450 lines
54 KiB
XML
450 lines
54 KiB
XML
<document id="E8FCF47E1C49599BF6D581D74B6316CF" ID-CLB-Dataset="54167" ID-DOI="10.6620/ZS.2021.60-35" ID-GBIF-Dataset="dab15414-b59f-48a1-96ce-2515b2143001" ID-PMC="PMC8652416" ID-PubMed="34963788" ID-Zenodo-Dep="8070503" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="tatiana" checkinTime="1687183603144" checkinUser="valdenar" docAuthor="Barahona-Segovia, Rodrigo M., Guzmán, Vicente Valdés & Pañinao-Monsálvez, Laura" docDate="2021" docId="03A087D3FFC5FF85B5E9FD57FAF2EF06" docLanguage="en" docName="ZoolStud.60.35.1-18.pdf" docOrigin="Zoological Studies 60 (35)" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2015-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2015-.journal_article" docStyleVersion="1" docTitle="Villalus inanis" docType="treatment" docVersion="7" lastPageNumber="14" masterDocId="FF99FFABFFCFFF88B50DFF8BFFB4EC30" masterDocTitle="Review of Chilean Cyrtinae (Diptera: Acroceridae) with the Description of Three New Species and the First Record of Villalus inanis from Argentina" masterLastPageNumber="18" masterPageNumber="1" pageNumber="11" updateTime="1716455461836" updateUser="tatiana">
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<mods:titleInfo id="C2E9FE63F75ED03A18F4CBDE553DB144">
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<mods:title id="AFB7D89CF5CAA4020AAB261D06266361">Review of Chilean Cyrtinae (Diptera: Acroceridae) with the Description of Three New Species and the First Record of Villalus inanis from Argentina</mods:title>
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<mods:namePart id="C07BE00EDB08E9E7EC9913CF21C74CCD">Barahona-Segovia, Rodrigo M.</mods:namePart>
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<mods:namePart id="D5931213DE6BF3E235FAA7CC70C4FD55">Guzmán, Vicente Valdés</mods:namePart>
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<mods:name id="722F933010494E57EFC497E298EC611E" type="personal">
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<mods:roleTerm id="BD828F8A6EBEBB52D53CD6656C9E8747">Author</mods:roleTerm>
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<mods:namePart id="D0891536918E680F4AAB3F6620AD01E3">Pañinao-Monsálvez, Laura</mods:namePart>
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<mods:typeOfResource id="59F7DD6AFAE635E98BA33BB1CAE67ED3">text</mods:typeOfResource>
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<mods:title id="2310F2EE89B8339A50B96B7130399A93">Zoological Studies</mods:title>
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<mods:part id="E512BDDF35A9521D08E1474E84FE884F">
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<mods:date id="22A5A9EF496FAD54CE5667EF674ECE04">2021</mods:date>
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<mods:number id="4EE5CA41FAD5232D84E1EB057FDF283F">2021-07-07</mods:number>
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<mods:number id="65A97CEDCC906AB3DB9D790D610E6322">60</mods:number>
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<mods:number id="A2BD7B6357769C689C222F3F3770CB16">35</mods:number>
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<treatment id="03A087D3FFC5FF85B5E9FD57FAF2EF06" ID-DOI="http://doi.org/10.5281/zenodo.8055927" ID-GBIF-Taxon="210383227" ID-Zenodo-Dep="8055927" LSID="urn:lsid:plazi:treatment:03A087D3FFC5FF85B5E9FD57FAF2EF06" httpUri="http://treatment.plazi.org/id/03A087D3FFC5FF85B5E9FD57FAF2EF06" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11">
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<subSubSection id="C313654EFFC5FF82B5E9FD57FD65EEC5" box="[228,721,732,757]" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="8BB636C5FFC5FF82B5E9FD57FD65EEC5" blockId="10.[228,721,732,789]" box="[228,721,732,757]" pageId="10" pageNumber="11">
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<heading id="D0FE81A9FFC5FF82B5E9FD57FD65EEC5" bold="true" box="[228,721,732,757]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="11" reason="2">
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<emphasis id="B97DEAD7FFC5FF82B5E9FD57FD65EEC5" bold="true" box="[228,721,732,757]" pageId="10" pageNumber="11">
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<taxonomicName id="4C094D46FFC5FF82B5E9FD57FDE9EEC5" ID-CoL="5BF4Q" ID-ENA="2566038" authority="(Philippi, 1865)" baseAuthorityName="Philippi" baseAuthorityYear="1865" box="[228,605,732,757]" class="Insecta" family="Acroceridae" genus="Villalus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="inanis">
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<emphasis id="B97DEAD7FFC5FF82B5E9FD57FE21EEC5" bold="true" box="[228,405,732,757]" italics="true" pageId="10" pageNumber="11">Villalus inanis</emphasis>
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(
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<bibRefCitation id="EF984B34FFC5FF82B4A9FD57FDE0EEC5" author="Philippi RA" box="[420,596,732,757]" pageId="10" pageNumber="11" pagination="595 - 782" refId="ref11962" refString="Philippi RA. 1865. Aufzahlung der chilenischen Dipteren. Verh Zool- Bot Ges Wien 15: 595 - 782." type="journal article" year="1865">Philippi, 1865</bibRefCitation>
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)
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</taxonomicName>
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: 645–646
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C313654EFFC5FF82B472FD76FD82EF25" box="[383,566,765,789]" pageId="10" pageNumber="11" type="description">
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||
<paragraph id="8BB636C5FFC5FF82B472FD76FD82EF25" blockId="10.[228,721,732,789]" box="[383,566,765,789]" pageId="10" pageNumber="11">
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||
(
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<figureCitation id="13322A40FFC5FF82B48BFD76FE54EF25" box="[390,480,765,789]" captionStart="Figs" captionStartId="11.[136,175,1897,1916]" captionTargetBox="[238,1303,1064,1871]" captionTargetId="figure-5@11.[133,1450,993,1938]" captionTargetPageId="11" captionText="Figs. 21–26. Genitalia of Holops and Villalus: (21) dorsal view of H. cyanea (Philippi), female; (22) dorsal view of H. pullomen sp. nov.; (23) dorsal and lateral view of H. frauenfeldi Schiner, male; (24) dorsal view of V. inanis (Philippi), female; (25) lateral view of H. pullomen, male; (26) dorsal view of gonocoxa of H. anarayae sp. nov. epa = epandrium; gncx = gonocoxa." figureDoi="http://doi.org/10.5281/zenodo.8070515" httpUri="https://zenodo.org/record/8070515/files/figure.png" pageId="10" pageNumber="11">Figs. 24</figureCitation>
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,
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<figureCitation id="13322A40FFC5FF82B4E6FD76FD9BEF25" box="[491,559,765,789]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="13.[137,177,1924,1943]" captionStartId-1="14.[176,215,1924,1943]" captionTargetBox-0="[136,1413,1027,1898]" captionTargetBox-1="[173,1451,951,1897]" captionTargetId-0="figure-369@13.[136,1413,1027,1898]" captionTargetId-1="figure-352@14.[163,1451,941,1897]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="Figs. 28–32. Villalus inanis (Philippi, 1865): (28) dorsal view; (29) lateral view; (30) in situ, lateral view (photo: William Tarrant); (31) in situ, dorsal view (photographed by Ignacio Valeria), and (33) dead, ventral view (photographed by Claudia Veas). Scale bars = 1 mm." captionText-1="Figs. 33–36. Villalus inanis (Philippi, 1865): (33) female, lateral view, holotype, (34) female, dorsal view, holotype, (35) male, lateral view and (36) male, frontal view. Scale bars are not available." figureDoi-0="http://doi.org/10.5281/zenodo.8070519" figureDoi-1="http://doi.org/10.5281/zenodo.8070521" httpUri-0="https://zenodo.org/record/8070519/files/figure.png" httpUri-1="https://zenodo.org/record/8070521/files/figure.png" pageId="10" pageNumber="11">28–36</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C313654EFFC5FF85B5A3FCB3FAF2EF06" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11" type="reference_group">
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||
<paragraph id="8BB636C5FFC5FF82B5A3FCB3FE4EEF7B" blockId="10.[174,603,824,870]" box="[174,506,824,843]" pageId="10" pageNumber="11">
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<treatmentCitationGroup id="AB1911EBFFC5FF82B5A3FCB3FE4EEF7B" box="[174,506,824,843]" pageId="10" pageNumber="11">
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=
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<treatmentCitation id="0AA810D4FFC5FF82B5B2FCB3FE43EF7B" author="Cole FR" box="[191,503,824,843]" page="64" pageId="10" pageNumber="11" year="1918">
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<taxonomicName id="4C094D46FFC5FF82B5B2FCB3FE43EF7B" ID-CoL="5BF4P" authority="Cole, 1918: 63 - 64" authorityName="Cole" authorityPageNumber="63 - 64" authorityYear="1918" box="[191,503,824,843]" class="Insecta" family="Acroceridae" genus="Villalus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="chilensis">
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<emphasis id="B97DEAD7FFC5FF82B5B2FCB3FEE4EF7B" box="[191,336,824,843]" italics="true" pageId="10" pageNumber="11">Villalus chilensis</emphasis>
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<bibRefCitation id="EF984B34FFC5FF82B458FCB3FE43EF7B" author="Cole FR" box="[341,503,824,843]" pageId="10" pageNumber="11" pagination="61 - 64" refId="ref11556" refString="Cole FR. 1918. A new genus of Cyrtidae from South America (Dip.). Entomol News 29: 61 - 64." type="journal article" year="1918">Cole, 1918: 63–64</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BB636C5FFC5FF82B5A3FCD8FDEFEF56" blockId="10.[174,603,824,870]" box="[174,603,850,870]" pageId="10" pageNumber="11">
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||
<treatmentCitationGroup id="AB1911EBFFC5FF82B5A3FCD8FDEFEF56" box="[174,603,850,870]" pageId="10" pageNumber="11">
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||
=
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<treatmentCitation id="0AA810D4FFC5FF82B5B2FCD8FE7DEF55" author="Cole FR" box="[191,457,850,870]" page="61" pageId="10" pageNumber="11" year="1918">
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<taxonomicName id="4C094D46FFC5FF82B5B2FCD8FE7DEF55" authority="Cole, 1918: 61" authorityName="Cole" authorityPageNumber="61" authorityYear="1918" box="[191,457,850,870]" class="Insecta" family="Acroceridae" genus="Lasia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dimidiata">
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<emphasis id="B97DEAD7FFC5FF82B5B2FCD8FEF2EF55" box="[191,326,850,869]" italics="true" pageId="10" pageNumber="11">Lasia dimidiata</emphasis>
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<bibRefCitation id="EF984B34FFC5FF82B446FCD9FE7DEF55" author="Cole FR" box="[331,457,850,870]" pageId="10" pageNumber="11" pagination="61 - 64" refId="ref11556" refString="Cole FR. 1918. A new genus of Cyrtidae from South America (Dip.). Entomol News 29: 61 - 64." type="journal article" year="1918">Cole, 1918: 61</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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||
(
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<emphasis id="B97DEAD7FFC5FF82B4DBFCD8FDFAEF55" box="[470,590,850,869]" italics="true" pageId="10" pageNumber="11">nomen nudum</emphasis>
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).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BB636C5FFC5FF83B5E7FC04FF5AEDA6" blockId="10.[174,774,910,966]" lastBlockId="11.[136,738,253,1014]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
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<emphasis id="B97DEAD7FFC5FF82B5E7FC04FE54EF96" box="[234,480,910,934]" italics="true" pageId="10" pageNumber="11">M a t e r i a l e x a m i n e d</emphasis>
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:
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<emphasis id="B97DEAD7FFC5FF82B4F5FC04FDD8EF96" box="[504,620,910,934]" italics="true" pageId="10" pageNumber="11">H o l o t y p e</emphasis>
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: “Valdivia, Chile” (A MNHNCL); other specimens:
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<collectingCountry id="F31E7655FFC5FF82B7A4FC25FCB6EFF6" box="[681,770,942,966]" name="Chile" pageId="10" pageNumber="11">CHILE</collectingCountry>
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.
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<emphasis id="B97DEAD7FFC5FF82B65FFF76FC66ED25" box="[850,978,253,277]" italics="true" pageId="10" pageNumber="11">Valparaíso</emphasis>
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: Parque Peñuelas,
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<date id="FFB71005FFC5FF82B1CDFF76FAF4ED25" box="[1216,1344,253,277]" pageId="10" pageNumber="11" value="1965-04-23">23.iv.1965</date>
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, Leg. N. Hichins (MNHNCL); Valparaíso,
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<date id="FFB71005FFC5FF82B1F6FE96FAD1ED05" box="[1275,1381,285,309]" pageId="10" pageNumber="11" value="1966-07">vii.1966</date>
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, Leg. W.H. Sielfield (MNHNCL);
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<emphasis id="B97DEAD7FFC5FF82B1A8FEB6FB45ED65" box="[1189,1265,317,341]" italics="true" pageId="10" pageNumber="11">Biobío</emphasis>
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: **Tirúa, Lago Lleu-Lleu,
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<date id="FFB71005FFC5FF82B6D6FED6FBEAED45" box="[987,1118,349,373]" pageId="10" pageNumber="11" value="2020-03-25">25.iii.2020</date>
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, phot. Leg. Ignacio Valeria (CSP, electronic voucher: https://figshare.com/ s/0e733c2cecb30b0de50b);
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<emphasis id="B97DEAD7FFC5FF82B19CFE15FA9BED85" box="[1169,1327,414,437]" italics="true" pageId="10" pageNumber="11">La Araucanía</emphasis>
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: **Parque Nacional Nahuelbuta,
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<date id="FFB71005FFC5FF82B15DFE36FB09EDE5" box="[1104,1213,445,469]" pageId="10" pageNumber="11" value="1982-03-07">7.iii.1982</date>
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, Leg. L.H. Pincheira (MZUC); **Fundo El Carmen, Temuco,
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<date id="FFB71005FFC5FF82B047FE56FA11EDC5" box="[1354,1445,477,501]" pageId="10" pageNumber="11" value="2016-04">iv.2016</date>
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, phot. Leg. Lucas Nuñez (CSP, electronic voucher: https://figshare.com/s/aba8eb49b2a1a1bf3999); **Gorbea,
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<date id="FFB71005FFC5FF82B6DAFDB6FBEFEE65" box="[983,1115,573,597]" pageId="10" pageNumber="11" value="2020-03-29">29.iii.2020</date>
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, phot. Leg. William Tarrant (CSP, electronic voucher: https://figshare.com/s/ c98c392e4847aedbe437);
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<emphasis id="B97DEAD7FFC5FF82B18AFDF5FB59EEA5" box="[1159,1261,638,661]" italics="true" pageId="10" pageNumber="11">Los Ríos</emphasis>
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: **Panguipulli, 1983, Leg. G. Piel (UACH); **Pichoy,
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<date id="FFB71005FFC5FF82B00DFD16FADBEE85" box="[1280,1391,669,693]" pageId="10" pageNumber="11" value="1991-02-30">30.ii.1991</date>
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, Leg. E. Krahmer (UACH); **Parque Llancahue,
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<date id="FFB71005FFC5FF82B03CFD36FA13EEE5" box="[1329,1447,701,725]" pageId="10" pageNumber="11" value="1978-03-25">25.iii.1978</date>
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, Leg. E. Krahmer (UACH); **Santo Domingo,
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<date id="FFB71005FFC5FF82B04FFD56FA12EEC5" box="[1346,1446,733,757]" pageId="10" pageNumber="11" value="1984-04-08">8.iv.1984</date>
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,
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<date id="FFB71005FFC5FF82B65FFD76FC73EF25" box="[850,967,765,789]" pageId="10" pageNumber="11" value="1987-04-03">3.iv.1987</date>
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,
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<date id="FFB71005FFC5FF82B6D4FD76FBFAEF25" box="[985,1102,765,789]" pageId="10" pageNumber="11" value="1989-04-02">2.iv.1989</date>
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, Leg. E. Krahmer (UACH); Valdivia, 5.IV.1977,
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<date id="FFB71005FFC5FF82B13BFC96FB29EF05" box="[1078,1181,797,821]" pageId="10" pageNumber="11" value="1981-04-03">3.iv.1981</date>
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,
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<date id="FFB71005FFC5FF82B1A7FC96FA96EF05" box="[1194,1314,797,821]" pageId="10" pageNumber="11" value="1983-03-30">30.iii.1983</date>
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,
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<date id="FFB71005FFC5FF82B03DFC96FA12EF05" box="[1328,1446,797,821]" pageId="10" pageNumber="11" value="1985-03-13">13.iii.1985</date>
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,
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<date id="FFB71005FFC5FF82B65FFCB5FC75EF65" box="[850,961,829,854]" pageId="10" pageNumber="11" value="1985-05-null">2. v. 1985</date>
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, Leg. Krahmer (UACH, MNHNCL); **Chahuilco,
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<date id="FFB71005FFC5FF82B6FCFCD6FBDFEF45" box="[1009,1131,861,885]" pageId="10" pageNumber="11" value="1990-03-11">11.iii.1990</date>
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, Leg. E. Krahmer (UACH); **Reumen,
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<date id="FFB71005FFC5FF82B6D5FCF6FBE4EFA5" box="[984,1104,893,917]" pageId="10" pageNumber="11" value="1991-03-15">15.iii.1991</date>
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, Leg. Lula Mata (UACH);
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<emphasis id="B97DEAD7FFC5FF82B089FCF5FC22EF85" italics="true" pageId="10" pageNumber="11">Los Lagos</emphasis>
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: **San Pablo, Osorno,
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<date id="FFB71005FFC5FF82B1A6FC16FA9CEF85" box="[1195,1320,925,949]" pageId="10" pageNumber="11" value="2020-03-24">24.iii.2020</date>
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, phot. Leg. Claudia Veas (CSP, electronic voucher: https://figshare. com/s/d771abe10446bd766c93).
|
||
<collectingCountry id="F31E7655FFC4FF83B726FF75FD6FED26" box="[555,731,254,278]" name="Argentina" pageId="11" pageNumber="12">ARGENTINA</collectingCountry>
|
||
.
|
||
<collectingRegion id="49CDF827FFC4FF83B585FE96FF56ED05" box="[136,226,285,309]" country="Argentina" name="Chubut" pageId="11" pageNumber="12">Chubut</collectingRegion>
|
||
: **S. Argentina, El Puelo, Topal, Nr. 354,
|
||
<date id="FFB71005FFC4FF83B585FEB6FEB3ED65" box="[136,263,317,341]" pageId="11" pageNumber="12" value="1961-03-25">25.iii.1961</date>
|
||
, Leg. NN (USNM); **S. Argentina,
|
||
<collectingRegion id="49CDF827FFC4FF83B7B4FEB6FF79ED46" country="Argentina" name="Rio Negro" pageId="11" pageNumber="12">Rio Negro</collectingRegion>
|
||
, El Bolson, Topal, Nr. 322,
|
||
<date id="FFB71005FFC4FF83B70DFED6FDC1ED45" box="[512,629,349,373]" pageId="11" pageNumber="12" value="1961-03-15">15.iii.1961</date>
|
||
, Leg. NN (USNM).
|
||
</paragraph>
|
||
<caption id="DF76664DFFC5FF82B5A3F80FFE5AEB84" ID-DOI="http://doi.org/10.5281/zenodo.8070513" ID-Zenodo-Dep="8070513" httpUri="https://zenodo.org/record/8070513/files/figure.png" pageId="10" pageNumber="11" startId="10.[174,213,1924,1943]" targetBox="[175,1451,1033,1895]" targetPageId="10" targetType="figure">
|
||
<paragraph id="8BB636C5FFC5FF82B5A3F80FFE5AEB84" blockId="10.[174,1453,1924,2039]" pageId="10" pageNumber="11">
|
||
<emphasis id="B97DEAD7FFC5FF82B5A3F80FFEAEEBA7" bold="true" box="[174,282,1924,1943]" pageId="10" pageNumber="11">Figs. 15–20.</emphasis>
|
||
<taxonomicName id="4C094D46FFC5FF82B42AF80FFED1EBA7" authorityName="Philippi" authorityYear="1865" box="[295,357,1924,1943]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97DEAD7FFC5FF82B42AF80FFED1EBA7" box="[295,357,1924,1943]" italics="true" pageId="10" pageNumber="11">Holops</emphasis>
|
||
</taxonomicName>
|
||
wing details: (15)
|
||
<taxonomicName id="4C094D46FFC5FF82B707F80EFDC2EBA7" authority="Barahona-Segovia & Guzmán & Pañinao-Monsálvez, 2021" authorityName="Barahona-Segovia & Guzmán & Pañinao-Monsálvez" authorityYear="2021" box="[522,630,1925,1943]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="anarayae" status="sp. nov.">
|
||
<emphasis id="B97DEAD7FFC5FF82B707F80EFDC2EBA7" box="[522,630,1925,1943]" italics="true" pageId="10" pageNumber="11">H. anarayae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57ACFFC5FF82B771F80EFD77EBA8" box="[636,707,1925,1944]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (16)
|
||
<taxonomicName id="4C094D46FFC5FF82B7F8F80EFC1CEBA7" authority="(Philippi)" baseAuthorityName="Philippi" box="[757,936,1924,1943]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cyanea">
|
||
<emphasis id="B97DEAD7FFC5FF82B7F8F80EFCF9EBA7" box="[757,845,1925,1943]" italics="true" pageId="10" pageNumber="11">H. cyanea</emphasis>
|
||
(Philippi)
|
||
</taxonomicName>
|
||
; (17)
|
||
<taxonomicName id="4C094D46FFC5FF82B6D6F80EFB2BEBA7" authority="Schiner" authorityName="Schiner" authorityYear="1868" box="[987,1183,1924,1943]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="frauenfeldi">
|
||
<emphasis id="B97DEAD7FFC5FF82B6D6F80EFBE1EBA7" box="[987,1109,1924,1943]" italics="true" pageId="10" pageNumber="11">H. frauenfeldi</emphasis>
|
||
Schiner
|
||
</taxonomicName>
|
||
; (18)
|
||
<taxonomicName id="4C094D46FFC5FF82B1DFF80EFAA3EBA7" authority="Barahona-Segovia & Guzmán & Pañinao-Monsálvez, 2021" authorityName="Barahona-Segovia & Guzmán & Pañinao-Monsálvez" authorityYear="2021" box="[1234,1303,1925,1943]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="grezi" status="sp. nov.">
|
||
<emphasis id="B97DEAD7FFC5FF82B1DFF80EFAA3EBA7" box="[1234,1303,1925,1943]" italics="true" pageId="10" pageNumber="11">H. grezi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57ACFFC5FF82B013F80EFAD0EBA8" box="[1310,1380,1925,1944]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (19)
|
||
<taxonomicName id="4C094D46FFC5FF82B09AF80EFF49EB83" authority="Barahona-Segovia & Guzmán & Pañinao-Monsálvez, 2021" authorityName="Barahona-Segovia & Guzmán & Pañinao-Monsálvez" authorityYear="2021" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="pullomen" status="sp. nov.">
|
||
<emphasis id="B97DEAD7FFC5FF82B09AF80EFF49EB83" italics="true" pageId="10" pageNumber="11">H. pullomen</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57ACFFC5FF82B40EF82AFEF3EB84" box="[259,327,1953,1972]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and (20)
|
||
<taxonomicName id="4C094D46FFC5FF82B491F82AFE5DEB83" box="[412,489,1953,1971]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="virens">
|
||
<emphasis id="B97DEAD7FFC5FF82B491F82AFE5DEB83" box="[412,489,1953,1971]" italics="true" pageId="10" pageNumber="11">H. virens</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<footnote id="E8122ACBFFC5FF82B191F86EFA1FEBC7" box="[1180,1451,2020,2039]" pageId="10" pageNumber="11">
|
||
<paragraph id="8BB636C5FFC5FF82B191F86EFA1FEBC7" blockId="10.[174,1453,1924,2039]" box="[1180,1451,2020,2039]" pageId="10" pageNumber="11">© 2021 Academia Sinica, Taiwan</paragraph>
|
||
</footnote>
|
||
<paragraph id="8BB636C5FFC4FF83B5C8FE15FF7AEDE6" blockId="11.[136,738,253,1014]" pageId="11" pageNumber="12">
|
||
<emphasis id="B97DEAD7FFC4FF83B5C8FE15FEF9ED85" box="[197,333,413,437]" italics="true" pageId="11" pageNumber="12">
|
||
<typeStatus id="54B28867FFC4FF83B5C8FE15FF43ED85" box="[197,247,414,437]" pageId="11" pageNumber="12">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
: Valdivia,
|
||
<collectingCountry id="F31E7655FFC4FF83B4CDFE16FE4EED85" box="[448,506,413,437]" name="Chile" pageId="11" pageNumber="12">Chile</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF984B34FFC4FF83B70AFE15FF76EDE6" author="Pape T & Thompson FC" pageId="11" pageNumber="12" refId="ref11932" refString="Pape T, Thompson FC. 2013. Systema Dipterorum, Version 1.5. http: // www. diptera. org /. Accessed on 5 January 2017." type="url" year="2013">Pape and Thompson 2013</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB636C5FFC4FF83B5C8FE55FE0AEF05" blockId="11.[136,738,253,1014]" pageId="11" pageNumber="12">
|
||
<emphasis id="B97DEAD7FFC4FF83B5C8FE55FE8BEDC5" box="[197,319,478,501]" italics="true" pageId="11" pageNumber="12">Diagnosis</emphasis>
|
||
: Orange-yellowish species. Head and eyes black. Head with half that maximum height of thorax. Scutum with two black, but very light longitudinal bands. Yellowish pilosity in all thoracic segments. Katepisternum brownish in ventral part. Meron grey and elongated with yellow margin. Legs completely yellowish. The cells sc, c, br, r
|
||
<subScript id="178D3480FFC4FF83B77FFD21FDCEEE88" attach="left" box="[626,634,682,696]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
|
||
, r
|
||
<subScript id="178D3480FFC4FF83B798FD21FD19EE88" attach="left" box="[661,685,682,696]" fontSize="6" pageId="11" pageNumber="12">2+3</subScript>
|
||
, r
|
||
<subScript id="178D3480FFC4FF83B7C5FD21FD55EE88" attach="left" box="[712,737,682,696]" fontSize="6" pageId="11" pageNumber="12">4+5</subScript>
|
||
and m
|
||
<subScript id="178D3480FFC4FF83B5C6FD41FF67EEE8" attach="left" box="[203,211,714,728]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
|
||
smoked with microtrichia. R
|
||
<subScript id="178D3480FFC4FF83B700FD41FDA1EEE8" attach="left" box="[525,533,714,728]" fontSize="6" pageId="11" pageNumber="12">4</subScript>
|
||
and R
|
||
<subScript id="178D3480FFC4FF83B751FD41FDD0EEE8" attach="left" box="[604,612,714,728]" fontSize="6" pageId="11" pageNumber="12">5</subScript>
|
||
not forked. Abdomen globose and slightly longer than thorax in dorsal view. Tergites and sclerites orange-yellowish with pilosity similar in color.
|
||
</paragraph>
|
||
<paragraph id="8BB636C5FFC4FF84B5C8FCB5FC03ED86" blockId="11.[136,738,253,1014]" lastBlockId="12.[850,1452,253,501]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
<emphasis id="B97DEAD7FFC4FF83B5C8FCB5FEFAEF65" box="[197,334,830,853]" italics="true" pageId="11" pageNumber="12">Description</emphasis>
|
||
: Length: 6.49 ±
|
||
<quantity id="4CF19B20FFC4FF83B711FCB5FD3CEF65" box="[540,648,830,854]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="11" pageNumber="12" unit="mm" value="0.18">0.18 mm</quantity>
|
||
; width: 1.78 ±
|
||
<quantity id="4CF19B20FFC4FF83B5D4FCD6FE88EF45" box="[217,316,861,885]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="mm" value="0.05">0.05 mm</quantity>
|
||
(head), 3.13 ±
|
||
<quantity id="4CF19B20FFC4FF83B4E1FCD5FDFBEF45" box="[492,591,862,886]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="12" unit="mm" value="0.09">0.09 mm</quantity>
|
||
(thorax) and 4.36 ±
|
||
<quantity id="4CF19B20FFC4FF83B5D1FCF6FEF5EFA5" box="[220,321,893,917]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="12" unit="mm" value="0.16">0.16 mm</quantity>
|
||
(abdomen);
|
||
<emphasis id="B97DEAD7FFC4FF83B4DAFCF5FDB9EFA5" box="[471,525,894,917]" italics="true" pageId="11" pageNumber="12">wing</emphasis>
|
||
: 6.96 ±
|
||
<quantity id="4CF19B20FFC4FF83B779FCF6FD68EFA5" box="[628,732,893,917]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="12" unit="mm" value="0.16">0.16 mm</quantity>
|
||
;
|
||
<emphasis id="B97DEAD7FFC4FF83B585FC16FF64EF85" box="[136,208,925,949]" italics="true" pageId="11" pageNumber="12">female</emphasis>
|
||
:
|
||
<emphasis id="B97DEAD7FFC4FF83B5EDFC16FEA0EF85" box="[224,276,925,949]" italics="true" pageId="11" pageNumber="12">head</emphasis>
|
||
: black; the size is approximately the half of the maximum height of the thorax; antenna inserted in the middle of the head; scape and pedicelum orange and similar in size among them; flagellum styliform and 3 times longer than pedicelum + scape together; antenna shorter than head length; eyes with short and yellowish pilosity; three ocelli brownish with ocellar triangle black, protuberant and with 6–8 yellow hairs; occiput black with long yellowish pilosity; mouth parts brownish;
|
||
<emphasis id="B97DEAD7FFC4FF83B6E9FE36FB98EDE5" box="[996,1068,445,469]" italics="true" pageId="11" pageNumber="12">thorax</emphasis>
|
||
: globose shape (
|
||
<figureCitation id="13322A40FFC4FF83B1E3FE36FACDEDE6" box="[1262,1401,445,470]" captionStart="Figs" captionStartId="13.[137,177,1924,1943]" captionTargetBox="[136,1413,1027,1898]" captionTargetId="figure-369@13.[136,1413,1027,1898]" captionTargetPageId="13" captionText="Figs. 28–32. Villalus inanis (Philippi, 1865): (28) dorsal view; (29) lateral view; (30) in situ, lateral view (photo: William Tarrant); (31) in situ, dorsal view (photographed by Ignacio Valeria), and (33) dead, ventral view (photographed by Claudia Veas). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8070519" httpUri="https://zenodo.org/record/8070519/files/figure.png" pageId="11" pageNumber="12">Figs. 28–29</figureCitation>
|
||
); orange scutum with two dark longitudinal bands slightly visible; scutum covered with long and dense orange-yellowish pilosity; pilosity sparse but uniformly distributed; postocellar lobe orange and yellowish pilosity; anterior spiracle light yellow; anepisternum globose and orange; katepisternum brownish in ventral part; anepisternum yellowish; meron grey and elongated with yellow margin; katepimeron and anatergite orange-yellow with few pilosity; scutellum orange and widest than long with yellowish pilosity; (
|
||
<figureCitation id="13322A40FFC4FF83B1F0FD76FA36EF26" box="[1277,1410,765,790]" captionStart="Figs" captionStartId="13.[137,177,1924,1943]" captionTargetBox="[136,1413,1027,1898]" captionTargetId="figure-369@13.[136,1413,1027,1898]" captionTargetPageId="13" captionText="Figs. 28–32. Villalus inanis (Philippi, 1865): (28) dorsal view; (29) lateral view; (30) in situ, lateral view (photo: William Tarrant); (31) in situ, dorsal view (photographed by Ignacio Valeria), and (33) dead, ventral view (photographed by Claudia Veas). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8070519" httpUri="https://zenodo.org/record/8070519/files/figure.png" pageId="11" pageNumber="12">Figs. 28–29</figureCitation>
|
||
,
|
||
<figureCitation id="13322A40FFC4FF83B620FC95FCC1EF06" box="[813,885,798,822]" captionStart="Figs" captionStartId="14.[176,215,1924,1943]" captionTargetBox="[173,1451,951,1897]" captionTargetId="figure-352@14.[163,1451,941,1897]" captionTargetPageId="14" captionText="Figs. 33–36. Villalus inanis (Philippi, 1865): (33) female, lateral view, holotype, (34) female, dorsal view, holotype, (35) male, lateral view and (36) male, frontal view. Scale bars are not available." figureDoi="http://doi.org/10.5281/zenodo.8070521" httpUri="https://zenodo.org/record/8070521/files/figure.png" pageId="11" pageNumber="12">33–34</figureCitation>
|
||
);
|
||
<emphasis id="B97DEAD7FFC4FF83B687FC96FC01EF05" box="[906,949,797,821]" italics="true" pageId="11" pageNumber="12">legs</emphasis>
|
||
: coxa, trochanters, femora, tibia and tarsi yellow covered by dense yellow pilosity; black claws;
|
||
<emphasis id="B97DEAD7FFC4FF83B620FCD5FCDFEF45" box="[813,875,862,885]" italics="true" pageId="11" pageNumber="12">wings</emphasis>
|
||
: all veins brownish; cell sc, c, br, r
|
||
<subScript id="178D3480FFC4FF83B1E3FCE1FB42EF48" attach="left" box="[1262,1270,874,888]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
|
||
, r
|
||
<subScript id="178D3480FFC4FF83B003FCE1FA92EF48" attach="left" box="[1294,1318,874,888]" fontSize="6" pageId="11" pageNumber="12">2+3</subScript>
|
||
, r
|
||
<subScript id="178D3480FFC4FF83B033FCE1FAE2EF48" attach="left" box="[1342,1366,874,888]" fontSize="6" pageId="11" pageNumber="12">4+5</subScript>
|
||
and m
|
||
<subScript id="178D3480FFC4FF83B64FFC01FCFEEFA8" attach="left" box="[834,842,906,920]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
|
||
smoked with microtrichia; other cell hyaline and membranous appearance; R
|
||
<subScript id="178D3480FFC4FF83B155FC21FBD4EF88" attach="left" box="[1112,1120,938,952]" fontSize="6" pageId="11" pageNumber="12">4</subScript>
|
||
and R
|
||
<subScript id="178D3480FFC4FF83B1A5FC21FB04EF88" attach="left" box="[1192,1200,938,952]" fontSize="6" pageId="11" pageNumber="12">5</subScript>
|
||
not forked; M
|
||
<subScript id="178D3480FFC4FF83B042FC21FAE3EF88" attach="left" box="[1359,1367,938,952]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
|
||
, M
|
||
<subScript id="178D3480FFC4FF83B073FC21FA32EF88" attach="left" box="[1406,1414,938,952]" fontSize="6" pageId="11" pageNumber="12">2</subScript>
|
||
and M
|
||
<subScript id="178D3480FFC4FF83B67FFC41FC3EEFE8" attach="left" box="[882,906,970,984]" fontSize="6" pageId="11" pageNumber="12">3+4</subScript>
|
||
are long, but not reaching the wing margin; all M veins similar in size; cell r
|
||
<subScript id="178D3480FFC4FF83B177FC61FB27EFC8" attach="left" box="[1146,1171,1002,1016]" fontSize="6" pageId="11" pageNumber="12">4+5</subScript>
|
||
elongated and width, with more than 1/2 length of R
|
||
<subScript id="178D3480FFC3FF84B70BFE81FDAAED28" attach="left" box="[518,542,266,280]" fontSize="6" pageId="12" pageNumber="13">2+3</subScript>
|
||
vein; cell m
|
||
<subScript id="178D3480FFC3FF84B7A7FE81FD06ED28" attach="left" box="[682,690,266,280]" fontSize="6" pageId="12" pageNumber="13">3</subScript>
|
||
absent; halters brownish; calypter rim blackish and interior part is brownish with dense and yellowish pilosity;
|
||
<emphasis id="B97DEAD7FFC3FF84B5A3FED6FEA6ED45" box="[174,274,349,373]" italics="true" pageId="12" pageNumber="13">abdomen</emphasis>
|
||
: globose and slightly longer than thorax; in dorsal view the width is the double of thorax and rough; all tergites orange with dense yellowish pilosity; tergites 1–3 similar in length; in dead animals, all sternites yellow with pilosity of same color and a big brown maculae in the middle (
|
||
<figureCitation id="13322A40FFC3FF84B166FF76FB70ED26" box="[1131,1220,253,278]" captionStart="Figs" captionStartId="13.[137,177,1924,1943]" captionTargetBox="[136,1413,1027,1898]" captionTargetId="figure-369@13.[136,1413,1027,1898]" captionTargetPageId="13" captionText="Figs. 28–32. Villalus inanis (Philippi, 1865): (28) dorsal view; (29) lateral view; (30) in situ, lateral view (photo: William Tarrant); (31) in situ, dorsal view (photographed by Ignacio Valeria), and (33) dead, ventral view (photographed by Claudia Veas). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8070519" httpUri="https://zenodo.org/record/8070519/files/figure.png" pageId="12" pageNumber="13">Fig. 32</figureCitation>
|
||
); in living animals, tergites 3–6 present a large brownish area in the middle (
|
||
<figureCitation id="13322A40FFC3FF84B657FEB6FC69ED66" box="[858,989,317,342]" captionStart="Figs" captionStartId="13.[137,177,1924,1943]" captionTargetBox="[136,1413,1027,1898]" captionTargetId="figure-369@13.[136,1413,1027,1898]" captionTargetPageId="13" captionText="Figs. 28–32. Villalus inanis (Philippi, 1865): (28) dorsal view; (29) lateral view; (30) in situ, lateral view (photo: William Tarrant); (31) in situ, dorsal view (photographed by Ignacio Valeria), and (33) dead, ventral view (photographed by Claudia Veas). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8070519" httpUri="https://zenodo.org/record/8070519/files/figure.png" pageId="12" pageNumber="13">Figs. 30–31</figureCitation>
|
||
); interior parts of tergites 4–6 covered with black and granulate plates;
|
||
<emphasis id="B97DEAD7FFC3FF84B173FED6FB69ED45" box="[1150,1245,349,373]" italics="true" pageId="12" pageNumber="13">genitalia</emphasis>
|
||
: cerci globose and yellow with pilosity of the same color in dorsal view (
|
||
<figureCitation id="13322A40FFC3FF84B657FE16FC1EED86" box="[858,938,413,438]" captionStart="Figs" captionStartId="11.[136,175,1897,1916]" captionTargetBox="[238,1303,1064,1871]" captionTargetId="figure-5@11.[133,1450,993,1938]" captionTargetPageId="11" captionText="Figs. 21–26. Genitalia of Holops and Villalus: (21) dorsal view of H. cyanea (Philippi), female; (22) dorsal view of H. pullomen sp. nov.; (23) dorsal and lateral view of H. frauenfeldi Schiner, male; (24) dorsal view of V. inanis (Philippi), female; (25) lateral view of H. pullomen, male; (26) dorsal view of gonocoxa of H. anarayae sp. nov. epa = epandrium; gncx = gonocoxa." figureDoi="http://doi.org/10.5281/zenodo.8070515" httpUri="https://zenodo.org/record/8070515/files/figure.png" pageId="12" pageNumber="13">Fig. 24</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF76664DFFC4FF83B585F8E2FC9DEB84" ID-DOI="http://doi.org/10.5281/zenodo.8070515" ID-Zenodo-Dep="8070515" httpUri="https://zenodo.org/record/8070515/files/figure.png" pageId="11" pageNumber="12" startId="11.[136,175,1897,1916]" targetBox="[238,1303,1064,1871]" targetPageId="11" targetType="figure">
|
||
<paragraph id="8BB636C5FFC4FF83B585F8E2FC9DEB84" blockId="11.[136,1413,1897,1972]" pageId="11" pageNumber="12">
|
||
<emphasis id="B97DEAD7FFC4FF83B585F8E2FF40EB4C" bold="true" box="[136,244,1897,1916]" pageId="11" pageNumber="12">Figs. 21–26.</emphasis>
|
||
Genitalia of
|
||
<taxonomicName id="4C094D46FFC4FF83B464F8E2FE13EB4C" authorityName="Philippi" authorityYear="1865" box="[361,423,1897,1916]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97DEAD7FFC4FF83B464F8E2FE13EB4C" box="[361,423,1897,1916]" italics="true" pageId="11" pageNumber="12">Holops</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C094D46FFC4FF83B4DCF8E2FDA6EB4C" authorityName="Cole" authorityYear="1918" box="[465,530,1897,1916]" class="Insecta" family="Acroceridae" genus="Villalus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97DEAD7FFC4FF83B4DCF8E2FDA6EB4C" box="[465,530,1897,1916]" italics="true" pageId="11" pageNumber="12">Villalus</emphasis>
|
||
</taxonomicName>
|
||
: (21) dorsal view of
|
||
<taxonomicName id="4C094D46FFC4FF83B7CBF8E1FCC2EB4C" authority="(Philippi)" baseAuthorityName="Philippi" box="[710,886,1897,1916]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cyanea">
|
||
<emphasis id="B97DEAD7FFC4FF83B7CBF8E1FCA8EB4C" box="[710,796,1898,1916]" italics="true" pageId="11" pageNumber="12">H. cyanea</emphasis>
|
||
(Philippi)
|
||
</taxonomicName>
|
||
, female; (22) dorsal view of
|
||
<taxonomicName id="4C094D46FFC4FF83B161F8E1FB61EB4C" authority="Barahona-Segovia & Guzmán & Pañinao-Monsálvez, 2021" authorityName="Barahona-Segovia & Guzmán & Pañinao-Monsálvez" authorityYear="2021" box="[1132,1237,1897,1916]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pullomen" status="sp. nov.">
|
||
<emphasis id="B97DEAD7FFC4FF83B161F8E1FB61EB4C" box="[1132,1237,1897,1916]" italics="true" pageId="11" pageNumber="12">H. pullomen</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57ACFFC4FF83B1D6F8E2FA95EB4C" box="[1243,1313,1897,1916]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (23) dorsal and lateral view of
|
||
<taxonomicName id="4C094D46FFC4FF83B43DF80DFE46EBA8" authority="Schiner" authorityName="Schiner" authorityYear="1868" box="[304,498,1925,1944]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="frauenfeldi">
|
||
<emphasis id="B97DEAD7FFC4FF83B43DF80DFE1DEBA8" box="[304,425,1925,1944]" italics="true" pageId="11" pageNumber="12">H. frauenfeldi</emphasis>
|
||
Schiner
|
||
</taxonomicName>
|
||
, male; (24) dorsal view of
|
||
<taxonomicName id="4C094D46FFC4FF83B7D1F80DFC34EBA8" authority="(Philippi)" baseAuthorityName="Philippi" baseAuthorityYear="1865" box="[732,896,1925,1944]" class="Insecta" family="Acroceridae" genus="Villalus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="inanis">
|
||
<emphasis id="B97DEAD7FFC4FF83B7D1F80DFC92EBA7" box="[732,806,1925,1944]" italics="true" pageId="11" pageNumber="12">V. inanis</emphasis>
|
||
(Philippi)
|
||
</taxonomicName>
|
||
, female; (25) lateral view of
|
||
<taxonomicName id="4C094D46FFC4FF83B171F80DFB52EBA8" authorityName="Barahona-Segovia & Guzmán & Pañinao-Monsálvez" authorityYear="2021" box="[1148,1254,1925,1944]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pullomen">
|
||
<emphasis id="B97DEAD7FFC4FF83B171F80DFB52EBA8" box="[1148,1254,1925,1944]" italics="true" pageId="11" pageNumber="12">H. pullomen</emphasis>
|
||
</taxonomicName>
|
||
, male; (26) dorsal view of gonocoxa of
|
||
<taxonomicName id="4C094D46FFC4FF83B430F829FE1CEB84" authority="Barahona-Segovia & Guzmán & Pañinao-Monsálvez, 2021" authorityName="Barahona-Segovia & Guzmán & Pañinao-Monsálvez" authorityYear="2021" box="[317,424,1954,1972]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="anarayae" status="sp. nov.">
|
||
<emphasis id="B97DEAD7FFC4FF83B430F829FE1CEB84" box="[317,424,1954,1972]" italics="true" pageId="11" pageNumber="12">H. anarayae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57ACFFC4FF83B4A0F82AFE44EB84" box="[429,496,1953,1972]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
epa = epandrium; gncx = gonocoxa.
|
||
</paragraph>
|
||
</caption>
|
||
<footnote id="E8122ACBFFC4FF83B585F86EFE2CEBC7" box="[136,408,2020,2039]" pageId="11" pageNumber="12">
|
||
<paragraph id="8BB636C5FFC4FF83B585F86EFE2CEBC7" blockId="11.[136,408,2020,2039]" box="[136,408,2020,2039]" pageId="11" pageNumber="12">© 2021 Academia Sinica, Taiwan</paragraph>
|
||
</footnote>
|
||
<paragraph id="8BB636C5FFC3FF85B682FE36FF58EE05" blockId="12.[850,1452,253,501]" lastBlockId="13.[136,737,253,981]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
|
||
<emphasis id="B97DEAD7FFC3FF84B682FE36FC72EDE5" box="[911,966,445,469]" italics="true" pageId="12" pageNumber="13">Male</emphasis>
|
||
:
|
||
<emphasis id="B97DEAD7FFC3FF84B6D8FE36FBBDEDE5" box="[981,1033,445,469]" italics="true" pageId="12" pageNumber="13">head</emphasis>
|
||
: similar to female;
|
||
<emphasis id="B97DEAD7FFC3FF84B1ECFE36FA93EDE5" box="[1249,1319,445,469]" italics="true" pageId="12" pageNumber="13">thorax</emphasis>
|
||
: postocellar lobe yellow; scutum and scutellum brownish with short yellowish pilosity; anterior spiracle yellow; superior part of anespisternum yellow and inferior brownish; katepisternum, anepimeron, meron, katatergite and anatergite brown (
|
||
<figureCitation id="13322A40FFC2FF85B440FED6FE65ED45" box="[333,465,349,373]" captionStart="Figs" captionStartId="14.[176,215,1924,1943]" captionTargetBox="[173,1451,951,1897]" captionTargetId="figure-352@14.[163,1451,941,1897]" captionTargetPageId="14" captionText="Figs. 33–36. Villalus inanis (Philippi, 1865): (33) female, lateral view, holotype, (34) female, dorsal view, holotype, (35) male, lateral view and (36) male, frontal view. Scale bars are not available." figureDoi="http://doi.org/10.5281/zenodo.8070521" httpUri="https://zenodo.org/record/8070521/files/figure.png" pageId="13" pageNumber="14">Figs. 35–36</figureCitation>
|
||
);
|
||
<emphasis id="B97DEAD7FFC2FF85B4EBFED6FD33ED45" box="[486,647,349,373]" italics="true" pageId="13" pageNumber="14">legs and wings</emphasis>
|
||
: similar to female;
|
||
<emphasis id="B97DEAD7FFC2FF85B40FFEF6FED2EDA5" box="[258,358,381,405]" italics="true" pageId="13" pageNumber="14">abdomen</emphasis>
|
||
: all tergites brown with long and yellow pilosity; sclerites yellowish; tergites 1–4 with same width; all sternites yellow with short pilosity of the same color.
|
||
<emphasis id="B97DEAD7FFC2FF85B43EFE56FE2CEDC5" box="[307,408,477,501]" italics="true" pageId="13" pageNumber="14">Genitalia</emphasis>
|
||
: not detached. However, cerci and epandrium have globose shape and yellow color (
|
||
<figureCitation id="13322A40FFC2FF85B59DFD96FF54EE05" box="[144,224,541,565]" captionStart="Figs" captionStartId="14.[176,215,1924,1943]" captionTargetBox="[173,1451,951,1897]" captionTargetId="figure-352@14.[163,1451,941,1897]" captionTargetPageId="14" captionText="Figs. 33–36. Villalus inanis (Philippi, 1865): (33) female, lateral view, holotype, (34) female, dorsal view, holotype, (35) male, lateral view and (36) male, frontal view. Scale bars are not available." figureDoi="http://doi.org/10.5281/zenodo.8070521" httpUri="https://zenodo.org/record/8070521/files/figure.png" pageId="13" pageNumber="14">Fig. 35</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF76664DFFC3FF84B5A2F82BFDF4EB83" ID-DOI="http://doi.org/10.5281/zenodo.8070517" ID-Zenodo-Dep="8070517" box="[175,576,1952,1971]" httpUri="https://zenodo.org/record/8070517/files/figure.png" pageId="12" pageNumber="13" startId="12.[175,206,1952,1971]" targetBox="[339,1292,552,1923]" targetPageId="12" targetType="figure">
|
||
<paragraph id="8BB636C5FFC3FF84B5A2F82BFDF4EB83" blockId="12.[175,576,1952,1971]" box="[175,576,1952,1971]" pageId="12" pageNumber="13">
|
||
<emphasis id="B97DEAD7FFC3FF84B5A2F82BFF46EB83" bold="true" box="[175,242,1952,1971]" pageId="12" pageNumber="13">Fig. 27.</emphasis>
|
||
Distributional map of
|
||
<taxonomicName id="4C094D46FFC3FF84B4B4F82BFDF4EB83" box="[441,576,1952,1971]" class="Insecta" family="Acroceridae" genus="Holops" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B97DEAD7FFC3FF84B4B4F82BFE43EB83" box="[441,503,1952,1971]" italics="true" pageId="12" pageNumber="13">Holops</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</caption>
|
||
<footnote id="E8122ACBFFC3FF84B191F86EFA1FEBC7" box="[1180,1451,2020,2039]" pageId="12" pageNumber="13">
|
||
<paragraph id="8BB636C5FFC3FF84B191F86EFA1FEBC7" blockId="12.[1180,1451,2020,2039]" box="[1180,1451,2020,2039]" pageId="12" pageNumber="13">© 2021 Academia Sinica, Taiwan</paragraph>
|
||
</footnote>
|
||
<paragraph id="8BB636C5FFC2FF85B5C8FDB6FE43EEA6" blockId="13.[136,737,253,981]" pageId="13" pageNumber="14">
|
||
<emphasis id="B97DEAD7FFC2FF85B5C8FDB6FEE5EE65" box="[197,337,573,597]" italics="true" pageId="13" pageNumber="14">Distribution</emphasis>
|
||
:
|
||
<collectingCountry id="F31E7655FFC2FF85B469FDB6FE11EE65" box="[356,421,573,597]" name="Chile" pageId="13" pageNumber="14">Chile</collectingCountry>
|
||
, from
|
||
<collectingRegion id="49CDF827FFC2FF85B4F9FDB6FD75EE65" box="[500,705,573,597]" country="Chile" name="Valparaiso" pageId="13" pageNumber="14">Valparaíso region</collectingRegion>
|
||
to
|
||
<collectingRegion id="49CDF827FFC2FF85B585FDD5FEF0EE45" box="[136,324,605,630]" country="Chile" name="Los Rios" pageId="13" pageNumber="14">Los Ríos region</collectingRegion>
|
||
and
|
||
<collectingCountry id="F31E7655FFC2FF85B48DFDD6FE45EE45" box="[384,497,605,629]" name="Argentina" pageId="13" pageNumber="14">Argentina</collectingCountry>
|
||
(first record; Shaun Winterton, pers. comm.) (
|
||
<figureCitation id="13322A40FFC2FF85B496FDF6FE5FEEA6" box="[411,491,637,662]" captionStart="Fig" captionStartId="15.[136,167,1954,1973]" captionTargetBox="[143,1406,932,1924]" captionTargetId="figure-317@15.[125,1416,930,1928]" captionTargetPageId="15" captionText="Fig. 37. Distributional map of Villalus inanis (Philippi, 1865)." figureDoi="http://doi.org/10.5281/zenodo.8070523" httpUri="https://zenodo.org/record/8070523/files/figure.png" pageId="13" pageNumber="14">Fig. 37</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB636C5FFC2FF85B5C8FD16FAA7ED45" blockId="13.[136,737,253,981]" lastBlockId="13.[813,1414,253,822]" pageId="13" pageNumber="14">
|
||
<emphasis id="B97DEAD7FFC2FF85B5C8FD16FE9EEE85" box="[197,298,669,693]" italics="true" pageId="13" pageNumber="14">Remarks</emphasis>
|
||
: native species distributed from the Santiagan province in central
|
||
<collectingCountry id="F31E7655FFC2FF85B4F0FD36FD89EEE5" box="[509,573,701,725]" name="Chile" pageId="13" pageNumber="14">Chile</collectingCountry>
|
||
subregion to Valdivian forest province in subantarctic subregion (
|
||
<bibRefCitation id="EF984B34FFC2FF85B582FD75FE85EF25" author="Morrone JJ" box="[143,305,765,790]" pageId="13" pageNumber="14" pagination="207 - 236" refId="ref11905" refString="Morrone JJ. 2015. Biogeographical regionalisation of the Andean region. Zootaxa 3936: 207 - 236. doi: 10.11646 / zootaxa. 3936.2.3." type="journal article" year="2015">Morrone 2015</bibRefCitation>
|
||
). We extend the historical distributional range
|
||
<quantity id="4CF19B20FFC2FF85B5C0FC95FE94EF05" box="[205,288,797,822]" metricMagnitude="5" metricUnit="m" metricValue="2.2399999999999998" pageId="13" pageNumber="14" unit="km" value="224.0">224 km</quantity>
|
||
to north from Valdivia (
|
||
<typeStatus id="54B28867FFC2FF85B73DFC95FDD4EF06" box="[560,608,798,822]" pageId="13" pageNumber="14">type</typeStatus>
|
||
locality) to Nahuelbuta National Park and we fill the gaps in the La
|
||
<collectingRegion id="49CDF827FFC2FF85B5A1FCD5FEDEEF45" box="[172,362,861,886]" country="Chile" name="Araucania" pageId="13" pageNumber="14">Araucanía region</collectingRegion>
|
||
and also, we extend in
|
||
<quantity id="4CF19B20FFC2FF85B774FCD6FD55EF45" box="[633,737,861,885]" metricMagnitude="4" metricUnit="m" metricValue="6.583" pageId="13" pageNumber="14" unit="km" value="65.83">65.83 km</quantity>
|
||
southeast from Valdivia to San Pablo,
|
||
<collectingRegion id="49CDF827FFC2FF85B728FCF5FD55EFA5" box="[549,737,893,918]" country="Chile" name="Los Lagos" pageId="13" pageNumber="14">Los Lagos region</collectingRegion>
|
||
(new distribution record). This species has been recently observed associated to semi-rural environments thanks to the citizen science program (
|
||
<figureCitation id="13322A40FFC2FF85B19EFF76FAA9ED26" box="[1171,1309,253,278]" captionStart="Figs" captionStartId="13.[137,177,1924,1943]" captionTargetBox="[136,1413,1027,1898]" captionTargetId="figure-369@13.[136,1413,1027,1898]" captionTargetPageId="13" captionText="Figs. 28–32. Villalus inanis (Philippi, 1865): (28) dorsal view; (29) lateral view; (30) in situ, lateral view (photo: William Tarrant); (31) in situ, dorsal view (photographed by Ignacio Valeria), and (33) dead, ventral view (photographed by Claudia Veas). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8070519" httpUri="https://zenodo.org/record/8070519/files/figure.png" pageId="13" pageNumber="14">Figs. 30–32</figureCitation>
|
||
).
|
||
<taxonomicName id="4C094D46FFC2FF85B03FFF76FCC5ED05" baseAuthorityName="Philippi" baseAuthorityYear="1865" class="Insecta" family="Acroceridae" genus="Villalus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="inanis">
|
||
<emphasis id="B97DEAD7FFC2FF85B03FFF76FCC5ED05" italics="true" pageId="13" pageNumber="14">Villalus inanis</emphasis>
|
||
</taxonomicName>
|
||
seems to be mainly distributed by the coastal forest which has been highly fragmented by forestry, urbanization and crops. The host is unknown.
|
||
</paragraph>
|
||
<paragraph id="8BB636C5FFC2FF85B664FEF6FB99EE85" blockId="13.[813,1414,253,822]" pageId="13" pageNumber="14">
|
||
The female
|
||
<typeStatus id="54B28867FFC2FF85B6FAFEF6FBEFEDA5" box="[1015,1115,381,405]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C094D46FFC2FF85B185FEF5FAB9EDA5" authorityName="Cole" authorityYear="1918" box="[1160,1293,381,405]" class="Insecta" family="Acroceridae" genus="Villalus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="chilensis">
|
||
<emphasis id="B97DEAD7FFC2FF85B185FEF5FAB9EDA5" box="[1160,1293,381,405]" italics="true" pageId="13" pageNumber="14">V. chilensis</emphasis>
|
||
</taxonomicName>
|
||
described by
|
||
<bibRefCitation id="EF984B34FFC2FF85B65EFE16FC6DED86" author="Cole FR" box="[851,985,413,438]" pageId="13" pageNumber="14" pagination="61 - 64" refId="ref11556" refString="Cole FR. 1918. A new genus of Cyrtidae from South America (Dip.). Entomol News 29: 61 - 64." type="journal article" year="1918">Cole (1918</bibRefCitation>
|
||
;
|
||
<figureCitation id="13322A40FFC2FF85B6E5FE16FBC3ED86" box="[1000,1143,413,438]" captionStart="Figs" captionStartId="14.[176,215,1924,1943]" captionTargetBox="[173,1451,951,1897]" captionTargetId="figure-352@14.[163,1451,941,1897]" captionTargetPageId="14" captionText="Figs. 33–36. Villalus inanis (Philippi, 1865): (33) female, lateral view, holotype, (34) female, dorsal view, holotype, (35) male, lateral view and (36) male, frontal view. Scale bars are not available." figureDoi="http://doi.org/10.5281/zenodo.8070521" httpUri="https://zenodo.org/record/8070521/files/figure.png" pageId="13" pageNumber="14">Figs. 33–34</figureCitation>
|
||
), looks as all
|
||
<taxonomicName id="4C094D46FFC2FF85B02DFE15FA32ED85" baseAuthorityName="Philippi" baseAuthorityYear="1865" box="[1312,1414,414,437]" class="Insecta" family="Acroceridae" genus="Villalus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="inanis">
|
||
<emphasis id="B97DEAD7FFC2FF85B02DFE15FA32ED85" box="[1312,1414,414,437]" italics="true" pageId="13" pageNumber="14">V. inanis</emphasis>
|
||
</taxonomicName>
|
||
examined for this work and therefore both species were considered the same. All morphological characteristics, especially wing veins are completely similar to specimens reviewed. According with Shaun Winterton (pers. comm.),
|
||
<specimenCount id="9D0FFD4CFFC2FF85B6DCFDB5FB8BEE65" box="[977,1087,573,597]" pageId="13" pageNumber="14" type="male">two males</specimenCount>
|
||
of this species possess sexual dimorphism because they are smaller than females with many brown (mainly thorax and tergites) and yellow segments (
|
||
<figureCitation id="13322A40FFC2FF85B693FD16FB95EE85" box="[926,1057,669,693]" captionStart="Figs" captionStartId="14.[176,215,1924,1943]" captionTargetBox="[173,1451,951,1897]" captionTargetId="figure-352@14.[163,1451,941,1897]" captionTargetPageId="14" captionText="Figs. 33–36. Villalus inanis (Philippi, 1865): (33) female, lateral view, holotype, (34) female, dorsal view, holotype, (35) male, lateral view and (36) male, frontal view. Scale bars are not available." figureDoi="http://doi.org/10.5281/zenodo.8070521" httpUri="https://zenodo.org/record/8070521/files/figure.png" pageId="13" pageNumber="14">Figs. 35–36</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB636C5FFC2FF85B664FD36FAF2EF06" blockId="13.[813,1414,253,822]" pageId="13" pageNumber="14">
|
||
<emphasis id="B97DEAD7FFC2FF85B664FD36FB94EEE5" box="[873,1056,701,725]" italics="true" pageId="13" pageNumber="14">Curatorial note</emphasis>
|
||
: According to
|
||
<bibRefCitation id="EF984B34FFC2FF85B1D8FD36FCC0EEC6" author="Gonzalez CR & Elgueta M & Ramirez F." pageId="13" pageNumber="14" pagination="427 - 440" refId="ref11831" refString="Gonzalez CR, Elgueta M, Ramirez F. 2018. A catalog of Acroceridae (Diptera) from Chile. Zootaxa 4374: 427 - 440. doi: 10.11646 / zootaxa. 4374.3.6." type="journal article" year="2018">González et al. (2018)</bibRefCitation>
|
||
the type specimen is not deposited in MNHNCL. Currently, the
|
||
<typeStatus id="54B28867FFC2FF85B6C7FD76FB93EF25" box="[970,1063,765,789]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
is in the California Department of Food and Agriculture and on loan from USNM.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |