treatments-xml/data/03/84/F6/0384F651FF988771FF2F0B78FCA11DF1.xml

<document id="E958513AF23A44469DD88C3BA3668810" ID-DOI="10.11646/zootaxa.5403.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10561629" ID-ZooBank="0C3D7428-8B33-46F5-AC53-2C4D96E0E4F5" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1706103575558" checkinUser="plazi" docAuthor="Silva, Marciara Lopes, Cohen, Simone Chinicz, Ottoni, Felipe Polivanov, Viana, Diego Carvalho, Meneses, Yuri Costa De &amp; Justo, Marcia Cristina Nascimento" docDate="2024" docId="0384F651FF988771FF2F0B78FCA11DF1" docLanguage="en" docName="zootaxa.5403.1.6.pdf" docOrigin="Zootaxa 5403 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5403.1.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diaphorocleidus munimensis Silva, Meneses, Cohen and Justo 2024, n. sp." docType="treatment" docVersion="3" lastPageNumber="97" masterDocId="FFBD8E29FF9E8776FFB8081EFF861C78" masterDocTitle="Two new species of Diaphorocleidus and new data on D. affinis (Monogenoidea: Dactylogyridae) parasitizing Bryconops cf. affinis (Characiformes: Iguanodectidae) from the Munim River basin, State of Maranhão, northeastern Brazil" masterLastPageNumber="103" masterPageNumber="91" pageNumber="97" updateTime="1706279861717" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="E289718E9EA2B111957881C34AEEA6AA">Two new species of Diaphorocleidus and new data on D. affinis (Monogenoidea: Dactylogyridae) parasitizing Bryconops cf. affinis (Characiformes: Iguanodectidae) from the Munim River basin, State of Maranhão, northeastern Brazil</mods:title>
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<mods:namePart id="6C3EC3B46FFDD0D7AD102B5BD514006C">Silva, Marciara Lopes</mods:namePart>
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<mods:affiliation id="B12A311A55BC9263B49F88358640D320">Programa de Pós-Graduação em Ciência Animal, Universidade Estadual do Maranhão, UEMA. Cidade Universitária Paulo VI, Avenida Lourenço Vieira da Silva 1000, 65055 - 310 São Luís, MA, Brazil</mods:affiliation>
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<mods:namePart id="B9E453D55DD445625DB740B9570623CA">Cohen, Simone Chinicz</mods:namePart>
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<mods:namePart id="237913AEE503749FF71DD13461F196EB">Ottoni, Felipe Polivanov</mods:namePart>
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<mods:affiliation id="E21A81D56C64584AEE68B4EED8827221">Laboratório de Sistemática e Ecologia de Organismos Aquáticos, Centro de Ciências de Chapadinha, Universidade Federal do Maranhão (UFMA), Campus de Chapadinha, BR- 222, KM 04, S / N, Chapadinha, MA, Brazil</mods:affiliation>
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<mods:namePart id="0967C0472AEE4CBFDDB76479D88E437E">Viana, Diego Carvalho</mods:namePart>
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<mods:affiliation id="CAD796715498A564A398596FD962CD78">Núcleo de Estudos Morfofisiológicos Avançados, Universidade Estadual da Região Tocantina do Maranhão. Campus Imperatriz, Rua Godofredo Viana 1300, 65901 - 480 Imperatriz, MA, Brazil</mods:affiliation>
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<mods:namePart id="77484D968363D2F5FDBE8D97A95B6269">Meneses, Yuri Costa De</mods:namePart>
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<mods:affiliation id="1EAF88071E6F4DC86005B31F368FA02F">Laboratório de Helmintos Parasitos de Peixes, Instituto Oswaldo Cruz, FIOCRUZ. Avenida Brasil 4365, 21040 - 360 Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:namePart id="A79BE6F1E5C893EFEE42969D72BE44A5">Justo, Marcia Cristina Nascimento</mods:namePart>
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<taxonomicName id="4C2D3CC4FF988770FF2F0B78FC1F1FF9" authority="Silva, Meneses, Cohen and Justo" authorityName="Silva, Meneses, Cohen and Justo" authorityYear="2024" box="[151,921,870,897]" class="Monogenea" family="Ancyrocephalidae" genus="Diaphorocleidus" kingdom="Animalia" order="Dactylogyridea" pageId="6" pageNumber="97" phylum="Platyhelminthes" rank="species" species="munimensis" status="sp. nov.">
<emphasis id="B9599B55FF988770FF2F0B78FE7C1FF9" bold="true" box="[151,506,870,897]" italics="true" pageId="6" pageNumber="97">Diaphorocleidus munimensis</emphasis>
Silva, Meneses, Cohen and Justo
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(
<figureCitation id="13165BC2FF988770FC490B78FBDB1FF8" box="[1009,1117,870,896]" captionStart="FIGURE 4" captionStartId="7.[151,250,1698,1722]" captionTargetBox="[306,1279,431,1668]" captionTargetId="figure-144@7.[298,1289,417,1675]" captionTargetPageId="7" captionText="FIGURE 4. Diaphorocleidus munimensis n. sp. from Bryconops cf. affinis (Characiformes, Iguanodectidae) from São José, municipality of Chapadinha and in Riacho Estrela, municipality of Anapurus, River Munim basin, Maranhão, Brazil: (a) total, ventral view; (b) copulatory complex, ventral view; (c) hook pairs 3, 4; (d) hook pairs 1,5; (e) ventral anchor; (f) dorsal anchor; (g) ventral bar; (h) dorsal bar. Scale bars: (a) 100 μm; (b) 15 μm; (c, d, g, h) 10 μm; (e, f) 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10561637" httpUri="https://zenodo.org/record/10561637/files/figure.png" pageId="6" pageNumber="97">Figure 4</figureCitation>
)
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Type-host: 
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cf. 
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(Günther)
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Type-locality: 
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of Chapadinha in the “Riacho Feio” (
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),
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and urban area of Anapurus in the “Riacho Estrela” (
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), tributaries of the 
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, 
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, 
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, 
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.
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<paragraph id="8B924747FF988770FF2F0C19FE0C1867" blockId="6.[151,1437,939,1178]" box="[151,394,1031,1055]" pageId="6" pageNumber="97">Site of infestation: Gills</paragraph>
<paragraph id="8B924747FF988770FF2F0C38FEE31825" blockId="6.[151,1437,939,1178]" pageId="6" pageNumber="97">Parasitological indexes: Total number of hosts: 64; number of infected hosts: 21; total number of parasites: 141; Range of intensity: 1–28.</paragraph>
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<materialsCitation id="3B454D1AFF988770FF2F0C7DFDF01803" box="[151,630,1123,1147]" collectionCode="CHIOC" pageId="6" pageNumber="97" specimenCode="CHIOC 40280" specimenCount="1" typeStatus="holotype">
Specimens deposited: 
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<specimenCode id="DB8BEF3CFF988770FE660C7DFDF01803" box="[478,630,1123,1147]" collectionCode="CHIOC" country="Brazil" httpUri="http://grbio.org/cool/sv7j-t59d" name="Helminthological Collection of Oswaldo Cruz Institute (Coleccion. Helmintologica del Instituto Oswaldo Cruz)" pageId="6" pageNumber="97">CHIOC 40280</specimenCode>
</materialsCitation>
, 
<materialsCitation id="3B454D1AFF988770FD380C7AFB931803" box="[640,1045,1123,1148]" collectionCode="CHIOC" isTailEnumeration="true" pageId="6" pageNumber="97" specimenCode="CHIOC 40281, 40282, 40283" specimenCount="1" typeStatus="paratype">
<typeStatus id="5496F9E5FF988770FD380C7AFD641804" box="[640,738,1124,1148]" pageId="6" pageNumber="97" type="paratype">Paratypes</typeStatus>
<specimenCode id="DB8BEF3CFF988770FD500C7DFC061803" box="[744,896,1123,1147]" collectionCode="CHIOC" country="Brazil" httpUri="http://grbio.org/cool/sv7j-t59d" name="Helminthological Collection of Oswaldo Cruz Institute (Coleccion. Helmintologica del Instituto Oswaldo Cruz)" pageId="6" pageNumber="97">CHIOC 40281</specimenCode>
, 
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, 
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a–b, 40284 a–b
</paragraph>
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<subSubSection id="C33714CCFF988770FF2F0C9CFB0718E2" box="[151,1153,1154,1178]" pageId="6" pageNumber="97" type="etymology">
<paragraph id="8B924747FF988770FF2F0C9CFB0718E2" blockId="6.[151,1437,939,1178]" box="[151,1153,1154,1178]" pageId="6" pageNumber="97">Etymology: The specific name is derived from the river basin where the specimens were collected.</paragraph>
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<emphasis id="B9599B55FF988770FF2F0CD6FEA7189A" bold="true" box="[151,289,1224,1250]" pageId="6" pageNumber="97">Description</emphasis>
(Based on 
<specimenCount id="9D2B8CCEFF988770FE170CD6FDC9189A" box="[431,591,1224,1250]" count="20" pageId="6" pageNumber="97" type="generic">20 specimens</specimenCount>
, mounted in Hoyer’s medium). Body fusiform, flattened dorsoventrally, 357 (174–644; n = 19) long by 155 (86–309, n = 20) greatest width. Tegument thin and smooth. Cephalic lobes moderately developed; three bilateral pairs of head organs; cephalic glands indistinct. Two pairs of eyes equidistant, anterior pair smaller than posterior. Pharynx spherical 20 (14–27; n = 3) in diameter. Two intestinal caeca, confluent posteriorly to gonads, without diverticula. Gonads overlapping, intercaecal; testis elongate, fusiform, dorsoposterior to germarium, 30 (24–33; n = 4) long by 26 (25–30; n = 4) wide; elongated seminal vesicle and single prostatic reservoir (
<figureCitation id="13165BC2FF988770FEB50DBEFEED19C2" box="[269,363,1440,1466]" captionStart="FIGURE 4" captionStartId="7.[151,250,1698,1722]" captionTargetBox="[306,1279,431,1668]" captionTargetId="figure-144@7.[298,1289,417,1675]" captionTargetPageId="7" captionText="FIGURE 4. Diaphorocleidus munimensis n. sp. from Bryconops cf. affinis (Characiformes, Iguanodectidae) from São José, municipality of Chapadinha and in Riacho Estrela, municipality of Anapurus, River Munim basin, Maranhão, Brazil: (a) total, ventral view; (b) copulatory complex, ventral view; (c) hook pairs 3, 4; (d) hook pairs 1,5; (e) ventral anchor; (f) dorsal anchor; (g) ventral bar; (h) dorsal bar. Scale bars: (a) 100 μm; (b) 15 μm; (c, d, g, h) 10 μm; (e, f) 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10561637" httpUri="https://zenodo.org/record/10561637/files/figure.png" pageId="6" pageNumber="97">Fig. 4A</figureCitation>
). Copulatory complex comprising male copulatory organ (MCO) and accessory piece. MCO sclerotized, tubular, 122 (109–130, n = 16) in total length, coiled with 2 counterclockwise rings, 22 (20–26, n = 14) diameter of the first ring; base forming a small tube with two circular flanges. Accessory piece composed of two large pieces, united in their proximal part, claw-shaped, distally hooked, with a small lateral structure serving as guide to MCO, 33 (28–40, n = 14) long by 22 (14–27; n = 11) wide (
<figureCitation id="13165BC2FF988770FC180E2EFC7F1A32" box="[928,1017,1584,1610]" captionStart="FIGURE 4" captionStartId="7.[151,250,1698,1722]" captionTargetBox="[306,1279,431,1668]" captionTargetId="figure-144@7.[298,1289,417,1675]" captionTargetPageId="7" captionText="FIGURE 4. Diaphorocleidus munimensis n. sp. from Bryconops cf. affinis (Characiformes, Iguanodectidae) from São José, municipality of Chapadinha and in Riacho Estrela, municipality of Anapurus, River Munim basin, Maranhão, Brazil: (a) total, ventral view; (b) copulatory complex, ventral view; (c) hook pairs 3, 4; (d) hook pairs 1,5; (e) ventral anchor; (f) dorsal anchor; (g) ventral bar; (h) dorsal bar. Scale bars: (a) 100 μm; (b) 15 μm; (c, d, g, h) 10 μm; (e, f) 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10561637" httpUri="https://zenodo.org/record/10561637/files/figure.png" pageId="6" pageNumber="97">Fig. 4B</figureCitation>
). Germarium 51 (34–62; n = 4) long, 30 (22–42; n = 4) wide. Vagina with submarginal vaginal opening, consisting of a thin sclerotized tube; seminal receptacle not observed. Vitelline follicles scattered throughout trunk but absent around the reproductive organs (
<figureCitation id="13165BC2FF988770FF270E82FF7F1ACE" box="[159,249,1692,1718]" captionStart="FIGURE 4" captionStartId="7.[151,250,1698,1722]" captionTargetBox="[306,1279,431,1668]" captionTargetId="figure-144@7.[298,1289,417,1675]" captionTargetPageId="7" captionText="FIGURE 4. Diaphorocleidus munimensis n. sp. from Bryconops cf. affinis (Characiformes, Iguanodectidae) from São José, municipality of Chapadinha and in Riacho Estrela, municipality of Anapurus, River Munim basin, Maranhão, Brazil: (a) total, ventral view; (b) copulatory complex, ventral view; (c) hook pairs 3, 4; (d) hook pairs 1,5; (e) ventral anchor; (f) dorsal anchor; (g) ventral bar; (h) dorsal bar. Scale bars: (a) 100 μm; (b) 15 μm; (c, d, g, h) 10 μm; (e, f) 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10561637" httpUri="https://zenodo.org/record/10561637/files/figure.png" pageId="6" pageNumber="97">Fig. 4A</figureCitation>
). Peduncle short. Haptor subhexagonal, 98 (78–118; n = 19) wide, with 7 pairs of hooks (
<figureCitation id="13165BC2FF988770FB6B0E82FAAB1ACE" box="[1235,1325,1692,1718]" captionStart="FIGURE 4" captionStartId="7.[151,250,1698,1722]" captionTargetBox="[306,1279,431,1668]" captionTargetId="figure-144@7.[298,1289,417,1675]" captionTargetPageId="7" captionText="FIGURE 4. Diaphorocleidus munimensis n. sp. from Bryconops cf. affinis (Characiformes, Iguanodectidae) from São José, municipality of Chapadinha and in Riacho Estrela, municipality of Anapurus, River Munim basin, Maranhão, Brazil: (a) total, ventral view; (b) copulatory complex, ventral view; (c) hook pairs 3, 4; (d) hook pairs 1,5; (e) ventral anchor; (f) dorsal anchor; (g) ventral bar; (h) dorsal bar. Scale bars: (a) 100 μm; (b) 15 μm; (c, d, g, h) 10 μm; (e, f) 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10561637" httpUri="https://zenodo.org/record/10561637/files/figure.png" pageId="6" pageNumber="97">Fig. 4A</figureCitation>
). Anchors dissimilar in size and shape. Ventral anchor with well-developed roots, superficial root rectangular and round short deep root, straight shaft and curved point, 32 (27–34, n = 15) long, base 20 (17–22, n = 15) wide (
<figureCitation id="13165BC2FF988770FB5E0EFAFAB81A86" box="[1254,1342,1764,1790]" captionStart="FIGURE 4" captionStartId="7.[151,250,1698,1722]" captionTargetBox="[306,1279,431,1668]" captionTargetId="figure-144@7.[298,1289,417,1675]" captionTargetPageId="7" captionText="FIGURE 4. Diaphorocleidus munimensis n. sp. from Bryconops cf. affinis (Characiformes, Iguanodectidae) from São José, municipality of Chapadinha and in Riacho Estrela, municipality of Anapurus, River Munim basin, Maranhão, Brazil: (a) total, ventral view; (b) copulatory complex, ventral view; (c) hook pairs 3, 4; (d) hook pairs 1,5; (e) ventral anchor; (f) dorsal anchor; (g) ventral bar; (h) dorsal bar. Scale bars: (a) 100 μm; (b) 15 μm; (c, d, g, h) 10 μm; (e, f) 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10561637" httpUri="https://zenodo.org/record/10561637/files/figure.png" pageId="6" pageNumber="97">Fig. 4E</figureCitation>
); Dorsal anchor with well-developed roots, superficial root elongated, deep root round and short, and a middle root between superficial and deep roots, straight shaft and short point, 21 (17–23, n = 15) long; base 15 (13–17, n = 15) wide (
<figureCitation id="13165BC2FF988770FF270F4EFF731B12" box="[159,245,1872,1898]" captionStart="FIGURE 4" captionStartId="7.[151,250,1698,1722]" captionTargetBox="[306,1279,431,1668]" captionTargetId="figure-144@7.[298,1289,417,1675]" captionTargetPageId="7" captionText="FIGURE 4. Diaphorocleidus munimensis n. sp. from Bryconops cf. affinis (Characiformes, Iguanodectidae) from São José, municipality of Chapadinha and in Riacho Estrela, municipality of Anapurus, River Munim basin, Maranhão, Brazil: (a) total, ventral view; (b) copulatory complex, ventral view; (c) hook pairs 3, 4; (d) hook pairs 1,5; (e) ventral anchor; (f) dorsal anchor; (g) ventral bar; (h) dorsal bar. Scale bars: (a) 100 μm; (b) 15 μm; (c, d, g, h) 10 μm; (e, f) 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10561637" httpUri="https://zenodo.org/record/10561637/files/figure.png" pageId="6" pageNumber="97">Fig. 4F</figureCitation>
). Ventral bar, robust, V-shaped, 37 (33–42, n = 12) long (
<figureCitation id="13165BC2FF988770FCC80F4EFC4C1B12" box="[880,970,1872,1898]" captionStart="FIGURE 4" captionStartId="7.[151,250,1698,1722]" captionTargetBox="[306,1279,431,1668]" captionTargetId="figure-144@7.[298,1289,417,1675]" captionTargetPageId="7" captionText="FIGURE 4. Diaphorocleidus munimensis n. sp. from Bryconops cf. affinis (Characiformes, Iguanodectidae) from São José, municipality of Chapadinha and in Riacho Estrela, municipality of Anapurus, River Munim basin, Maranhão, Brazil: (a) total, ventral view; (b) copulatory complex, ventral view; (c) hook pairs 3, 4; (d) hook pairs 1,5; (e) ventral anchor; (f) dorsal anchor; (g) ventral bar; (h) dorsal bar. Scale bars: (a) 100 μm; (b) 15 μm; (c, d, g, h) 10 μm; (e, f) 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10561637" httpUri="https://zenodo.org/record/10561637/files/figure.png" pageId="6" pageNumber="97">Fig. 4G</figureCitation>
); dorsal bar thin, long, slightly curved, 39 (33–46, n = 12) long (
<figureCitation id="13165BC2FF988770FE340F6AFE611BF6" box="[396,487,1908,1934]" captionStart="FIGURE 4" captionStartId="7.[151,250,1698,1722]" captionTargetBox="[306,1279,431,1668]" captionTargetId="figure-144@7.[298,1289,417,1675]" captionTargetPageId="7" captionText="FIGURE 4. Diaphorocleidus munimensis n. sp. from Bryconops cf. affinis (Characiformes, Iguanodectidae) from São José, municipality of Chapadinha and in Riacho Estrela, municipality of Anapurus, River Munim basin, Maranhão, Brazil: (a) total, ventral view; (b) copulatory complex, ventral view; (c) hook pairs 3, 4; (d) hook pairs 1,5; (e) ventral anchor; (f) dorsal anchor; (g) ventral bar; (h) dorsal bar. Scale bars: (a) 100 μm; (b) 15 μm; (c, d, g, h) 10 μm; (e, f) 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10561637" httpUri="https://zenodo.org/record/10561637/files/figure.png" pageId="6" pageNumber="97">Fig. 4H</figureCitation>
). Hooks similar in shape, with subunits of equal size, protruding thumb; hook pairs 1 and 5 reduced in size, 15 (15–16; n = 14) (
<figureCitation id="13165BC2FF988770FDCF0F86FD571BCA" box="[631,721,1944,1970]" captionStart="FIGURE 4" captionStartId="7.[151,250,1698,1722]" captionTargetBox="[306,1279,431,1668]" captionTargetId="figure-144@7.[298,1289,417,1675]" captionTargetPageId="7" captionText="FIGURE 4. Diaphorocleidus munimensis n. sp. from Bryconops cf. affinis (Characiformes, Iguanodectidae) from São José, municipality of Chapadinha and in Riacho Estrela, municipality of Anapurus, River Munim basin, Maranhão, Brazil: (a) total, ventral view; (b) copulatory complex, ventral view; (c) hook pairs 3, 4; (d) hook pairs 1,5; (e) ventral anchor; (f) dorsal anchor; (g) ventral bar; (h) dorsal bar. Scale bars: (a) 100 μm; (b) 15 μm; (c, d, g, h) 10 μm; (e, f) 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10561637" httpUri="https://zenodo.org/record/10561637/files/figure.png" pageId="6" pageNumber="97">Fig. 4D</figureCitation>
), pairs 3 and 4 longer than other pairs, 22 (20–22; n = 14) (
<figureCitation id="13165BC2FF988770FAD70F86FF3A1BAE" captionStart="FIGURE 4" captionStartId="7.[151,250,1698,1722]" captionTargetBox="[306,1279,431,1668]" captionTargetId="figure-144@7.[298,1289,417,1675]" captionTargetPageId="7" captionText="FIGURE 4. Diaphorocleidus munimensis n. sp. from Bryconops cf. affinis (Characiformes, Iguanodectidae) from São José, municipality of Chapadinha and in Riacho Estrela, municipality of Anapurus, River Munim basin, Maranhão, Brazil: (a) total, ventral view; (b) copulatory complex, ventral view; (c) hook pairs 3, 4; (d) hook pairs 1,5; (e) ventral anchor; (f) dorsal anchor; (g) ventral bar; (h) dorsal bar. Scale bars: (a) 100 μm; (b) 15 μm; (c, d, g, h) 10 μm; (e, f) 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10561637" httpUri="https://zenodo.org/record/10561637/files/figure.png" pageId="6" pageNumber="97">Fig. 4C</figureCitation>
). Pairs 2, 6 and 7 18 (17–19; n = 21) long. FH loop approximately 30% of shank length.
</paragraph>
</subSubSection>
<subSubSection id="C33714CCFF988771FF7F0FFEFCA11DF1" lastPageId="7" lastPageNumber="98" pageId="6" pageNumber="97" type="discussion">
<paragraph id="8B924747FF988771FF7F0FFEFCA11DF1" blockId="6.[151,1437,1224,2043]" lastBlockId="7.[151,1437,151,393]" lastPageId="7" lastPageNumber="98" pageId="6" pageNumber="97">
<emphasis id="B9599B55FF988770FF7F0FFEFEB21B82" bold="true" box="[199,308,2016,2042]" pageId="6" pageNumber="97">Remarks</emphasis>
: 
<taxonomicName id="4C2D3CC4FF988770FEFF0FFEFD141B82" authority="Silva &amp; Cohen &amp; Ottoni &amp; Viana &amp; Meneses &amp; Justo, 2024" authorityName="Silva &amp; Cohen &amp; Ottoni &amp; Viana &amp; Meneses &amp; Justo" authorityYear="2024" box="[327,658,2016,2042]" class="Monogenea" family="Ancyrocephalidae" genus="Diaphorocleidus" kingdom="Animalia" order="Dactylogyridea" pageId="6" pageNumber="97" phylum="Platyhelminthes" rank="species" species="munimensis" status="sp. nov.">
<emphasis id="B9599B55FF988770FEFF0FFEFD141B82" box="[327,658,2016,2042]" italics="true" pageId="6" pageNumber="97">Diaphorocleidus munimensis</emphasis>
</taxonomicName>
<emphasis id="B9599B55FF988770FD250FFEFD661B82" bold="true" box="[669,736,2016,2042]" pageId="6" pageNumber="97">
<taxonomicNameLabel id="A26A262EFF988770FD250FFEFD661B82" box="[669,736,2016,2042]" pageId="6" pageNumber="97" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
resembles 
<taxonomicName id="4C2D3CC4FF988770FCD00FFFFB871B82" authorityName="Jogunoori, Kritsky &amp; Venkatanarasaiah" authorityYear="2004" box="[872,1025,2016,2042]" class="Monogenea" family="Ancyrocephalidae" genus="Diaphorocleidus" kingdom="Animalia" order="Dactylogyridea" pageId="6" pageNumber="97" phylum="Platyhelminthes" rank="species" species="armillatus">
<emphasis id="B9599B55FF988770FCD00FFFFB871B82" box="[872,1025,2016,2042]" italics="true" pageId="6" pageNumber="97">D. armillatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2D3CC4FF988770FBF90FFFFB7E1B82" authority="Silva &amp; Cohen &amp; Ottoni &amp; Viana &amp; Meneses &amp; Justo, 2024" authorityName="Silva &amp; Cohen &amp; Ottoni &amp; Viana &amp; Meneses &amp; Justo" authorityYear="2024" box="[1089,1272,2016,2043]" class="Monogenea" family="Ancyrocephalidae" genus="Diaphorocleidus" kingdom="Animalia" order="Dactylogyridea" pageId="6" pageNumber="97" phylum="Platyhelminthes" rank="species" species="forcipiformis" status="n. sp.">
<emphasis id="B9599B55FF988770FBF90FFFFBD31B82" box="[1089,1109,2017,2042]" italics="true" pageId="6" pageNumber="97">D</emphasis>
. 
<emphasis id="B9599B55FF988770FBDF0FFEFB7E1B82" box="[1127,1272,2016,2042]" italics="true" pageId="6" pageNumber="97">forcipiformis</emphasis>
</taxonomicName>
<emphasis id="B9599B55FF988770FABC0FFEFAC11B82" bold="true" box="[1284,1351,2016,2042]" pageId="6" pageNumber="97">
<taxonomicNameLabel id="A26A262EFF988770FABC0FFEFAC11B82" box="[1284,1351,2016,2042]" pageId="6" pageNumber="97" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by the number of the rings of the MCO (2 counterclockwise). However, the new species differs from these mainly by the morphology of the accessory piece (plate-like in 
<taxonomicName id="4C2D3CC4FF998771FD7A08A5FCDE1CAD" authorityName="Jogunoori, Kritsky &amp; Venkatanarasaiah" authorityYear="2004" box="[706,856,187,213]" class="Monogenea" family="Ancyrocephalidae" genus="Diaphorocleidus" kingdom="Animalia" order="Dactylogyridea" pageId="7" pageNumber="98" phylum="Platyhelminthes" rank="species" species="armillatus">
<emphasis id="B9599B55FF998771FD7A08A5FCDE1CAD" box="[706,856,187,213]" italics="true" pageId="7" pageNumber="98">D. armillatus</emphasis>
</taxonomicName>
, and bifurcated, claw-shaped in 
<taxonomicName id="4C2D3CC4FF998771FB7108A5FAF81CAD" authority="Silva &amp; Cohen &amp; Ottoni &amp; Viana &amp; Meneses &amp; Justo, 2024" authorityName="Silva &amp; Cohen &amp; Ottoni &amp; Viana &amp; Meneses &amp; Justo" authorityYear="2024" box="[1225,1406,187,213]" class="Monogenea" family="Ancyrocephalidae" genus="Diaphorocleidus" kingdom="Animalia" order="Dactylogyridea" pageId="7" pageNumber="98" phylum="Platyhelminthes" rank="species" species="forcipiformis" status="n. sp.">
<emphasis id="B9599B55FF998771FB7108A5FB5B1CAC" box="[1225,1245,187,212]" italics="true" pageId="7" pageNumber="98">D</emphasis>
. 
<emphasis id="B9599B55FF998771FB5508A5FAF81CAD" box="[1261,1406,187,213]" italics="true" pageId="7" pageNumber="98">forcipiformis</emphasis>
</taxonomicName>
<emphasis id="B9599B55FF998771FA3E08A5FF3E1C81" bold="true" pageId="7" pageNumber="98">
<taxonomicNameLabel id="A26A262EFF998771FA3E08A5FF3E1C81" pageId="7" pageNumber="98" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
). 
<taxonomicName id="4C2D3CC4FF998771FF6808C1FD911C80" authority="Silva &amp; Cohen &amp; Ottoni &amp; Viana &amp; Meneses &amp; Justo, 2024" authorityName="Silva &amp; Cohen &amp; Ottoni &amp; Viana &amp; Meneses &amp; Justo" authorityYear="2024" box="[208,535,223,249]" class="Monogenea" family="Ancyrocephalidae" genus="Diaphorocleidus" kingdom="Animalia" order="Dactylogyridea" pageId="7" pageNumber="98" phylum="Platyhelminthes" rank="species" species="munimensis" status="sp. nov.">
<emphasis id="B9599B55FF998771FF6808C1FD911C80" box="[208,535,223,249]" italics="true" pageId="7" pageNumber="98">Diaphorocleidus munimensis</emphasis>
</taxonomicName>
<emphasis id="B9599B55FF998771FDA608C1FDDB1C81" bold="true" box="[542,605,223,249]" pageId="7" pageNumber="98">
<taxonomicNameLabel id="A26A262EFF998771FDA608C1FDDB1C81" box="[542,605,223,249]" pageId="7" pageNumber="98" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
also differs from 
<taxonomicName id="4C2D3CC4FF998771FC9F08C1FC3B1C81" authorityName="Jogunoori, Kritsky &amp; Venkatanarasaiah" authorityYear="2004" box="[807,957,223,249]" class="Monogenea" family="Ancyrocephalidae" genus="Diaphorocleidus" kingdom="Animalia" order="Dactylogyridea" pageId="7" pageNumber="98" phylum="Platyhelminthes" rank="species" species="armillatus">
<emphasis id="B9599B55FF998771FC9F08C1FC3B1C81" box="[807,957,223,249]" italics="true" pageId="7" pageNumber="98">D. armillatus</emphasis>
</taxonomicName>
by the body length (357 long by 155 wide in the new species 
<emphasis id="B9599B55FF998771FED6091DFE031D64" box="[366,389,259,284]" italics="true" pageId="7" pageNumber="98">vs</emphasis>
280 long by 78 wide in 
<taxonomicName id="4C2D3CC4FF998771FD23091DFCB71D65" authorityName="Jogunoori, Kritsky &amp; Venkatanarasaiah" authorityYear="2004" box="[667,817,259,285]" class="Monogenea" family="Ancyrocephalidae" genus="Diaphorocleidus" kingdom="Animalia" order="Dactylogyridea" pageId="7" pageNumber="98" phylum="Platyhelminthes" rank="species" species="armillatus">
<emphasis id="B9599B55FF998771FD23091DFCB71D65" box="[667,817,259,285]" italics="true" pageId="7" pageNumber="98">D. armillatus</emphasis>
</taxonomicName>
). By the similarity in the morphology of the accessory piece (pincer-shaped, formed by two support processes), the new species resembles 
<taxonomicName id="4C2D3CC4FF998771FB8D0939FB3A1D38" authorityName="Mendoza-Franco, Aguirre-Macedo &amp; Vidal-Martinez" authorityYear="2007" box="[1077,1212,295,320]" class="Monogenea" family="Ancyrocephalidae" genus="Diaphorocleidus" kingdom="Animalia" order="Dactylogyridea" pageId="7" pageNumber="98" phylum="Platyhelminthes" rank="species" species="petrosusi">
<emphasis id="B9599B55FF998771FB8D0939FB3A1D38" box="[1077,1212,295,320]" italics="true" pageId="7" pageNumber="98">D. petrosusi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2D3CC4FF998771FB570939FD371D1D" authority="Santos Neto, Costa, Soares &amp; Domingues, 2018" authorityName="Santos Neto, Costa, Soares &amp; Domingues" authorityYear="2018" class="Monogenea" family="Ancyrocephalidae" genus="Diaphorocleidus" kingdom="Animalia" order="Dactylogyridea" pageId="7" pageNumber="98" phylum="Platyhelminthes" rank="species" species="sclerocolpus">
<emphasis id="B9599B55FF998771FB570939FA1B1D39" box="[1263,1437,295,321]" italics="true" pageId="7" pageNumber="98">D. sclerocolpus</emphasis>
<bibRefCitation id="EFBC3AB6FF998771FF2F0955FD371D1D" author="Santos Neto, J. F. &amp; Costa, N. G. S. &amp; Soares, G. B. &amp; Domingues, M. V." box="[151,689,331,357]" pageId="7" pageNumber="98" pagination="208 - 219" refId="ref9811" refString="Santos Neto, J. F., Costa, N. G. S., Soares, G. B. &amp; Domingues, M. V. (2018) Monogenoidean parasites of Acestrorhynchus falcatus (Characiformes: Acestrorhynchidae) from Para, Brazil: species of Diaphorocleidus and Rhinoxenoides n. gen. (Monogenoidea: Dactylogyridae). Journal of Helminthology, 93 (2), 208 - 219. https: // doi. org / 10.1017 / S 0022149 X 18000019" type="journal article" year="2018">Santos Neto, Costa, Soares &amp; Domingues, 2018</bibRefCitation>
</taxonomicName>
, but differs by the width of the two support processes, which in the new species is very robust, one being wider than the other.
</paragraph>
</subSubSection>
</treatment>
</document>