444 lines
56 KiB
XML
444 lines
56 KiB
XML
<document id="0DAD06F181CAF4103B83CF77E67B5A7A" ID-CLB-Dataset="45914" ID-DOI="10.5281/zenodo.207028" ID-GBIF-Dataset="77b6eef4-11f7-4dd3-a6f4-c28e5655e2bc" ID-ISSN="1175-5326" ID-Zenodo-Dep="207028" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460413407798" checkinUser="plazi" docAuthor="Venegas, Pablo J., Duran, Vilma, Landauro, Caroll Z. & Lujan, Lesly" docDate="2011" docId="03838E2C0A40FFE7FF0B8379FD1FF82D" docLanguage="en" docName="zt03109p048.pdf" docOrigin="Zootaxa 3109" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Enyalioides rudolfarndti Venegas, Duran, Landauro & Lujan, 2011, sp. nov." docType="treatment" docVersion="10" lastPageNumber="45" masterDocId="FFBAF6540A41FFE1FF9C813FFFBFFFEC" masterDocTitle="A distinctive new species of wood lizard (Hoplocercinae, Enyalioides) from the Yanachaga Chemillen National Park in central Peru" masterLastPageNumber="48" masterPageNumber="39" pageNumber="40" updateTime="1698328212273" updateUser="ExternalLinkService">
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<mods:title id="735FC98E52675C41B6A99F881716B33A">A distinctive new species of wood lizard (Hoplocercinae, Enyalioides) from the Yanachaga Chemillen National Park in central Peru</mods:title>
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<mods:name id="BCDE560F276906A57071D576EC3C4AA7" type="personal">
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<mods:namePart id="73DB4328F1CC4B1E7AB033AC59139D91">Venegas, Pablo J.</mods:namePart>
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<mods:roleTerm id="EBBD3CDAE091A285E873E9E45C80040D">Author</mods:roleTerm>
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<mods:namePart id="D518EC03AC3E40304D00F4C76FBEB9E8">Duran, Vilma</mods:namePart>
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<mods:roleTerm id="B314657B1F7C511E4CEC51F5CEF9C61B">Author</mods:roleTerm>
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<mods:namePart id="84D345D3436D972ADE2F11A2005DDFD3">Landauro, Caroll Z.</mods:namePart>
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<mods:namePart id="FB32BF7EFD5D73D1CAF317E45C5F0924">Lujan, Lesly</mods:namePart>
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<mods:title id="F0A2FEF0BECA3D23959EC90498641D28">Zootaxa</mods:title>
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<mods:part id="5A6008207039933942EFBD57A0A0D61B">
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<mods:date id="53ACCA58F855E53575538FC188F49A33">2011</mods:date>
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<treatment id="03838E2C0A40FFE7FF0B8379FD1FF82D" ID-DOI="http://doi.org/10.5281/zenodo.6192050" ID-GBIF-Taxon="119424172" ID-Zenodo-Dep="6192050" LSID="urn:lsid:plazi:treatment:03838E2C0A40FFE7FF0B8379FD1FF82D" httpUri="http://treatment.plazi.org/id/03838E2C0A40FFE7FF0B8379FD1FF82D" lastPageId="6" lastPageNumber="45" pageId="1" pageNumber="40">
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<subSubSection id="C3306CB10A40FFE0FF0B8379FC12FD01" pageId="1" pageNumber="40" type="nomenclature">
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<paragraph id="8B953F3A0A40FFE0FF0B8379FD9BFD8C" blockId="1.[151,548,582,641]" box="[151,548,582,608]" pageId="1" pageNumber="40">
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<heading id="D0DD88560A40FFE0FF0B8379FD9BFD8C" bold="true" box="[151,548,582,608]" fontSize="11" level="1" pageId="1" pageNumber="40" reason="1">
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<emphasis id="B95EE3280A40FFE0FF0B8379FD9BFD8C" bold="true" box="[151,548,582,608]" pageId="1" pageNumber="40">
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<taxonomicName id="4C2A44B90A40FFE0FF0B8379FE7FFD8C" box="[151,448,582,608]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="rudolfarndti" status="sp. nov.">
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<emphasis id="B95EE3280A40FFE0FF0B8379FE7FFD8C" bold="true" box="[151,448,582,608]" italics="true" pageId="1" pageNumber="40">Enyalioides rudolfarndti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26D5E530A40FFE0FE5B8379FD9BFD8C" box="[455,548,582,608]" pageId="1" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B953F3A0A40FFE0FF0B8356FEAEFD6D" blockId="1.[151,548,582,641]" box="[151,273,617,641]" pageId="1" pageNumber="40">
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(
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<figureCitation id="131123BF0A40FFE0FF038356FEB6FD6D" box="[159,265,617,641]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1903,1926]" captionStartId-1="3.[151,250,1927,1950]" captionStartId-2="4.[151,250,1752,1775]" captionTargetBox-0="[151,1436,916,1879]" captionTargetBox-1="[151,1436,193,1904]" captionTargetBox-2="[188,1407,203,1689]" captionTargetId-0="figure@2.[151,1436,916,1880]" captionTargetId-1="figure@3.[151,1436,193,1906]" captionTargetId-2="figure@4.[151,1436,174,1749]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Dorsolateral view of the holotype of Enyalioides rudolfarndti sp. n. (CORBIDI 0 7209, adult male, SVL 116 mm). Photograph by P. J. Venegas." captionText-1="FIGURE 2. Close-up of head of Enyalioides rudolfarndti sp. n. (CORBIDI 07209) showing the orange anthehumeral blotch. Photograph by P. J. Venegas." captionText-2="FIGURE 3. Dorsolateral and ventral views of the paratypes of Enyalioides rudolfarndti sp. n. Adult male (A, B, CORBIDI 07212), adult female (C, D, CORBIDI 07210), and juvenile (E, F, CORBIDI 07213). Photographs by P. J. Venegas." httpUri-0="https://zenodo.org/record/207029/files/figure.png" httpUri-1="https://zenodo.org/record/207030/files/figure.png" httpUri-2="https://zenodo.org/record/207031/files/figure.png" pageId="1" pageNumber="40">Figs. 1–3</figureCitation>
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)
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</paragraph>
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<paragraph id="8B953F3A0A40FFE0FF0B838EFC93FD25" blockId="1.[151,941,689,750]" box="[151,812,689,713]" pageId="1" pageNumber="40">Proposed standard English name: throat-sliced wood lizard</paragraph>
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<paragraph id="8B953F3A0A40FFE0FF0B83EAFC12FD01" blockId="1.[151,941,689,750]" box="[151,941,725,750]" pageId="1" pageNumber="40">Proposed standard Spanish name: lagartija de palo de garganta cortada</paragraph>
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</subSubSection>
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<subSubSection id="C3306CB10A40FFE0FF0B8222FCDFFC06" pageId="1" pageNumber="40" type="materials_examined">
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<paragraph id="8B953F3A0A40FFE0FF0B8222FC93FC92" blockId="1.[151,1437,797,2009]" pageId="1" pageNumber="40">
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<emphasis id="B95EE3280A40FFE0FF0B8222FEB4FCD9" bold="true" box="[151,267,797,821]" pageId="1" pageNumber="40">
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<typeStatus id="549181980A40FFE0FF0B8222FEB9FCD9" box="[151,262,797,821]" pageId="1" pageNumber="40" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED3BA7FF0A40FFE0FE888222FE38FCD9" box="[276,391,797,821]" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="1" pageNumber="40">CORBIDI</collectionCode>
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0 7209 (
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<figureCitation id="131123BF0A40FFE0FE758222FDE8FCD9" box="[489,599,797,821]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1903,1926]" captionStartId-1="3.[151,250,1927,1950]" captionTargetBox-0="[151,1436,916,1879]" captionTargetBox-1="[151,1436,193,1904]" captionTargetId-0="figure@2.[151,1436,916,1880]" captionTargetId-1="figure@3.[151,1436,193,1906]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Dorsolateral view of the holotype of Enyalioides rudolfarndti sp. n. (CORBIDI 0 7209, adult male, SVL 116 mm). Photograph by P. J. Venegas." captionText-1="FIGURE 2. Close-up of head of Enyalioides rudolfarndti sp. n. (CORBIDI 07209) showing the orange anthehumeral blotch. Photograph by P. J. Venegas." httpUri-0="https://zenodo.org/record/207029/files/figure.png" httpUri-1="https://zenodo.org/record/207030/files/figure.png" pageId="1" pageNumber="40">Figs. 1–2</figureCitation>
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), an adult male from the Pan de Azucar trail near the Puesto de Control Huampal in the
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<collectionCode id="ED3BA7FF0A40FFE0FED0827EFE28FCB5" box="[332,407,833,857]" pageId="1" pageNumber="40">YCNP</collectionCode>
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(
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<geoCoordinate id="EE1E59FD0A40FFE0FE3B827EFD87FCB5" box="[423,568,833,857]" direction="south" orientation="latitude" pageId="1" pageNumber="40" precision="15" value="-10.184167">10°11´03´´ S</geoCoordinate>
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<geoCoordinate id="EE1E59FD0A40FFE0FDDC827EFD60FCB5" box="[576,735,833,857]" direction="west" orientation="longitude" pageId="1" pageNumber="40" precision="15" value="-75.574165">75°34´27´´ W</geoCoordinate>
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; WGS 84) at
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<quantity id="4CD292DF0A40FFE0FCE5827EFC6AFCB5" box="[889,981,833,857]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="40" unit="m" value="1050.0">1050 m</quantity>
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.a.s.l., collected on
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<date id="FF9419FA0A40FFE0FB34827EFA83FCB5" box="[1192,1340,833,857]" pageId="1" pageNumber="40" value="2010-08-16">16.VIII.2010</date>
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, Provincia de Oxapampa, Región de Pasco,
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<collectingCountry id="F33D7FAA0A40FFE0FDB38259FDDAFC91" box="[559,613,870,893]" name="Peru" pageId="1" pageNumber="40">Peru</collectingCountry>
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, by P. J. Venegas.
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</paragraph>
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<paragraph id="8B953F3A0A40FFE0FF5B82B5FCDFFC06" blockId="1.[151,1437,797,2009]" pageId="1" pageNumber="40">
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<emphasis id="B95EE3280A40FFE0FF5B82B5FEF9FC4E" bold="true" box="[199,326,906,930]" pageId="1" pageNumber="40">
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<typeStatus id="549181980A40FFE0FF5B82B5FEFEFC4E" box="[199,321,906,930]" pageId="1" pageNumber="40" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED3BA7FF0A40FFE0FED182B6FE7FFC4D" box="[333,448,905,929]" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="1" pageNumber="40">CORBIDI</collectionCode>
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0 7210 and 0 7213, an adult female and a juvenile, respectively, collected with the
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<typeStatus id="549181980A40FFE0FAFD82B6FF78FC2A" pageId="1" pageNumber="40" type="holotype">holotype</typeStatus>
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by P. J. Venegas, V. Duran, C. Z. Landauro, and L. Lujan.
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<collectionCode id="ED3BA7FF0A40FFE0FCC88292FC78FC29" box="[852,967,941,965]" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="1" pageNumber="40">CORBIDI</collectionCode>
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0 7212, an adult male from the same location of the
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<typeStatus id="549181980A40FFE0FE8D82EEFECCFC05" box="[273,371,977,1001]" pageId="1" pageNumber="40" type="holotype">holotype</typeStatus>
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but taken on
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<date id="FF9419FA0A40FFE0FD9782EEFD21FC05" box="[523,670,977,1001]" pageId="1" pageNumber="40" value="2010-08-19">19.VIII.2010</date>
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by P. J. Venegas.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3306CB10A40FFE0FF5B82CAFBD3F979" pageId="1" pageNumber="40" type="diagnosis">
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<paragraph id="8B953F3A0A40FFE0FF5B82CAFBD3F979" blockId="1.[151,1437,797,2009]" pageId="1" pageNumber="40">
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<emphasis id="B95EE3280A40FFE0FF5B82CAFEFEFBE2" bold="true" box="[199,321,1013,1038]" pageId="1" pageNumber="40">Diagnosis.</emphasis>
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<taxonomicName id="4C2A44B90A40FFE0FED782CAFDDDFBE1" box="[331,610,1013,1037]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="rudolfarndti">
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<emphasis id="B95EE3280A40FFE0FED782CAFDDDFBE1" box="[331,610,1013,1037]" italics="true" pageId="1" pageNumber="40">Enyalioides rudolfarndti</emphasis>
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</taxonomicName>
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can be easily distinguished from other species of
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<taxonomicName id="4C2A44B90A40FFE0FB3782CAFA90FBE1" box="[1195,1327,1013,1037]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="40" phylum="Chordata" rank="genus">
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<emphasis id="B95EE3280A40FFE0FB3782CAFA90FBE1" box="[1195,1327,1013,1037]" italics="true" pageId="1" pageNumber="40">Enyalioides</emphasis>
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</taxonomicName>
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from the Amazon basin by the combination of the following characters: (1) scales posterior to the superciliaries enlarged (relative to adjacent scales), forming a well defined longitudinal row of distinctly raised scales across the lateral edge of the head in juveniles and adults of both sexes; (2) 30 or fewer longitudinal rows of dorsals in a transverse line between the dorsolateral crests at midbody; (3) a distinct orange round blotch on the antehumeral region in adult males; (4) ventral scales strongly keeled; (5) caudal scales heterogeneous in size on each autotomic segment. The orange round blotch on the antehumeral region in adult males of
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<taxonomicName id="4C2A44B90A40FFE0FC2785F2FB6EFB09" box="[955,1233,1229,1253]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="rudolfarndti">
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<emphasis id="B95EE3280A40FFE0FC2785F2FB6EFB09" box="[955,1233,1229,1253]" italics="true" pageId="1" pageNumber="40">Enyalioides rudolfarndti</emphasis>
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</taxonomicName>
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is present also in some male individuals of
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<taxonomicName id="4C2A44B90A40FFE0FE2685CDFDE5FAE5" box="[442,602,1265,1289]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="palpebralis">
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<emphasis id="B95EE3280A40FFE0FE2685CDFDE5FAE5" box="[442,602,1265,1289]" italics="true" pageId="1" pageNumber="40">E. palpebralis</emphasis>
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</taxonomicName>
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(
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<figureCitation id="131123BF0A40FFE0FDF685CEFD12FAE5" box="[618,685,1265,1289]" captionStart="FIGURE 4" captionStartId="6.[151,250,1071,1094]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@6.[151,1436,193,1050]" captionTargetPageId="6" captionText="FIGURE 4. Close-up of head of Enyalioides palpebralis (CORBIDI 0 8347, adult male, SVL 112 mm) showing the orange antehumeral blotch. Photograph by G. Chávez." httpUri="https://zenodo.org/record/207032/files/figure.png" pageId="1" pageNumber="40">Fig. 4</figureCitation>
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). Furthermore, the new species also shares with
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<taxonomicName id="4C2A44B90A40FFE0FB4F85CDFACDFAE5" box="[1235,1394,1265,1289]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="palpebralis">
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<emphasis id="B95EE3280A40FFE0FB4F85CDFACDFAE5" box="[1235,1394,1265,1289]" italics="true" pageId="1" pageNumber="40">E. palpebralis</emphasis>
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</taxonomicName>
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the presence of enlarged scales posterior to the superciliaries and the presence of strongly keeled ventral scales; however, the latter species can be easily distinguished from
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<taxonomicName id="4C2A44B90A40FFE0FC9B8405FC10FABD" box="[775,943,1337,1361]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="rudolfarndti">
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<emphasis id="B95EE3280A40FFE0FC9B8405FC10FABD" box="[775,943,1337,1361]" italics="true" pageId="1" pageNumber="40">E. rudolfarndti</emphasis>
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</taxonomicName>
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by having a superciliary triangular flap that projects posterolaterally over each eye. In addition,
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<taxonomicName id="4C2A44B90A40FFE0FD7F8461FC3CFA99" box="[739,899,1373,1397]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="palpebralis">
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<emphasis id="B95EE3280A40FFE0FD7F8461FC3CFA99" box="[739,899,1373,1397]" italics="true" pageId="1" pageNumber="40">E. palpebralis</emphasis>
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</taxonomicName>
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is the only species that has (most specimens) a discontinuous vertebral crest, having a small gap on the neck, and lacks femoral pores (
|
||
<bibRefCitation id="EFBB42CB0A40FFE0FBC384BEFA2FFA75" author="Torres-Carvajal" box="[1119,1424,1409,1433]" pageId="1" pageNumber="40" refString="Torres-Carvajal, O., Etheridge, R. & de Queiroz, K. (2011) A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania). Zootaxa, 2752, 1 - 44." type="journal article" year="2011">
|
||
Torres-Carvajal
|
||
<emphasis id="B95EE3280A40FFE0FA8B84BDFAF1FA75" box="[1303,1358,1409,1433]" italics="true" pageId="1" pageNumber="40">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C2A44B90A40FFE0FF0B849AFE24FA51" box="[151,411,1445,1469]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="cofanorum">
|
||
<emphasis id="B95EE3280A40FFE0FF0B849AFE24FA51" box="[151,411,1445,1469]" italics="true" pageId="1" pageNumber="40">Enyalioides cofanorum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2A44B90A40FFE0FE4E8499FDD9FA51" box="[466,614,1445,1469]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="microlepis">
|
||
<emphasis id="B95EE3280A40FFE0FE4E8499FDD9FA51" box="[466,614,1445,1469]" italics="true" pageId="1" pageNumber="40">E. microlepis</emphasis>
|
||
</taxonomicName>
|
||
share with the new species the presence of strongly keeled ventral scales and caudal scales of heterogeneous size on each autotomic segment. However,
|
||
<taxonomicName id="4C2A44B90A40FFE0FB9784F5FB1CFA0D" box="[1035,1187,1481,1505]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="cofanorum">
|
||
<emphasis id="B95EE3280A40FFE0FB9784F5FB1CFA0D" box="[1035,1187,1481,1505]" italics="true" pageId="1" pageNumber="40">E. cofanorum</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C2A44B90A40FFE0FAAB84F5FF5CF9E9" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="rudolfarndti">
|
||
<emphasis id="B95EE3280A40FFE0FAAB84F5FF5CF9E9" italics="true" pageId="1" pageNumber="40">E. rudolfarndti</emphasis>
|
||
</taxonomicName>
|
||
(characters in parentheses) by having with more than 33 dorsal scales in a transverse line between the dorsolateral crests at midbody (30 or fewer) and scattered, projecting, large dorsal scales (absent).
|
||
<taxonomicName id="4C2A44B90A40FFE0FA84872EFEB3F9A1" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="microlepis">
|
||
<emphasis id="B95EE3280A40FFE0FA84872EFEB3F9A1" italics="true" pageId="1" pageNumber="40">Enyalioides microlepis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C2A44B90A40FFE0FE028709FDF9F9A1" box="[414,582,1589,1613]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="rudolfarndti">
|
||
<emphasis id="B95EE3280A40FFE0FE028709FDF9F9A1" box="[414,582,1589,1613]" italics="true" pageId="1" pageNumber="40">E. rudolfarndti</emphasis>
|
||
</taxonomicName>
|
||
by having more than 40 dorsal scales in a transverse line between the dorsolateral crest at midbody (30 or fewer dorsal scales), and a low vertebral crest (high). In addition, adult males of
|
||
<taxonomicName id="4C2A44B90A40FFE0FA188765FEAFF979" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="cofanorum">
|
||
<emphasis id="B95EE3280A40FFE0FA188765FEAFF979" italics="true" pageId="1" pageNumber="40">E. cofanorum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2A44B90A40FFE0FEDA8741FE64F979" box="[326,475,1661,1685]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="microlepis">
|
||
<emphasis id="B95EE3280A40FFE0FEDA8741FE64F979" box="[326,475,1661,1685]" italics="true" pageId="1" pageNumber="40">E. microlepis</emphasis>
|
||
</taxonomicName>
|
||
have a black patch that covering the gular region (absent).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3306CB10A40FFE4FF59879EFF52F874" lastPageId="5" lastPageNumber="44" pageId="1" pageNumber="40" type="description">
|
||
<paragraph id="8B953F3A0A40FFE3FF59879EFDADFEF0" blockId="1.[151,1437,797,2009]" lastBlockId="2.[151,1437,151,860]" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40">
|
||
<emphasis id="B95EE3280A40FFE0FF59879EFE5CF956" bold="true" box="[197,483,1697,1722]" pageId="1" pageNumber="40">
|
||
Description of
|
||
<typeStatus id="549181980A40FFE0FEEA879DFE60F956" box="[374,479,1698,1722]" pageId="1" pageNumber="40" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Male (
|
||
<figureCitation id="131123BF0A40FFE0FDA8879EFD1EF955" box="[564,673,1697,1721]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1903,1926]" captionStartId-1="3.[151,250,1927,1950]" captionTargetBox-0="[151,1436,916,1879]" captionTargetBox-1="[151,1436,193,1904]" captionTargetId-0="figure@2.[151,1436,916,1880]" captionTargetId-1="figure@3.[151,1436,193,1906]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Dorsolateral view of the holotype of Enyalioides rudolfarndti sp. n. (CORBIDI 0 7209, adult male, SVL 116 mm). Photograph by P. J. Venegas." captionText-1="FIGURE 2. Close-up of head of Enyalioides rudolfarndti sp. n. (CORBIDI 07209) showing the orange anthehumeral blotch. Photograph by P. J. Venegas." httpUri-0="https://zenodo.org/record/207029/files/figure.png" httpUri-1="https://zenodo.org/record/207030/files/figure.png" pageId="1" pageNumber="40">Figs. 1–2</figureCitation>
|
||
); SVL=
|
||
<quantity id="4CD292DF0A40FFE0FC9F879EFCDDF955" box="[771,866,1697,1721]" metricMagnitude="-1" metricUnit="m" metricValue="1.16" pageId="1" pageNumber="40" unit="mm" value="116.0">116 mm</quantity>
|
||
; TL =
|
||
<quantity id="4CD292DF0A40FFE0FC33879EFBAFF955" box="[943,1040,1697,1721]" metricMagnitude="-1" metricUnit="m" metricValue="1.91" pageId="1" pageNumber="40" unit="mm" value="191.0">191 mm</quantity>
|
||
; maximum head width =
|
||
<quantity id="4CD292DF0A40FFE0FAAC879EFA27F955" box="[1328,1432,1697,1721]" metricMagnitude="-2" metricUnit="m" metricValue="2.32" pageId="1" pageNumber="40" unit="mm" value="23.2">23.2 mm</quantity>
|
||
; head length =
|
||
<quantity id="4CD292DF0A40FFE0FEAB87FAFE21F931" box="[311,414,1733,1757]" metricMagnitude="-2" metricUnit="m" metricValue="3.16" pageId="1" pageNumber="40" unit="mm" value="31.6">31.6 mm</quantity>
|
||
; head height =
|
||
<quantity id="4CD292DF0A40FFE0FDD587FAFD24F931" box="[585,667,1733,1757]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="40" unit="mm" value="23.0">23 mm</quantity>
|
||
; dorsal head scales including parietal region multicarinate; scales on lateral edge of head just posterior to superciliaries enlarged, forming a well defined longitudinal row of 7 (left) or 6 (right) distinctly raised scales; temporal scales small, conical, juxtaposed, nearly homogeneous in size; 1 large, pretympanic conical scale in anterodorsal margin of the tympanum; superciliaries 17; canthals 4; postrostrals 4; left supralabials 13 if counted to a point right below middle of eye, and 16 if counted to commisure of mouth (13 and 16 on right side, respectively); rostral (2.92 ×
|
||
<quantity id="4CD292DF0A40FFE0FD378646FCADF87D" box="[683,786,1913,1937]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="1" pageNumber="40" unit="mm" value="1.16">1.16 mm</quantity>
|
||
) about twice as wide as the adjacent supralabials; single longitudinal row of lorilabials between suboculars and supralabials at level of middle of the eye, longitudinal rows of lorilabials anterior to this point 2–3; loreal region broken into small, smooth, and juxtaposed scales; nasal at the level of supralabial IV; left infralabials 10 if counted to a point right below middle of eye, and 14 if counted to commisure of mouth (11 and 16 on right side, respectively); mental (2.77 ×
|
||
<quantity id="4CD292DF0A43FFE3FC598183FB94FF38" box="[965,1067,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" pageId="2" pageNumber="41" unit="mm" value="1.39">1.39 mm</quantity>
|
||
) wider and higher than the adjacent infralabials; postmentals 2; gulars ventrally projected; gular fold weakly defined, complete midventrally; neck with only 2 distinct oblique folds.
|
||
</paragraph>
|
||
<paragraph id="8B953F3A0A43FFE3FF598018FDF3FD68" blockId="2.[151,1437,151,860]" pageId="2" pageNumber="41">Vertebral crest strongly projecting, decreasing in size posteriorly, with vertebrals on the neck at least four times higher than vertebrals between the hind limbs; crest bifurcates posteriorly at base of the tail and extends onto the tail for less than one-third of its length; flanks between forelimbs and hind limbs with dorsolateral and ventrolateral folds, as well as some oblique folds; scales on dorsolateral folds slightly larger than on the adjacent scales and forming a distinct row of raised scales; dorsal scales between dorsolateral fold and vertebral crest heterogeneous in size with small and large, strongly keeled, and imbricate scales; scales on flanks (i.e., ventral to dorsolateral fold) heterogeneous in size with some scales similar in size to dorsals surrounded by minute keeled scales, and some scattered enlarged scales with projecting keels that are two to three times larger than the adjacent scales on the near hind limb insertion; ventral scales imbricate, strongly keeled, rectangular, without a posterolateral mucron; ventrals more than twice the size of the dorsals.</paragraph>
|
||
<paragraph id="8B953F3A0A43FFE3FF5983B0FCF0FCB0" blockId="2.[151,1437,151,860]" pageId="2" pageNumber="41">
|
||
Limb scales keeled dorsally and ventrally; thigh scales homogeneous in size dorsally and heterogeneous in size posteriorly, with most scales less than half of the size of those scales on the anterior and ventral aspects; subdigitals on Finger
|
||
<date id="FF9419FA0A43FFE3FE9783E7FEEFFD03" box="[267,336,727,751]" pageId="2" pageNumber="41" value="1919-04">IV 19</date>
|
||
; subdigitals on Toe
|
||
<date id="FF9419FA0A43FFE3FDAE83E7FDC9FD03" box="[562,630,727,751]" pageId="2" pageNumber="41" value="1925-04">IV 25</date>
|
||
; femoral pore on each side 1; tail compressed and gradually decreasing in height towards tip; caudal scales strongly keeled and imbricate, increasing in size posteriorly on lateral and dorsal aspects of each autotomic segment; ventral caudals larger than dorsal caudals, with individual autotomic segments being 3 scales long ventrally and 5 scales long dorsally.
|
||
</paragraph>
|
||
<caption id="DF556FB20A43FFE3FF0B8650FE03F848" httpUri="https://zenodo.org/record/207029/files/figure.png" pageId="2" pageNumber="41" targetBox="[151,1436,916,1879]" targetPageId="2">
|
||
<paragraph id="8B953F3A0A43FFE3FF0B8650FE03F848" blockId="2.[151,1436,1903,1956]" pageId="2" pageNumber="41">
|
||
<emphasis id="B95EE3280A43FFE3FF0B8650FEACF869" bold="true" box="[151,275,1903,1926]" pageId="2" pageNumber="41">FIGURE 1.</emphasis>
|
||
Dorsolateral view of the holotype of
|
||
<taxonomicName id="4C2A44B90A43FFE3FD118650FC38F869" box="[653,903,1903,1925]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="41" phylum="Chordata" rank="species" species="rudolfarndti" status="sp. nov.">
|
||
<emphasis id="B95EE3280A43FFE3FD118650FC38F869" box="[653,903,1903,1925]" italics="true" pageId="2" pageNumber="41">Enyalioides rudolfarndti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EE3280A43FFE3FC11864FFC79F869" bold="true" box="[909,966,1904,1925]" pageId="2" pageNumber="41">
|
||
<taxonomicNameLabel id="A26D5E530A43FFE3FC11864FFC79F869" box="[909,966,1904,1925]" pageId="2" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(CORBIDI 0 7209, adult male, SVL 116 mm). Photograph by P. J. Venegas.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF556FB20A42FFE2FF0B86B8FE04F850" httpUri="https://zenodo.org/record/207030/files/figure.png" pageId="3" pageNumber="42" targetBox="[151,1436,193,1904]" targetPageId="3">
|
||
<paragraph id="8B953F3A0A42FFE2FF0B86B8FE04F850" blockId="3.[151,1437,1927,1980]" pageId="3" pageNumber="42">
|
||
<emphasis id="B95EE3280A42FFE2FF0B86B8FEACF871" bold="true" box="[151,275,1927,1950]" pageId="3" pageNumber="42">FIGURE 2.</emphasis>
|
||
Close-up of head of
|
||
<taxonomicName id="4C2A44B90A42FFE2FE7186B7FD57F872" box="[493,744,1928,1950]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="rudolfarndti" status="sp. nov.">
|
||
<emphasis id="B95EE3280A42FFE2FE7186B7FD57F872" box="[493,744,1928,1950]" italics="true" pageId="3" pageNumber="42">Enyalioides rudolfarndti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EE3280A42FFE2FD7286B7FC97F871" bold="true" box="[750,808,1928,1949]" pageId="3" pageNumber="42">
|
||
<taxonomicNameLabel id="A26D5E530A42FFE2FD7286B7FC97F871" box="[750,808,1928,1949]" pageId="3" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(CORBIDI 07209) showing the orange anthehumeral blotch. Photograph by P. J. Venegas.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF556FB20A45FFE5FF0B87E7FA97F8E1" httpUri="https://zenodo.org/record/207031/files/figure.png" pageId="4" pageNumber="43" targetBox="[188,1407,203,1689]" targetPageId="4">
|
||
<paragraph id="8B953F3A0A45FFE5FF0B87E7FA97F8E1" blockId="4.[151,1436,1752,1805]" pageId="4" pageNumber="43">
|
||
<emphasis id="B95EE3280A45FFE5FF0B87E7FEABF901" bold="true" box="[151,276,1752,1775]" pageId="4" pageNumber="43">FIGURE 3.</emphasis>
|
||
Dorsolateral and ventral views of the paratypes of
|
||
<taxonomicName id="4C2A44B90A45FFE5FCB687E7FB9AF902" box="[810,1061,1752,1774]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="43" phylum="Chordata" rank="species" species="rudolfarndti" status="sp. nov.">
|
||
<emphasis id="B95EE3280A45FFE5FCB687E7FB9AF902" box="[810,1061,1752,1774]" italics="true" pageId="4" pageNumber="43">Enyalioides rudolfarndti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EE3280A45FFE5FBB187E6FBD7F902" bold="true" box="[1069,1128,1753,1774]" pageId="4" pageNumber="43">
|
||
<taxonomicNameLabel id="A26D5E530A45FFE5FBB187E6FBD7F902" box="[1069,1128,1753,1774]" pageId="4" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Adult male (A, B, CORBIDI 07212), adult female (C, D, CORBIDI 07210), and juvenile (E, F, CORBIDI 07213). Photographs by P. J. Venegas.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B953F3A0A45FFE4FF59860BFDC5FF1B" blockId="4.[151,1436,1844,2012]" lastBlockId="5.[151,1437,151,500]" lastPageId="5" lastPageNumber="44" pageId="4" pageNumber="43">
|
||
Color in life of
|
||
<typeStatus id="549181980A45FFE5FEE8860BFE69F8A0" box="[372,470,1844,1868]" pageId="4" pageNumber="43" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="131123BF0A45FFE5FE79860BFDEDF8A0" box="[485,594,1844,1868]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1903,1926]" captionStartId-1="3.[151,250,1927,1950]" captionTargetBox-0="[151,1436,916,1879]" captionTargetBox-1="[151,1436,193,1904]" captionTargetId-0="figure@2.[151,1436,916,1880]" captionTargetId-1="figure@3.[151,1436,193,1906]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Dorsolateral view of the holotype of Enyalioides rudolfarndti sp. n. (CORBIDI 0 7209, adult male, SVL 116 mm). Photograph by P. J. Venegas." captionText-1="FIGURE 2. Close-up of head of Enyalioides rudolfarndti sp. n. (CORBIDI 07209) showing the orange anthehumeral blotch. Photograph by P. J. Venegas." httpUri-0="https://zenodo.org/record/207029/files/figure.png" httpUri-1="https://zenodo.org/record/207030/files/figure.png" pageId="4" pageNumber="43">Figs. 1–2</figureCitation>
|
||
): scales on dorsal and lateral surface of head, including labials, rostral, and mental, green; gulars greenish white; skin between gulars black; small dark gray and diffuse gular patch immediately anterior to gular fold; one big orange blotch on each side of the neck, extending posteriorly onto the antehumeral region and ventrally near the dark gular patch; dorsal background green; pale gray round blotches covering dorsolateral region from the neck to base of the tail; fine black reticulation on dorsal aspect of the body, limbs, flanks, and the proximal portion of the tail; ventrolateral region sky blue; ventral surface of body, limbs, and tail white; lateral borders of venter and ventral surface of thighs and shanks sky blue; the iris is gray with a fine brown radiation; pupil round with a white margin.
|
||
</paragraph>
|
||
<paragraph id="8B953F3A0A44FFE4FF59803BFB37FE88" blockId="5.[151,1437,151,500]" pageId="5" pageNumber="44">Color in ethanol after 10 months of preservation: dorsal background pale turquoise, gulars grayish turquoise; antehumeral blotch yellow; reticulation on the dorsal aspect of body, limbs, and base of the tail dark gray; gray blotches on dorsolateral region absent; ventral surface of body, limbs, and tail dark cream.</paragraph>
|
||
<paragraph id="8B953F3A0A44FFE4FF598050FE4DFE18" blockId="5.[151,1437,151,500]" pageId="5" pageNumber="44">
|
||
<emphasis id="B95EE3280A44FFE4FF598050FE86FE64" bold="true" box="[197,313,367,392]" pageId="5" pageNumber="44">Variation.</emphasis>
|
||
Meristic and morphometric characters of
|
||
<taxonomicName id="4C2A44B90A44FFE4FCBE804FFB86FE64" box="[802,1081,368,392]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="44" phylum="Chordata" rank="species" species="rudolfarndti">
|
||
<emphasis id="B95EE3280A44FFE4FCBE804FFB86FE64" box="[802,1081,368,392]" italics="true" pageId="5" pageNumber="44">Enyalioides rudolfarndti</emphasis>
|
||
</taxonomicName>
|
||
are summarized in
|
||
<tableCitation id="C6A80A810A44FFE4FA838050FACAFE6B" box="[1311,1397,367,391]" captionStart="TABLE 1" captionStartId="5.[151,235,548,569]" captionText="TABLE 1. Summary of morphological characters and measurements (mm) of E. rudolfarndti sp. n. Range follow by mean ± standard deviation." pageId="5" pageNumber="44">Table 1</tableCitation>
|
||
. In life, the single adult male
|
||
<typeStatus id="549181980A44FFE4FE2080AAFDA3FE41" box="[444,540,405,429]" pageId="5" pageNumber="44" type="paratype">paratype</typeStatus>
|
||
CORBIDI 0 7212 (
|
||
<figureCitation id="131123BF0A44FFE4FD6880ABFC8AFE40" box="[756,821,404,428]" captionStart="FIGURE 3" captionStartId="4.[151,250,1752,1775]" captionTargetBox="[188,1407,203,1689]" captionTargetId="figure@4.[151,1436,174,1749]" captionTargetPageId="4" captionText="FIGURE 3. Dorsolateral and ventral views of the paratypes of Enyalioides rudolfarndti sp. n. Adult male (A, B, CORBIDI 07212), adult female (C, D, CORBIDI 07210), and juvenile (E, F, CORBIDI 07213). Photographs by P. J. Venegas." httpUri="https://zenodo.org/record/207031/files/figure.png" pageId="5" pageNumber="44">Fig. 3</figureCitation>
|
||
A–B) has the same background coloration as the
|
||
<typeStatus id="549181980A44FFE4FAFD80ABFF78FE3C" pageId="5" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
but has the dorsum, flanks, dorsal surface of limbs and tail covered by black flecks and not by a fine black reticulation as on the
|
||
<typeStatus id="549181980A44FFE4FE1480E3FE51FE18" box="[392,494,476,500]" pageId="5" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF556FB20A44FFE4FF0B831BFEE7FDB4" pageId="5" pageNumber="44">
|
||
<paragraph id="8B953F3A0A44FFE4FF0B831BFEE7FDB4" blockId="5.[151,1436,547,1311]" pageId="5" pageNumber="44">
|
||
<emphasis id="B95EE3280A44FFE4FF0B831BFF41FDD5" bold="true" box="[151,254,548,569]" pageId="5" pageNumber="44">TABLE 1</emphasis>
|
||
. Summary of morphological characters and measurements (mm) of
|
||
<taxonomicName id="4C2A44B90A44FFE4FC20831AFBEAFDD6" box="[956,1109,548,570]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="44" phylum="Chordata" rank="species" species="rudolfarndti" status="sp. nov.">
|
||
<emphasis id="B95EE3280A44FFE4FC20831AFBEAFDD6" box="[956,1109,548,570]" italics="true" pageId="5" pageNumber="44">E. rudolfarndti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95EE3280A44FFE4FBC0831BFB29FDD5" bold="true" box="[1116,1174,548,569]" pageId="5" pageNumber="44">
|
||
<taxonomicNameLabel id="A26D5E530A44FFE4FBC0831BFB29FDD5" box="[1116,1174,548,569]" pageId="5" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Range follow by mean ± standard deviation.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B953F3A0A44FFE4FF00834CFEB7FD65" blockId="5.[151,1436,547,1311]" box="[156,264,627,649]" pageId="5" pageNumber="44">Characters</paragraph>
|
||
<paragraph id="8B953F3A0A44FFE4FF0083A2FBEBFCB4" blockId="5.[151,1436,547,1311]" pageId="5" pageNumber="44">Dorsal in transverse row between dorsolateral crests at midbody 27–30, 28.25 ± 1.5 Ventrals in transverse row at midbody 28–32, 30.25 ± 3.69 Vertebrals from occiput to base of tail 42–47, 45.5 ± 2.38 Gulars 33–36, 34.5 ± 1.29 Infralabials 11, 11</paragraph>
|
||
<paragraph id="8B953F3A0A44FFE4FF008254FB82FB95" blockId="5.[151,1436,547,1311]" pageId="5" pageNumber="44">
|
||
Supralabials 11–14, 12.5 ± 1.29 Canthals 3–4, 3.75 Superciliaries 14–18, 15.25 ± 1.89 Transverse rows of ventrals between fore and hind limb 35–38, 36.75 ± 1.5 Subdigitals fingers
|
||
<date id="FF9419FA0A44FFE4FEFD852DFBC4FBCB" box="[353,1147,1042,1063]" pageId="5" pageNumber="44" value="1920-04-19" valueMax="1920-04-23" valueMin="1920-04-19">IV 19–23, 20</date>
|
||
± 2 Subdigitals toe
|
||
<date id="FF9419FA0A44FFE4FEA68504FBE7FBBC" box="[314,1112,1082,1104]" pageId="5" pageNumber="44" value="1925-04-28">IV 25–28</date>
|
||
, 26.5 ± 1.29 Femoral pores 1, 1
|
||
</paragraph>
|
||
<paragraph id="8B953F3A0A44FFE4FF0085B2FB44FAF3" blockId="5.[151,1436,547,1311]" pageId="5" pageNumber="44">
|
||
Head length/Head width 1.30–1.43, 1.35 ± 0.06 Fore limb length/SVL 0.43–0.56, 0.50 ± 0.05
|
||
<collectingCountry id="F33D7FAA0A44FFE4FF0085DFFF70FB1A" box="[156,207,1248,1270]" name="India" pageId="5" pageNumber="44">Hind</collectingCountry>
|
||
limb length/SVL 0.76–0.87, 0.80 ± 0.05 Tail length/Total length 0.60–1.49, 0.83 ± 0.44
|
||
</paragraph>
|
||
<paragraph id="8B953F3A0A44FFE4FF59845AFDD3F92C" blockId="5.[151,1436,1380,1944]" pageId="5" pageNumber="44">
|
||
In life, the single female
|
||
<typeStatus id="549181980A44FFE4FE46845AFD84FA91" box="[474,571,1381,1405]" pageId="5" pageNumber="44" type="paratype">paratype</typeStatus>
|
||
CORBIDI 0 7210 (
|
||
<figureCitation id="131123BF0A44FFE4FC93845BFCF0FA90" box="[783,847,1380,1404]" captionStart="FIGURE 3" captionStartId="4.[151,250,1752,1775]" captionTargetBox="[188,1407,203,1689]" captionTargetId="figure@4.[151,1436,174,1749]" captionTargetPageId="4" captionText="FIGURE 3. Dorsolateral and ventral views of the paratypes of Enyalioides rudolfarndti sp. n. Adult male (A, B, CORBIDI 07212), adult female (C, D, CORBIDI 07210), and juvenile (E, F, CORBIDI 07213). Photographs by P. J. Venegas." httpUri="https://zenodo.org/record/207031/files/figure.png" pageId="5" pageNumber="44">Fig. 3</figureCitation>
|
||
C–D) has the head mossy green with black markings including flecks on the dorsal surface of head; a narrow stripe posterior to the superciliaries extends along the lateral edge of the skull roof; a broad postocular stripe extends to the commisure of the mouth and the anterior border of the tympanum, and an infraorbital stripe is present between the eyes and the labials; the dorsal surface of the neck has a white blotch on each side behind the tympanum, followed by a longitudinal pale stripe that extends to the scapular region; the gular region is dark gray, darker on the gular fold, with dark brown spots; the dorsal background is mossy green with transverse diffuse black bars on the body, limbs, and tail; the ventral surface of the body, limbs and tail is grayish white with black spots in the lateral borders of the belly, ventral surface of the limbs, and the tail; iris gray with a brown reticulation. The vertebral crest and the enlarged and raised scales posterior to the superciliaries are high as in the males.
|
||
</paragraph>
|
||
<paragraph id="8B953F3A0A44FFE4FF5987F2FF52F874" blockId="5.[151,1436,1380,1944]" pageId="5" pageNumber="44">
|
||
In life, the juvenile
|
||
<typeStatus id="549181980A44FFE4FE3F87F2FDBCF909" box="[419,515,1741,1765]" pageId="5" pageNumber="44" type="paratype">paratype</typeStatus>
|
||
CORBIDI 0 7213 (
|
||
<figureCitation id="131123BF0A44FFE4FD4687F3FCA4F908" box="[730,795,1740,1764]" captionStart="FIGURE 3" captionStartId="4.[151,250,1752,1775]" captionTargetBox="[188,1407,203,1689]" captionTargetId="figure@4.[151,1436,174,1749]" captionTargetPageId="4" captionText="FIGURE 3. Dorsolateral and ventral views of the paratypes of Enyalioides rudolfarndti sp. n. Adult male (A, B, CORBIDI 07212), adult female (C, D, CORBIDI 07210), and juvenile (E, F, CORBIDI 07213). Photographs by P. J. Venegas." httpUri="https://zenodo.org/record/207031/files/figure.png" pageId="5" pageNumber="44">Fig. 3</figureCitation>
|
||
E–F) has the head copper with black flecks on the dorsal surface and a broad dark brown subocular stripe; the dorsal background is dark brown with copper transverse bars on the body, limbs, and tail; the belly is copper with scattered dark copper and brown spots; the ventral surfaces of the limbs and the tail are dark brown with copper spots; the gular region is copper and darker than the belly, with some white spots on the ventral border of the snout; the iris is reddish brown and the pupil is round with a golden margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF556FB20A47FFE7FF0B8510FDCFFB88" httpUri="https://zenodo.org/record/207032/files/figure.png" pageId="6" pageNumber="45" targetBox="[151,1436,193,1049]" targetPageId="6">
|
||
<paragraph id="8B953F3A0A47FFE7FF0B8510FDCFFB88" blockId="6.[151,1436,1071,1124]" pageId="6" pageNumber="45">
|
||
<emphasis id="B95EE3280A47FFE7FF0B8510FEAAFBA9" bold="true" box="[151,277,1071,1094]" pageId="6" pageNumber="45">FIGURE 4.</emphasis>
|
||
Close-up of head of
|
||
<taxonomicName id="4C2A44B90A47FFE7FE698510FD57FBA9" box="[501,744,1071,1093]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="palpebralis">
|
||
<emphasis id="B95EE3280A47FFE7FE698510FD57FBA9" box="[501,744,1071,1093]" italics="true" pageId="6" pageNumber="45">Enyalioides palpebralis</emphasis>
|
||
</taxonomicName>
|
||
(CORBIDI 0 8347, adult male, SVL 112 mm) showing the orange antehumeral blotch. Photograph by G. Chávez.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3306CB10A47FFE7FF5985AFFE8BFA6C" pageId="6" pageNumber="45" type="distribution">
|
||
<paragraph id="8B953F3A0A47FFE7FF5985AFFE8BFA6C" blockId="6.[151,1437,1168,1985]" pageId="6" pageNumber="45">
|
||
<emphasis id="B95EE3280A47FFE7FF5985AFFDF5FB45" bold="true" box="[197,586,1168,1193]" pageId="6" pageNumber="45">Distribution and natural history.</emphasis>
|
||
<taxonomicName id="4C2A44B90A47FFE7FDCC85AEFCDBFB45" box="[592,868,1169,1193]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="rudolfarndti">
|
||
<emphasis id="B95EE3280A47FFE7FDCC85AEFCDBFB45" box="[592,868,1169,1193]" italics="true" pageId="6" pageNumber="45">Enyalioides rudolfarndti</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="549181980A47FFE7FBE885AEFB1BFB45" box="[1140,1188,1169,1193]" pageId="6" pageNumber="45">type</typeStatus>
|
||
locality in the Región de Pasco, at an elevation of
|
||
<quantity id="4CD292DF0A47FFE7FE40858AFD86FB21" box="[476,569,1205,1229]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="6" pageNumber="45" unit="m" value="1050.0">1050 m</quantity>
|
||
.a.s.l., on the upper Amazon basin of central
|
||
<collectingCountry id="F33D7FAA0A47FFE7FBAB8589FBD5FB21" box="[1079,1130,1206,1229]" name="Peru" pageId="6" pageNumber="45">Peru</collectingCountry>
|
||
(
|
||
<figureCitation id="131123BF0A47FFE7FBE7858AFB7DFB21" box="[1147,1218,1205,1229]" captionStart="FIGURE 5" captionStartId="7.[151,250,1879,1902]" captionTargetBox="[174,1414,222,1763]" captionTargetId="figure@7.[151,1436,195,1858]" captionTargetPageId="7" captionText="FIGURE 5. Distribution of Enyalioides rudolfarndti sp. nov. in Peru. The red triangle indicates the type locality, the only locality where the species is known to date. White circles are important cities for reference." httpUri="https://zenodo.org/record/207033/files/figure.png" pageId="6" pageNumber="45">Fig. 5</figureCitation>
|
||
). This new species inhabit the premontane forest of the Río Huancabamba canyon that lies within the YCNP in the Pasco region (
|
||
<figureCitation id="131123BF0A47FFE7FAF285E7FF17FAF9" captionStart="FIGURE 6" captionStartId="8.[151,250,1178,1201]" captionTargetBox="[151,1436,193,1156]" captionTargetId="figure@8.[151,1436,193,1157]" captionTargetPageId="8" captionText="FIGURE 6. Habitat at the type locality of Enyalioides rudolfarndti sp. nov. on August of 2010. Photograph by P. J. Venegas." httpUri="https://zenodo.org/record/207034/files/figure.png" pageId="6" pageNumber="45">Fig. 6</figureCitation>
|
||
). All individuals of
|
||
<taxonomicName id="4C2A44B90A47FFE7FE1285C1FD88FAF9" box="[398,567,1277,1301]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="rudolfarndti">
|
||
<emphasis id="B95EE3280A47FFE7FE1285C1FD88FAF9" box="[398,567,1277,1301]" italics="true" pageId="6" pageNumber="45">E. rudolfarndti</emphasis>
|
||
</taxonomicName>
|
||
were collected at night sleeping on horizontal stems of bushes up to
|
||
<quantity id="4CD292DF0A47FFE7FAD185C2FA23FAF9" box="[1357,1436,1277,1301]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="45" unit="m" value="1.5">1.50 m</quantity>
|
||
above the ground. Sympatric species of reptiles collected with
|
||
<taxonomicName id="4C2A44B90A47FFE7FCC7841EFBCFFAD5" box="[859,1136,1313,1337]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="rudolfarndti">
|
||
<emphasis id="B95EE3280A47FFE7FCC7841EFBCFFAD5" box="[859,1136,1313,1337]" italics="true" pageId="6" pageNumber="45">Enyalioides rudolfarndti</emphasis>
|
||
</taxonomicName>
|
||
were
|
||
<emphasis id="B95EE3280A47FFE7FB29841EFB44FAB1" italics="true" pageId="6" pageNumber="45">
|
||
<taxonomicName id="4C2A44B90A47FFE7FB29841EFF64FAB1" authority="Clelia" authorityName="Clelia" class="Reptilia" family="Gymnophthalmidae" genus="Cercosaura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="argulus">Cercosaura argulus, Clelia</taxonomicName>
|
||
clelia, Dipsas catesbyi, D. indica, D. schunkii, Micrurus annellatus, Oxyrhopus melanogenys,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C2A44B90A47FFE7FAAE8479FE92FA6D" class="Reptilia" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="torquatus">
|
||
<emphasis id="B95EE3280A47FFE7FAAE8479FE92FA6D" italics="true" pageId="6" pageNumber="45">Stenocercus torquatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3306CB10A47FFE7FF5984B2FE37FA01" pageId="6" pageNumber="45" type="etymology">
|
||
<paragraph id="8B953F3A0A47FFE7FF5984B2FE37FA01" blockId="6.[151,1437,1168,1985]" pageId="6" pageNumber="45">
|
||
<emphasis id="B95EE3280A47FFE7FF5984B2FEF3FA49" bold="true" box="[197,332,1421,1445]" pageId="6" pageNumber="45">Etymology.</emphasis>
|
||
The specific name is a patronym for Dr. Rudolf G. Arndt of Pomona, New
|
||
<collectingCountry id="F33D7FAA0A47FFE7FB7884B1FA94FA4A" box="[1252,1323,1422,1446]" name="Jersey" pageId="6" pageNumber="45">Jersey</collectingCountry>
|
||
,
|
||
<collectingCountry id="F33D7FAA0A47FFE7FAA784B2FAC9FA49" box="[1339,1398,1421,1445]" name="United States of America" pageId="6" pageNumber="45">USA</collectingCountry>
|
||
, in recognition of his financial support for the improvement of the herpetological collection of CORBIDI through the BIOPAT-Programme.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3306CB10A47FFE7FF5984C6FD1FF82D" pageId="6" pageNumber="45" type="discussion">
|
||
<paragraph id="8B953F3A0A47FFE7FF5984C6FD1FF82D" blockId="6.[151,1437,1168,1985]" pageId="6" pageNumber="45">
|
||
<emphasis id="B95EE3280A47FFE7FF5984C6FE86F9FD" bold="true" box="[197,313,1529,1553]" pageId="6" pageNumber="45">Remarks.</emphasis>
|
||
Although
|
||
<taxonomicName id="4C2A44B90A47FFE7FE2884C6FD75F9FD" box="[436,714,1529,1553]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="rudolfarndti">
|
||
<emphasis id="B95EE3280A47FFE7FE2884C6FD75F9FD" box="[436,714,1529,1553]" italics="true" pageId="6" pageNumber="45">Enyalioides rudolfarndti</emphasis>
|
||
</taxonomicName>
|
||
and a new species of
|
||
<taxonomicName id="4C2A44B90A47FFE7FC5784C6FBE5F9FD" box="[971,1114,1529,1553]" class="Reptilia" family="Gymnophthalmidae" genus="Euspondylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="45" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95EE3280A47FFE7FC5784C6FBE5F9FD" box="[971,1114,1529,1553]" italics="true" pageId="6" pageNumber="45">Euspondylus</emphasis>
|
||
</taxonomicName>
|
||
are the only new species of reptile discovered in the Cordillera Yanachaga in the two last decades, the recent taxonomic work on amphibians and herpetological inventories in this region has resulted in the description of 27 new species of amphibians, of which 10 were discovered inside YCNP (
|
||
<bibRefCitation id="EFBB42CB0A47FFE7FDF6875AFCE1F991" author="Duellman" box="[618,862,1637,1661]" pageId="6" pageNumber="45" refString="Duellman, W. E., Trueb, L. & Lehr, E. (2006) A new species of marsupial frog (Anura: Hylidae: Gastrotheca) from the Amazon slopes of the Cordillera Oriental in Peru. Copeia, 2006, 595 - 603." type="journal article" year="2006">
|
||
Duellman
|
||
<emphasis id="B95EE3280A47FFE7FD7D8759FCA5F991" box="[737,794,1637,1661]" italics="true" pageId="6" pageNumber="45">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBB42CB0A47FFE7FCF5875AFBF8F991" author="Lehr" box="[873,1095,1637,1661]" pageId="6" pageNumber="45" refString="Lehr, E. & Trueb, L. (2007) Diversity among New World microhylid frogs (Anura: Microhylidae): morphological and osteological comparisons between Nelsonophryne (Gunther 1901) and a new genus from Peru. Zoological Journal of the Linnean Society, 149, 583 - 609." type="journal article" year="2007">Lehr & Trueb 2007</bibRefCitation>
|
||
; Lehr
|
||
<emphasis id="B95EE3280A47FFE7FB0C8759FB76F991" box="[1168,1225,1637,1661]" italics="true" pageId="6" pageNumber="45">et al.</emphasis>
|
||
2007; Chaparro
|
||
<emphasis id="B95EE3280A47FFE7FA148759FF0BF94D" italics="true" pageId="6" pageNumber="45">et al.</emphasis>
|
||
2008;
|
||
<bibRefCitation id="EFBB42CB0A47FFE7FE9D87B6FE75F94C" author="Boano" box="[257,458,1672,1697]" pageId="6" pageNumber="45" refString="Boano, G., Mazzotti, S. & Sindaco, R. (2008) A new peculiar frog species of the genus Pristimantis from Yanachaga-Chemillen National Park, Peru. Zootaxa, 1674, 51 - 57." type="journal article" year="2008">
|
||
Boano
|
||
<emphasis id="B95EE3280A47FFE7FECD87B5FE36F94D" box="[337,393,1673,1697]" italics="true" pageId="6" pageNumber="45">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBB42CB0A47FFE7FE4A87B7FD41F94C" author="Duellman" box="[470,766,1672,1696]" pageId="6" pageNumber="45" refString="Duellman, W. E. & Hedges, S. B. (2008) Two minute species of Phrynopus (Lissamphibia: Anura) from the Cordillera Oriental in Peru. Zootaxa, 1675, 59 - 66." type="journal article" year="2008">Duellman & Hedges 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBB42CB0A47FFE7FC9587B7FBF8F94C" author="Duellman" box="[777,1095,1672,1696]" pageId="6" pageNumber="45" refString="Duellman, W. E. & Chaparro, J. C. (2008) Two distinctive new species of Pristimantis (Anura: Strabomantidae) from the Cordillera Oriental with a distributional synopsis of strabomantids in Central Peru. Zootaxa, 1918, 13 - 25." type="journal article" year="2008">Duellman & Chaparro 2008</bibRefCitation>
|
||
). The lack of new reptiles taxa discoveries in this region probably relates to an absence of taxonomic work in this group in general. Moreover, due to its complex topography, the Cordillera Yanachaga has not been adequately sampled and some areas, such as YCNP, have been only slightly explored. However, in a rapid biological inventory carried out in the northern portion of YCNP during
|
||
<date id="FF9419FA0A47FFE7FE0D8626FD99F8DC" box="[401,550,1816,1841]" pageId="6" pageNumber="45" value="2010-08">August 2010</date>
|
||
, we collected the new lizard species described herein; a new species of gymnophthalmid lizard, genus
|
||
<taxonomicName id="4C2A44B90A47FFE7FE4F8602FCD7F8B9" authority="Chavez et al. 2011" authorityName="Chavez et al." authorityYear="2011" box="[467,872,1853,1877]" class="Reptilia" family="Gymnophthalmidae" genus="Euspondylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="45" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95EE3280A47FFE7FE4F8602FDDDF8B9" box="[467,610,1853,1877]" italics="true" pageId="6" pageNumber="45">Euspondylus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBB42CB0A47FFE7FDEB8602FCE0F8B9" author="Chavez" box="[631,863,1853,1877]" pageId="6" pageNumber="45" refString="Chavez, G., Siu-Ting, K., Duran, V. & Venegas, P. J. (2011) Two new species of Andean gymnophthalmid lizards of the genus Euspondylus (Reptilia, Squamata) from central and southern Peru. Zookeys, 109, 1 - 17." type="journal article" year="2011">
|
||
Chavez
|
||
<emphasis id="B95EE3280A47FFE7FD458601FCA8F8B9" box="[729,791,1853,1877]" italics="true" pageId="6" pageNumber="45">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; two new species of strabomantid frogs, genus
|
||
<emphasis id="B95EE3280A47FFE7FF0B865EFEA6F895" box="[151,281,1889,1913]" italics="true" pageId="6" pageNumber="45">
|
||
<taxonomicName id="4C2A44B90A47FFE7FF0B865EFEACF895" box="[151,275,1889,1913]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="45" phylum="Chordata" rank="genus">Phrynopus</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
and one new species of casqued tree frog, genus
|
||
<taxonomicName id="4C2A44B90A47FFE7FCD8865EFC56F895" box="[836,1001,1889,1913]" class="Amphibia" family="Hylidae" genus="Osteocephalus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="45" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95EE3280A47FFE7FCD8865EFC56F895" box="[836,1001,1889,1913]" italics="true" pageId="6" pageNumber="45">Osteocephalus</emphasis>
|
||
</taxonomicName>
|
||
, all of which are now being described. These findings highlight the importance of collecting in poorly explored areas of
|
||
<collectingCountry id="F33D7FAA0A47FFE7FBBE86B9FBE7F871" box="[1058,1112,1926,1949]" name="Peru" pageId="6" pageNumber="45">Peru</collectingCountry>
|
||
, a country that awaits a large number of discoveries in herpetological terms.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |