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<document id="A2771DA3460C2B0561FFE7AC8D6A5825" ID-CLB-Dataset="9276" ID-DOI="10.11646/zootaxa.4895.2.2" ID-GBIF-Dataset="0ffdae09-7a8d-4c69-8372-0c84e7e72605" ID-ISSN="1175-5326" ID-Zenodo-Dep="4322455" ID-ZooBank="8F5516C4-D6EC-46C1-8008-23683AB5DF6F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1608024653419" checkinUser="plazi" docAuthor="Makarchenko, Eugenyi A., Semenchenko, Alexander A. &amp; Palatov, Dmitry M." docDate="2020" docId="03830441FFEDFFD3FF57F9234D9AFEF4" docLanguage="en" docName="zootaxa.4895.2.2.pdf" docOrigin="Zootaxa 4895 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Shilovia Makarchenko" docType="key" docVersion="7" lastPageNumber="208" masterDocId="FFBA7C39FFE6FFDFFFC0FFB04806FFD6" masterDocTitle="Review of the genus Shilovia Makarchenko (Diptera: Chironomidae: Diamesinae Boreoheptagyiini) from the mountains of Central Asia, with morphological description and DNA barcoding of known species" masterLastPageNumber="210" masterPageNumber="196" pageNumber="207" updateTime="1698847560547" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8AC43E7471DD88F4064C0DC0B3B5B466">Review of the genus Shilovia Makarchenko (Diptera: Chironomidae: Diamesinae Boreoheptagyiini) from the mountains of Central Asia, with morphological description and DNA barcoding of known species</mods:title>
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<mods:namePart id="798786CD01BB69997D37684971835527">Makarchenko, Eugenyi A.</mods:namePart>
<mods:affiliation id="99D7ECF7EDED3F81FE754A226AD760AB">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, 100 let Vladivostoku 159, 690022 Vladivostok, Russia.</mods:affiliation>
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<mods:namePart id="22960FC4F76CAF7F0AECB191E9BF6B49">Semenchenko, Alexander A.</mods:namePart>
<mods:affiliation id="63B08098B89CD4309536170867F6F08B">Laboratory of Ecology and Evolutionary Biology of Aquatic Organisms, Far Eastern Federal University, Suhanova St. 8, 690950 Vladivostok, Russia.</mods:affiliation>
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<mods:namePart id="9601635BEB9F5E528273176DFB63B578">Palatov, Dmitry M.</mods:namePart>
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<mods:affiliation id="27FC003F0C4CCBDB0A1479A0CCD8675E">A. N. Severtsov Institute of Ecology and Evolution of the Russian Academy of Sciences, Leninskij prosp. 33, 119071 Moscow, Russia. triops @ yandex. ru; https: // orcid. org / 0000 - 0002 - 8826 - 9316</mods:affiliation>
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<treatment id="03830441FFEDFFD3FF57F9234D9AFEF4" ID-DOI="http://doi.org/10.5281/zenodo.4329120" ID-GBIF-Taxon="176092605" ID-Zenodo-Dep="4329120" LSID="urn:lsid:plazi:treatment:03830441FFEDFFD3FF57F9234D9AFEF4" httpUri="http://treatment.plazi.org/id/03830441FFEDFFD3FF57F9234D9AFEF4" lastPageId="12" lastPageNumber="208" pageId="11" pageNumber="207">
<subSubSection id="C330E6DCFFEDFFD4FF57F92348E4F923" pageId="11" pageNumber="207" type="nomenclature">
<paragraph id="8B95B557FFEDFFD4FF57F9234B15F97B" blockId="11.[151,787,1682,1709]" box="[151,787,1682,1709]" pageId="11" pageNumber="207">
<heading id="D0DD023BFFEDFFD4FF57F9234B15F97B" bold="true" box="[151,787,1682,1709]" fontSize="11" level="1" pageId="11" pageNumber="207" reason="1">
<emphasis id="B95E6945FFEDFFD4FF57F9234B15F97B" bold="true" box="[151,787,1682,1709]" pageId="11" pageNumber="207">
Key to the known species of
<taxonomicName id="4C2ACED4FFEDFFD4FE39F9224B15F97B" ID-CoL="64529" ID-ENA="2045254" authority="Makarchenko" authorityName="Makarchenko" box="[505,787,1682,1709]" class="Insecta" family="Chironomidae" genus="Shilovia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis id="B95E6945FFEDFFD4FE39F9224A5AF97B" bold="true" box="[505,604,1682,1709]" italics="true" pageId="11" pageNumber="207">Shilovia</emphasis>
Makarchenko
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<paragraph id="8B95B557FFEDFFD4FF57F96B48E4F923" blockId="11.[151,226,1755,1781]" box="[151,226,1755,1781]" pageId="11" pageNumber="207">
<heading id="D0DD023BFFEDFFD4FF57F96B48E4F923" bold="true" box="[151,226,1755,1781]" fontSize="11" level="1" pageId="11" pageNumber="207" reason="1">
<emphasis id="B95E6945FFEDFFD4FF57F96B48E4F923" bold="true" box="[151,226,1755,1781]" pageId="11" pageNumber="207">Males</emphasis>
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<subSubSection id="C330E6DCFFEDFFD3FF57F8904D9AFEF4" lastPageId="12" lastPageNumber="208" pageId="11" pageNumber="207" type="key">
<keyStep id="30DEADF2FFEDFFD3FF57F8904D9AFF7B" lastPageId="12" lastPageNumber="208" pageId="11" pageNumber="207">
<paragraph id="8B95B557FFEDFFD4FF57F8904D9AF858" blockId="11.[151,1437,1823,2022]" pageId="11" pageNumber="207">
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1. Antenna with 5 flagellomeres. Legs spotted: basal 1/3 or half of femur and tibia yellow, distal 2/3 or half brown; ta
<subScript id="17AEB712FFEDFFD4FAF0F89F4D31F8ED" attach="left" box="[1328,1335,1839,1851]" fontSize="5" pageId="11" pageNumber="207">1</subScript>
ta
<subScript id="17AEB712FFEDFFD4FA94F89F4D5DF8ED" attach="left" box="[1364,1371,1839,1851]" fontSize="5" pageId="11" pageNumber="207">5</subScript>
brown (
<figureCitation id="1311A9D2FFEDFFD4FF29F88D4923F885" box="[233,293,1853,1875]" captionStart="FIGURES 19" captionStartId="4.[151,264,1824,1848]" captionTargetBox="[151,1432,172,1788]" captionTargetId="figure-18@4.[151,1435,170,1800]" captionTargetPageId="4" captionText="FIGURES 19. Shilovia rara Makarchenko, male. 1, head; 2, legs; 3, basal part of wing; 4, antepronotum and mesonotum, dorsal view; 5, sternapodeme and inferior volsella; 6, antenna; 7, distal part of gonostylus; 89, hypopygium in dorsal view. 12, 7, 9, male from Kyrgyzstan; 36, 8, male from Tajikistan (holotype). P1 fore leg, P2 mid leg, P3 hind leg." figureDoi="http://doi.org/10.5281/zenodo.4322457" httpUri="https://zenodo.org/record/4322457/files/figure.png" pageId="11" pageNumber="207">Fig. 2</figureCitation>
). Dorsocentrals in anterior part of mesonotum not reach preanal setae (
<figureCitation id="1311A9D2FFEDFFD4FC13F88D4C08F885" box="[979,1038,1853,1875]" captionStart="FIGURES 19" captionStartId="4.[151,264,1824,1848]" captionTargetBox="[151,1432,172,1788]" captionTargetId="figure-18@4.[151,1435,170,1800]" captionTargetPageId="4" captionText="FIGURES 19. Shilovia rara Makarchenko, male. 1, head; 2, legs; 3, basal part of wing; 4, antepronotum and mesonotum, dorsal view; 5, sternapodeme and inferior volsella; 6, antenna; 7, distal part of gonostylus; 89, hypopygium in dorsal view. 12, 7, 9, male from Kyrgyzstan; 36, 8, male from Tajikistan (holotype). P1 fore leg, P2 mid leg, P3 hind leg." figureDoi="http://doi.org/10.5281/zenodo.4322457" httpUri="https://zenodo.org/record/4322457/files/figure.png" pageId="11" pageNumber="207">Fig. 4</figureCitation>
). Posterior margin of tergite IX concave; gonostylus in subapical part along outer edge with rounded extension; inferior volsella and basal lobe of gonocoxite present; sternapodeme long and narrow (
<figureCitation id="1311A9D2FFEDFFD4FDD2F8C84A82F85B" box="[530,644,1911,1934]" captionStart="FIGURES 19" captionStartId="4.[151,264,1824,1848]" captionTargetBox="[151,1432,172,1788]" captionTargetId="figure-18@4.[151,1435,170,1800]" captionTargetPageId="4" captionText="FIGURES 19. Shilovia rara Makarchenko, male. 1, head; 2, legs; 3, basal part of wing; 4, antepronotum and mesonotum, dorsal view; 5, sternapodeme and inferior volsella; 6, antenna; 7, distal part of gonostylus; 89, hypopygium in dorsal view. 12, 7, 9, male from Kyrgyzstan; 36, 8, male from Tajikistan (holotype). P1 fore leg, P2 mid leg, P3 hind leg." figureDoi="http://doi.org/10.5281/zenodo.4322457" httpUri="https://zenodo.org/record/4322457/files/figure.png" pageId="11" pageNumber="207">Figs. 5, 79</figureCitation>
,
<figureCitation id="1311A9D2FFEDFFD4FD4DF8C84AEBF85B" box="[653,749,1911,1933]" captionStart="FIGURES 2127" captionStartId="9.[151,264,1737,1761]" captionTargetBox="[160,1436,195,1669]" captionTargetId="figure-21@9.[151,1436,181,1713]" captionTargetPageId="9" captionText="FIGURES 2127. Shilovia rara Makarchenko (21, 2425), S. xinhuawangi sp. nov. (22, 27) and S. yakovlevi sp. nov. (23, 26), males. 2123, gonostylus; 24, inferior volsella and basal lobe of gonocoxite; 2527, sternapodeme, phallapodeme and aedeagal lobe. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4322465" httpUri="https://zenodo.org/record/4322465/files/figure.png" pageId="11" pageNumber="207">21, 2425</figureCitation>
) …................................
<taxonomicName id="4C2ACED4FFEDFFD4FB58F8C84D9AF858" ID-CoL="4X4NR" authority="Makarchenko" authorityName="Makarchenko" authorityYear="1989" box="[1176,1436,1912,1934]" class="Insecta" family="Chironomidae" genus="Shilovia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="207" phylum="Arthropoda" rank="species" species="rara">
<emphasis id="B95E6945FFEDFFD4FB58F8C84D12F85B" box="[1176,1300,1912,1933]" italics="true" pageId="11" pageNumber="207">Shilovia rara</emphasis>
Makarchenko
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</paragraph>
<paragraph id="8B95B557FFEDFFD3FF57F8254D9AFF7B" blockId="11.[151,1437,1823,2022]" lastBlockId="12.[151,1437,151,291]" lastPageId="12" lastPageNumber="208" pageId="11" pageNumber="207">
<keyLead id="30DB1662FFEDFFD3FF57F8254D9AFF7B" lastPageId="12" lastPageNumber="208" pageId="11" pageNumber="207">
Antenna with 6 flagellomeres. Legs not spotted: at least basal 3/4 of femur yellowish or yellow, distal 1/4 dark brown; tibia, ta
<subScript id="17AEB712FFEDFFD4FF33F87248FCF818" attach="left" box="[243,250,1986,1998]" fontSize="5" pageId="11" pageNumber="207">1</subScript>
ta
<subScript id="17AEB712FFEDFFD4FED7F8714918F81B" attach="left" box="[279,286,1985,1997]" fontSize="5" pageId="11" pageNumber="207">5</subScript>
dark brown (
<figureCitation id="1311A9D2FFEDFFD4FE66F80249F6F81E" box="[422,496,1970,1992]" captionStart="FIGURES 1620" captionStartId="8.[151,264,1844,1868]" captionTargetBox="[151,1435,181,1821]" captionTargetId="figure-18@8.[151,1435,181,1821]" captionTargetPageId="8" captionText="FIGURES 1620. Shilovia xinhuawangi sp. nov. (16) and S. yakovlevi sp. nov. (1720), males. 16, legs; 17, basal part of wing; 18, dorsal part of mesonotum; 19, gonocoxite and gonostylus; 20, part of hypopygium in dorsal view. Designations are the same as in Figures 19." figureDoi="http://doi.org/10.5281/zenodo.4322461" httpUri="https://zenodo.org/record/4322461/files/figure.png" pageId="11" pageNumber="207">Fig. 16</figureCitation>
). Dorsocentrals in anterior part of mesonotum reach preanal setae (
<figureCitation id="1311A9D2FFEDFFD4FB42F8024CCCF81E" box="[1154,1226,1970,1992]" captionStart="FIGURES 1620" captionStartId="8.[151,264,1844,1868]" captionTargetBox="[151,1435,181,1821]" captionTargetId="figure-18@8.[151,1435,181,1821]" captionTargetPageId="8" captionText="FIGURES 1620. Shilovia xinhuawangi sp. nov. (16) and S. yakovlevi sp. nov. (1720), males. 16, legs; 17, basal part of wing; 18, dorsal part of mesonotum; 19, gonocoxite and gonostylus; 20, part of hypopygium in dorsal view. Designations are the same as in Figures 19." figureDoi="http://doi.org/10.5281/zenodo.4322461" httpUri="https://zenodo.org/record/4322461/files/figure.png" pageId="11" pageNumber="207">Fig. 18</figureCitation>
). Posterior margin of tergite IX straight or slightly concave; gonostylus another shape, in subapical part along outer edge without rounded extension; basal lobe of gonocoxite reduced; sternapodeme wide basally and narrow distally (
<figureCitation id="1311A9D2FFEAFFD3FC2DFF274C59FF7B" box="[1005,1119,151,173]" captionStart="FIGURES 1015" captionStartId="5.[151,264,1824,1848]" captionTargetBox="[156,1426,193,1790]" captionTargetId="figure-21@5.[151,1436,181,1800]" captionTargetPageId="5" captionText="FIGURES 1015. Shilovia xinhuawangi sp. nov., male. 10, basal part of wing; 11, antenna; 12, dorsal part of mesonotum; 13, scutellum and postnotum; 14, hypopygium in dorsal view; 15, hypopygium without tergite IX in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4322459" httpUri="https://zenodo.org/record/4322459/files/figure.png" pageId="12" pageNumber="208">Figs. 1415</figureCitation>
,
<figureCitation id="1311A9D2FFEAFFD3FBA8FF284CA3FF7B" box="[1128,1189,152,173]" captionStart="FIGURES 1620" captionStartId="8.[151,264,1844,1868]" captionTargetBox="[151,1435,181,1821]" captionTargetId="figure-18@8.[151,1435,181,1821]" captionTargetPageId="8" captionText="FIGURES 1620. Shilovia xinhuawangi sp. nov. (16) and S. yakovlevi sp. nov. (1720), males. 16, legs; 17, basal part of wing; 18, dorsal part of mesonotum; 19, gonocoxite and gonostylus; 20, part of hypopygium in dorsal view. Designations are the same as in Figures 19." figureDoi="http://doi.org/10.5281/zenodo.4322461" httpUri="https://zenodo.org/record/4322461/files/figure.png" pageId="12" pageNumber="208">1920</figureCitation>
,
<figureCitation id="1311A9D2FFEAFFD3FB6FFF284D35FF7B" box="[1199,1331,151,173]" captionStart="FIGURES 2127" captionStartId="9.[151,264,1737,1761]" captionTargetBox="[160,1436,195,1669]" captionTargetId="figure-21@9.[151,1436,181,1713]" captionTargetPageId="9" captionText="FIGURES 2127. Shilovia rara Makarchenko (21, 2425), S. xinhuawangi sp. nov. (22, 27) and S. yakovlevi sp. nov. (23, 26), males. 2123, gonostylus; 24, inferior volsella and basal lobe of gonocoxite; 2527, sternapodeme, phallapodeme and aedeagal lobe. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4322465" httpUri="https://zenodo.org/record/4322465/files/figure.png" pageId="12" pageNumber="208">2223, 2627</figureCitation>
).... … 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DEADF2FFEAFFD3FF57FF054D9AFEF4" pageId="12" pageNumber="208">
<paragraph id="8B95B557FFEAFFD3FF57FF054D9AFF31" blockId="12.[151,1437,151,291]" pageId="12" pageNumber="208">
<keyLead id="30DB1662FFEAFFD3FF57FF054D9AFF31" pageId="12" pageNumber="208">
2. Dorsocentrals in posterior part converge with acrostichal setae (
<figureCitation id="1311A9D2FFEAFFD3FCFDFF054B82FF1D" box="[829,900,181,203]" captionStart="FIGURES 1015" captionStartId="5.[151,264,1824,1848]" captionTargetBox="[156,1426,193,1790]" captionTargetId="figure-21@5.[151,1436,181,1800]" captionTargetPageId="5" captionText="FIGURES 1015. Shilovia xinhuawangi sp. nov., male. 10, basal part of wing; 11, antenna; 12, dorsal part of mesonotum; 13, scutellum and postnotum; 14, hypopygium in dorsal view; 15, hypopygium without tergite IX in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4322459" httpUri="https://zenodo.org/record/4322459/files/figure.png" pageId="12" pageNumber="208">Fig. 12</figureCitation>
). Postnotum with setae (
<figureCitation id="1311A9D2FFEAFFD3FBADFF054CB5FF1D" box="[1133,1203,181,203]" captionStart="FIGURES 1015" captionStartId="5.[151,264,1824,1848]" captionTargetBox="[156,1426,193,1790]" captionTargetId="figure-21@5.[151,1436,181,1800]" captionTargetPageId="5" captionText="FIGURES 1015. Shilovia xinhuawangi sp. nov., male. 10, basal part of wing; 11, antenna; 12, dorsal part of mesonotum; 13, scutellum and postnotum; 14, hypopygium in dorsal view; 15, hypopygium without tergite IX in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4322459" httpUri="https://zenodo.org/record/4322459/files/figure.png" pageId="12" pageNumber="208">Fig. 13</figureCitation>
). Wing with setae on Cu and An (
<figureCitation id="1311A9D2FFEAFFD3FEF4FF62497AFF3E" box="[308,380,210,232]" captionStart="FIGURES 1015" captionStartId="5.[151,264,1824,1848]" captionTargetBox="[156,1426,193,1790]" captionTargetId="figure-21@5.[151,1436,181,1800]" captionTargetPageId="5" captionText="FIGURES 1015. Shilovia xinhuawangi sp. nov., male. 10, basal part of wing; 11, antenna; 12, dorsal part of mesonotum; 13, scutellum and postnotum; 14, hypopygium in dorsal view; 15, hypopygium without tergite IX in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4322459" httpUri="https://zenodo.org/record/4322459/files/figure.png" pageId="12" pageNumber="208">Fig. 10</figureCitation>
). Gonostylus length/gonostylus width 2.863.67..................... …
<taxonomicName id="4C2ACED4FFEAFFD3FBB9FF624D4CFF31" ID-CoL="9HKD9" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2020" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2020" box="[1145,1354,210,231]" class="Insecta" family="Chironomidae" genus="Shilovia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="208" phylum="Arthropoda" rank="species" species="xinhuawangi" status="sp. nov.">
<emphasis id="B95E6945FFEAFFD3FBB9FF624D4CFF31" box="[1145,1354,210,231]" italics="true" pageId="12" pageNumber="208">Shilovia xinhuawangi</emphasis>
</taxonomicName>
<emphasis id="B95E6945FFEAFFD3FA90FF624D9AFF31" bold="true" box="[1360,1436,210,231]" pageId="12" pageNumber="208">
<taxonomicNameLabel id="A26DD43EFFEAFFD3FA90FF624D9AFF31" box="[1360,1436,210,231]" pageId="12" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B95B557FFEAFFD3FF57FF404D9AFEF4" blockId="12.[151,1437,151,291]" pageId="12" pageNumber="208">
<keyLead id="30DB1662FFEAFFD3FF57FF404D9AFEF4" pageId="12" pageNumber="208">
Dorsocentrals in posterior part not converge with acrostichal setae (
<figureCitation id="1311A9D2FFEAFFD3FCA4FF5F4BADFED0" box="[868,939,239,262]" captionStart="FIGURES 1620" captionStartId="8.[151,264,1844,1868]" captionTargetBox="[151,1435,181,1821]" captionTargetId="figure-18@8.[151,1435,181,1821]" captionTargetPageId="8" captionText="FIGURES 1620. Shilovia xinhuawangi sp. nov. (16) and S. yakovlevi sp. nov. (1720), males. 16, legs; 17, basal part of wing; 18, dorsal part of mesonotum; 19, gonocoxite and gonostylus; 20, part of hypopygium in dorsal view. Designations are the same as in Figures 19." figureDoi="http://doi.org/10.5281/zenodo.4322461" httpUri="https://zenodo.org/record/4322461/files/figure.png" pageId="12" pageNumber="208">Fig. 18</figureCitation>
). Postnotum, Cu and An of wing without setae (
<figureCitation id="1311A9D2FFEAFFD3FAB5FF5F48FAFEF5" captionStart="FIGURES 1620" captionStartId="8.[151,264,1844,1868]" captionTargetBox="[151,1435,181,1821]" captionTargetId="figure-18@8.[151,1435,181,1821]" captionTargetPageId="8" captionText="FIGURES 1620. Shilovia xinhuawangi sp. nov. (16) and S. yakovlevi sp. nov. (1720), males. 16, legs; 17, basal part of wing; 18, dorsal part of mesonotum; 19, gonocoxite and gonostylus; 20, part of hypopygium in dorsal view. Designations are the same as in Figures 19." figureDoi="http://doi.org/10.5281/zenodo.4322461" httpUri="https://zenodo.org/record/4322461/files/figure.png" pageId="12" pageNumber="208">Fig. 17</figureCitation>
). Gonostylus length/gonostylus width 5.227.20 …...................................
<taxonomicName id="4C2ACED4FFEAFFD3FB5AFEBD4D4CFEF4" ID-CoL="9HKDB" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2020" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2020" box="[1178,1354,269,290]" class="Insecta" family="Chironomidae" genus="Shilovia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="208" phylum="Arthropoda" rank="species" species="yakovlevi" status="sp. nov.">
<emphasis id="B95E6945FFEAFFD3FB5AFEBD4D4CFEF4" box="[1178,1354,269,290]" italics="true" pageId="12" pageNumber="208">Shilovia yakovlevi</emphasis>
</taxonomicName>
<emphasis id="B95E6945FFEAFFD3FA90FEBD4D9AFEF4" bold="true" box="[1360,1436,269,290]" pageId="12" pageNumber="208">
<taxonomicNameLabel id="A26DD43EFFEAFFD3FA90FEBD4D9AFEF4" box="[1360,1436,269,290]" pageId="12" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>