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<document id="4594C8DFD0443D3ED25831CB71054F6B" ID-CLB-Dataset="4646" ID-DOI="10.1515/aemnp-2017-0074" ID-GBIF-Dataset="89a4f3b6-aa27-4ae2-b818-66024ba993d6" ID-ISSN="0374-1036" ID-Zenodo-Dep="5378027" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630131332669" checkinUser="marcus" docAuthor="Weidenhoffer, Zdeněk &amp; Šumpich, Jan" docDate="2017" docId="039E4266FFAAFF9E871BAB70FF5FFC10" docLanguage="en" docName="ActaEntMusNatPra.2017.57.1.275-278.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 57 (1)" docSource="https://www.aemnp.eu/acta-entomologica/volume-57-1/1679/neolycaena-rhymnaria-arcana-sp-nov-from-tibet-lepidoptera-lycaenidae.html" docStyle="DocumentStyle:60E99616ED1F2F5EE637DA5E5309A4D2.7:ActaEntMusNatPra.2017.journal_article" docStyleId="60E99616ED1F2F5EE637DA5E5309A4D2" docStyleName="ActaEntMusNatPra.2017.journal_article" docStyleVersion="7" docTitle="Neolycaena (Rhymnaria) arcana Weidenhoffer &amp; Šumpich 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="278" masterDocId="FFA73A1EFFABFF9D866DA849FFE3FFA2" masterDocTitle="Neolycaena (Rhymnaria) arcana sp. nov. from Tibet (Lepidoptera: Lycaenidae)" masterLastPageNumber="278" masterPageNumber="275" pageNumber="276" updateTime="1699166944546" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="401E8D4338B9422D406DFFF5305C0F93" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FB180E131B2D87AB2AF86265F690B5B6">
<mods:title id="03F25D133C083A676C22462DB927FCD0">Neolycaena (Rhymnaria) arcana sp. nov. from Tibet (Lepidoptera: Lycaenidae)</mods:title>
</mods:titleInfo>
<mods:name id="5DCB7BB83A14679F5D523A6621D9118D" type="personal">
<mods:role id="52CFF00387116516631C1EE1DF4EF732">
<mods:roleTerm id="86273EB17E987C5B107B413FB65611BF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8A323CF0A69323D966AB8BBE89AD05FA">Weidenhoffer, Zdeněk</mods:namePart>
<mods:affiliation id="3BFE800A83E41E724833C14FA14ED23A">) &amp;) Praha 10, Vyžlovská 36, CZ- 100 00, Czech Republic; e-mail: wff @ upcmail. cz</mods:affiliation>
</mods:name>
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<mods:role id="1B6953F5B32C389E9702C10BA7C9799E">
<mods:roleTerm id="7B029BF3F3E51C5BD664714454FEA9D2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="154B9ECF1FDC30325129CE64B375BCCC">Šumpich, Jan</mods:namePart>
<mods:affiliation id="4B48209B4072ED3E0ACE761831EFFD35">) &amp;) Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic; e-mail: jansumpich @ seznam. cz</mods:affiliation>
</mods:name>
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<mods:title id="E274F2AB959F84280082D7522153CE85">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="A57D9C246D9556AD40FA36F261C9E5E8">
<mods:date id="539FEF9C46AB0C01311923FCCAF9A1C7">2017</mods:date>
<mods:detail id="F05805150C8E9452BEC59E2AECDC5E59" type="pubDate">
<mods:number id="5DC99261856D2AF5751F2AAABF7E20D0">2017-06-30</mods:number>
</mods:detail>
<mods:detail id="04798F052F3605409DE364F3636A25A8" type="volume">
<mods:number id="0506D4BAD51309543F00F873B132C0FC">57</mods:number>
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<mods:number id="AB1A28BDC9D117B4F4787D57F02872DF">1</mods:number>
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<mods:start id="912865D2803A2A9A4025218E1EA49DD1">275</mods:start>
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<mods:location id="7B8B3036D6C5FE351257799016FB61E6">
<mods:url id="9A6C6E1E8891A86F638F6CA5FF50984A">https://www.aemnp.eu/acta-entomologica/volume-57-1/1679/neolycaena-rhymnaria-arcana-sp-nov-from-tibet-lepidoptera-lycaenidae.html</mods:url>
</mods:location>
<mods:classification id="24AF83FF5E91734CCE4951E147E28144">journal article</mods:classification>
<mods:identifier id="408D05003A12A8831D5BEA8B68C1B52C" type="CLB-Dataset">4646</mods:identifier>
<mods:identifier id="6C0A4CA838531648609B8BA70C3C91E5" type="DOI">10.1515/aemnp-2017-0074</mods:identifier>
<mods:identifier id="9F24AAD1251A5E26A7E96B708C416EA0" type="GBIF-Dataset">89a4f3b6-aa27-4ae2-b818-66024ba993d6</mods:identifier>
<mods:identifier id="6E29321C527EC1663C2F82FFC34F58B2" type="ISSN">0374-1036</mods:identifier>
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<mods:identifier id="F2C9044ABD9E6B067A4FF817AE766F0C" type="ZooBank">F45E0B46-4538-4F41-8336-971105422A61</mods:identifier>
</mods:mods>
<treatment id="039E4266FFAAFF9E871BAB70FF5FFC10" ID-DOI="http://doi.org/10.5281/zenodo.5335483" ID-GBIF-Taxon="185200975" ID-Zenodo-Dep="5335483" LSID="urn:lsid:plazi:treatment:039E4266FFAAFF9E871BAB70FF5FFC10" httpUri="http://treatment.plazi.org/id/039E4266FFAAFF9E871BAB70FF5FFC10" lastPageId="3" lastPageNumber="278" pageId="1" pageNumber="276">
<subSubSection id="C32DA0FBFFAAFF9C871BAB70FCDFFCF3" box="[374,828,825,849]" pageId="1" pageNumber="276" type="nomenclature">
<paragraph id="8B88F370FFAAFF9C871BAB70FCDFFCF3" blockId="1.[374,828,825,849]" box="[374,828,825,849]" pageId="1" pageNumber="276">
<heading id="D0C0441CFFAAFF9C871BAB70FCDFFCF3" bold="true" box="[374,828,825,849]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="276" reason="3">
<emphasis id="B9432F62FFAAFF9C871BAB70FCDFFCF3" bold="true" box="[374,828,825,849]" pageId="1" pageNumber="276">
<taxonomicName id="4C3788F3FFAAFF9C871BAB70FD03FCF3" authority="Weidenhoffer &amp; Šumpich, 2017" authorityName="Weidenhoffer &amp; Šumpich" authorityYear="2017" box="[374,736,825,849]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="276" phylum="Arthropoda" rank="species" species="arcana" status="sp. nov." subGenus="Rhymnaria">
<emphasis id="B9432F62FFAAFF9C871BAB70FE14FCF3" bold="true" box="[374,503,825,849]" italics="true" pageId="1" pageNumber="276">Neolycaena</emphasis>
(
<emphasis id="B9432F62FFAAFF9C846AAB70FD60FCF3" bold="true" box="[519,643,825,849]" italics="true" pageId="1" pageNumber="276">Rhymnaria</emphasis>
)
<emphasis id="B9432F62FFAAFF9C84FEAB73FD03FCF3" bold="true" box="[659,736,826,849]" italics="true" pageId="1" pageNumber="276">arcana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2709219FFAAFF9C848BAB73FCDFFCF3" box="[742,828,826,849]" pageId="1" pageNumber="276" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32DA0FBFFAAFF9C8440AB11FD66FCC9" box="[557,645,856,875]" pageId="1" pageNumber="276" type="description">
<paragraph id="8B88F370FFAAFF9C8440AB11FD66FCC9" blockId="1.[557,645,856,875]" box="[557,645,856,875]" pageId="1" pageNumber="276">
(
<figureCitation id="130CEFF5FFAAFF9C845EAB11FD9DFCC9" box="[563,638,856,875]" captionStart="Figs 1–7" captionStartId="2.[151,187,1519,1538]" captionTargetBox="[162,1115,226,1462]" captionTargetId="figure-10@2.[162,1115,219,1480]" captionTargetPageId="2" captionText="Figs 1–7. Neolycaena (Rhymnaria) arcana sp. nov.1–2 – male (holotype):1 – dorsal view; 2 – ventral view;3 – original label; 4–7 – male genitalia: 4 – ventral view; 5 – lateral view; 6 – detail of spine in distal part of valvae; 7 – aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5378031" httpUri="https://zenodo.org/record/5378031/files/figure.png" pageId="1" pageNumber="276">Figs 1–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32DA0FBFFAAFF9C861CABCBFE21FC69" pageId="1" pageNumber="276" type="materials_examined">
<paragraph id="8B88F370FFAAFF9C861CABCBFDD7FC37" blockId="1.[113,1089,898,971]" box="[113,564,898,917]" pageId="1" pageNumber="276">
<emphasis id="B9432F62FFAAFF9C861CABCBFF09FC37" bold="true" box="[113,234,898,917]" pageId="1" pageNumber="276">
<typeStatus id="548C4DD2FFAAFF9C861CABCBFF7DFC37" box="[113,158,898,917]" pageId="1" pageNumber="276">Type</typeStatus>
locality.
</emphasis>
<collectingRegion id="49F33D92FFAAFF9C8682ABCBFEF8FC37" box="[239,283,898,917]" country="China" name="Xizang" pageId="1" pageNumber="276">Tibet</collectingRegion>
(in its wider, pre-1950 concept).
</paragraph>
<paragraph id="8B88F370FFAAFF9C861CABD4FE21FC69" blockId="1.[113,1089,898,971]" pageId="1" pageNumber="276">
<emphasis id="B9432F62FFAAFF9C861CABD4FF10FC12" bold="true" box="[113,243,925,944]" pageId="1" pageNumber="276">Type material.</emphasis>
<typeStatus id="548C4DD2FFAAFF9C869BABD4FEABFC0C" box="[246,328,925,944]" pageId="1" pageNumber="276" type="holotype">HOLOTYPE</typeStatus>
: J, ‘
<collectingRegion id="49F33D92FFAAFF9C8700ABD4FE7CFC12" box="[365,415,925,944]" country="China" name="Xizang" pageId="1" pageNumber="276">Tibet</collectingRegion>
. [handwritten]’, ‘HolotypusJ /
<taxonomicName id="4C3788F3FFAAFF9C84CAABD4FC9AFC12" authority="" authorityName="Zhdanko" authorityYear="1983" box="[679,889,925,944]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="276" phylum="Arthropoda" rank="subGenus" subGenus="Rhymnaria">
<emphasis id="B9432F62FFAAFF9C84CAABD4FC9AFC12" box="[679,889,925,944]" italics="true" pageId="1" pageNumber="276">Neolycaena (Rhymnaria)</emphasis>
</taxonomicName>
/
<taxonomicName id="4C3788F3FFAAFF9C85EBABD4FC22FC12" authorityName="Weidenhoffer &amp; Šumpich" authorityYear="2017" box="[902,961,925,944]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="276" phylum="Arthropoda" rank="species" species="arcana">
<emphasis id="B9432F62FFAAFF9C85EBABD4FC22FC12" box="[902,961,925,944]" italics="true" pageId="1" pageNumber="276">arcana</emphasis>
</taxonomicName>
/ Weidenhoffer &amp; Šumpich, 2017 [red label]’ (
<collectionCode id="ED266BB5FFAAFF9C871AABF1FE5BFC69" box="[375,440,952,971]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="276">NMPC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C32DA0FBFFAAFF9E861CABAAFE2BFEF0" lastPageId="3" lastPageNumber="278" pageId="1" pageNumber="276" type="description">
<paragraph id="8B88F370FFAAFF9C861CABAAFCA2FA78" blockId="1.[113,1089,994,1562]" pageId="1" pageNumber="276">
<emphasis id="B9432F62FFAAFF9C861CABAAFEDDFC58" bold="true" box="[113,318,994,1018]" pageId="1" pageNumber="276">
Description.
<emphasis id="B9432F62FFAAFF9C876CABABFEDDFC58" bold="true" box="[257,318,994,1018]" italics="true" pageId="1" pageNumber="276">Adult</emphasis>
</emphasis>
(
<figureCitation id="130CEFF5FFAAFF9C8726ABAAFE4FFC58" box="[331,428,995,1018]" captionStart="Figs 1–7" captionStartId="2.[151,187,1519,1538]" captionTargetBox="[162,1115,226,1462]" captionTargetId="figure-10@2.[162,1115,219,1480]" captionTargetPageId="2" captionText="Figs 1–7. Neolycaena (Rhymnaria) arcana sp. nov.1–2 – male (holotype):1 – dorsal view; 2 – ventral view;3 – original label; 4–7 – male genitalia: 4 – ventral view; 5 – lateral view; 6 – detail of spine in distal part of valvae; 7 – aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5378031" httpUri="https://zenodo.org/record/5378031/files/figure.png" pageId="1" pageNumber="276">Figs 1–2</figureCitation>
). Forewing length (from apex to point of attachment to thorax)
<quantity id="4CCF5E95FFAAFF9C861CAC4AFF32FBB8" box="[113,209,1027,1050]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="1" pageNumber="276" unit="mm" value="14.6">14.6 mm</quantity>
, length/width ratio 1: 0.6. Ground colour of upperside of wings brown, slightly paler in basal and apical part of forewing and anal part of hindwing. Weakly visible dark brown row of spots in submarginal part of hindwings. Narrow marginal line of both wings blackish brown, fringes brown. Underside of forewing dark brown, lighter in costal and marginal part of wings; narrow marginal line and fringes dark brown, white antemarginal line absent, submarginal dark spots indistinctive. White postdiscal band not reaching inner margin but well developed and formed by row of crescent spots. Wing drawings completed by thin white line at discocellular vein. Underside of hindwing in basal and discal parts light brown, in postdiscal part even paler. Marginal line and fringes dark brown, white antemarginal line weakly developed. Submarginal row formed by twofold row of rounded black spots joined with inconspicuous yellow link; inner spots of twofold row markedly larger than outer spots. White postdiscal band very prominent and formed by wide crescent spots bordered by blackish brown crescents from inner side. Regular bow of postdiscal spot interrupted only in field 6; this spot markedly shifted to base of wing. Discocellular vein white-bordered. Antennal club ventrally black with ochreous tip, dorsally fully ochreous-coloured.
</paragraph>
<paragraph id="8B88F370FFAAFF9E86FDADABFE7CFE90" blockId="1.[113,1089,994,1562]" lastBlockId="3.[113,1089,219,946]" lastPageId="3" lastPageNumber="278" pageId="1" pageNumber="276">
<emphasis id="B9432F62FFAAFF9C86FDADABFECDFA58" bold="true" box="[144,302,1506,1530]" italics="true" pageId="1" pageNumber="276">Male genitalia</emphasis>
(
<figureCitation id="130CEFF5FFAAFF9C8751ADAAFE7DFA58" box="[316,414,1507,1530]" captionStart="Figs 1–7" captionStartId="2.[151,187,1519,1538]" captionTargetBox="[162,1115,226,1462]" captionTargetId="figure-10@2.[162,1115,219,1480]" captionTargetPageId="2" captionText="Figs 1–7. Neolycaena (Rhymnaria) arcana sp. nov.1–2 – male (holotype):1 – dorsal view; 2 – ventral view;3 – original label; 4–7 – male genitalia: 4 – ventral view; 5 – lateral view; 6 – detail of spine in distal part of valvae; 7 – aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5378031" httpUri="https://zenodo.org/record/5378031/files/figure.png" pageId="1" pageNumber="276">Figs 4–7</figureCitation>
). In ventral projection valvae distally terminated by short transparent spine, basal part of valvae equal to three fifth of total length, narrowing distally. Saccus well developed, laterally curved. Falces long, arcuately saggy, distally narrowed till spear shaped ending. Saccus well developed, in lateral projection curved. Aedeagus with two long cornuti and sclerotized keel.
</paragraph>
<caption id="DF48A3F8FFA9FF9F86FAADA6FB8BF9BF" ID-DOI="http://doi.org/10.5281/zenodo.5378031" ID-Zenodo-Dep="5378031" httpUri="https://zenodo.org/record/5378031/files/figure.png" pageId="2" pageNumber="277" startId="2.[151,187,1519,1538]" targetBox="[162,1115,226,1462]" targetPageId="2">
<paragraph id="8B88F370FFA9FF9F86FAADA6FB8BF9BF" blockId="2.[151,1128,1519,1565]" pageId="2" pageNumber="277">
Figs 1–7.
<taxonomicName id="4C3788F3FFA9FF9F868AADA6FE16F9A0" authority="Weidenhoffer &amp; Šumpich, 2017" authorityName="Weidenhoffer &amp; Šumpich" authorityYear="2017" box="[231,501,1519,1538]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="277" phylum="Arthropoda" rank="species" species="arcana" status="sp. nov." subGenus="Rhymnaria">
<emphasis id="B9432F62FFA9FF9F868AADA6FEA9F9A0" box="[231,330,1519,1538]" italics="true" pageId="2" pageNumber="277">Neolycaena</emphasis>
(
<emphasis id="B9432F62FFA9FF9F8739ADA6FE52F9A0" box="[340,433,1519,1538]" italics="true" pageId="2" pageNumber="277">Rhymnaria</emphasis>
)
<emphasis id="B9432F62FFA9FF9F87D6ADA6FE16F9A0" box="[443,501,1519,1538]" italics="true" pageId="2" pageNumber="277">arcana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2709219FFA9FF9F8794ADA6FDDAF9A0" box="[505,569,1519,1538]" pageId="2" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
1–2 – male (holotype):1 – dorsal view; 2 – ventral view;3 – original label; 4–7 – male genitalia: 4 – ventral view; 5 – lateral view; 6 – detail of spine in distal part of valvae; 7 – aedeagus.
</paragraph>
</caption>
<paragraph id="8B88F370FFA8FF9E86FDA973FE2BFEF0" blockId="3.[113,1089,219,946]" box="[144,456,314,338]" pageId="3" pageNumber="278">
<emphasis id="B9432F62FFA8FF9E86FDA973FEB2FEF0" bold="true" box="[144,337,314,338]" pageId="3" pageNumber="278">
<emphasis id="B9432F62FFA8FF9E86FDA973FEAAFEF0" bold="true" box="[144,329,314,338]" italics="true" pageId="3" pageNumber="278">Female genitalia</emphasis>
.
</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32DA0FBFFA8FF9E861CA912FC42FD70" pageId="3" pageNumber="278" type="diagnosis">
<paragraph id="8B88F370FFA8FF9E861CA912FC42FD70" blockId="3.[113,1089,219,946]" pageId="3" pageNumber="278">
<emphasis id="B9432F62FFA8FF9E861CA912FE88FED0" bold="true" box="[113,363,347,370]" pageId="3" pageNumber="278">Differential diagnosis.</emphasis>
Only two species (
<taxonomicName id="4C3788F3FFA8FF9E8459A912FCB7FED0" authority="Zhdanko, 1994" authorityName="Zhdanko" authorityYear="1994" box="[564,852,347,370]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="aeto" subGenus="Rhymnaria">
<emphasis id="B9432F62FFA8FF9E8459A912FDAEFED0" box="[564,589,347,370]" italics="true" pageId="3" pageNumber="278">N.</emphasis>
(
<emphasis id="B9432F62FFA8FF9E8436A912FD91FED0" box="[603,626,347,370]" italics="true" pageId="3" pageNumber="278">R.</emphasis>
)
<emphasis id="B9432F62FFA8FF9E84EDA912FD4DFED0" box="[640,686,347,370]" italics="true" pageId="3" pageNumber="278">aeto</emphasis>
Zhdanko, 1994
</taxonomicName>
and
<taxonomicName id="4C3788F3FFA8FF9E85EBA912FBFEFED0" authority="Weidenhoffer &amp; Šumpich, 2017" authorityName="Weidenhoffer &amp; Šumpich" authorityYear="2017" box="[902,1053,347,370]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="arcana" status="sp. nov." subGenus="Rhymnaria">
<emphasis id="B9432F62FFA8FF9E85EBA912FC7CFED0" box="[902,927,347,370]" italics="true" pageId="3" pageNumber="278">N.</emphasis>
(
<emphasis id="B9432F62FFA8FF9E85C0A912FC27FED0" box="[941,964,347,370]" italics="true" pageId="3" pageNumber="278">R.</emphasis>
)
<emphasis id="B9432F62FFA8FF9E85BFA912FBFEFED0" box="[978,1053,347,370]" italics="true" pageId="3" pageNumber="278">arcana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2709219FFA8FF9E824FA912FF43FE30" pageId="3" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
) from the whole subgenus
<taxonomicName id="4C3788F3FFA8FF9E87D2A932FDD5FE30" authorityName="Zhdanko" authorityYear="1983" box="[447,566,379,402]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="subGenus" subGenus="Rhymnaria">
<emphasis id="B9432F62FFA8FF9E87D2A932FDD5FE30" box="[447,566,379,402]" italics="true" pageId="3" pageNumber="278">Rhymnaria</emphasis>
</taxonomicName>
have a white row of postdiscal spots in the shape of broad crescents on the underside of hindwings. All other species of the subgenus have postdiscal spots in the form of white lines or white dots. From
<taxonomicName id="4C3788F3FFA8FF9E84BFA9F2FCAFFE70" authorityName="Zhdanko" authorityYear="1994" box="[722,844,443,466]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="aeto" subGenus="Rhymnaria">
<emphasis id="B9432F62FFA8FF9E84BFA9F2FD08FE70" box="[722,747,443,466]" italics="true" pageId="3" pageNumber="278">N.</emphasis>
(
<emphasis id="B9432F62FFA8FF9E8494A9F2FCF3FE70" box="[761,784,443,466]" italics="true" pageId="3" pageNumber="278">R.</emphasis>
)
<emphasis id="B9432F62FFA8FF9E8573A9F2FCAFFE70" box="[798,844,443,466]" italics="true" pageId="3" pageNumber="278">aeto</emphasis>
</taxonomicName>
the new species differs in having dark brown crescents proximally adjacent to the white ones; in twofold submarginal row of black spots with the internal spots always significantly larger than the external ones (same size by
<taxonomicName id="4C3788F3FFA8FF9E8767AA52FE65FD90" authorityName="Zhdanko" authorityYear="1994" box="[266,390,539,562]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="aeto" subGenus="Rhymnaria">
<emphasis id="B9432F62FFA8FF9E8767AA52FEC0FD90" box="[266,291,539,562]" italics="true" pageId="3" pageNumber="278">N.</emphasis>
(
<emphasis id="B9432F62FFA8FF9E875FAA52FEAAFD90" box="[306,329,539,562]" italics="true" pageId="3" pageNumber="278">R.</emphasis>
)
<emphasis id="B9432F62FFA8FF9E8735AA52FE65FD90" box="[344,390,539,562]" italics="true" pageId="3" pageNumber="278">aeto</emphasis>
</taxonomicName>
); in the length of forewing (
<quantity id="4CCF5E95FFA8FF9E84DBAA52FCCAFD90" box="[694,809,539,562]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="3" pageNumber="278" unit="mm" value="17.5" valueMax="18.0" valueMin="17.0">17–18 mm</quantity>
at
<taxonomicName id="4C3788F3FFA8FF9E8527AA52FC24FD90" authorityName="Zhdanko" authorityYear="1994" box="[842,967,539,562]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="aeto" subGenus="Rhymnaria">
<emphasis id="B9432F62FFA8FF9E8527AA52FC87FD90" box="[842,868,539,562]" italics="true" pageId="3" pageNumber="278">N.</emphasis>
(
<emphasis id="B9432F62FFA8FF9E851EAA52FC69FD90" box="[883,906,539,562]" italics="true" pageId="3" pageNumber="278">R.</emphasis>
)
<emphasis id="B9432F62FFA8FF9E85F4AA52FC24FD90" box="[921,967,539,562]" italics="true" pageId="3" pageNumber="278">aeto</emphasis>
</taxonomicName>
). The morphology of the male genitalia, especially the apical part of valvae (either rounded or sharply pointed) has a diagnostic relevance at a species level as well.
<taxonomicName id="4C3788F3FFA8FF9E849DAA12FC0FFDD0" authority="Weidenhoffer &amp; Šumpich, 2017" authorityName="Weidenhoffer &amp; Šumpich" authorityYear="2017" box="[752,1004,603,626]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="arcana" status="sp. nov." subGenus="Rhymnaria">
<emphasis id="B9432F62FFA8FF9E849DAA12FC8DFDD0" box="[752,878,603,626]" italics="true" pageId="3" pageNumber="278">Neolycaena</emphasis>
(
<emphasis id="B9432F62FFA8FF9E8511AA12FC70FDD0" box="[892,915,603,626]" italics="true" pageId="3" pageNumber="278">R.</emphasis>
)
<emphasis id="B9432F62FFA8FF9E85CCAA12FC0FFDD0" box="[929,1004,603,626]" italics="true" pageId="3" pageNumber="278">arcana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2709219FFA8FF9E859DAA12FBA2FDD0" box="[1008,1089,603,626]" pageId="3" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
together with seven other species (
<taxonomicName id="4C3788F3FFA8FF9E8788AA32FD80FD30" authorityName="Zhdanko" authorityYear="1994" box="[485,611,635,658]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="aeto" subGenus="Rhymnaria">
<emphasis id="B9432F62FFA8FF9E8788AA32FE1DFD30" box="[485,510,635,658]" italics="true" pageId="3" pageNumber="278">N.</emphasis>
(
<emphasis id="B9432F62FFA8FF9E8463AA32FDC6FD30" box="[526,549,635,658]" italics="true" pageId="3" pageNumber="278">R.</emphasis>
)
<emphasis id="B9432F62FFA8FF9E8458AA32FD80FD30" box="[565,611,635,658]" italics="true" pageId="3" pageNumber="278">aeto</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3788F3FFA8FF9E841DAA32FCE0FD30" authorityName="Oberthur" authorityYear="1881" box="[624,771,635,658]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="davidi" subGenus="Rhymnaria">
<emphasis id="B9432F62FFA8FF9E841DAA32FD69FD30" box="[624,650,635,658]" italics="true" pageId="3" pageNumber="278">N.</emphasis>
(
<emphasis id="B9432F62FFA8FF9E84F4AA32FD53FD30" box="[665,688,635,658]" italics="true" pageId="3" pageNumber="278">R.</emphasis>
)
<emphasis id="B9432F62FFA8FF9E84ADAA32FCE0FD30" box="[704,771,635,658]" italics="true" pageId="3" pageNumber="278">davidi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3788F3FFA8FF9E857DAA32FEA6FD10" authority="Zhdanko &amp; Korshunov, 1985" authorityName="Zhdanko &amp; Korshunov" authorityYear="1985" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="falkovitschi" subGenus="Rhymnaria">
<emphasis id="B9432F62FFA8FF9E857DAA32FCC9FD30" box="[784,810,635,658]" italics="true" pageId="3" pageNumber="278">N.</emphasis>
(
<emphasis id="B9432F62FFA8FF9E8554AA32FCB3FD30" box="[825,848,635,658]" italics="true" pageId="3" pageNumber="278">R.</emphasis>
)
<emphasis id="B9432F62FFA8FF9E850DAA32FC38FD30" box="[864,987,635,658]" italics="true" pageId="3" pageNumber="278">falkovitschi</emphasis>
Zhdanko &amp; Korshunov, 1985
</taxonomicName>
,
<taxonomicName id="4C3788F3FFA8FF9E8722AAD2FD9DFD10" authority="Zhdanko, 1998" authorityName="Zhdanko" authorityYear="1998" box="[335,638,667,690]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="oschi" subGenus="Rhymnaria">
<emphasis id="B9432F62FFA8FF9E8722AAD2FE8AFD10" box="[335,361,667,690]" italics="true" pageId="3" pageNumber="278">N.</emphasis>
(
<emphasis id="B9432F62FFA8FF9E871AAAD2FE6DFD10" box="[375,398,667,690]" italics="true" pageId="3" pageNumber="278">R.</emphasis>
)
<emphasis id="B9432F62FFA8FF9E87F1AAD2FE37FD10" box="[412,468,667,690]" italics="true" pageId="3" pageNumber="278">oschi</emphasis>
Zhdanko, 1998
</taxonomicName>
,
<taxonomicName id="4C3788F3FFA8FF9E84E5AAD2FCD3FD10" authorityName="Eversmann" authorityYear="1832" box="[648,816,667,690]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="rhymnus" subGenus="Rhymnaria">
<emphasis id="B9432F62FFA8FF9E84E5AAD2FD42FD10" box="[648,673,667,690]" italics="true" pageId="3" pageNumber="278">N.</emphasis>
(
<emphasis id="B9432F62FFA8FF9E84C2AAD2FD24FD10" box="[687,711,667,690]" italics="true" pageId="3" pageNumber="278">R.</emphasis>
)
<emphasis id="B9432F62FFA8FF9E84B8AAD2FCD3FD10" box="[725,816,667,690]" italics="true" pageId="3" pageNumber="278">rhymnus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3788F3FFA8FF9E8550AAD2FC0CFD10" authorityName="Grum-Grshimailo" authorityYear="1891" box="[829,1007,667,690]" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="tangutica" subGenus="Rhymnaria">
<emphasis id="B9432F62FFA8FF9E8550AAD2FCB5FD10" box="[829,854,667,690]" italics="true" pageId="3" pageNumber="278">N.</emphasis>
(
<emphasis id="B9432F62FFA8FF9E8509AAD2FC9FFD10" box="[868,892,667,690]" italics="true" pageId="3" pageNumber="278">R.</emphasis>
)
<emphasis id="B9432F62FFA8FF9E85E7AAD2FC0CFD10" box="[906,1007,667,690]" italics="true" pageId="3" pageNumber="278">tangutica</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C3788F3FFA8FF9E8245AAD2FE40FD70" authority="Churkin, 2006" authorityName="Churkin" authorityYear="2006" class="Insecta" family="Lycaenidae" genus="Neolycaena" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="zhdankoi" subGenus="Rhymnaria">
<emphasis id="B9432F62FFA8FF9E8245AAD2FBA2FD10" box="[1064,1089,667,690]" italics="true" pageId="3" pageNumber="278">N.</emphasis>
(
<emphasis id="B9432F62FFA8FF9E8617AAF2FF71FD70" box="[122,146,699,722]" italics="true" pageId="3" pageNumber="278">R.</emphasis>
)
<emphasis id="B9432F62FFA8FF9E86CCAAF2FEE2FD70" box="[161,257,699,722]" italics="true" pageId="3" pageNumber="278">zhdankoi</emphasis>
Churkin, 2006
</taxonomicName>
) belongs to a species group with spined valvae.
</paragraph>
</subSubSection>
<subSubSection id="C32DA0FBFFA8FF9E861CAA92FF22FCB0" pageId="3" pageNumber="278" type="etymology">
<paragraph id="8B88F370FFA8FF9E861CAA92FF22FCB0" blockId="3.[113,1089,219,946]" pageId="3" pageNumber="278">
<emphasis id="B9432F62FFA8FF9E861CAA92FF11FD50" bold="true" box="[113,242,731,754]" pageId="3" pageNumber="278">Etymology.</emphasis>
The species epithet is the Latin adjective
<emphasis id="B9432F62FFA8FF9E84CFAA92FD14FD50" box="[674,759,731,754]" italics="true" pageId="3" pageNumber="278">arcanus</emphasis>
(-
<emphasis id="B9432F62FFA8FF9E8560AA92FCF9FD50" box="[781,794,731,754]" italics="true" pageId="3" pageNumber="278">a</emphasis>
, -
<emphasis id="B9432F62FFA8FF9E8542AA92FCB3FD50" box="[815,848,731,754]" italics="true" pageId="3" pageNumber="278">um</emphasis>
), meaning mysterious, hidden.
</paragraph>
</subSubSection>
<subSubSection id="C32DA0FBFFA8FF9E861CAB52FEA7FC90" box="[113,324,795,818]" pageId="3" pageNumber="278" type="biology_ecology">
<paragraph id="8B88F370FFA8FF9E861CAB52FEA7FC90" blockId="3.[113,1089,219,946]" box="[113,324,795,818]" pageId="3" pageNumber="278">
<emphasis id="B9432F62FFA8FF9E861CAB52FF2EFC90" bold="true" box="[113,205,795,818]" pageId="3" pageNumber="278">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32DA0FBFFA8FF9E861CAB72FD1CFCF0" box="[113,767,827,850]" pageId="3" pageNumber="278" type="distribution">
<paragraph id="8B88F370FFA8FF9E861CAB72FD1CFCF0" blockId="3.[113,1089,219,946]" box="[113,767,827,850]" pageId="3" pageNumber="278">
<emphasis id="B9432F62FFA8FF9E861CAB72FEE7FCF0" bold="true" box="[113,260,827,850]" pageId="3" pageNumber="278">Distribution.</emphasis>
<collectingRegion id="49F33D92FFA8FF9E8764AB72FEA3FCF0" box="[265,320,827,850]" country="China" name="Xizang" pageId="3" pageNumber="278">Tibet</collectingRegion>
(in its wide pre-1950 extent) (this paper).
</paragraph>
</subSubSection>
<subSubSection id="C32DA0FBFFA8FF9E861CAB12FF5FFC10" pageId="3" pageNumber="278" type="discussion">
<paragraph id="8B88F370FFA8FF9E861CAB12FF5FFC10" blockId="3.[113,1089,219,946]" pageId="3" pageNumber="278">
<emphasis id="B9432F62FFA8FF9E861CAB12FF35FCD0" bold="true" box="[113,214,859,882]" pageId="3" pageNumber="278">Remark.</emphasis>
Unfortunatelly, due to the ageing process the specimen suffered certain change in colouration and therefore it is not possible to establish the exact original tint of the brown colour.
</paragraph>
</subSubSection>
</treatment>
</document>
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