs form rectangular-shaped deeper-laying microcells. Spaces between cross ribs are approximately
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<quantity id="4CF5D1EAFF820D15FE91FBC77AD8FBCC" box="[339,422,1025,1051]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="2940" unit="mm" value="0.5">0.5 mm</quantity>
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, and regular. The microcell openings are circular and pepper-pot nanostructures can be seen through the microcell openings (
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<figureCitation id="1336608AFF820D15FCA9FBE7789BFBEC" box="[875,997,1059,1083]" captionStart="Figure 3" captionStartId="8.[138,198,910,929]" captionTargetBox="[138,1186,133,874]" captionTargetId="figure-209@8.[138,1186,133,874]" captionTargetPageId="8" captionText="Figure 3. Scanning electron micrographs of Penaincisalia perezi scale microstructures. (A) Tile- and shovel-shaped scales with dentate anterior margins; (B) low longitudinal ridges with flutes, straight cross ribs, open microcells, deep-laying pepper-pot structures; (C) moderate longitudinal ridges, curved cross-ribs, pepper-pot layers; (D) high longitudinal ridges, v-shaped cross ribs, small circular opening in microcell abyss." figureDoi="http://doi.org/10.5281/zenodo.5220000" httpUri="https://zenodo.org/record/5220000/files/figure.png" pageId="7" pageNumber="2940">Figure 3C</figureCitation>
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).
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</paragraph>
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<caption id="DF722C87FF820D15FFBFFC4A7F5AFC01" ID-DOI="http://doi.org/10.5281/zenodo.5219998" ID-Zenodo-Dep="5219998" httpUri="https://zenodo.org/record/5219998/files/figure.png" pageId="7" pageNumber="2940" startId="7.[125,185,910,929]" targetBox="[125,1173,133,874]" targetPageId="7">
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<paragraph id="8BB27C0FFF820D15FFBFFC4A7F5AFC01" blockId="7.[125,1173,910,982]" pageId="7" pageNumber="2940">
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Figure 2. Scanning electron micrographs of
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<taxonomicName id="4C0D078CFF820D15FDE1FC4A798EFC76" authorityName="Johnson" authorityYear="1990" box="[547,752,910,929]" class="Insecta" family="Lycaenidae" genus="Penaincisalia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="2940" phylum="Arthropoda" rank="species" species="aurulenta">
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<emphasis id="B979A01DFF820D15FDE1FC4A798EFC76" box="[547,752,910,929]" italics="true" pageId="7" pageNumber="2940">Penaincisalia aurulenta</emphasis>
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</taxonomicName>
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scale microstructures. (A) Tile- and shovelshaped scales with dentate anterior margins; (B) longitudinal ridges with flutes, straight cross ribs; (C) widely open microcells, wispy internal structures; (D) widely open microcells, deep-laying pepper-pot structures.
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</paragraph>
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</caption>
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<paragraph id="8BB27C0FFF820D15FFBFFB817A47FB08" blockId="7.[125,1172,1025,1247]" pageId="7" pageNumber="2940">
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3. The longitudinal ridges with flutes are high and the cross ribs are v-shaped and form rectangular, very narrow, almost completely closed microcells with a single circular opening at the bottom. The spaces between the cross ribs are less than
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<quantity id="4CF5D1EAFF820D15FC3FFB427F2EFB49" box="[1021,1104,1156,1182]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="2940" unit="mm" value="0.1">0.1 mm</quantity>
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; their sequence is regular. Pepper-pot nanostructures can be seen beneath the openings (
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<figureCitation id="1336608AFF820D15FF72FB037A55FB08" box="[176,299,1223,1247]" captionStart="Figure 3" captionStartId="8.[138,198,910,929]" captionTargetBox="[138,1186,133,874]" captionTargetId="figure-209@8.[138,1186,133,874]" captionTargetPageId="8" captionText="Figure 3. Scanning electron micrographs of Penaincisalia perezi scale microstructures. (A) Tile- and shovel-shaped scales with dentate anterior margins; (B) low longitudinal ridges with flutes, straight cross ribs, open microcells, deep-laying pepper-pot structures; (C) moderate longitudinal ridges, curved cross-ribs, pepper-pot layers; (D) high longitudinal ridges, v-shaped cross ribs, small circular opening in microcell abyss." figureDoi="http://doi.org/10.5281/zenodo.5220000" httpUri="https://zenodo.org/record/5220000/files/figure.png" pageId="7" pageNumber="2940">Figure 3D</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |