272 lines
36 KiB
XML
272 lines
36 KiB
XML
<document id="D403CF11D60DBC5208E9C14F560667C3" ID-ISSN="1175-5326" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629742108138" checkinUser="felipe" docAuthor="Murányi, Dávid" docDate="2011" docId="03903327FF80FFC7FF6DFF05FDF0D184" docLanguage="en" docName="zt03049p046.pdf" docOrigin="Zootaxa 3049" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Isoperla vevcianensis Ikonomov 1980" docType="treatment" docVersion="5" lastPageNumber="22" masterDocId="FFA94B5FFF94FFD2FFFAFF92FD0BD04B" masterDocTitle="Balkanian species of the genus Isoperla Banks, 1906 (Plecoptera: Perlodidae)" masterLastPageNumber="46" masterPageNumber="1" pageNumber="21" updateTime="1699105341794" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="EFD9947ACBCD1552E9542767B5D2E8CF">Balkanian species of the genus Isoperla Banks, 1906 (Plecoptera: Perlodidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="3734C0D75CD704F911F97A8BF866DB1B" type="personal">
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<mods:namePart id="9E66D4169132C0A2541ED4342F77E269">Murányi, Dávid</mods:namePart>
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<mods:typeOfResource id="BBDE54B82E31FD5608364DAC4CFD532E">text</mods:typeOfResource>
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<mods:title id="0C391DBE8EC20DB76C47FCFEBB625C18">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A668871E5874D655EBA7BF4684EABF12">
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<mods:date id="B7D6ADE7F4118F44D1296EB25B92BD12">2011</mods:date>
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<mods:detail id="2D587BEB43E4AA14F5B6B71024E747D4" type="pubDate">
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<mods:number id="3A13D6722B06169BD9932BC165DEA68A">2011-10-05</mods:number>
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</mods:detail>
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<mods:detail id="979D4EA540EBE711459262FD1EAD480F" type="volume">
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<mods:number id="DEA221F97042F3A89DB8D2C733C27380">3049</mods:number>
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</mods:detail>
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<mods:extent id="9CCF30A6FC02A87BCCCD33D2724C9D22" unit="page">
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<mods:start id="DC068E5F26B4F0E0736ED099D40831CE">1</mods:start>
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<mods:classification id="705885FA78E22A27ED24480285D7FC86">journal article</mods:classification>
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<mods:identifier id="DE4B0D3CB0C3C3294D6A789340DD17D4" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="03903327FF80FFC7FF6DFF05FDF0D184" ID-DOI="http://doi.org/10.5281/zenodo.5282969" ID-Zenodo-Dep="5282969" LSID="urn:lsid:plazi:treatment:03903327FF80FFC7FF6DFF05FDF0D184" httpUri="http://treatment.plazi.org/id/03903327FF80FFC7FF6DFF05FDF0D184" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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<subSubSection id="C323D1BAFF80FFC6FF6DFF05FF6DD0FA" box="[151,614,151,177]" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="8B868231FF80FFC6FF6DFF05FF6DD0FA" blockId="20.[151,614,151,210]" box="[151,614,151,177]" pageId="20" pageNumber="21">
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<heading id="D0CE355DFF80FFC6FF6DFF05FF6DD0FA" bold="true" box="[151,614,151,177]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
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<taxonomicName id="4C39F9B2FF80FFC6FF6DFF05FF6DD0FA" ID-CoL="3QB2L" authority="Ikonomov, 1980" authorityName="Ikonomov" authorityYear="1980" box="[151,614,151,177]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="vevcianensis">
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<emphasis id="B94D5E23FF80FFC6FF6DFF05FF6DD0FA" bold="true" box="[151,614,151,177]" pageId="20" pageNumber="21">
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<emphasis id="B94D5E23FF80FFC6FF6DFF05FC93D0FA" bold="true" box="[151,408,151,177]" italics="true" pageId="20" pageNumber="21">Isoperla vevcianensis</emphasis>
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<bibRefCitation id="EFA8FFC0FF80FFC6FE5AFF05FF6DD0FA" author="Ikonomov" box="[416,614,151,177]" firstAuthor="Ikonomov" pageId="20" pageNumber="21" pagination="19 - 31" refId="ref28351" refString="Ikonomov, P. (1980) Nouvelles especes de Plecopteres (Insecta, Plecoptera) de Macedoine. II. Fragmenta Balcanica, 11, 19 - 31." type="journal article" year="1980">Ikonomov, 1980</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C323D1BAFF80FFC6FF6DFF28FC6ED099" box="[151,357,186,210]" pageId="20" pageNumber="21" type="description">
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||
<paragraph id="8B868231FF80FFC6FF6DFF28FC6ED099" blockId="20.[151,614,151,210]" box="[151,357,186,210]" pageId="20" pageNumber="21">
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(
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<figureCitation id="13029EB4FF80FFC6FF65FF28FC2DD099" box="[159,294,186,210]" captionStart="FIGURES 62–66" captionStartId="21.[151,264,1794,1817]" captionTargetBox="[187,1426,511,1780]" captionTargetId="figure-197@21.[151,1436,487,1791]" captionTargetPageId="21" captionText="FIGURES 62–66. Isoperla vevcianensis Ikonomov, 1980 — 62: head and pronotum; 63: male terminalia, dorsal view; 64: male terminalia, ventral view; 65: meso- and metathorax, ventral view; 66: medial penial armatures — scale 1 mm: 62–65; 0.5 mm: 66." pageId="20" pageNumber="21">Figs. 62–66</figureCitation>
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,
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<figureCitation id="13029EB4FF80FFC6FEC8FF28FC56D099" box="[306,349,186,210]" captionStart="FIGURES 126–127" captionStartId="41.[151,264,2001,2024]" captionTargetBox="[374,1212,188,1983]" captionTargetId="figure-29@41.[374,1212,188,1984]" captionTargetPageId="41" captionText="FIGURES 126–127. Distribution of the Balkanian endemic Isoperla Banks, 1906 — grey areas are above 1000 meters." pageId="20" pageNumber="21">127</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C323D1BAFF80FFC6FF6DFE90F9FCD153" box="[151,1271,258,280]" pageId="20" pageNumber="21" type="reference_group">
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<paragraph id="8B868231FF80FFC6FF6DFE90F9FCD153" blockId="20.[151,1271,258,280]" box="[151,1271,258,280]" pageId="20" pageNumber="21">
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<treatmentCitationGroup id="AB29A51FFF80FFC6FF6DFE90F9FCD153" box="[151,1271,258,280]" pageId="20" pageNumber="21">
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<taxonomicName id="4C39F9B2FF80FFC6FF6DFE90FF16D153" ID-CoL="3QB2L" authority="Ikonomov, 1980" authorityName="Ikonomov" authorityYear="1980" box="[151,541,258,280]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="vevcianensis">
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<emphasis id="B94D5E23FF80FFC6FF6DFE90FC79D153" box="[151,370,258,280]" italics="true" pageId="20" pageNumber="21">Isoperla vevcianensis</emphasis>
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<bibRefCitation id="EFA8FFC0FF80FFC6FE82FE90FF16D153" author="Ikonomov" box="[376,541,258,280]" firstAuthor="Ikonomov" pageId="20" pageNumber="21" pagination="19 - 31" refId="ref28351" refString="Ikonomov, P. (1980) Nouvelles especes de Plecopteres (Insecta, Plecoptera) de Macedoine. II. Fragmenta Balcanica, 11, 19 - 31." type="journal article" year="1980">Ikonomov, 1980</bibRefCitation>
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||
</taxonomicName>
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||
—
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||
<treatmentCitation id="0A98A420FF80FFC6FDBEFE90FE00D153" author="Ikonomov, P." box="[580,779,258,280]" page="26" pageId="20" pageNumber="21" year="1980">
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||
<bibRefCitation id="EFA8FFC0FF80FFC6FDBEFE90FE00D153" author="Ikonomov" box="[580,779,258,280]" firstAuthor="Ikonomov" page="26" pageId="20" pageNumber="21" pagination="19 - 31" refId="ref28351" refString="Ikonomov, P. (1980) Nouvelles especes de Plecopteres (Insecta, Plecoptera) de Macedoine. II. Fragmenta Balcanica, 11, 19 - 31." type="journal article" year="1980">Ikonomov 1980: 26</bibRefCitation>
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</treatmentCitation>
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(original description of male, female and larva).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C323D1BAFF80FFC6FF6DFED7FCD5D1A6" pageId="20" pageNumber="21" type="materials_examined">
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||
<paragraph id="8B868231FF80FFC6FF6DFED7FCD5D1A6" blockId="20.[151,1437,325,1969]" pageId="20" pageNumber="21">
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<emphasis id="B94D5E23FF80FFC6FF6DFED7FC89D115" bold="true" box="[151,386,325,350]" pageId="20" pageNumber="21">Material examined:</emphasis>
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<materialsCitation id="3B51886CFF80FFC6FE73FED7FCD1D1A6" collectingDate="2008-03-07" collectingDateMax="2008-04-07" collectingDateMin="2008-03-07" collectionCode="HNHM, CGV" collectorName="Z. Barina & Barina, D & Pifko, A" country="Albania" county="Mts Jablanica" elevation="1900" latitude="41.27905" location="Librazhd district" longLatPrecision="1" longitude="20.50103" municipality="Mts Jablanica" pageId="20" pageNumber="21" specimenCount="1">
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||
<collectingCountry id="F32EC2A1FF80FFC6FE73FED7FF00D116" box="[393,523,325,349]" name="Albania" pageId="20" pageNumber="21">ALBANIA</collectingCountry>
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||
:
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||
<location id="8EE6D4EAFF80FFC6FDECFED7FFDFD116" LSID="urn:lsid:plazi:treatment:03903327FF80FFC7FF6DFF05FDF0D184:8EE6D4EAFF80FFC6FDECFED7FFDFD116" box="[534,724,325,349]" country="Albania" county="Mts Jablanica" latitude="41.27905" longLatPrecision="1" longitude="20.50103" municipality="Mts Jablanica" name="Librazhd district" pageId="20" pageNumber="21">Librazhd district</location>
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||
:
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||
<location id="8EE6D4EAFF80FFC6FD24FED7FE4BD116" LSID="urn:lsid:plazi:treatment:03903327FF80FFC7FF6DFF05FDF0D184:8EE6D4EAFF80FFC6FD24FED7FE4BD116" box="[734,832,325,349]" country="Albania" county="Mts Jablanica" latitude="41.27905" longLatPrecision="1" longitude="20.50103" municipality="Mts Jablanica" name="Qarishte" pageId="20" pageNumber="21">Qarishtë</location>
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,
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||
<collectingMunicipality id="6BE2184BFF80FFC6FCB1FED4FEE3D116" box="[843,1000,325,349]" pageId="20" pageNumber="21">Mts Jablanica</collectingMunicipality>
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, brook E of the village,
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||
<geoCoordinate id="EE0DE4F6FF80FFC6FB0FFED4F888D116" box="[1269,1411,325,349]" degrees="41.24569" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="41.24569">N 41.24569°</geoCoordinate>
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||
<geoCoordinate id="EE0DE4F6FF80FFC6FA71FED4FC05D1CA" degrees="20.51238" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="1" value="20.51238">E 20.51238°</geoCoordinate>
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,
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||
<quantity id="4CC12FD4FF80FFC6FEE0FEFBFC7AD1CA" box="[282,369,361,385]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="20" pageNumber="21" unit="m" value="1900.0">
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||
<elevation id="00146502FF80FFC6FEE0FEFBFC7AD1CA" box="[282,369,361,385]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="20" pageNumber="21" unit="m" value="1900.0">1900 m</elevation>
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</quantity>
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,
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||
<date id="FF87A4F1FF80FFC6FE87FEFBFCF5D1CA" box="[381,510,361,385]" pageId="20" pageNumber="21" value="2008-04-07">
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||
<collectingDate id="EFC35D19FF80FFC6FE87FEFBFCF5D1CA" box="[381,510,361,385]" pageId="20" pageNumber="21" value="2008-04-07">04.07.2008</collectingDate>
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||
</date>
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||
, leg.
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||
<collectorName id="26CCE7E7FF80FFC6FDC0FEF8FFACD1CA" box="[570,679,361,385]" pageId="20" pageNumber="21">Z. Barina</collectorName>
|
||
, D.
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||
<collectorName id="26CCE7E7FF80FFC6FD2CFEFBFE33D1CA" box="[726,824,361,385]" pageId="20" pageNumber="21">Pifkó, A</collectorName>
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||
. Vojtkó:
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||
<quantity id="4CC12FD4FF80FFC6FC67FEFBFECAD1CA" box="[925,961,361,385]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="m" value="1.0">
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||
<elevation id="00146502FF80FFC6FC67FEFBFECAD1CA" box="[925,961,361,385]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="m" value="1.0">1m</elevation>
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||
</quantity>
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||
(
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||
<collectionCode id="ED281AF4FF80FFC6FC2AFEFBF920D1CA" box="[976,1067,361,385]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="20" pageNumber="21">HNHM</collectionCode>
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||
; used for drawings, penial armatures prepared on slide); Steblevë,
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||
<collectingCounty id="62E7FABDFF80FFC6FDD6FE1CFFC6D1EE" box="[556,717,397,421]" pageId="20" pageNumber="21">Mts Jablanica</collectingCounty>
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||
, brook SE of the village,
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||
<geoCoordinate id="EE0DE4F6FF80FFC6FC04FE1CF99AD1EE" box="[1022,1169,397,421]" degrees="41.27905" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="41.27905">N 41.27905°</geoCoordinate>
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||
<geoCoordinate id="EE0DE4F6FF80FFC6FB66FE1CF825D1EE" box="[1180,1326,397,421]" degrees="20.50103" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="1" value="20.50103">E 20.50103°</geoCoordinate>
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,
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||
<quantity id="4CC12FD4FF80FFC6FAC7FE1FF89CD1EE" box="[1341,1431,397,421]" metricMagnitude="3" metricUnit="m" metricValue="1.8649999999999998" pageId="20" pageNumber="21" unit="m" value="1865.0">
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<elevation id="00146502FF80FFC6FAC7FE1FF89CD1EE" box="[1341,1431,397,421]" metricMagnitude="3" metricUnit="m" metricValue="1.8649999999999998" pageId="20" pageNumber="21" unit="m" value="1865.0">1865 m</elevation>
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||
</quantity>
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,
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||
<date id="FF87A4F1FF80FFC6FF6DFE23FC13D182" box="[151,280,433,457]" pageId="20" pageNumber="21" value="2008-03-07">
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||
<collectingDate id="EFC35D19FF80FFC6FF6DFE23FC13D182" box="[151,280,433,457]" pageId="20" pageNumber="21" value="2008-03-07">03.07.2008</collectingDate>
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||
</date>
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||
, leg.
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||
<collectorName id="26CCE7E7FF80FFC6FEA8FE20FCB5D182" box="[338,446,433,457]" pageId="20" pageNumber="21">Z. Barina</collectorName>
|
||
, D.
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||
<collectorName id="26CCE7E7FF80FFC6FE11FE23FF47D182" box="[491,588,433,457]" pageId="20" pageNumber="21">Pifkó, A</collectorName>
|
||
. Vojtkó:
|
||
<quantity id="4CC12FD4FF80FFC6FD55FE23FFD9D182" box="[687,722,433,457]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="m" value="1.0">
|
||
<elevation id="00146502FF80FFC6FD55FE23FFD9D182" box="[687,722,433,457]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="m" value="1.0">1m</elevation>
|
||
</quantity>
|
||
(
|
||
<collectionCode id="ED281AF4FF80FFC6FD1BFE23FE37D182" box="[737,828,433,457]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="20" pageNumber="21">HNHM</collectionCode>
|
||
; penial armatures prepared on slide),
|
||
<quantity id="4CC12FD4FF80FFC6FB27FE23F80BD182" box="[1245,1280,433,457]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="m" value="1.0">
|
||
<elevation id="00146502FF80FFC6FB27FE23F80BD182" box="[1245,1280,433,457]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="m" value="1.0">1m</elevation>
|
||
</quantity>
|
||
(
|
||
<collectionCode id="ED281AF4FF80FFC6FAF5FE23F846D182" box="[1295,1357,433,457]" pageId="20" pageNumber="21">CGV</collectionCode>
|
||
; penial armatures prepared on slide)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C323D1BAFF80FFC6FF3DFE6AFEE5D212" pageId="20" pageNumber="21" type="diagnosis">
|
||
<paragraph id="8B868231FF80FFC6FF3DFE6AFEE5D212" blockId="20.[151,1437,325,1969]" pageId="20" pageNumber="21">
|
||
<emphasis id="B94D5E23FF80FFC6FF3DFE6AFC48D25A" bold="true" box="[199,323,504,529]" pageId="20" pageNumber="21">Diagnosis:</emphasis>
|
||
This broadheaded species is characterized by the presence of a medial penial armature that is covered in spike-like scale and divided into a trapezoidal upper and a rounded lower part, by the absences of lateral penial armatures, and by the presence of an enlarged vesicle of sternite VIII.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C323D1BAFF80FFC6FF3DFDF7FE3CD7FA" pageId="20" pageNumber="21" type="description">
|
||
<paragraph id="8B868231FF80FFC6FF3DFDF7FEEDD442" blockId="20.[151,1437,325,1969]" pageId="20" pageNumber="21">
|
||
<emphasis id="B94D5E23FF80FFC6FF3DFDF7FC50D235" bold="true" box="[199,347,613,638]" pageId="20" pageNumber="21">Description:</emphasis>
|
||
Large-sized species, macropterous. Body length: males
|
||
<quantity id="4CC12FD4FF80FFC6FC35FDF7F94CD236" box="[975,1095,613,637]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="20" pageNumber="21" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
|
||
(n=3); forewing length: males
|
||
<quantity id="4CC12FD4FF80FFC6FF6DFD1BFC2FD2EA" box="[151,292,649,673]" metricMagnitude="-2" metricUnit="m" metricValue="1.325" metricValueMax="1.35" metricValueMin="1.3" pageId="20" pageNumber="21" unit="mm" value="13.25" valueMax="13.5" valueMin="13.0">13–13.5 mm</quantity>
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||
(n=3). General colour yellowish but the meso- and metanotum and the abdomen mostly brown; pilosity of the body and legs short and dense. Head broad and rounded, yellow with a brown patch between the three ocelli; tentorial callosities and M-line more or less distinct, a few wrinkles present between the M-line and the lateral margins; the occiput lacks rugosities but is brown laterally (
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||
<figureCitation id="13029EB4FF80FFC6FC8BFD67FECCD346" box="[881,967,757,781]" captionStart="FIGURES 62–66" captionStartId="21.[151,264,1794,1817]" captionTargetBox="[187,1426,511,1780]" captionTargetId="figure-197@21.[151,1436,487,1791]" captionTargetPageId="21" captionText="FIGURES 62–66. Isoperla vevcianensis Ikonomov, 1980 — 62: head and pronotum; 63: male terminalia, dorsal view; 64: male terminalia, ventral view; 65: meso- and metathorax, ventral view; 66: medial penial armatures — scale 1 mm: 62–65; 0.5 mm: 66." pageId="20" pageNumber="21">Fig. 62</figureCitation>
|
||
). Eyes smaller than the area delimited by the three ocelli. Scape dark brown, pedicel and the following three or four antennomeres are brown but distal part of the antenna is dark brown; palpi brown. Pronotum yellow, rectangular, edges angled; rugosities are numerous and brown coloured, anterior and posterior lines brown under and above the rugosities. Mesonotum dark brown but yellow anteriorly, sometimes medially also; metanotum dark brown but yellow anteriorly. Wings yellowish, venation pale in the anterior two thirds, brownish in the posterior third. Ventral surface of thorax mostly pale, meso- and metabasisternum mostly dark brown; furcasternites light brown, furcal pits black (
|
||
<figureCitation id="13029EB4FF80FFC6FBB2FC5FF995D3AE" box="[1096,1182,973,997]" captionStart="FIGURES 62–66" captionStartId="21.[151,264,1794,1817]" captionTargetBox="[187,1426,511,1780]" captionTargetId="figure-197@21.[151,1436,487,1791]" captionTargetPageId="21" captionText="FIGURES 62–66. Isoperla vevcianensis Ikonomov, 1980 — 62: head and pronotum; 63: male terminalia, dorsal view; 64: male terminalia, ventral view; 65: meso- and metathorax, ventral view; 66: medial penial armatures — scale 1 mm: 62–65; 0.5 mm: 66." pageId="20" pageNumber="21">Fig. 65</figureCitation>
|
||
). Femora pale but the dorsal surface and the ventral edges brown. Tibiae brown, tarsi dark brown.
|
||
</paragraph>
|
||
<paragraph id="8B868231FF80FFC6FF3DFB87FFA9D53A" blockId="20.[151,1437,325,1969]" pageId="20" pageNumber="21">
|
||
Male abdomen: First tergite dark brown but yellowish medially. Tergites II–VI entirely dark brown, tergite VII entirely dark brown or dark brown with two pale patches posteriorly. Pale markings become more extensive on tergites VIII–IX, but a transverse anterior dark brown stripe and a medial dark brown arrow-like mark always present; tergite X yellowish with two brown patches medially (
|
||
<figureCitation id="13029EB4FF80FFC6FCCEFB13FE83D4D2" box="[820,904,1153,1177]" captionStart="FIGURES 62–66" captionStartId="21.[151,264,1794,1817]" captionTargetBox="[187,1426,511,1780]" captionTargetId="figure-197@21.[151,1436,487,1791]" captionTargetPageId="21" captionText="FIGURES 62–66. Isoperla vevcianensis Ikonomov, 1980 — 62: head and pronotum; 63: male terminalia, dorsal view; 64: male terminalia, ventral view; 65: meso- and metathorax, ventral view; 66: medial penial armatures — scale 1 mm: 62–65; 0.5 mm: 66." pageId="20" pageNumber="21">Fig. 63</figureCitation>
|
||
). Transverse row of four pigmented spots seen on all but tergite X. Sternites II–VI brown with a medial transverse row of four spots; posterior part of sternite VII with a more or less extensive pale pattern. Sternite VIII paler and with a dark pattern, but only two big spots. Vesicle of sternite VIII brown, as wide as long, its posterior margin is weakly rounded; as long as half the segment’s length (
|
||
<figureCitation id="13029EB4FF80FFC6FF17FA83FC48D562" box="[237,323,1297,1321]" captionStart="FIGURES 62–66" captionStartId="21.[151,264,1794,1817]" captionTargetBox="[187,1426,511,1780]" captionTargetId="figure-197@21.[151,1436,487,1791]" captionTargetPageId="21" captionText="FIGURES 62–66. Isoperla vevcianensis Ikonomov, 1980 — 62: head and pronotum; 63: male terminalia, dorsal view; 64: male terminalia, ventral view; 65: meso- and metathorax, ventral view; 66: medial penial armatures — scale 1 mm: 62–65; 0.5 mm: 66." pageId="20" pageNumber="21">Fig. 64</figureCitation>
|
||
). Sternite IX yellowish brown but darker posteriorly, the medial penial armatures barely visible in the posterior half. Paraprocts dark brown, blunt, strong and slightly recurved; cerci dark brown in the basal segments, distal part of the cerci are usually paler.
|
||
</paragraph>
|
||
<paragraph id="8B868231FF80FFC6FF3DFAEFFC09D692" blockId="20.[151,1437,325,1969]" pageId="20" pageNumber="21">
|
||
Penis: Lobes and the distribution of non coloured scales, hairs and sensilla cannot be studied because the specimens were collected without evertion of the penis. The medial penial armature is divided into an upper and a lower part and probably located on the ventral lobe (
|
||
<figureCitation id="13029EB4FF80FFC6FD52FA57FFF6D596" box="[680,765,1477,1501]" captionStart="FIGURES 62–66" captionStartId="21.[151,264,1794,1817]" captionTargetBox="[187,1426,511,1780]" captionTargetId="figure-197@21.[151,1436,487,1791]" captionTargetPageId="21" captionText="FIGURES 62–66. Isoperla vevcianensis Ikonomov, 1980 — 62: head and pronotum; 63: male terminalia, dorsal view; 64: male terminalia, ventral view; 65: meso- and metathorax, ventral view; 66: medial penial armatures — scale 1 mm: 62–65; 0.5 mm: 66." pageId="20" pageNumber="21">Fig. 66</figureCitation>
|
||
), lateral penial armatures lacking. The upper medial penial armature is trapezoidal, tapered towards the basal section; length 500 µm, width 450 µm. The lower medial penial armature is rounded, slightly elongated; length 200 µm, width 120 µm. The scales are spike-like, those on the upper part are longer but of the same width. The length of the longer ones of the upper part is 150 µm, of the shortest ones in the lower part is 30 µm; width 15 µm. The penis is bald between the upper and lower parts, but bears quite large triangular scales laterally to the lower part; these are grading into smaller ones, shown on
|
||
<figureCitation id="13029EB4FF80FFC6FB77F9EBF9EBD6DA" box="[1165,1248,1657,1681]" captionStart="FIGURES 62–66" captionStartId="21.[151,264,1794,1817]" captionTargetBox="[187,1426,511,1780]" captionTargetId="figure-197@21.[151,1436,487,1791]" captionTargetPageId="21" captionText="FIGURES 62–66. Isoperla vevcianensis Ikonomov, 1980 — 62: head and pronotum; 63: male terminalia, dorsal view; 64: male terminalia, ventral view; 65: meso- and metathorax, ventral view; 66: medial penial armatures — scale 1 mm: 62–65; 0.5 mm: 66." pageId="20" pageNumber="21">Fig. 66</figureCitation>
|
||
, to the right side of the lower medial penial armature. Hydra-like scales occur above a bald line apically to the upper medial penial armature.
|
||
</paragraph>
|
||
<paragraph id="8B868231FF80FFC6FF3DF977FE0FD6B6" blockId="20.[151,1437,325,1969]" box="[199,772,1765,1790]" pageId="20" pageNumber="21">Female and larva not studied herein, egg unknown.</paragraph>
|
||
<paragraph id="8B868231FF80FFC6FF3DF89BFE3CD7FA" blockId="20.[151,1437,325,1969]" pageId="20" pageNumber="21">
|
||
<emphasis id="B94D5E23FF80FFC6FF3DF89BFC34D769" bold="true" box="[199,319,1801,1826]" pageId="20" pageNumber="21">Affinities:</emphasis>
|
||
The species was originally not assigned to a species group sensu
|
||
<bibRefCitation id="EFA8FFC0FF80FFC6FBE7F89BF9C2D76A" author="Consiglio" box="[1053,1225,1801,1825]" firstAuthor="Consiglio" pageId="20" pageNumber="21" pagination="67 - 75" refId="ref27914" refString="Consiglio, C. (1967) Una nuova specie Italiana e considerazioni sui gruppi di specie nel genere Isoperla (Plecoptera, Perlodidae). Fragmenta Entomologica, Roma, 5, 67 - 75." type="journal article" year="1967">Consiglio 1967</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="EFA8FFC0FF80FFC6FB2DF89BF884D76A" author="Ikonomov" box="[1239,1423,1801,1825]" firstAuthor="Ikonomov" pageId="20" pageNumber="21" pagination="19 - 31" refId="ref28351" refString="Ikonomov, P. (1980) Nouvelles especes de Plecopteres (Insecta, Plecoptera) de Macedoine. II. Fragmenta Balcanica, 11, 19 - 31." type="journal article" year="1980">Ikonomov 1980</bibRefCitation>
|
||
); herein it is assigned to the
|
||
<taxonomicName id="4C39F9B2FF80FFC6FE32F8BFFF17D70E" authorityName="Illies" authorityYear="1952" box="[456,540,1837,1861]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="silesica">
|
||
<emphasis id="B94D5E23FF80FFC6FE32F8BFFF17D70E" box="[456,540,1837,1861]" italics="true" pageId="20" pageNumber="21">silesica</emphasis>
|
||
</taxonomicName>
|
||
group. It differs from
|
||
<taxonomicName id="4C39F9B2FF80FFC6FCE7F8BCFEA2D70E" authorityName="Ikonomov" authorityYear="1980" box="[797,937,1837,1861]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="breviptera">
|
||
<emphasis id="B94D5E23FF80FFC6FCE7F8BCFEA2D70E" box="[797,937,1837,1861]" italics="true" pageId="20" pageNumber="21">I. breviptera</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C39F9B2FF80FFC6FC42F8BCF92ED70E" authorityName="Illies" authorityYear="1952" box="[952,1061,1837,1861]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="silesica">
|
||
<emphasis id="B94D5E23FF80FFC6FC42F8BCF92ED70E" box="[952,1061,1837,1861]" italics="true" pageId="20" pageNumber="21">I. silesica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C39F9B2FF80FFC6FBA7F8BCF9B7D70E" authorityName="Tierno de Figueroa & Fochetti" authorityYear="2001" box="[1117,1212,1837,1861]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="zwicki">
|
||
<emphasis id="B94D5E23FF80FFC6FBA7F8BCF9B7D70E" box="[1117,1212,1837,1861]" italics="true" pageId="20" pageNumber="21">I. zwicki</emphasis>
|
||
</taxonomicName>
|
||
by the shape of the upper medial penial armature which is trapezoidal contrary to the triangular armature of
|
||
<taxonomicName id="4C39F9B2FF80FFC6FB56F8C0F830D722" authorityName="Ikonomov" authorityYear="1980" box="[1196,1339,1873,1897]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="breviptera">
|
||
<emphasis id="B94D5E23FF80FFC6FB56F8C0F830D722" box="[1196,1339,1873,1897]" italics="true" pageId="20" pageNumber="21">I. breviptera</emphasis>
|
||
</taxonomicName>
|
||
and the rounded armature of
|
||
<taxonomicName id="4C39F9B2FF80FFC6FE73F8E4FCFDD7C6" authorityName="Illies" authorityYear="1952" box="[393,502,1909,1933]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="silesica">
|
||
<emphasis id="B94D5E23FF80FFC6FE73F8E4FCFDD7C6" box="[393,502,1909,1933]" italics="true" pageId="20" pageNumber="21">I. silesica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C39F9B2FF80FFC6FDCBF8E4FF9BD7C6" authorityName="Tierno de Figueroa & Fochetti" authorityYear="2001" box="[561,656,1909,1933]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="zwicki">
|
||
<emphasis id="B94D5E23FF80FFC6FDCBF8E4FF9BD7C6" box="[561,656,1909,1933]" italics="true" pageId="20" pageNumber="21">I. zwicki</emphasis>
|
||
</taxonomicName>
|
||
; furthermore,
|
||
<taxonomicName id="4C39F9B2FF80FFC6FCCDF8E4FEC8D7C6" authorityName="Ikonomov" authorityYear="1980" box="[823,963,1909,1933]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="breviptera">
|
||
<emphasis id="B94D5E23FF80FFC6FCCDF8E4FEC8D7C6" box="[823,963,1909,1933]" italics="true" pageId="20" pageNumber="21">I. breviptera</emphasis>
|
||
</taxonomicName>
|
||
is micropterous. The broad head and the large vesicle of sternite VIII is diagnostic of
|
||
<emphasis id="B94D5E23FF80FFC6FD71F808FE3CD7FA" box="[651,823,1946,1969]" italics="true" pageId="20" pageNumber="21">
|
||
<taxonomicName id="4C39F9B2FF80FFC6FD71F808FE38D7FA" authorityName="Ikonomov" authorityYear="1980" box="[651,819,1946,1969]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="vevcianensis">I. vevcianensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C323D1BAFF81FFC7FF3DFF0AFDF0D184" pageId="21" pageNumber="22" type="biology_ecology">
|
||
<paragraph id="8B868231FF81FFC7FF3DFF0AFDF0D184" blockId="21.[151,1436,151,463]" pageId="21" pageNumber="22">
|
||
<emphasis id="B94D5E23FF81FFC7FF3DFF0AFCF7D0FB" bold="true" box="[199,508,151,176]" pageId="21" pageNumber="22">Ecology and distribution:</emphasis>
|
||
The species was originally described from the Macedonian side of the Jablanica Mountains (
|
||
<bibRefCitation id="EFA8FFC0FF81FFC7FEE6FF2EFCD8D09F" author="Ikonomov" box="[284,467,188,212]" firstAuthor="Ikonomov" pageId="21" pageNumber="22" pagination="19 - 31" refId="ref28351" refString="Ikonomov, P. (1980) Nouvelles especes de Plecopteres (Insecta, Plecoptera) de Macedoine. II. Fragmenta Balcanica, 11, 19 - 31." type="journal article" year="1980">Ikonomov 1980</bibRefCitation>
|
||
), only a few kilometers from the present Albanian localities (
|
||
<figureCitation id="13029EB4FF81FFC7FB81FF2EF9D6D09F" box="[1147,1245,188,212]" captionStart="FIGURES 126–127" captionStartId="41.[151,264,2001,2024]" captionTargetBox="[374,1212,188,1983]" captionTargetId="figure-29@41.[374,1212,188,1984]" captionTargetPageId="41" captionText="FIGURES 126–127. Distribution of the Balkanian endemic Isoperla Banks, 1906 — grey areas are above 1000 meters." pageId="21" pageNumber="22">Fig. 127</figureCitation>
|
||
). Hitherto, it was known only from the
|
||
<typeStatus id="54823C93FF81FFC7FE74FF72FCB5D0B3" box="[398,446,224,248]" pageId="21" pageNumber="22">type</typeStatus>
|
||
locality (
|
||
<bibRefCitation id="EFA8FFC0FF81FFC7FDD0FF4DFFE8D0BC" author="Ikonomov" box="[554,739,223,247]" firstAuthor="Ikonomov" pageId="21" pageNumber="22" pagination="81 - 124" refId="ref28381" refString="Ikonomov, P. (1986) Plecopteres de Macedoine (Insecta, Plecoptera). Acta Musei Macedonici Scientiarum Naturalium, 18, 81 - 124." type="journal article" year="1986">Ikonomov 1986</bibRefCitation>
|
||
); consequently, the species is new for
|
||
<collectingCountry id="F32EC2A1FF81FFC7FB6CFF4DF9F9D0BC" box="[1174,1266,223,247]" name="Albania" pageId="21" pageNumber="22">Albania</collectingCountry>
|
||
. It seems to be an endemic of the Jablanica Mountains as it was not found in adjacent Macedonian or Albanian mountain systems. In
|
||
<collectingCountry id="F32EC2A1FF81FFC7FF4FFEB5FC1BD174" box="[181,272,295,319]" name="Albania" pageId="21" pageNumber="22">Albania</collectingCountry>
|
||
, it was collected from alpine grassland brooks above
|
||
<quantity id="4CC12FD4FF81FFC7FC9FFEB5FEB1D174" box="[869,954,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="21" pageNumber="22" unit="m" value="1850.0">1850 m</quantity>
|
||
during early July. Associates include
|
||
<taxonomicName id="4C39F9B2FF81FFC7FA9BFEBBFC5AD12F" authority="tripartita" authorityName="tripartita Illies" authorityYear="1954" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="tripartita">
|
||
<emphasis id="B94D5E23FF81FFC7FA9BFEBBFC5AD12F" italics="true" pageId="21" pageNumber="22">I. tripartita tripartita</emphasis>
|
||
</taxonomicName>
|
||
at the Qarishtë brook and
|
||
<emphasis id="B94D5E23FF81FFC7FD85FEDFFE78D12F" box="[639,883,333,356]" italics="true" pageId="21" pageNumber="22">
|
||
<taxonomicName id="4C39F9B2FF81FFC7FD85FEDFFE65D12F" authority="tripartita" authorityName="tripartita Illies" authorityYear="1954" box="[639,878,333,356]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="tripartita">I. tripartita tripartita</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicName id="4C39F9B2FF81FFC7FC81FEDFF82CD12F" authority="Aubert, 1956" authorityName="Aubert" authorityYear="1956" box="[891,1319,332,356]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="metsovonica">
|
||
<emphasis id="B94D5E23FF81FFC7FC81FEDFF982D12F" box="[891,1161,332,356]" italics="true" pageId="21" pageNumber="22">Leuctra cf. metsovonica</emphasis>
|
||
<bibRefCitation id="EFA8FFC0FF81FFC7FB6AFEDEF82CD12F" author="Aubert" box="[1168,1319,332,356]" firstAuthor="Aubert" pageId="21" pageNumber="22" pagination="187 - 213" refId="ref27586" refString="Aubert, J. (1956) Contribution a l'etude des Plecopteres de Grece. Mitteilungen der Schweizerischen entomologischen Gesellschaft, 29, 187 - 213." type="journal article" year="1956">Aubert, 1956</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C39F9B2FF81FFC7FACEFEDFFC83D1CC" authority="Zwick, 1978" authorityName="Zwick" authorityYear="1978" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="caligula">
|
||
<emphasis id="B94D5E23FF81FFC7FACEFEDFFDF8D1C3" italics="true" pageId="21" pageNumber="22">Nemoura caligula</emphasis>
|
||
<bibRefCitation id="EFA8FFC0FF81FFC7FF00FEFDFC83D1CC" author="Zwick" box="[250,392,367,391]" firstAuthor="Zwick" pageId="21" pageNumber="22" pagination="213 - 239" refId="ref30884" refString="Zwick, P. (1978) Steinfliegen (Plecoptera) aus Griechenland und benachbarten Landern. Mitteilungen der Schweizerischen entomologischen Gesellschaft, 51, 21 - 38, 213 - 239." type="journal article" year="1978">Zwick, 1978</bibRefCitation>
|
||
</taxonomicName>
|
||
and a deformed
|
||
<taxonomicName id="4C39F9B2FF81FFC7FDBFFEE3FFD2D1C3" authorityName="Kempny" authorityYear="1898" box="[581,729,369,392]" class="Insecta" family="Nemouridae" genus="Protonemura" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D5E23FF81FFC7FDBFFEE3FFD2D1C3" box="[581,729,369,392]" italics="true" pageId="21" pageNumber="22">Protonemura</emphasis>
|
||
</taxonomicName>
|
||
female at the Steblevë brook.
|
||
<taxonomicName id="4C39F9B2FF81FFC7FBCAFEE2F862D1CC" authority="tripartita" authorityName="tripartita Illies" authorityYear="1954" box="[1072,1385,368,392]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="tripartita">
|
||
<emphasis id="B94D5E23FF81FFC7FBCAFEE2F862D1CC" box="[1072,1385,368,392]" italics="true" pageId="21" pageNumber="22">Isoperla tripartita tripartita</emphasis>
|
||
</taxonomicName>
|
||
was more numerous than
|
||
<taxonomicName id="4C39F9B2FF81FFC7FE79FE06FF2CD1E7" authorityName="Ikonomov" authorityYear="1980" box="[387,551,404,428]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="21" pageNumber="21" phylum="Arthropoda" rank="species" species="vevcianensis">
|
||
I.
|
||
<emphasis id="B94D5E23FF81FFC7FE60FE07FF2CD1E7" box="[410,551,405,428]" italics="true" pageId="21" pageNumber="22">vevcianensis</emphasis>
|
||
</taxonomicName>
|
||
at either site. At the
|
||
<typeStatus id="54823C93FF81FFC7FCF5FE06FE34D1E7" box="[783,831,404,428]" pageId="21" pageNumber="22">type</typeStatus>
|
||
locality it occurred between 1100 and
|
||
<quantity id="4CC12FD4FF81FFC7FB0BFE06F84ED1E0" box="[1265,1349,404,428]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="21" pageNumber="22" unit="m" value="1450.0">1450 m</quantity>
|
||
in June and July.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |